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<document id="4F75F64B345125DC6BB2D976E8BCEBFD" ID-CLB-Dataset="199795" ID-DOI="10.5852/ejt.2022.835.1909" ID-GBIF-Dataset="31a2a3ea-9d40-4178-8629-da4b275f37dc" ID-ISSN="2118-9773" ID-Zenodo-Dep="7033400" ID-ZooBank="8B8432B1-C714-4179-8687-66902F4CBF53" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1661823783728" checkinUser="felipe" docAuthor="Groll, Elisa Von &amp; Lopes-Andrade, Cristiano" docDate="2022" docId="A5415E5FF414FFC8985BF980FD6EFA25" docLanguage="en" docName="ejt-835_1-97.pdf" docOrigin="European Journal of Taxonomy 835 (1)" docSource="http://dx.doi.org/10.5852/ejt.2022.835.1909" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cyparium achardi von Groll &amp; Lopes-Andrade 2022, sp. nov." docType="treatment" docUuid="ABFE4E1F-DB31-46C1-BDC7-BDD1C4FF81E3" docUuidSource="ZooBank" docVersion="11" lastPageNumber="25" masterDocId="59782627F41AFFD09A20FFFAFFCCFFB2" masterDocTitle="Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species" masterLastPageNumber="97" masterPageNumber="1" pageNumber="15" updateTime="1699374449524" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="CD3C568FEE83CA1C8A50B4BB2F98B5A1">Contributions to the taxonomy of Neotropical Cyparium Erichson (Coleoptera: Staphylinidae: Scaphidiinae), with the description of five new species</mods:title>
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<mods:namePart id="CE3A02E08060A559084D778E60CF81C9">Groll, Elisa Von</mods:namePart>
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<mods:affiliation id="FE05AEDFD3968C367B8BC5F7F5CA14CD">Programa de Pós-Graduação em Biologia Animal, Universidade Federal de Viçosa, Av. Peter Henry Rolfs, s / n, 36570 - 900, Viçosa, MG, Brasil. &amp; Laboratório de Sistemática e Biologia de Coleoptera, Departamento de Biologia Animal, Universidade Federal de Viçosa, Av. Peter Henry Rolfs, s / n, 36570 - 900, Viçosa, MG, Brasil.</mods:affiliation>
<mods:nameIdentifier id="E89D383B293BA4EFD4FE7FC121E05DEC" type="email">elisavgroll@gmail.com</mods:nameIdentifier>
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<mods:namePart id="440FB9FA18A0178EE43DC4004FC45807">Lopes-Andrade, Cristiano</mods:namePart>
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<mods:affiliation id="9CCC12A41B326BB74F37C1EFD144B758">Programa de Pós-Graduação em Biologia Animal, Universidade Federal de Viçosa, Av. Peter Henry Rolfs, s / n, 36570 - 900, Viçosa, MG, Brasil. &amp; Laboratório de Sistemática e Biologia de Coleoptera, Departamento de Biologia Animal, Universidade Federal de Viçosa, Av. Peter Henry Rolfs, s / n, 36570 - 900, Viçosa, MG, Brasil.</mods:affiliation>
<mods:nameIdentifier id="B9E13A43F51638974EEEDA8E8B7C825E" type="email">ciidae@gmail.com</mods:nameIdentifier>
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<mods:title id="4509617239116E12438363493A22C02A">European Journal of Taxonomy</mods:title>
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<mods:date id="20D578D2E07AE2B40F809E5286FC5A2D">2022</mods:date>
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<mods:number id="5E6E73E7F11F3EB0D14E1F728C1AA52F">2022-08-24</mods:number>
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<mods:number id="6DDBA2F36C9B6C91F74E21FF561D97F0">835</mods:number>
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<treatment id="A5415E5FF414FFC8985BF980FD6EFA25" ID-DOI="http://doi.org/10.5281/zenodo.7392074" ID-GBIF-Taxon="205699035" ID-Zenodo-Dep="7392074" LSID="urn:lsid:zoobank.org:act:ABFE4E1F-DB31-46C1-BDC7-BDD1C4FF81E3" httpUri="http://treatment.plazi.org/id/A5415E5FF414FFC8985BF980FD6EFA25" lastPageId="24" lastPageNumber="25" pageId="14" pageNumber="15">
<subSubSection id="65F2BCC2F414FFDE985BF980FC75F926" box="[635,953,1657,1684]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="2D57EF49F414FFDE985BF980FC75F926" blockId="14.[351,1237,1657,1754]" box="[635,953,1657,1684]" pageId="14" pageNumber="15">
<heading id="761F5825F414FFDE985BF980FC75F926" box="[635,953,1657,1684]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="15" reason="2">
<taxonomicName id="EAE894CAF414FFDE985BF980FC9BF926" ID-CoL="9GKWB" authority="Groll &amp; Lopes-Andrade, 2022" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[635,855,1657,1684]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="achardi" status="sp. nov.">
<emphasis id="1F9C335BF414FFDE985BF980FC9BF926" bold="true" box="[635,855,1657,1684]" italics="true" pageId="14" pageNumber="15">Cyparium achardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AF8E20F414FFDE997FF980FC75F926" box="[863,953,1658,1684]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="65F2BCC2F414FFDE9B7FF967FC03F968" pageId="14" pageNumber="15" type="description">
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urn:lsid:zoobank.org:act:
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</uri>
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<paragraph id="2D57EF49F414FFDE9846F93AFC03F968" blockId="14.[351,1237,1657,1754]" box="[614,975,1728,1754]" pageId="14" pageNumber="15">
<figureCitation id="B5D3F3CCF414FFDE9846F93AFD7DF968" box="[614,689,1728,1754]" captionStart="Fig" captionStartId="7.[189,232,1799,1825]" captionTargetBox="[193,1384,280,1696]" captionTargetId="figure-11@7.[189,1398,265,1704]" captionTargetPageId="7" captionText="Fig. 4. Cyparium Erichson, 1845. Neotropical species. Illustrations based on the original descriptions. *Illustrations made based on photographs, specimens, or illustrations made by other authors; **size not provided in original description. Model: Cyparium collare Pic, 1920. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7033408" httpUri="https://zenodo.org/record/7033408/files/figure.png" pageId="14" pageNumber="15">Figs 4</figureCitation>
,
<figureCitation id="B5D3F3CCF414FFDE989DF93AFD37F968" box="[701,763,1728,1754]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStartId-0="16.[189,232,1814,1840]" captionStartId-1="18.[189,232,1831,1857]" captionStartId-2="19.[189,232,1773,1799]" captionStartId-3="20.[189,232,1771,1797]" captionStartId-4="21.[189,232,1831,1857]" captionStartId-5="22.[189,232,1831,1857]" captionStartId-6="23.[189,232,1808,1834]" captionTargetBox-0="[189,1398,273,1759]" captionTargetBox-1="[192,1393,273,1749]" captionTargetBox-2="[189,1398,285,1696]" captionTargetBox-3="[189,1392,273,1719]" captionTargetBox-4="[189,1392,266,1756]" captionTargetBox-5="[236,1394,275,1759]" captionTargetBox-6="[189,1398,266,1752]" captionTargetId-0="figure-16@16.[189,1398,265,1759]" captionTargetId-1="figure-17@18.[189,1399,265,1759]" captionTargetId-2="figure-11@19.[189,1398,265,1700]" captionTargetId-3="figure-16@20.[189,1398,265,1727]" captionTargetId-4="figure-11@21.[189,1398,266,1760]" captionTargetId-5="figure-17@22.[189,1399,265,1759]" captionTargetId-6="figure-12@23.[189,1398,264,1758]" captionTargetPageId-0="16" captionTargetPageId-1="18" captionTargetPageId-2="19" captionTargetPageId-3="20" captionTargetPageId-4="21" captionTargetPageId-5="22" captionTargetPageId-6="23" captionText-0="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." captionText-1="Fig. 8. Cyparium achardi sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D. Holotype, ♂ (CELC), pronotum, dorsal view. E–G. Paratype, ♂ (CELC), prothorax. E. Frontal view. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: ns = notosternal suture; pr = profurca; st = sclerotized plate. Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.5 mm." captionText-2="Fig. 9. Cyparium achardi sp. nov., paratype, ♂ (CELC). A. Prothorax, lateral view. B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: al = alacrista; ap = anterior phragma; ip = intercoxal plate; ml = median line; mma = median membranous area; mp = mesoventral process; mvl = mesoventral line; mxl = mesocoxal line; pp = prosternal process; ps = prescutellar line; px = procoxal rest; sss = scutoscutellar suture. Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." captionText-3="Fig. 10. Cyparium achardi sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: CAS = cubitoanal strut; fa = furcal arms; MP 1+2 = medial field; MSP = medial spur; RA 4 = radius anterior; RP 3+4 = radius posterior; sr = ‘stalk ridge’; vlf = ventral longitudinal flange. Scale bars:A–D, F–H = 0.2 mm; E = 0.5 mm." captionText-4="Fig. 11. Cyparium achardi sp. nov. A, C. Holotype, ♂ (CELC). B, D–G. Paratype ♂ (CELC). A–B. Abdomen, dorsal view (A) and ventral view (B). C. Ventrite 1. D. Tergite VIII. E. Terminalia. F. Tergite IX. G. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: st = sternite; tg = tergite; vs = ventral strut. Scale bars: A–B = 0.4 mm; C–G = 0.1 mm." captionText-5="Fig. 12. Cyparium achardi sp. nov., paratype, ♂ (CELC): aedeagi. A. Lateral view. B. Frontal view. C. Dorsal view. D. Internal sac, frontal view. E. Internal sac, dorsal view. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: hk = hook; op = opening; pa = parameres. Scale bars = 0.2 mm." captionText-6="Fig. 13. Cyparium achardi sp. nov., paratype, ♀ (CELC). A. Tergite VIII.B. Sternite VIII.C. Terminalia. D. Sclerite of vaginal plate. E. Ovipositor. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: bc = bursa copulatrix; dg = distal gonocoxites; gs = gonostyli; sc = sclerite; sp = spermatheca. Scale bars: A–C = 0.2 mm; D–E = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7033414" figureDoi-1="http://doi.org/10.5281/zenodo.7033416" figureDoi-2="http://doi.org/10.5281/zenodo.7033418" figureDoi-3="http://doi.org/10.5281/zenodo.7033420" figureDoi-4="http://doi.org/10.5281/zenodo.7033422" figureDoi-5="http://doi.org/10.5281/zenodo.7033424" figureDoi-6="http://doi.org/10.5281/zenodo.7033426" httpUri-0="https://zenodo.org/record/7033414/files/figure.png" httpUri-1="https://zenodo.org/record/7033416/files/figure.png" httpUri-2="https://zenodo.org/record/7033418/files/figure.png" httpUri-3="https://zenodo.org/record/7033420/files/figure.png" httpUri-4="https://zenodo.org/record/7033422/files/figure.png" httpUri-5="https://zenodo.org/record/7033424/files/figure.png" httpUri-6="https://zenodo.org/record/7033426/files/figure.png" pageId="14" pageNumber="15">7–13</figureCitation>
