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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.863.33183" ID-GBIF-Dataset="659f8e05-f9a9-4621-a07b-47f47822a694" ID-PMC="PMC6639350" ID-Pensoft-Pub="1313-2970-863-1" ID-PubMed="31341391" ID-ZBK="7F518F005EBB4F3DA2AD324B1760F3FB" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-863-1" ModsDocOrigin="ZooKeys 863" ModsDocTitle="Rainforest and cloud forest Scolytodes (Curculionidae, Scolytinae, Hexacolini) from the Arthropods of La Selva inventory in Costa Rica: new species, new synonymy, new records" checkinTime="1562893184001" checkinUser="pensoft" docAuthor="Jordal, Bjarte H. &amp; Kirkendall, Lawrence R." docDate="2019" docId="A57A489C7FD3B87158105148C3316009" docLanguage="en" docName="ZooKeys 863: 1-34" docOrigin="ZooKeys 863" docSource="http://dx.doi.org/10.3897/zookeys.863.33183" docTitle="Scolytodes niger Jordal &amp; Kirkendall, sp. nov." docType="treatment" docUuid="C160D713-112C-45F3-8FCB-B3CE555CBA7C" docUuidSource="ZooBank" docVersion="5" lastPageNumber="1" masterDocId="FFF6300DFF864305FFECE10BD91CFFA6" masterDocTitle="Rainforest and cloud forest Scolytodes (Curculionidae, Scolytinae, Hexacolini) from the Arthropods of La Selva inventory in Costa Rica: new species, new synonymy, new records" masterLastPageNumber="34" masterPageNumber="1" pageNumber="1" updateTime="1668167558736" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Rainforest and cloud forest Scolytodes (Curculionidae, Scolytinae, Hexacolini) from the Arthropods of La Selva inventory in Costa Rica: new species, new synonymy, new records</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jordal, Bjarte H.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kirkendall, Lawrence R.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>863</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>34</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.863.33183</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.863.33183</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-863-1</mods:identifier>
<mods:identifier type="ZBK">7F518F005EBB4F3DA2AD324B1760F3FB</mods:identifier>
<mods:identifier type="ZooBank">7F518F005EBB4F3DA2AD324B1760F3FB</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="158605318" LSID="urn:lsid:zoobank.org:act:C160D713-112C-45F3-8FCB-B3CE555CBA7C" httpUri="http://treatment.plazi.org/id/A57A489C7FD3B87158105148C3316009" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/C160D713-112C-45F3-8FCB-B3CE555CBA7C" authority="Jordal &amp; Kirkendall" class="Insecta" family="Curculionidae" genus="Scolytodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolytodes niger" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="niger">Scolytodes niger Jordal &amp; Kirkendall</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
Figs 2, 5, 8
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
Holotype, female: Costa Rica, Guanacaste prov., Rincon de La Vieja, Las Pailas, 18.II.1996-020A, R. Anderson,
<taxonomicName class="Magnoliopsida" family="Clusiaceae" genus="Clusia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Clusia rosea" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="rosea">Clusia rosea</taxonomicName>
forest litter. Allotype male: Costa Rica, Puntarenas, 11 km SW Est. Biol. Las Cruces, 1450 m,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="8.778611">08°46'43&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-83.030556">83°01'50&quot;W</geoCoordinate>
, 9.VII.1999, R. Anderson, wet cloud forest litter, 99-124E. Paratypes: Prov. Heredia, 6 km ENE Vara Blanca, 1950-2050 m,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="10.183333">10°11'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-84.11667">84°07'W</geoCoordinate>
, 21 Feb. 2002,
<normalizedToken originalValue="INBioOETALAS">INBio-OET-ALAS</normalizedToken>
transect, 20/TN/12/005 (INB0003223381) (1); San
<normalizedToken originalValue="José">Jose</normalizedToken>
prov.,
<normalizedToken originalValue="Zurquí">Zurqui</normalizedToken>
de Moravia, 1600 m, 30.6.1997, L. Kirkendall, H. Lezama, flight intercept trap (1). Holotype and allotype deposited in CMNC, 1 paratype in MNCR, and 1 paratype in ZMBN.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Interstriae 10 carinate to level of ventrite 2; protibiae with an additional mesal tooth near tarsal insertion. Metanepisternum covered with white plumose setae. Very similar to
<taxonomicName lsidName="S. clusiae" pageId="0" pageNumber="1" rank="species" species="clusiae">S. clusiae</taxonomicName>
