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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.564.6819" ID-GBIF-Dataset="2b9eca3b-58b9-4499-bbb4-64b8881e6a3f" ID-PMC="PMC4820093" ID-Pensoft-Pub="1313-2970-564-121" ID-PubMed="27081334" ID-ZBK="8D6563D67C4F4435BE6C19CCE2F9882F" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-564-121" ModsDocOrigin="ZooKeys 564" ModsDocTitle="Contributions to the knowledge of subterranean trechine beetles in southern Chinas karsts: five new genera (Insecta, Coleoptera, Carabidae, Trechinae)" checkinTime="1455646106532" checkinUser="pensoft" docAuthor="Tian, Mingyi, Huang, Sunbin, Wang, Xinhui &amp; Tang, Mingruo" docDate="2016" docId="A609C42B76B0B164CAF52AF51935432C" docLanguage="en" docName="ZooKeys 564: 121-156" docOrigin="ZooKeys 564" docSource="http://dx.doi.org/10.3897/zookeys.564.6819" docTitle="Huoyanodytes Tian &amp; Huang, gen. n." docType="treatment" docUuid="1A50BCBA-6A05-4D77-8E78-3EC6C157CAC6" docUuidSource="ZooBank" docVersion="5" lastPageNumber="138" masterDocId="646BFFE3E466300A6C0AFFE5FFBA1245" masterDocTitle="Contributions to the knowledge of subterranean trechine beetles in southern China's karsts: five new genera (Insecta, Coleoptera, Carabidae, Trechinae)" masterLastPageNumber="156" masterPageNumber="121" pageNumber="137" updateTime="1668162679916" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Contributions to the knowledge of subterranean trechine beetles in southern China's karsts: five new genera (Insecta, Coleoptera, Carabidae, Trechinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tian, Mingyi</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Huang, Sunbin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wang, Xinhui</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tang, Mingruo</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>564</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>121</mods:start>
<mods:end>156</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.564.6819</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.564.6819</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-564-121</mods:identifier>
<mods:identifier type="ZBK">8D6563D67C4F4435BE6C19CCE2F9882F</mods:identifier>
<mods:identifier type="ZooBank">8D6563D67C4F4435BE6C19CCE2F9882F</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127880886" LSID="urn:lsid:zoobank.org:act:1A50BCBA-6A05-4D77-8E78-3EC6C157CAC6" httpUri="http://treatment.plazi.org/id/A609C42B76B0B164CAF52AF51935432C" lastPageId="17" lastPageNumber="138" pageId="16" pageNumber="137">
<subSubSection pageId="16" pageNumber="137" type="multiple">
<paragraph pageId="16" pageNumber="137">
<pageBreakToken pageId="16" pageNumber="137" start="start">Taxon</pageBreakToken>
classification Animalia Coleoptera Carabidae
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="137" type="nomenclature">
<paragraph pageId="16" pageNumber="137">
<taxonomicName LSID="http://zoobank.org/1A50BCBA-6A05-4D77-8E78-3EC6C157CAC6" authority="Tian &amp; Huang" class="Insecta" family="Carabidae" genus="Huoyanodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Huoyanodytes" order="Coleoptera" pageId="16" pageNumber="137" phylum="Arthropoda" rank="genus">Huoyanodytes Tian &amp; Huang</taxonomicName>
<taxonomicNameLabel pageId="16" pageNumber="137">gen. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="137" type="type species">
<paragraph pageId="16" pageNumber="137">Type species.</paragraph>
<paragraph pageId="16" pageNumber="137">
<taxonomicName class="Insecta" family="Carabidae" genus="Huoyanodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Huoyanodytes tujiaphilus" order="Coleoptera" pageId="16" pageNumber="137" phylum="Arthropoda" rank="species" species="tujiaphilus">Huoyanodytes tujiaphilus</taxonomicName>
Tian &amp; Huang, sp. n. (Cave Tujiamei Dong, Longshan, Hunan)
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="138" type="diagnosis">
<paragraph pageId="17" pageNumber="138">
<pageBreakToken pageId="17" pageNumber="138" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph pageId="17" pageNumber="138">Large-sized, semi-aphaenopsian beetles, with elongated and tube-like head, long fore body, bidentate right mandible, bisetose mentum, well defined labial suture, tubiform and tumid prothorax, five pores in the humeral group of the marginal umbilicate series and disappeared elytral striae.</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="138" type="generic characteristics">
<paragraph pageId="17" pageNumber="138">Generic characteristics.</paragraph>
<paragraph pageId="17" pageNumber="138">Large-sized, semi-aphaenopsian trechine, eyeless, unpigmented and apterous; fore body longer than elytra; head tube-like, parallel-sided, without neck constriction; much longer than wide, head (including mandible) as long as prothorax; mandible elongate, right mandible bidentate; ligula multisetose; submentum 10-setose, mentum bisetose, each of abdominal ventrites IV-VII 4-setose; labial suture clear, well separating mentum and submentum; frontal furrows short, subparallel-sided, two pairs of supra-orbital pores present; antennae long, extending to a little before elytral apex; prothorax elongate, somewhat tubiform, propleura distinctly tumid and thus visible from above; both fore and hind pronotal angles obtusely rounded; elytra ovate, strongly convex, making marginal side partly concealed and invisible from above; humeral angles rounded, widest at about middle, striae completely missing; apex broadly rounded; two dorsal and the pre-apical pores present on each elytron, humeral group of umbilicate marginal pores composed of five pores, middle group backwardly located, at about apical third of elytra; femora more dilated near subapex; tibiae long and slender, without longitudinal grooves externally.</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="138" type="remarks">
<paragraph pageId="17" pageNumber="138">Remarks.</paragraph>
<paragraph pageId="17" pageNumber="138">
Again, the affinities of
<taxonomicName class="Insecta" family="Carabidae" genus="Huoyanodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Huoyanodytes" order="Coleoptera" pageId="17" pageNumber="138" phylum="Arthropoda" rank="genus">Huoyanodytes</taxonomicName>
gen. n. are bound to remain obscure. Its tube-like head, the more dilated subapically femora, the very convex pronotum and elytra, and the peculiar elytral chaetotaxy are the apomorphies that make it unrelated to any other genera so far known in China. It must be pointed out that it is the first example in a trechine beetle which humeral group of umbilicate marginal pores as composed of five pores, instead of four.
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="138" type="etymology">
<paragraph pageId="17" pageNumber="138">Etymology.</paragraph>
<paragraph pageId="17" pageNumber="138">Huoyan+dytes, to refer to this genus occurring in Huoyan Karst. Gender masculine.</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="138" type="range">
<paragraph pageId="17" pageNumber="138">Range.</paragraph>
<paragraph pageId="17" pageNumber="138">China (northwestern Hunan) (Fig. 5e).</paragraph>
</subSubSection>
</treatment>
</document>