,
<figureCitation id="B5D3F3CCF414FFDE9927F93AFCE4F968" box="[775,808,1728,1754]" captionStart="Fig" captionStartId="65.[189,232,1952,1978]" captionTargetBox="[189,1398,1082,1910]" captionTargetId="figure-297@65.[189,1398,1082,1910]" captionTargetPageId="65" captionText="Fig. 46. A. Distribution map of species of Cyparium Erichson, 1845, South America. B. Mata da Biologia. C. Mata do Paraíso." figureDoi="http://doi.org/10.5281/zenodo.7033501" httpUri="https://zenodo.org/record/7033501/files/figure.png" pageId="14" pageNumber="15">46</figureCitation>
; Supp. file 1A
</paragraph>
</subSubSection>
<subSubSection id="65F2BCC2F414FFDE9A9DF8FBFC5BF865" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="2D57EF49F414FFDE9A9DF8FBFEFAF8A9" blockId="14.[189,310,1793,1819]" box="[189,310,1793,1819]" pageId="14" pageNumber="15">
<heading id="761F5825F414FFDE9A9DF8FBFEFAF8A9" bold="true" box="[189,310,1793,1819]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="1F9C335BF414FFDE9A9DF8FBFEFAF8A9" bold="true" box="[189,310,1793,1819]" pageId="14" pageNumber="15">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="2D57EF49F414FFDE9A9DF8CBFC5BF865" blockId="14.[189,1401,1840,2007]" pageId="14" pageNumber="15">
TL: 3.00–
<quantity id="EA1042ACF414FFDE9B15F8CAFE53F8F9" box="[309,415,1840,1867]" metricMagnitude="-3" metricUnit="m" metricValue="3.28" pageId="14" pageNumber="15" unit="mm" value="3.28">3.28 mm</quantity>
in males and
<quantity id="EA1042ACF414FFDE9869F8CAFD39F8F9" box="[585,757,1840,1867]" metricMagnitude="-3" metricUnit="m" metricValue="3.36" metricValueMax="3.4" metricValueMin="3.32" pageId="14" pageNumber="15" unit="mm" value="3.36" valueMax="3.4" valueMin="3.32">3.32–3.40 mm</quantity>
in females. Entirely black, with lateral areas of some ventral sclerites lighter (
<figureCitation id="B5D3F3CCF414FFDE9BD0F8A9FDB5F8DF" box="[496,633,1875,1901]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="14" pageNumber="15">Fig. 7A–C</figureCitation>
). Eyes long (
<figureCitation id="B5D3F3CCF414FFDE9906F8A9FC47F8DF" box="[806,907,1875,1901]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="14" pageNumber="15">Fig. 7D</figureCitation>
). Hypomeron and metaventrite smooth. Metanepisternum and metepimeron with imbricate microsculpture. Tergite VIII of males coarsely punctate (
<figureCitation id="B5D3F3CCF414FFDE9B10F860FE57F806" box="[304,411,1946,1972]" captionStart="Fig" captionStartId="21.[189,232,1831,1857]" captionTargetBox="[189,1392,266,1756]" captionTargetId="figure-11@21.[189,1398,266,1760]" captionTargetPageId="21" captionText="Fig. 11. Cyparium achardi sp. nov. A, C. Holotype, ♂ (CELC). B, D–G. Paratype ♂ (CELC). A–B. Abdomen, dorsal view (A) and ventral view (B). C. Ventrite 1. D. Tergite VIII. E. Terminalia. F. Tergite IX. G. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: st = sternite; tg = tergite; vs = ventral strut. Scale bars: A–B = 0.4 mm; C–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033422" httpUri="https://zenodo.org/record/7033422/files/figure.png" pageId="14" pageNumber="15">Fig. 11D</figureCitation>
). Aedeagus strongly sclerotized (
<figureCitation id="B5D3F3CCF414FFDE993FF860FC62F806" box="[799,942,1946,1972]" captionStart="Fig" captionStartId="22.[189,232,1831,1857]" captionTargetBox="[236,1394,275,1759]" captionTargetId="figure-17@22.[189,1399,265,1759]" captionTargetPageId="22" captionText="Fig. 12. Cyparium achardi sp. nov., paratype, ♂ (CELC): aedeagi. A. Lateral view. B. Frontal view. C. Dorsal view. D. Internal sac, frontal view. E. Internal sac, dorsal view. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: hk = hook; op = opening; pa = parameres. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7033424" httpUri="https://zenodo.org/record/7033424/files/figure.png" pageId="14" pageNumber="15">Fig. 12A–C</figureCitation>
); apex and parameres short (
<figureCitation id="B5D3F3CCF414FFDE9EDCF860FAA5F806" box="[1276,1385,1946,1972]" captionStart="Fig" captionStartId="22.[189,232,1831,1857]" captionTargetBox="[236,1394,275,1759]" captionTargetId="figure-17@22.[189,1399,265,1759]" captionTargetPageId="22" captionText="Fig. 12. Cyparium achardi sp. nov., paratype, ♂ (CELC): aedeagi. A. Lateral view. B. Frontal view. C. Dorsal view. D. Internal sac, frontal view. E. Internal sac, dorsal view. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: hk = hook; op = opening; pa = parameres. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7033424" httpUri="https://zenodo.org/record/7033424/files/figure.png" pageId="14" pageNumber="15">Fig. 12A</figureCitation>
). Distal gonocoxites of females straight and thick (
<figureCitation id="B5D3F3CCF414FFDE98DDF847FC46F865" box="[765,906,1981,2007]" captionStart="Fig" captionStartId="23.[189,232,1808,1834]" captionTargetBox="[189,1398,266,1752]" captionTargetId="figure-12@23.[189,1398,264,1758]" captionTargetPageId="23" captionText="Fig. 13. Cyparium achardi sp. nov., paratype, ♀ (CELC). A. Tergite VIII.B. Sternite VIII.C. Terminalia. D. Sclerite of vaginal plate. E. Ovipositor. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: bc = bursa copulatrix; dg = distal gonocoxites; gs = gonostyli; sc = sclerite; sp = spermatheca. Scale bars: A–C = 0.2 mm; D–E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033426" httpUri="https://zenodo.org/record/7033426/files/figure.png" pageId="14" pageNumber="15">Fig. 13C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="65F2BCC2F415FFDF9A9DFEF3FE22FEC6" pageId="15" pageNumber="16" type="etymology">
<paragraph id="2D57EF49F415FFDF9A9DFEF3FE88FE91" blockId="15.[189,324,265,291]" box="[189,324,265,291]" pageId="15" pageNumber="16">
<heading id="761F5825F415FFDF9A9DFEF3FE88FE91" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="1F9C335BF415FFDF9A9DFEF3FE88FE91" bold="true" box="[189,324,265,291]" pageId="15" pageNumber="16">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="2D57EF49F415FFDF9A9DFEC2FE22FEC6" blockId="15.[189,1399,311,372]" pageId="15" pageNumber="16">
In homage to the entomologist Julien Achard (1881–1925) for his significant contribution on the Neotropical
<taxonomicName id="EAE894CAF415FFDF9B6FFEA0FE26FEC6" authorityName="Latreille" authorityYear="1806" box="[335,490,346,372]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="subFamily" subFamily="Scaphidiinae">Scaphidiinae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="65F2BCC2F415FFDF9A9DFE61FEB2FB83" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="2D57EF49F415FFDF9A9DFE61FE66FE07" blockId="15.[189,426,411,437]" box="[189,426,411,437]" pageId="15" pageNumber="16">
<heading id="761F5825F415FFDF9A9DFE61FE66FE07" bold="true" box="[189,426,411,437]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="1F9C335BF415FFDF9A9DFE61FE66FE07" bold="true" box="[189,426,411,437]" pageId="15" pageNumber="16">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="2D57EF49F415FFDF9ACAFE30FE0AFDFC" blockId="15.[189,1399,458,590]" pageId="15" pageNumber="16">
<materialsCitation id="9D80E514F415FFDF9ACAFE30FE0AFDFC" ID-GBIF-Occurrence="3983014382" collectingDate="2019-11-19" collectionCode="EPTEA" collectorName="LabCol" country="Brazil" location="Mata do Paraiso" municipality="Vicosa" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="holotype">
<emphasis id="1F9C335BF415FFDF9ACAFE30FE90FE56" bold="true" box="[234,348,458,484]" pageId="15" pageNumber="16">
<typeStatus id="F25351EBF415FFDF9ACAFE30FE90FE56" box="[234,348,458,484]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="55FFAFD9F415FFDF9A9DFE14FEEAFDBA" box="[189,294,494,520]" name="Brazil" pageId="15" pageNumber="16">BRAZIL</collectingCountry>
•
<specimenCount id="3BEE24C0F415FFDF9B1EFE14FE9BFDBA" box="[318,343,494,520]" pageId="15" pageNumber="16" type="male">♂</specimenCount>
;
<collectingRegion id="EF2C21ABF415FFDF9B44FE17FDC8FDB5" box="[356,516,493,519]" country="Brazil" name="Minas Gerais" pageId="15" pageNumber="16">Minas Gerais</collectingRegion>
,
<collectingMunicipality id="CD337533F415FFDF982FFE17FDAEFDB5" box="[527,610,493,519]" pageId="15" pageNumber="16">Viçosa</collectingMunicipality>
,
<collectionCode id="4BF9778CF415FFDF984DFE14FD0BFDBA" box="[621,711,494,520]" pageId="15" pageNumber="16">EPTEA</collectionCode>
<location id="2837B992F415FFDF98EEFE14FC5CFDBA" LSID="urn:lsid:plazi:treatment:A5415E5FF414FFC8985BF980FD6EFA25:2837B992F415FFDF98EEFE14FC5CFDBA" box="[718,912,493,520]" country="Brazil" municipality="Vicosa" name="Mata do Paraiso" pageId="15" pageNumber="16" stateProvince="Minas Gerais">Mata do Paraíso</location>
;
<date id="5956C989F415FFDF99BBFE14FBF1FDBA" box="[923,1085,494,520]" pageId="15" pageNumber="16" value="2019-11-19">
<collectingDate id="49123061F415FFDF99BBFE14FBF1FDBA" box="[923,1085,494,520]" pageId="15" pageNumber="16" value="2019-11-19">19 Nov. 2019</collectingDate>
</date>
;
<collectorName id="801D8A9FF415FFDF9E69FE17FB6DFDB5" box="[1097,1185,493,519]" pageId="15" pageNumber="16">LabCol</collectorName>
leg.; “Fungo 27 \\ Em
<taxonomicName id="EAE894CAF415FFDF9ACFFDEBFE0BFD99" box="[239,455,529,555]" class="Agaricomycetes" family="Agaricaceae" genus="Leucoagaricus" kingdom="Fungi" order="Agaricales" pageId="15" pageNumber="16" phylum="Basidiomycota" rank="species" species="undetermined">
<emphasis id="1F9C335BF415FFDF9ACFFDEBFE51FD98" box="[239,413,529,554]" italics="true" pageId="15" pageNumber="16">Leucoagaricus</emphasis>
sp.
</taxonomicName>
\\
<taxonomicName id="EAE894CAF415FFDF9BCAFDEBFBD1FD99" authority="von Groll &amp; Lopes-Andrade" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[490,1053,529,555]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="achardi">
<emphasis id="1F9C335BF415FFDF9BCAFDEBFD0EFD99" box="[490,706,529,555]" italics="true" pageId="15" pageNumber="16">Cyparium achardi</emphasis>
von Groll &amp; Lopes-Andrade
</taxonomicName>
<typeStatus id="F25351EBF415FFDF9E07FDEBFB14FD99" box="[1063,1240,529,555]" pageId="15" pageNumber="16" type="holotype">HOLOTYPUS</typeStatus>
” [red paper];
<collectionCode id="4BF9778CF415FFDF9A9FFDCEFEC2FDFC" box="[191,270,564,590]" pageId="15" pageNumber="16">CELC</collectionCode>
. (Supp. file1A).