Wood, 1969, but differs by the smooth and shiny pronotum, the shorter setae in a smaller area in the female frons, and by the distinct, albeit tiny, interstrial punctures.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description female.</paragraph>
<paragraph pageId="0" pageNumber="1">
Length 1.5 mm, 2.1
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide; color black. Head. Eyes sinuate, separated above by 2.0
<normalizedToken originalValue="×">x</normalizedToken>
their width. Frons weakly impressed from just above level of antennal insertion to epistoma; surface smooth, densely punctured in impressed area, scattered punctures above, shiny. Vestiture consisting of short setae in impressed area and on epistoma. Antennal club with two sutures marked by shorter setae, segment 1 and 2 corneous, sutures slightly constricted; funiculus 6-segmented. Pronotum shiny, with shallow, small, elongated punctures spaced by 2-3
<normalizedToken originalValue="×">x</normalizedToken>
their diameter (length). Vestiture consisting of 8 erect long setae (4
<normalizedToken originalValue="2">-2-</normalizedToken>
2). Elytra generally smooth and shiny; striae not impressed, punctures shallow, tiny, appear elongated but composed by two punctures in one, each pair separated in rows by their length; interstriae 4
<normalizedToken originalValue="×">x</normalizedToken>
as wide as striae, punctures of same size as in striae, much more separated, mainly in rows. Interstriae 10 carinate to level of ventrite 2. Vestiture consisting of regular rows of erect, interstrial setae of variable thickness, and regularly placed, fine, short, semi-recumbent setae in striae and interstriae, mainly on posterior half. Legs. Procoxae broadly separated by 0.9
<normalizedToken originalValue="×">x</normalizedToken>
and mesocoxae 1.2
<normalizedToken originalValue="×">x</normalizedToken>
the width of one procoxa. Protibiae slightly broadening distally, lateral teeth 1 and 2 of similar size, with a faint extension of cuticle between them, and with 3-4 additional small, sharp spines or granules along the lateral edge towards base; an additional mesal tooth present near tarsal insertion; protibial mucro obtuse. Meso- and metatibiae with 5 and 6 lateral, socketed, small teeth on distal half and third, respectively. Ventral vestiture. Setae on metanepisternum quadrifid to strongly plumose and densely placed, on mesanepisternum mainly trifid, on metasternum mainly simple, some bifid. Sclerolepidia scale-like or plumose.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="male">
<paragraph pageId="0" pageNumber="1">Male.</paragraph>
<paragraph pageId="0" pageNumber="1">
1.4-1.6 mm long, 2.0-2.1
<normalizedToken originalValue="×">x</normalizedToken>
as wide as long. Identical to female, except frons slightly more convex on lower part, with less dense vestiture.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="key">
<paragraph pageId="0" pageNumber="1">Key</paragraph>
<paragraph pageId="0" pageNumber="1">
(
<bibRefCitation author="Wood, SL" journalOrPublisher="Great Basin Naturalist Memoirs" pageId="0" pageNumber="1" pagination="1 - 1359" title="The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph." volume="6" year="1982">Wood 1982</bibRefCitation>
). Keys to couplet 25 with some uncertainty;
<taxonomicName lsidName="S. clusiae" pageId="0" pageNumber="1" rank="species" species="clusiae">S. clusiae</taxonomicName>
in couplet 24 is a better match.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The Latin name
<taxonomicName lsidName="niger" pageId="0" pageNumber="1" rank="species" species="niger">niger</taxonomicName>
is a masculine adjective, meaning shining black, referring to the dark and shiny appearance of this species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
<paragraph pageId="0" pageNumber="1">Biology and distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is known from four Costa Rican cloud forest localities - in the northern and southern slopes of Volcan Barva (Braulio Carrillo), the southern slope of Rincon de La Vieja, and close to the Panama border (Las Cruces). Two specimens were collected by flight intercept trapping, and two by leaf litter sifting, one of which from
<taxonomicName class="Insecta" family="Clusiidae" genus="Clusia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clusia" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Clusia</taxonomicName>
litter.
</paragraph>
</subSubSection>
</treatment>
</document>