</materialsCitation>
</paragraph>
<paragraph id="2D57EF49F415FFDF9ACAFD8FFB02FC7E" blockId="15.[189,1398,629,973]" pageId="15" pageNumber="16">
<materialsCitation id="9D80E514F415FFDF9ACAFD8FFE4BFD4B" ID-GBIF-Occurrence="3983014372" collectingDate="2016-11-08" collectionCode="T, CELC" collectorName="C. Lopes-Andrade" country="Brazil" location="Pesquisador" municipality="Vicosa" pageId="15" pageNumber="16" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Minas Gerais" typeStatus="paratype">
<emphasis id="1F9C335BF415FFDF9ACAFD8FFEABFD3D" bold="true" box="[234,359,629,655]" pageId="15" pageNumber="16">
<typeStatus id="F25351EBF415FFDF9ACAFD8FFEABFD3D" box="[234,359,629,655]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="55FFAFD9F415FFDF9A9DFD63FEEAFD01" box="[189,294,665,691]" name="Brazil" pageId="15" pageNumber="16">BRAZIL</collectingCountry>
•
<specimenCount id="3BEE24C0F415FFDF9B68FD63FE5DFD01" box="[328,401,665,691]" pageId="15" pageNumber="16" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="3BEE24C0F415FFDF9B83FD63FE1FFD01" box="[419,467,665,691]" pageId="15" pageNumber="16" type="female">1 ♀</specimenCount>
(
<specimenCount id="3BEE24C0F415FFDF9BCBFD63FDD6FD01" box="[491,538,665,691]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
entirely dissected, preserved in glycerin); same collection data as for holotype; “
<collectionCode id="4BF9778CF415FFDF9B65FD46FE95FD64" box="[325,345,700,726]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T</collectionCode>
. do
<location id="2837B992F415FFDF9BB6FD46FDE6FD64" LSID="urn:lsid:plazi:treatment:A5415E5FF414FFC8985BF980FD6EFA25:2837B992F415FFDF9BB6FD46FDE6FD64" box="[406,554,700,726]" country="Brazil" name="Pesquisador" pageId="15" pageNumber="16">Pesquisador</location>
”;
<date id="5956C989F415FFDF9866FD46FD2AFD67" box="[582,742,699,726]" pageId="15" pageNumber="16" value="2016-11-08">
<collectingDate id="49123061F415FFDF9866FD46FD2AFD67" box="[582,742,699,726]" pageId="15" pageNumber="16" value="2016-11-08">8 Nov. 2016</collectingDate>
</date>
;
<collectorName id="801D8A9FF415FFDF98D8FD46FC1BFD64" box="[760,983,700,726]" pageId="15" pageNumber="16">C. Lopes-Andrade</collectorName>
<emphasis id="1F9C335BF415FFDF99C5FD46FBEAFD64" box="[997,1062,700,726]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
leg.; “\\ Em
<taxonomicName id="EAE894CAF415FFDF9EF5FD46FED3FD4B" class="Agaricomycetes" family="Omphalotaceae" genus="Marasmiellus" kingdom="Fungi" order="Agaricales" pageId="15" pageNumber="16" phylum="Basidiomycota" rank="species" species="volvatus">
<emphasis id="1F9C335BF415FFDF9EF5FD46FED3FD4B" italics="true" pageId="15" pageNumber="16">Marasmiellus volvatus</emphasis>
</taxonomicName>
”;
<collectionCode id="4BF9778CF415FFDF9B1CFD25FE4BFD4B" box="[316,391,735,761]" pageId="15" pageNumber="16">CELC</collectionCode>
</materialsCitation>
•
<materialsCitation id="9D80E514F415FFDF9B83FD25FC28FCAE" ID-GBIF-Occurrence="3983014330" collectingDate="2019-11-14" collectionCode="CELC" collectorName="LabCol" country="Brazil" location="Mata do Paraiso" municipality="Vicosa" pageId="15" pageNumber="16" specimenCount="3" specimenCount-female="3" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCount id="3BEE24C0F415FFDF9B83FD25FE2AFD4B" box="[419,486,735,761]" pageId="15" pageNumber="16" type="female">2 ♀♀</specimenCount>
(
<specimenCount id="3BEE24C0F415FFDF9BDAFD25FDE9FD4B" box="[506,549,735,761]" pageId="15" pageNumber="16" type="female">1 ♀</specimenCount>
, abdomen dissected, preserved in glycerin); same collection data as for holotype;
<date id="5956C989F415FFDF9B14FCF8FE1BFCAE" box="[308,471,770,797]" pageId="15" pageNumber="16" value="2019-11-14">
<collectingDate id="49123061F415FFDF9B14FCF8FE1BFCAE" box="[308,471,770,797]" pageId="15" pageNumber="16" value="2019-11-14">14 Nov. 2019</collectingDate>
</date>
; “Fungo 22 \\ Em
<taxonomicName id="EAE894CAF415FFDF9892FCF8FCB3FCAE" box="[690,895,770,796]" class="Agaricomycetes" family="Omphalotaceae" genus="Marasmiellus" kingdom="Fungi" order="Agaricales" pageId="15" pageNumber="16" phylum="Basidiomycota" rank="species" species="undetermined">
<emphasis id="1F9C335BF415FFDF9892FCF8FC9FFCAE" box="[690,851,770,796]" italics="true" pageId="15" pageNumber="16">Marasmiellus</emphasis>
sp.
</taxonomicName>
”;
<collectionCode id="4BF9778CF415FFDF99B9FCF8FC28FCAE" box="[921,996,770,796]" pageId="15" pageNumber="16">CELC</collectionCode>
</materialsCitation>
•
<materialsCitation id="9D80E514F415FFDF99DFFCF8FCDDFC8D" ID-GBIF-Occurrence="3983014364" collectingDate="2019-11-19" collectionCode="CELC" collectorName="LabCol" country="Brazil" location="Mata do Paraiso" municipality="Vicosa" pageId="15" pageNumber="16" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCount id="3BEE24C0F415FFDF99DFFCF8FBE0FCAF" box="[1023,1068,770,797]" pageId="15" pageNumber="16" type="female">1 ♀</specimenCount>
; same collection data as for holotype; “Fungo 26 \\ Em
<taxonomicName id="EAE894CAF415FFDF9820FCDFFD67FC8D" box="[512,683,805,831]" class="Agaricomycetes" family="Hygrophoraceae" genus="Hygrocybe" kingdom="Fungi" order="Agaricales" pageId="15" pageNumber="16" phylum="Basidiomycota" rank="species" species="undetermined">
<emphasis id="1F9C335BF415FFDF9820FCDFFD4CFC8D" box="[512,640,805,831]" italics="true" pageId="15" pageNumber="16">Hygrocybe</emphasis>
sp.
</taxonomicName>
”;
<collectionCode id="4BF9778CF415FFDF98E6FCDFFCDDFC8D" box="[710,785,805,831]" pageId="15" pageNumber="16">CELC</collectionCode>
</materialsCitation>
•
<materialsCitation id="9D80E514F415FFDF990AFCDFFD72FCD1" ID-GBIF-Occurrence="3983014376" collectingDate="2019-11-19" collectionCode="CELC" collectorName="LabCol" country="Brazil" location="Mata do Paraiso" municipality="Vicosa" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCount id="3BEE24C0F415FFDF990AFCDFFC96FCF2" box="[810,858,805,832]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
; same collection data as for holotype; “Fungo 28 \\ Em
<taxonomicName id="EAE894CAF415FFDF9B10FCB3FD98FCD0" box="[304,596,840,866]" class="Agaricomycetes" family="Agaricaceae" genus="Leucocoprinus" kingdom="Fungi" order="Agaricales" pageId="15" pageNumber="16" phylum="Basidiomycota" rank="species" species="ianthinus">
<emphasis id="1F9C335BF415FFDF9B10FCB3FD98FCD0" box="[304,596,840,866]" italics="true" pageId="15" pageNumber="16">Leucocoprinus ianthinus</emphasis>
</taxonomicName>
”;
<collectionCode id="4BF9778CF415FFDF9853FCB3FD72FCD1" box="[627,702,841,867]" pageId="15" pageNumber="16">CELC</collectionCode>
</materialsCitation>
•
<materialsCitation id="9D80E514F415FFDF98FBFCB3FD7EFC34" ID-GBIF-Occurrence="3983014329" collectingDate="2019-11-19" collectionCode="CELC" collectorName="LabCol" country="Brazil" location="Mata do Paraiso" municipality="Vicosa" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCount id="3BEE24C0F415FFDF98FBFCB3FCC0FCD1" box="[731,780,841,867]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
; same collection data as for holotype; “Fungo 12 \\ Em
<taxonomicName id="EAE894CAF415FFDF9ACFFC96FD85FC34" box="[239,585,876,902]" class="Agaricomycetes" family="Marasmiaceae" genus="Marasmius" kingdom="Fungi" order="Agaricales" pageId="15" pageNumber="16" phylum="Basidiomycota" rank="species" species="haematocephalus">
<emphasis id="1F9C335BF415FFDF9ACFFC96FD85FC34" box="[239,585,876,902]" italics="true" pageId="15" pageNumber="16">Marasmius haematocephalus</emphasis>
</taxonomicName>
”;
<collectionCode id="4BF9778CF415FFDF9847FC96FD7EFC34" box="[615,690,876,902]" pageId="15" pageNumber="16">CELC</collectionCode>
</materialsCitation>
•
<materialsCitation id="9D80E514F415FFDF98EEFC96FC92FC1B" ID-GBIF-Occurrence="3983014365" collectingDate="2019-11-26" collectionCode="CELC" collectorName="LabCol" country="Brazil" location="Mata do Paraiso" municipality="Vicosa" pageId="15" pageNumber="16" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCount id="3BEE24C0F415FFDF98EEFC96FD31FC34" box="[718,765,876,902]" pageId="15" pageNumber="16" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<date id="5956C989F415FFDF9EECFC91FABEFC34" box="[1228,1394,875,902]" pageId="15" pageNumber="16" value="2019-11-26">
<collectingDate id="49123061F415FFDF9EECFC91FABEFC34" box="[1228,1394,875,902]" pageId="15" pageNumber="16" value="2019-11-26">26 Nov. 2019</collectingDate>
</date>
;
<collectorName id="801D8A9FF415FFDF9A9FFC75FEDBFC1B" box="[191,279,911,937]" pageId="15" pageNumber="16">LabCol</collectorName>
leg.; “Fungo 53 \\ Em
<taxonomicName id="EAE894CAF415FFDF9800FC6AFD35FC1B" box="[544,761,911,937]" class="Agaricomycetes" family="Agaricaceae" genus="Leucocoprinus" kingdom="Fungi" order="Agaricales" pageId="15" pageNumber="16" phylum="Basidiomycota" rank="species" species="undetermined">
<emphasis id="1F9C335BF415FFDF9800FC6AFD02FC1B" box="[544,718,912,937]" italics="true" pageId="15" pageNumber="16">Leucocoprinus</emphasis>
sp.
</taxonomicName>
”;
<collectionCode id="4BF9778CF415FFDF9933FC75FC92FC1B" box="[787,862,911,937]" pageId="15" pageNumber="16">CELC</collectionCode>
</materialsCitation>
•
<materialsCitation id="9D80E514F415FFDF9956FC75FB07FC7E" ID-GBIF-Occurrence="3983014384" collectingDate="2019-11-20" collectionCode="CELC" collectorName="Recanto das Cigarras &amp; LabCol" country="Brazil" location="Vicosa" municipality="Mata da Biol." pageId="15" pageNumber="16" specimenCount="1" typeStatus="paratype">
<specimenCount id="3BEE24C0F415FFDF9956FC75FC06FC1B" box="[886,970,911,937]" pageId="15" pageNumber="16" type="generic">1 spec.</specimenCount>
;
<location id="2837B992F415FFDF99F6FC75FBE5FC1B" LSID="urn:lsid:plazi:treatment:A5415E5FF414FFC8985BF980FD6EFA25:2837B992F415FFDF99F6FC75FBE5FC1B" box="[982,1065,911,937]" country="Brazil" municipality="Mata da Biol." name="Vicosa" pageId="15" pageNumber="16">Viçosa</location>
,
<collectorName id="801D8A9FF415FFDF9E13FC75FAE3FC1B" box="[1075,1327,911,937]" pageId="15" pageNumber="16">Recanto das Cigarras</collectorName>
,
<collectingMunicipality id="CD337533F415FFDF9F1AFC75FED1FC7E" pageId="15" pageNumber="16">Mata da Biol.</collectingMunicipality>
;
<date id="5956C989F415FFDF9B09FC48FE07FC7E" box="[297,459,946,973]" pageId="15" pageNumber="16" value="2019-11-20">
<collectingDate id="49123061F415FFDF9B09FC48FE07FC7E" box="[297,459,946,973]" pageId="15" pageNumber="16" value="2019-11-20">20 Nov. 2019</collectingDate>
</date>
;
<collectorName id="801D8A9FF415FFDF9BF7FC48FDE3FC7E" box="[471,559,946,972]" pageId="15" pageNumber="16">LabCol</collectorName>
leg; “Fungo 19 \\ Em
<taxonomicName id="EAE894CAF415FFDF9916FC49FB93FC7E" box="[822,1119,947,972]" class="Agaricomycetes" family="Agaricaceae" genus="Leucocoprinus" kingdom="Fungi" order="Agaricales" pageId="15" pageNumber="16" phylum="Basidiomycota" rank="species" species="cepistipes">
<emphasis id="1F9C335BF415FFDF9916FC49FB93FC7E" box="[822,1119,947,972]" italics="true" pageId="15" pageNumber="16">Leucocoprinus cepistipes</emphasis>
</taxonomicName>
”;
<collectionCode id="4BF9778CF415FFDF9E5CFC48FB07FC7E" box="[1148,1227,946,972]" pageId="15" pageNumber="16">CELC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="2D57EF49F415FFDF9A9DFC09FEB2FB83" blockId="15.[189,1398,1011,1073]" pageId="15" pageNumber="16">
All
<typeStatus id="F25351EBF415FFDF9ACFFC0EFE93FBBC" box="[239,351,1012,1038]" pageId="15" pageNumber="16" type="paratype">paratypes</typeStatus>
additionally labelled “
<taxonomicName id="EAE894CAF415FFDF985DFC0EFB08FBBF" authority="von Groll &amp; Lopes-Andrade" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[637,1220,1011,1038]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="achardi">
<emphasis id="1F9C335BF415FFDF985DFC0EFC95FBBF" box="[637,857,1011,1038]" italics="true" pageId="15" pageNumber="16">Cyparium achardi</emphasis>
von Groll &amp; Lopes-Andrade
</taxonomicName>
<typeStatus id="F25351EBF415FFDF9EF1FC0EFABAFBBC" box="[1233,1398,1012,1038]" pageId="15" pageNumber="16" type="paratype">PARATYPUS</typeStatus>
[yellow paper]”.
</paragraph>
</subSubSection>
<subSubSection id="65F2BCC2F415FFC89A9DFBA2FB9AFCB8" lastPageId="24" lastPageNumber="25" pageId="15" pageNumber="16" type="description">
<paragraph id="2D57EF49F415FFDF9A9DFBA2FE82FBC0" blockId="15.[189,334,1112,1138]" box="[189,334,1112,1138]" pageId="15" pageNumber="16">
<heading id="761F5825F415FFDF9A9DFBA2FE82FBC0" bold="true" box="[189,334,1112,1138]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="1F9C335BF415FFDF9A9DFBA2FE82FBC0" bold="true" box="[189,334,1112,1138]" pageId="15" pageNumber="16">Description</emphasis>
</heading>
</paragraph>
<paragraph id="2D57EF49F415FFDF9A9DFB7DFEFCFB76" blockId="15.[189,1399,1159,1220]" pageId="15" pageNumber="16">
MEASUREMENTS (
<typeStatus id="F25351EBF415FFDF9BA2FB7DFE21FB13" box="[386,493,1159,1185]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
, in mm). TL 3.12, PL 1.31, PA 0.80, PB 1.90, EW 1.20, EL 2.17, IS 0.18, HW 0.80.
</paragraph>
<paragraph id="2D57EF49F415FFDF9A9DFB11FC6BFA20" blockId="15.[188,1399,1259,1427]" pageId="15" pageNumber="16">
COLORATION. Black; iridescent (
<figureCitation id="B5D3F3CCF415FFDF9809FB11FD66FAB7" box="[553,682,1259,1285]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="15" pageNumber="16">Fig. 7A–C</figureCitation>
). Clypeus yellowish-brown; mouth parts yellowish (
<figureCitation id="B5D3F3CCF415FFDF9F2BFB11FAA4FAB7" box="[1291,1384,1259,1285]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="15" pageNumber="16">Fig. 7D</figureCitation>
); antennomeres I–VI and apical half of XI yellow; VII–X dark yellow (
<figureCitation id="B5D3F3CCF415FFDF9E61FAF4FB0BFA9B" box="[1089,1223,1294,1321]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="15" pageNumber="16">Fig. 7D–F</figureCitation>
). Hypomeron, epipleuron, and metepimeron brown to black, lighter laterally (
<figureCitation id="B5D3F3CCF415FFDF99BDFAC8FC36FAFE" box="[925,1018,1330,1356]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="15" pageNumber="16">Fig. 7B</figureCitation>
). Thorax and abdomen brown to black, lighter laterally (
<figureCitation id="B5D3F3CCF415FFDF9BF0FAAFFDE1FADD" box="[464,557,1365,1391]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="15" pageNumber="16">Fig. 7B</figureCitation>
). Coxae and femora light to chestnut brown, lighter posteriorly in some specimens; tibiae light to dark brown; tarsi yellow (
<figureCitation id="B5D3F3CCF415FFDF993AFA82FC56FA20" box="[794,922,1400,1426]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="15" pageNumber="16">Fig. 7B–C</figureCitation>
).
</paragraph>
<paragraph id="2D57EF49F415FFDF9A9DFA40FB39F95F" blockId="15.[189,1401,1465,1773]" pageId="15" pageNumber="16">
HEAD. Punctation dense, fine (
<figureCitation id="B5D3F3CCF415FFDF980EFA43FD5CFA66" box="[558,656,1465,1492]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="15" pageNumber="16">Fig. 7D</figureCitation>
). Eyes tapered apically (
<figureCitation id="B5D3F3CCF415FFDF99E0FA43FBEEFA66" box="[960,1058,1465,1492]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="15" pageNumber="16">Fig. 7D</figureCitation>
). Labrum transverse; lateral areas rounded, smoothly delimited apically; central margin slightly curved; sclerotized portion rounded with a row of short setae; lateral setae slightly exceeding margins of labrum; slightly porose centrally (
<figureCitation id="B5D3F3CCF415FFDF9AE7F9D9FEE9F98F" box="[199,293,1571,1597]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="15" pageNumber="16">Fig. 7G</figureCitation>
). Mandibles curved, subapical serrations on left mandible conspicuous (
<figureCitation id="B5D3F3CCF415FFDF9EA0F9D9FB35F98F" box="[1152,1273,1571,1597]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="15" pageNumber="16">Fig. 7H–I</figureCitation>
). Maxillae with palpomere II thickened; galea and lacinia densely pubescent, narrow (
<figureCitation id="B5D3F3CCF415FFDF9E0EF9BCFB4CF9D2" box="[1070,1152,1606,1632]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="15" pageNumber="16">Fig. 7J</figureCitation>
). Mentum with sides distinctly angulated, curved apically (
<figureCitation id="B5D3F3CCF415FFDF985EF993FD12F936" box="[638,734,1641,1668]" captionStart="Fig" captionStartId="18.[189,232,1831,1857]" captionTargetBox="[192,1393,273,1749]" captionTargetId="figure-17@18.[189,1399,265,1759]" captionTargetPageId="18" captionText="Fig. 8. Cyparium achardi sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D. Holotype, ♂ (CELC), pronotum, dorsal view. E–G. Paratype, ♂ (CELC), prothorax. E. Frontal view. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: ns = notosternal suture; pr = profurca; st = sclerotized plate. Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033416" httpUri="https://zenodo.org/record/7033416/files/figure.png" pageId="15" pageNumber="16">Fig. 8A</figureCitation>
). Setae of labial palpomere II exceeding palpomere III; palpomere III thick, with long apical setae (
<figureCitation id="B5D3F3CCF415FFDF98EBF977FCE7F915" box="[715,811,1677,1703]" captionStart="Fig" captionStartId="18.[189,232,1831,1857]" captionTargetBox="[192,1393,273,1749]" captionTargetId="figure-17@18.[189,1399,265,1759]" captionTargetPageId="18" captionText="Fig. 8. Cyparium achardi sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D. Holotype, ♂ (CELC), pronotum, dorsal view. E–G. Paratype, ♂ (CELC), prothorax. E. Frontal view. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: ns = notosternal suture; pr = profurca; st = sclerotized plate. Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033416" httpUri="https://zenodo.org/record/7033416/files/figure.png" pageId="15" pageNumber="16">Fig. 8A</figureCitation>
). Hypopharynx with sclerotized plate cup-shaped (
<figureCitation id="B5D3F3CCF415FFDF9AE7F94AFE80F978" box="[199,332,1712,1738]" captionStart="Fig" captionStartId="18.[189,232,1831,1857]" captionTargetBox="[192,1393,273,1749]" captionTargetId="figure-17@18.[189,1399,265,1759]" captionTargetPageId="18" captionText="Fig. 8. Cyparium achardi sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D. Holotype, ♂ (CELC), pronotum, dorsal view. E–G. Paratype, ♂ (CELC), prothorax. E. Frontal view. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: ns = notosternal suture; pr = profurca; st = sclerotized plate. Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033416" httpUri="https://zenodo.org/record/7033416/files/figure.png" pageId="15" pageNumber="16">Fig. 8A–B</figureCitation>
). Postgena with strigulate microsculpture; gular pores sparse; gula small and rounded at anterior region (
<figureCitation id="B5D3F3CCF415FFDF9B5BF929FE1BF95F" box="[379,471,1747,1773]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="15" pageNumber="16">Fig. 7C</figureCitation>
). Antennae variable in size between males and females (
<figureCitation id="B5D3F3CCF415FFDF9E4DF929FB24F95F" box="[1133,1256,1747,1773]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="15" pageNumber="16">Fig. 7E–F</figureCitation>
).
</paragraph>
<paragraph id="2D57EF49F415FFDF9A9DF8EFFA92F82A" blockId="15.[187,1398,1812,1945]" pageId="15" pageNumber="16">
PROTHORAX. Pronotum smooth, shining; punctation somewhat dense, fine; pubescence short, fine (
<figureCitation id="B5D3F3CCF415FFDF9AE6F8C2FE88F8E0" box="[198,324,1848,1874]" captionStart="Fig" captionStartId="18.[189,232,1831,1857]" captionTargetBox="[192,1393,273,1749]" captionTargetId="figure-17@18.[189,1399,265,1759]" captionTargetPageId="18" captionText="Fig. 8. Cyparium achardi sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D. Holotype, ♂ (CELC), pronotum, dorsal view. E–G. Paratype, ♂ (CELC), prothorax. E. Frontal view. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: ns = notosternal suture; pr = profurca; st = sclerotized plate. Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033416" httpUri="https://zenodo.org/record/7033416/files/figure.png" pageId="15" pageNumber="16">Fig. 8D–E</figureCitation>
); somewhat squared, lateral areas curved, forming an obtuse angle at lateral areas of posterior margin (
<figureCitation id="B5D3F3CCF415FFDF9B3EF8A1FEB6F8C7" box="[286,378,1883,1909]" captionStart="Fig" captionStartId="18.[189,232,1831,1857]" captionTargetBox="[192,1393,273,1749]" captionTargetId="figure-17@18.[189,1399,265,1759]" captionTargetPageId="18" captionText="Fig. 8. Cyparium achardi sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D. Holotype, ♂ (CELC), pronotum, dorsal view. E–G. Paratype, ♂ (CELC), prothorax. E. Frontal view. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: ns = notosternal suture; pr = profurca; st = sclerotized plate. Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033416" httpUri="https://zenodo.org/record/7033416/files/figure.png" pageId="15" pageNumber="16">Fig. 8E</figureCitation>
). Hypomeron smooth (
<figureCitation id="B5D3F3CCF415FFDF98ABF8A1FD2BF8C7" box="[651,743,1883,1909]" captionStart="Fig" captionStartId="18.[189,232,1831,1857]" captionTargetBox="[192,1393,273,1749]" captionTargetId="figure-17@18.[189,1399,265,1759]" captionTargetPageId="18" captionText="Fig. 8. Cyparium achardi sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D. Holotype, ♂ (CELC), pronotum, dorsal view. E–G. Paratype, ♂ (CELC), prothorax. E. Frontal view. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: ns = notosternal suture; pr = profurca; st = sclerotized plate. Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033416" httpUri="https://zenodo.org/record/7033416/files/figure.png" pageId="15" pageNumber="16">Fig. 8F</figureCitation>
). Notosternal suture straight, inward-directed (
<figureCitation id="B5D3F3CCF415FFDF9F2FF8A1FAA5F8C7" box="[1295,1385,1883,1909]" captionStart="Fig" captionStartId="18.[189,232,1831,1857]" captionTargetBox="[192,1393,273,1749]" captionTargetId="figure-17@18.[189,1399,265,1759]" captionTargetPageId="18" captionText="Fig. 8. Cyparium achardi sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D. Holotype, ♂ (CELC), pronotum, dorsal view. E–G. Paratype, ♂ (CELC), prothorax. E. Frontal view. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: ns = notosternal suture; pr = profurca; st = sclerotized plate. Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033416" httpUri="https://zenodo.org/record/7033416/files/figure.png" pageId="15" pageNumber="16">Fig. 8F</figureCitation>
). Profurca elongated, exceeding half of foramen (
<figureCitation id="B5D3F3CCF415FFDF98CFF884FC82F82A" box="[751,846,1918,1944]" captionStart="Fig" captionStartId="18.[189,232,1831,1857]" captionTargetBox="[192,1393,273,1749]" captionTargetId="figure-17@18.[189,1399,265,1759]" captionTargetPageId="18" captionText="Fig. 8. Cyparium achardi sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D. Holotype, ♂ (CELC), pronotum, dorsal view. E–G. Paratype, ♂ (CELC), prothorax. E. Frontal view. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: ns = notosternal suture; pr = profurca; st = sclerotized plate. Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033416" httpUri="https://zenodo.org/record/7033416/files/figure.png" pageId="15" pageNumber="16">Fig. 8G</figureCitation>
). Prosternal process acute apically (
<figureCitation id="B5D3F3CCF415FFDF9ED2F884FA9DF82A" box="[1266,1361,1918,1944]" captionStart="Fig" captionStartId="19.[189,232,1773,1799]" captionTargetBox="[189,1398,285,1696]" captionTargetId="figure-11@19.[189,1398,265,1700]" captionTargetPageId="19" captionText="Fig. 9. Cyparium achardi sp. nov., paratype, ♂ (CELC). A. Prothorax, lateral view. B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: al = alacrista; ap = anterior phragma; ip = intercoxal plate; ml = median line; mma = median membranous area; mp = mesoventral process; mvl = mesoventral line; mxl = mesocoxal line; pp = prosternal process; ps = prescutellar line; px = procoxal rest; sss = scutoscutellar suture. Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033418" httpUri="https://zenodo.org/record/7033418/files/figure.png" pageId="15" pageNumber="16">Fig. 9A</figureCitation>
).
</paragraph>
<caption id="7997BFC1F40AFFC09A9DF8ECFAB5F828" ID-DOI="http://doi.org/10.5281/zenodo.7033414" ID-Zenodo-Dep="7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="16" pageNumber="17" startId="16.[189,232,1814,1840]" targetBox="[189,1398,273,1759]" targetPageId="16">
<paragraph id="2D57EF49F40AFFC09A9DF8ECFAB5F828" blockId="16.[189,1401,1814,1947]" pageId="16" pageNumber="17">
<emphasis id="1F9C335BF40AFFC09A9DF8ECFEDDF883" bold="true" box="[189,273,1814,1841]" pageId="16" pageNumber="17">Fig. 7.</emphasis>
<taxonomicName id="EAE894CAF40AFFC09B3FF8EDFE30F882" authority="Groll &amp; Lopes-Andrade, 2022" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[287,508,1814,1841]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="achardi" status="sp. nov.">
<emphasis id="1F9C335BF40AFFC09B3FF8EDFE30F882" box="[287,508,1814,1841]" italics="true" pageId="16" pageNumber="17">Cyparium achardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AF8E20F40AFFC09829F8EDFDA6F883" box="[521,618,1815,1841]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="1F9C335BF40AFFC09857F8ECFD7CF882" bold="true" box="[631,688,1814,1840]" pageId="16" pageNumber="17">A–D</emphasis>
. Holotype, ♂ (CELC).
<emphasis id="1F9C335BF40AFFC099FDF8ECFC3EF882" bold="true" box="[989,1010,1814,1840]" pageId="16" pageNumber="17">A</emphasis>
. Dorsal view.
<emphasis id="1F9C335BF40AFFC09E8EF8EDFB0EF883" bold="true" box="[1198,1218,1815,1841]" pageId="16" pageNumber="17">B</emphasis>
. Ventral view.
<emphasis id="1F9C335BF40AFFC09A9DF8C3FF1EF8E1" bold="true" box="[189,210,1849,1875]" pageId="16" pageNumber="17">C</emphasis>
. Lateral view.
<emphasis id="1F9C335BF40AFFC09BAAF8C0FE53F8E6" bold="true" box="[394,415,1850,1876]" pageId="16" pageNumber="17">D</emphasis>
. Frontal view.
<emphasis id="1F9C335BF40AFFC09878F8C0FD47F8E6" bold="true" box="[600,651,1850,1876]" pageId="16" pageNumber="17">E–F</emphasis>
. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F).
<emphasis id="1F9C335BF40AFFC09AD2F8A7FEEAF8C5" bold="true" box="[242,294,1885,1911]" pageId="16" pageNumber="17">G–J</emphasis>
. Paratype, ♂ (CELC).
<emphasis id="1F9C335BF40AFFC0981DF8A7FD98F8C5" bold="true" box="[573,596,1885,1911]" pageId="16" pageNumber="17">G</emphasis>
. Labrum.
<emphasis id="1F9C335BF40AFFC098F4F8A7FCCEF8C5" bold="true" box="[724,770,1885,1911]" pageId="16" pageNumber="17">H–I</emphasis>
. Mandible.
<emphasis id="1F9C335BF40AFFC099B7F8A7FC60F8C5" bold="true" box="[919,940,1885,1911]" pageId="16" pageNumber="17">H</emphasis>
. Left.
<emphasis id="1F9C335BF40AFFC09E20F8A7FBC5F8C5" bold="true" box="[1024,1033,1885,1911]" pageId="16" pageNumber="17">I</emphasis>
. Right.
<emphasis id="1F9C335BF40AFFC09E4EF8A7FBB7F8C5" bold="true" box="[1134,1147,1885,1911]" pageId="16" pageNumber="17">J</emphasis>
. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1.
</paragraph>
</caption>
<paragraph id="2D57EF49F40BFFC19A9DFEF3FAA6FE02" blockId="17.[189,1399,265,432]" pageId="17" pageNumber="18">
MESOTHORAX. Mesonotum with prescutellar suture (= scutellar lines,
<bibRefCitation id="497992B8F40BFFC19E21FEF3FAD9FE96" author="Leschen R. A. B. &amp; Lobl I." box="[1025,1301,265,292]" pageId="17" pageNumber="18" pagination="1 - 63" refId="ref32588" refString="Leschen R. A. B. &amp; Lobl I. 2005. Phylogeny and classification of Scaphisomatini Staphylinidae: Scaphidiinae with notes on mycophagy, termitophily, and functional morphology. Coleopterists Society Monographs 3: 1 - 63. https: // doi. org / 10.1649 / 0010 - 065 X (2005) 059 [0001: PACOSS] 2.0. CO; 2" type="journal article" year="2005">Leschen &amp; Löbl, 2005</bibRefCitation>
) almost straight (
<figureCitation id="B5D3F3CCF40BFFC19B09FED6FE45FEF4" box="[297,393,300,326]" captionStart="Fig" captionStartId="19.[189,232,1773,1799]" captionTargetBox="[189,1398,285,1696]" captionTargetId="figure-11@19.[189,1398,265,1700]" captionTargetPageId="19" captionText="Fig. 9. Cyparium achardi sp. nov., paratype, ♂ (CELC). A. Prothorax, lateral view. B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: al = alacrista; ap = anterior phragma; ip = intercoxal plate; ml = median line; mma = median membranous area; mp = mesoventral process; mvl = mesoventral line; mxl = mesocoxal line; pp = prosternal process; ps = prescutellar line; px = procoxal rest; sss = scutoscutellar suture. Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033418" httpUri="https://zenodo.org/record/7033418/files/figure.png" pageId="17" pageNumber="18">Fig. 9B</figureCitation>
). Scutellum rounded posteriorly (
<figureCitation id="B5D3F3CCF40BFFC19901FED6FC4EFEF4" box="[801,898,300,326]" captionStart="Fig" captionStartId="19.[189,232,1773,1799]" captionTargetBox="[189,1398,285,1696]" captionTargetId="figure-11@19.[189,1398,265,1700]" captionTargetPageId="19" captionText="Fig. 9. Cyparium achardi sp. nov., paratype, ♂ (CELC). A. Prothorax, lateral view. B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: al = alacrista; ap = anterior phragma; ip = intercoxal plate; ml = median line; mma = median membranous area; mp = mesoventral process; mvl = mesoventral line; mxl = mesocoxal line; pp = prosternal process; ps = prescutellar line; px = procoxal rest; sss = scutoscutellar suture. Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033418" httpUri="https://zenodo.org/record/7033418/files/figure.png" pageId="17" pageNumber="18">Fig. 9B</figureCitation>
). Anterior phragma wide, slightly curved (
<figureCitation id="B5D3F3CCF40BFFC19AE6FEB5FEEAFEDB" box="[198,294,335,361]" captionStart="Fig" captionStartId="19.[189,232,1773,1799]" captionTargetBox="[189,1398,285,1696]" captionTargetId="figure-11@19.[189,1398,265,1700]" captionTargetPageId="19" captionText="Fig. 9. Cyparium achardi sp. nov., paratype, ♂ (CELC). A. Prothorax, lateral view. B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: al = alacrista; ap = anterior phragma; ip = intercoxal plate; ml = median line; mma = median membranous area; mp = mesoventral process; mvl = mesoventral line; mxl = mesocoxal line; pp = prosternal process; ps = prescutellar line; px = procoxal rest; sss = scutoscutellar suture. Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033418" httpUri="https://zenodo.org/record/7033418/files/figure.png" pageId="17" pageNumber="18">Fig. 9C</figureCitation>
). Mesanepisternum smooth (
<figureCitation id="B5D3F3CCF40BFFC198A4FEB5FD2AFEDB" box="[644,742,335,361]" captionStart="Fig" captionStartId="19.[189,232,1773,1799]" captionTargetBox="[189,1398,285,1696]" captionTargetId="figure-11@19.[189,1398,265,1700]" captionTargetPageId="19" captionText="Fig. 9. Cyparium achardi sp. nov., paratype, ♂ (CELC). A. Prothorax, lateral view. B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: al = alacrista; ap = anterior phragma; ip = intercoxal plate; ml = median line; mma = median membranous area; mp = mesoventral process; mvl = mesoventral line; mxl = mesocoxal line; pp = prosternal process; ps = prescutellar line; px = procoxal rest; sss = scutoscutellar suture. Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033418" httpUri="https://zenodo.org/record/7033418/files/figure.png" pageId="17" pageNumber="18">Fig. 9D</figureCitation>
). Procoxal rest rounded, curved posteriorly (
<figureCitation id="B5D3F3CCF40BFFC19F27FEB5FAA5FEDB" box="[1287,1385,335,361]" captionStart="Fig" captionStartId="19.[189,232,1773,1799]" captionTargetBox="[189,1398,285,1696]" captionTargetId="figure-11@19.[189,1398,265,1700]" captionTargetPageId="19" captionText="Fig. 9. Cyparium achardi sp. nov., paratype, ♂ (CELC). A. Prothorax, lateral view. B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: al = alacrista; ap = anterior phragma; ip = intercoxal plate; ml = median line; mma = median membranous area; mp = mesoventral process; mvl = mesoventral line; mxl = mesocoxal line; pp = prosternal process; ps = prescutellar line; px = procoxal rest; sss = scutoscutellar suture. Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033418" httpUri="https://zenodo.org/record/7033418/files/figure.png" pageId="17" pageNumber="18">Fig. 9D</figureCitation>
). Mesoventral line rounded; median line moderately curved; area between median line and mesocoxal line longer laterally (
<figureCitation id="B5D3F3CCF40BFFC19B93FE6CFDDDFE02" box="[435,529,406,432]" captionStart="Fig" captionStartId="19.[189,232,1773,1799]" captionTargetBox="[189,1398,285,1696]" captionTargetId="figure-11@19.[189,1398,265,1700]" captionTargetPageId="19" captionText="Fig. 9. Cyparium achardi sp. nov., paratype, ♂ (CELC). A. Prothorax, lateral view. B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: al = alacrista; ap = anterior phragma; ip = intercoxal plate; ml = median line; mma = median membranous area; mp = mesoventral process; mvl = mesoventral line; mxl = mesocoxal line; pp = prosternal process; ps = prescutellar line; px = procoxal rest; sss = scutoscutellar suture. Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033418" httpUri="https://zenodo.org/record/7033418/files/figure.png" pageId="17" pageNumber="18">Fig. 9D</figureCitation>
). Mesoventral process large and straight at base in lateral view (
<figureCitation id="B5D3F3CCF40BFFC19F21FE6CFA91FE02" box="[1281,1373,406,432]" captionStart="Fig" captionStartId="19.[189,232,1773,1799]" captionTargetBox="[189,1398,285,1696]" captionTargetId="figure-11@19.[189,1398,265,1700]" captionTargetPageId="19" captionText="Fig. 9. Cyparium achardi sp. nov., paratype, ♂ (CELC). A. Prothorax, lateral view. B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: al = alacrista; ap = anterior phragma; ip = intercoxal plate; ml = median line; mma = median membranous area; mp = mesoventral process; mvl = mesoventral line; mxl = mesocoxal line; pp = prosternal process; ps = prescutellar line; px = procoxal rest; sss = scutoscutellar suture. Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033418" httpUri="https://zenodo.org/record/7033418/files/figure.png" pageId="17" pageNumber="18">Fig. 9E</figureCitation>
).
</paragraph>
<paragraph id="2D57EF49F40BFFC19A9DFE22FDA5FD5A" blockId="17.[189,1400,471,744]" pageId="17" pageNumber="18">
METATHORAX. Metanotum with wide alacrista, rounded and somewhat turned to the posterior end; scutoscutellar suture slightly straight; median membranous area wide and very short (
<figureCitation id="B5D3F3CCF40BFFC19F27FE00FAA5FDA7" box="[1287,1385,506,533]" captionStart="Fig" captionStartId="19.[189,232,1773,1799]" captionTargetBox="[189,1398,285,1696]" captionTargetId="figure-11@19.[189,1398,265,1700]" captionTargetPageId="19" captionText="Fig. 9. Cyparium achardi sp. nov., paratype, ♂ (CELC). A. Prothorax, lateral view. B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: al = alacrista; ap = anterior phragma; ip = intercoxal plate; ml = median line; mma = median membranous area; mp = mesoventral process; mvl = mesoventral line; mxl = mesocoxal line; pp = prosternal process; ps = prescutellar line; px = procoxal rest; sss = scutoscutellar suture. Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033418" httpUri="https://zenodo.org/record/7033418/files/figure.png" pageId="17" pageNumber="18">Fig. 9F</figureCitation>
). Metaventrite smooth; punctation sparse, fine; some specimens with faint lateral imbricate microsculpture (
<figureCitation id="B5D3F3CCF40BFFC19AE6FDBBFEEEFDE9" box="[198,290,577,603]" captionStart="Fig" captionStartId="19.[189,232,1773,1799]" captionTargetBox="[189,1398,285,1696]" captionTargetId="figure-11@19.[189,1398,265,1700]" captionTargetPageId="19" captionText="Fig. 9. Cyparium achardi sp. nov., paratype, ♂ (CELC). A. Prothorax, lateral view. B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: al = alacrista; ap = anterior phragma; ip = intercoxal plate; ml = median line; mma = median membranous area; mp = mesoventral process; mvl = mesoventral line; mxl = mesocoxal line; pp = prosternal process; ps = prescutellar line; px = procoxal rest; sss = scutoscutellar suture. Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033418" httpUri="https://zenodo.org/record/7033418/files/figure.png" pageId="17" pageNumber="18">Fig. 9D</figureCitation>
). Mesocoxal line forming an open angle next to each coxal cavity and finely punctate under coxal cavity (
<figureCitation id="B5D3F3CCF40BFFC19B36FD9EFEBAFDCC" box="[278,374,612,638]" captionStart="Fig" captionStartId="19.[189,232,1773,1799]" captionTargetBox="[189,1398,285,1696]" captionTargetId="figure-11@19.[189,1398,265,1700]" captionTargetPageId="19" captionText="Fig. 9. Cyparium achardi sp. nov., paratype, ♂ (CELC). A. Prothorax, lateral view. B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: al = alacrista; ap = anterior phragma; ip = intercoxal plate; ml = median line; mma = median membranous area; mp = mesoventral process; mvl = mesoventral line; mxl = mesocoxal line; pp = prosternal process; ps = prescutellar line; px = procoxal rest; sss = scutoscutellar suture. Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033418" httpUri="https://zenodo.org/record/7033418/files/figure.png" pageId="17" pageNumber="18">Fig. 9D</figureCitation>
). Metanepisternum and metepimeron (
<figureCitation id="B5D3F3CCF40BFFC19962FD9EFC0AFDCC" box="[834,966,612,638]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="17" pageNumber="18">Figs 7B–C</figureCitation>
,
<figureCitation id="B5D3F3CCF40BFFC199F4FD9EFC35FDCC" box="[980,1017,612,638]" captionStart="Fig" captionStartId="19.[189,232,1773,1799]" captionTargetBox="[189,1398,285,1696]" captionTargetId="figure-11@19.[189,1398,265,1700]" captionTargetPageId="19" captionText="Fig. 9. Cyparium achardi sp. nov., paratype, ♂ (CELC). A. Prothorax, lateral view. B–C. Scutellar plate. B. Dorsal view. C. Apically slanted view. D–E. Meso- and metathorax. D. Ventral view. E. Lateral view. F. Metanotum. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: al = alacrista; ap = anterior phragma; ip = intercoxal plate; ml = median line; mma = median membranous area; mp = mesoventral process; mvl = mesoventral line; mxl = mesocoxal line; pp = prosternal process; ps = prescutellar line; px = procoxal rest; sss = scutoscutellar suture. Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033418" httpUri="https://zenodo.org/record/7033418/files/figure.png" pageId="17" pageNumber="18">9D</figureCitation>
) with imbricate microsculpture, but almost inconspicuous in some specimens. Intercoxal plates smooth (
<figureCitation id="B5D3F3CCF40BFFC19E2DFD7DFBA5FD13" box="[1037,1129,647,673]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="17" pageNumber="18">Fig. 7B</figureCitation>
). Metendosternite with straight furcal arms; ‘stalk ridge’ not exceeding half of stalk length (
<figureCitation id="B5D3F3CCF40BFFC19E23FD50FBBFFD77" box="[1027,1139,682,709]" captionStart="Fig" captionStartId="20.[189,232,1771,1797]" captionTargetBox="[189,1392,273,1719]" captionTargetId="figure-16@20.[189,1398,265,1727]" captionTargetPageId="20" captionText="Fig. 10. Cyparium achardi sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: CAS = cubitoanal strut; fa = furcal arms; MP 1+2 = medial field; MSP = medial spur; RA 4 = radius anterior; RP 3+4 = radius posterior; sr = ‘stalk ridge’; vlf = ventral longitudinal flange. Scale bars:A–D, F–H = 0.2 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033420" httpUri="https://zenodo.org/record/7033420/files/figure.png" pageId="17" pageNumber="18">Fig. 10A</figureCitation>
); ventral longitudinal flange flat in lateral view (
<figureCitation id="B5D3F3CCF40BFFC19BD1FD34FD90FD5A" box="[497,604,718,744]" captionStart="Fig" captionStartId="20.[189,232,1771,1797]" captionTargetBox="[189,1392,273,1719]" captionTargetId="figure-16@20.[189,1398,265,1727]" captionTargetPageId="20" captionText="Fig. 10. Cyparium achardi sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: CAS = cubitoanal strut; fa = furcal arms; MP 1+2 = medial field; MSP = medial spur; RA 4 = radius anterior; RP 3+4 = radius posterior; sr = ‘stalk ridge’; vlf = ventral longitudinal flange. Scale bars:A–D, F–H = 0.2 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033420" httpUri="https://zenodo.org/record/7033420/files/figure.png" pageId="17" pageNumber="18">Fig. 10B</figureCitation>
).
</paragraph>
<paragraph id="2D57EF49F40BFFC19A9DFCEAFCD2FC6B" blockId="17.[189,1400,783,985]" pageId="17" pageNumber="18">
WINGS. Elytra slightly longer than wide, partially covering tergite VI (
<figureCitation id="B5D3F3CCF40BFFC199DFFCF5FB93FC9B" box="[1023,1119,783,809]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="17" pageNumber="18">Fig. 7A</figureCitation>
); basal and lateral lines punctate (
<figureCitation id="B5D3F3CCF40BFFC19B13FCC8FE5EFCFE" box="[307,402,818,844]" captionStart="Fig" captionStartId="18.[189,232,1831,1857]" captionTargetBox="[192,1393,273,1749]" captionTargetId="figure-17@18.[189,1399,265,1759]" captionTargetPageId="18" captionText="Fig. 8. Cyparium achardi sp. nov. A–C. Paratype, ♂ (CELC). A. Labium. B. Hypopharynx. C. Head, ventral view. D. Holotype, ♂ (CELC), pronotum, dorsal view. E–G. Paratype, ♂ (CELC), prothorax. E. Frontal view. F. Ventral view. G. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: ns = notosternal suture; pr = profurca; st = sclerotized plate. Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033416" httpUri="https://zenodo.org/record/7033416/files/figure.png" pageId="17" pageNumber="18">Fig. 8D</figureCitation>
); sutural line dashed; adsutural area with a row of short setae (
<figureCitation id="B5D3F3CCF40BFFC19E5DFCC8FB11FCFE" box="[1149,1245,818,844]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="17" pageNumber="18">Fig. 7A</figureCitation>
); six rows of coarse punctures (not including sutural line), but row 5 and 6 somewhat intermixed (
<figureCitation id="B5D3F3CCF40BFFC19EE1FCAFFAEAFCC2" box="[1217,1318,853,880]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="17" pageNumber="18">Figs 7A</figureCitation>
,
<figureCitation id="B5D3F3CCF40BFFC19F15FCACFAA6FCC2" box="[1333,1386,854,880]" captionStart="Fig" captionStartId="20.[189,232,1771,1797]" captionTargetBox="[189,1392,273,1719]" captionTargetId="figure-16@20.[189,1398,265,1727]" captionTargetPageId="20" captionText="Fig. 10. Cyparium achardi sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: CAS = cubitoanal strut; fa = furcal arms; MP 1+2 = medial field; MSP = medial spur; RA 4 = radius anterior; RP 3+4 = radius posterior; sr = ‘stalk ridge’; vlf = ventral longitudinal flange. Scale bars:A–D, F–H = 0.2 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033420" httpUri="https://zenodo.org/record/7033420/files/figure.png" pageId="17" pageNumber="18">10C</figureCitation>
); apical coarse punctation moderately sparse; apical serrations moderately large, well visible (
<figureCitation id="B5D3F3CCF40BFFC19EDBFC83FAA5FC21" box="[1275,1385,889,915]" captionStart="Fig" captionStartId="20.[189,232,1771,1797]" captionTargetBox="[189,1392,273,1719]" captionTargetId="figure-16@20.[189,1398,265,1727]" captionTargetPageId="20" captionText="Fig. 10. Cyparium achardi sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: CAS = cubitoanal strut; fa = furcal arms; MP 1+2 = medial field; MSP = medial spur; RA 4 = radius anterior; RP 3+4 = radius posterior; sr = ‘stalk ridge’; vlf = ventral longitudinal flange. Scale bars:A–D, F–H = 0.2 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033420" httpUri="https://zenodo.org/record/7033420/files/figure.png" pageId="17" pageNumber="18">Fig. 10D</figureCitation>
); pubescence short, fine, denser on disc. Epipleuron with a row of punctures (poorly marked in some specimens). Hind wings fully developed (
<figureCitation id="B5D3F3CCF40BFFC19886FC45FCDDFC6B" box="[678,785,959,985]" captionStart="Fig" captionStartId="20.[189,232,1771,1797]" captionTargetBox="[189,1392,273,1719]" captionTargetId="figure-16@20.[189,1398,265,1727]" captionTargetPageId="20" captionText="Fig. 10. Cyparium achardi sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: CAS = cubitoanal strut; fa = furcal arms; MP 1+2 = medial field; MSP = medial spur; RA 4 = radius anterior; RP 3+4 = radius posterior; sr = ‘stalk ridge’; vlf = ventral longitudinal flange. Scale bars:A–D, F–H = 0.2 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033420" httpUri="https://zenodo.org/record/7033420/files/figure.png" pageId="17" pageNumber="18">Fig. 10E</figureCitation>
).
</paragraph>
<paragraph id="2D57EF49F40BFFC19A9DFBFBFE08FB36" blockId="17.[189,1399,1024,1156]" pageId="17" pageNumber="18">
LEGS. Pro-, meso- and metacoxae, and femora with strigulate microsculpture. Pro- and mesofemora somewhat fusiform (
<figureCitation id="B5D3F3CCF40BFFC19B8FFBDEFDF0FB8C" box="[431,572,1060,1086]" captionStart="Fig" captionStartId="20.[189,232,1771,1797]" captionTargetBox="[189,1392,273,1719]" captionTargetId="figure-16@20.[189,1398,265,1727]" captionTargetPageId="20" captionText="Fig. 10. Cyparium achardi sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: CAS = cubitoanal strut; fa = furcal arms; MP 1+2 = medial field; MSP = medial spur; RA 4 = radius anterior; RP 3+4 = radius posterior; sr = ‘stalk ridge’; vlf = ventral longitudinal flange. Scale bars:A–D, F–H = 0.2 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033420" httpUri="https://zenodo.org/record/7033420/files/figure.png" pageId="17" pageNumber="18">Fig. 10F–G</figureCitation>
); punctation moderately sparse and coarse. Metafemora straight, finely punctate (
<figureCitation id="B5D3F3CCF40BFFC19B12FBBDFE6EFBD3" box="[306,418,1095,1121]" captionStart="Fig" captionStartId="20.[189,232,1771,1797]" captionTargetBox="[189,1392,273,1719]" captionTargetId="figure-16@20.[189,1398,265,1727]" captionTargetPageId="20" captionText="Fig. 10. Cyparium achardi sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: CAS = cubitoanal strut; fa = furcal arms; MP 1+2 = medial field; MSP = medial spur; RA 4 = radius anterior; RP 3+4 = radius posterior; sr = ‘stalk ridge’; vlf = ventral longitudinal flange. Scale bars:A–D, F–H = 0.2 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033420" httpUri="https://zenodo.org/record/7033420/files/figure.png" pageId="17" pageNumber="18">Fig. 10H</figureCitation>
). Mesotibiae densely spinose, spines fine (
<figureCitation id="B5D3F3CCF40BFFC19981FBBDFBDCFBD3" box="[929,1040,1095,1121]" captionStart="Fig" captionStartId="20.[189,232,1771,1797]" captionTargetBox="[189,1392,273,1719]" captionTargetId="figure-16@20.[189,1398,265,1727]" captionTargetPageId="20" captionText="Fig. 10. Cyparium achardi sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: CAS = cubitoanal strut; fa = furcal arms; MP 1+2 = medial field; MSP = medial spur; RA 4 = radius anterior; RP 3+4 = radius posterior; sr = ‘stalk ridge’; vlf = ventral longitudinal flange. Scale bars:A–D, F–H = 0.2 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033420" httpUri="https://zenodo.org/record/7033420/files/figure.png" pageId="17" pageNumber="18">Fig. 10G</figureCitation>
). Metatibiae sparsely spinose, spines fine (
<figureCitation id="B5D3F3CCF40BFFC19B6AFB90FE7BFB36" box="[330,439,1130,1156]" captionStart="Fig" captionStartId="20.[189,232,1771,1797]" captionTargetBox="[189,1392,273,1719]" captionTargetId="figure-16@20.[189,1398,265,1727]" captionTargetPageId="20" captionText="Fig. 10. Cyparium achardi sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: CAS = cubitoanal strut; fa = furcal arms; MP 1+2 = medial field; MSP = medial spur; RA 4 = radius anterior; RP 3+4 = radius posterior; sr = ‘stalk ridge’; vlf = ventral longitudinal flange. Scale bars:A–D, F–H = 0.2 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033420" httpUri="https://zenodo.org/record/7033420/files/figure.png" pageId="17" pageNumber="18">Fig. 10H</figureCitation>
).
</paragraph>
<paragraph id="2D57EF49F40BFFC19A9DFB56FD05FAE0" blockId="17.[187,1399,1195,1362]" pageId="17" pageNumber="18">
ABDOMEN. Tergites VI–VIII with imbricate microsculpture (
<figureCitation id="B5D3F3CCF40BFFC1994BFB51FC19FB74" box="[875,981,1195,1222]" captionStart="Fig" captionStartId="21.[189,232,1831,1857]" captionTargetBox="[189,1392,266,1756]" captionTargetId="figure-11@21.[189,1398,266,1760]" captionTargetPageId="21" captionText="Fig. 11. Cyparium achardi sp. nov. A, C. Holotype, ♂ (CELC). B, D–G. Paratype ♂ (CELC). A–B. Abdomen, dorsal view (A) and ventral view (B). C. Ventrite 1. D. Tergite VIII. E. Terminalia. F. Tergite IX. G. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: st = sternite; tg = tergite; vs = ventral strut. Scale bars: A–B = 0.4 mm; C–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033422" httpUri="https://zenodo.org/record/7033422/files/figure.png" pageId="17" pageNumber="18">Fig. 11A</figureCitation>
). Tergite VII trapezoidal; punctation dense, coarse; pubescence sparse, fine (
<figureCitation id="B5D3F3CCF40BFFC198A5FB35FD3EFB5B" box="[645,754,1231,1257]" captionStart="Fig" captionStartId="21.[189,232,1831,1857]" captionTargetBox="[189,1392,266,1756]" captionTargetId="figure-11@21.[189,1398,266,1760]" captionTargetPageId="21" captionText="Fig. 11. Cyparium achardi sp. nov. A, C. Holotype, ♂ (CELC). B, D–G. Paratype ♂ (CELC). A–B. Abdomen, dorsal view (A) and ventral view (B). C. Ventrite 1. D. Tergite VIII. E. Terminalia. F. Tergite IX. G. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: st = sternite; tg = tergite; vs = ventral strut. Scale bars: A–B = 0.4 mm; C–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033422" httpUri="https://zenodo.org/record/7033422/files/figure.png" pageId="17" pageNumber="18">Fig. 11A</figureCitation>
). Ventrite 1 sparsely and coarsely punctate; pubescence sparse, fine; strigulate microsculpture (
<figureCitation id="B5D3F3CCF40BFFC198B9FB08FCE7FABE" box="[665,811,1266,1292]" captionStart="Fig" captionStartId="21.[189,232,1831,1857]" captionTargetBox="[189,1392,266,1756]" captionTargetId="figure-11@21.[189,1398,266,1760]" captionTargetPageId="21" captionText="Fig. 11. Cyparium achardi sp. nov. A, C. Holotype, ♂ (CELC). B, D–G. Paratype ♂ (CELC). A–B. Abdomen, dorsal view (A) and ventral view (B). C. Ventrite 1. D. Tergite VIII. E. Terminalia. F. Tergite IX. G. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: st = sternite; tg = tergite; vs = ventral strut. Scale bars: A–B = 0.4 mm; C–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033422" httpUri="https://zenodo.org/record/7033422/files/figure.png" pageId="17" pageNumber="18">Fig. 11B–C</figureCitation>
). Metacoxal lines finely punctate. Ventrites 2–5 sparsely and finely punctate; pubescence sparse, fine; imbricate microsculpture anteriorly and with strigulate microsculpture middle-posteriorly.
</paragraph>
<paragraph id="2D57EF49F40BFFC19ACAFA80FEF9FA26" blockId="17.[187,1399,1402,1710]" box="[234,309,1402,1428]" pageId="17" pageNumber="18">
<emphasis id="1F9C335BF40BFFC19ACAFA80FEF9FA26" bold="true" box="[234,309,1402,1428]" pageId="17" pageNumber="18">Males</emphasis>
</paragraph>
<paragraph id="2D57EF49F40BFFC19A9DFA67FE92F91F" blockId="17.[187,1399,1402,1710]" pageId="17" pageNumber="18">
MEASUREMENTS (n = 1,
<typeStatus id="F25351EBF40BFFC19BCDFA67FD9BFA05" box="[493,599,1437,1463]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
; in mm). Antennomeres (length(width)): 0.18(0.06), 0.11(0.05), 0.10(0.04), 0.09(0.04), 0.10(0.05), 0.07(0.06), 0.09(0.10), 0.09(0.11), 0.10(0.12), 0.10(0.13), 0.17(0.14); (n = 5, including the
<typeStatus id="F25351EBF40BFFC19B8BFA19FDD9FA4F" box="[427,533,1507,1533]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
; in mm): TL 3.00–3.28 (mean = 3.14, standard deviation ± 0.10), PL 1.20– 1.31 (1.25 ± 0.04), PA 0.80–0.97 (0.89 ± 0.06), PB 1.72–2.07 (1.93 ± 0.13), SL 0.13–0.17 (0.14 ± 0.01), SW 0.15–0.20 (0.18 ± 0.02), EI 1.72–1.92 (1.81 ± 0.07), EL 2.07–2.32 (2.18 ± 0.09), EW 1.07–1.25 (1.19 ± 0.07), EH 0.70–0.80 (0.75 ± 0.05), HW 0.74–0.83 (0.78 ± 0.03), IS 0.15–0.24 (0.17 ± 0.03), WA 0.14–0.17 (0.16 ± 0.01), MC 0.72–0.92 (0.81 ± 0.08), MB 0.42–0.50 (0.46 ± 0.04), VL 0.62–0.70 (0.64 ± 0.03).
</paragraph>
<paragraph id="2D57EF49F40BFFC19A9DF92FFB87F857" blockId="17.[189,1399,1749,2022]" pageId="17" pageNumber="18">
Antennae almost reaching level of mesanepisternum, smaller than in females (
<figureCitation id="B5D3F3CCF40BFFC19EB0F92FFB3EF95D" box="[1168,1266,1749,1775]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="17" pageNumber="18">Fig. 7E</figureCitation>
). Pro- and mesotarsomeres I–III enlarged, with tenent setae (
<figureCitation id="B5D3F3CCF40BFFC1991DF902FC62F8A0" box="[829,942,1784,1810]" captionStart="Fig" captionStartId="20.[189,232,1771,1797]" captionTargetBox="[189,1392,273,1719]" captionTargetId="figure-16@20.[189,1398,265,1727]" captionTargetPageId="20" captionText="Fig. 10. Cyparium achardi sp. nov., paratype, ♂ (CELC). A–B. Metendosternite. A. Dorsal view. B. Lateral view. C–D. Elytron. C. Entire. D. Apex. E. Hind wing. F–H. Legs. F. Fore. G. Middle. H. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: CAS = cubitoanal strut; fa = furcal arms; MP 1+2 = medial field; MSP = medial spur; RA 4 = radius anterior; RP 3+4 = radius posterior; sr = ‘stalk ridge’; vlf = ventral longitudinal flange. Scale bars:A–D, F–H = 0.2 mm; E = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7033420" httpUri="https://zenodo.org/record/7033420/files/figure.png" pageId="17" pageNumber="18">Fig. 10F</figureCitation>
). Tergite VIII triangular; punctation dense, coarse, subglabrous (
<figureCitation id="B5D3F3CCF40BFFC1982AF8E1FDBBF887" box="[522,631,1819,1845]" captionStart="Fig" captionStartId="21.[189,232,1831,1857]" captionTargetBox="[189,1392,266,1756]" captionTargetId="figure-11@21.[189,1398,266,1760]" captionTargetPageId="21" captionText="Fig. 11. Cyparium achardi sp. nov. A, C. Holotype, ♂ (CELC). B, D–G. Paratype ♂ (CELC). A–B. Abdomen, dorsal view (A) and ventral view (B). C. Ventrite 1. D. Tergite VIII. E. Terminalia. F. Tergite IX. G. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: st = sternite; tg = tergite; vs = ventral strut. Scale bars: A–B = 0.4 mm; C–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033422" httpUri="https://zenodo.org/record/7033422/files/figure.png" pageId="17" pageNumber="18">Fig. 11D</figureCitation>
). Tergite IX with rectangular ventral struts (
<figureCitation id="B5D3F3CCF40BFFC19EA3F8E1FAC9F887" box="[1155,1285,1819,1845]" captionStart="Fig" captionStartId="21.[189,232,1831,1857]" captionTargetBox="[189,1392,266,1756]" captionTargetId="figure-11@21.[189,1398,266,1760]" captionTargetPageId="21" captionText="Fig. 11. Cyparium achardi sp. nov. A, C. Holotype, ♂ (CELC). B, D–G. Paratype ♂ (CELC). A–B. Abdomen, dorsal view (A) and ventral view (B). C. Ventrite 1. D. Tergite VIII. E. Terminalia. F. Tergite IX. G. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: st = sternite; tg = tergite; vs = ventral strut. Scale bars: A–B = 0.4 mm; C–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033422" httpUri="https://zenodo.org/record/7033422/files/figure.png" pageId="17" pageNumber="18">Fig. 11E,F</figureCitation>
). Sternite VIII rectangular, longish; finely punctate and with strigulate microsculpture (
<figureCitation id="B5D3F3CCF40BFFC19E4AF8C4FB15F8EA" box="[1130,1241,1854,1880]" captionStart="Fig" captionStartId="21.[189,232,1831,1857]" captionTargetBox="[189,1392,266,1756]" captionTargetId="figure-11@21.[189,1398,266,1760]" captionTargetPageId="21" captionText="Fig. 11. Cyparium achardi sp. nov. A, C. Holotype, ♂ (CELC). B, D–G. Paratype ♂ (CELC). A–B. Abdomen, dorsal view (A) and ventral view (B). C. Ventrite 1. D. Tergite VIII. E. Terminalia. F. Tergite IX. G. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: st = sternite; tg = tergite; vs = ventral strut. Scale bars: A–B = 0.4 mm; C–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033422" httpUri="https://zenodo.org/record/7033422/files/figure.png" pageId="17" pageNumber="18">Fig. 11G</figureCitation>
). Sternite IX thicker posteriorly and triangular anteriorly (
<figureCitation id="B5D3F3CCF40BFFC198FEF89BFC85F8CE" box="[734,841,1889,1916]" captionStart="Fig" captionStartId="21.[189,232,1831,1857]" captionTargetBox="[189,1392,266,1756]" captionTargetId="figure-11@21.[189,1398,266,1760]" captionTargetPageId="21" captionText="Fig. 11. Cyparium achardi sp. nov. A, C. Holotype, ♂ (CELC). B, D–G. Paratype ♂ (CELC). A–B. Abdomen, dorsal view (A) and ventral view (B). C. Ventrite 1. D. Tergite VIII. E. Terminalia. F. Tergite IX. G. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: st = sternite; tg = tergite; vs = ventral strut. Scale bars: A–B = 0.4 mm; C–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033422" httpUri="https://zenodo.org/record/7033422/files/figure.png" pageId="17" pageNumber="18">Fig. 11F</figureCitation>
). Aedeagus strongly sclerotized, enlarged and darker basely and narrowed apically, apex of median lobe short (
<figureCitation id="B5D3F3CCF40BFFC1999DF87FFB8AF82D" box="[957,1094,1925,1951]" captionStart="Fig" captionStartId="22.[189,232,1831,1857]" captionTargetBox="[236,1394,275,1759]" captionTargetId="figure-17@22.[189,1399,265,1759]" captionTargetPageId="22" captionText="Fig. 12. Cyparium achardi sp. nov., paratype, ♂ (CELC): aedeagi. A. Lateral view. B. Frontal view. C. Dorsal view. D. Internal sac, frontal view. E. Internal sac, dorsal view. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: hk = hook; op = opening; pa = parameres. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7033424" httpUri="https://zenodo.org/record/7033424/files/figure.png" pageId="17" pageNumber="18">Fig. 12A,B</figureCitation>
); openings in dorsal view somewhat enlarged, forming an acute angle (
<figureCitation id="B5D3F3CCF40BFFC198FAF852FC84F870" box="[730,840,1960,1986]" captionStart="Fig" captionStartId="22.[189,232,1831,1857]" captionTargetBox="[236,1394,275,1759]" captionTargetId="figure-17@22.[189,1399,265,1759]" captionTargetPageId="22" captionText="Fig. 12. Cyparium achardi sp. nov., paratype, ♂ (CELC): aedeagi. A. Lateral view. B. Frontal view. C. Dorsal view. D. Internal sac, frontal view. E. Internal sac, dorsal view. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: hk = hook; op = opening; pa = parameres. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7033424" httpUri="https://zenodo.org/record/7033424/files/figure.png" pageId="17" pageNumber="18">Fig. 12C</figureCitation>
); internal sac with irregular sclerites, with two hooks (
<figureCitation id="B5D3F3CCF40BFFC19B35F831FE57F857" box="[277,411,1995,2021]" captionStart="Fig" captionStartId="22.[189,232,1831,1857]" captionTargetBox="[236,1394,275,1759]" captionTargetId="figure-17@22.[189,1399,265,1759]" captionTargetPageId="22" captionText="Fig. 12. Cyparium achardi sp. nov., paratype, ♂ (CELC): aedeagi. A. Lateral view. B. Frontal view. C. Dorsal view. D. Internal sac, frontal view. E. Internal sac, dorsal view. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: hk = hook; op = opening; pa = parameres. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7033424" httpUri="https://zenodo.org/record/7033424/files/figure.png" pageId="17" pageNumber="18">Fig. 12D,E</figureCitation>
), parameres short, somewhat enlarged apically (
<figureCitation id="B5D3F3CCF40BFFC199F0F831FBF2F857" box="[976,1086,1995,2021]" captionStart="Fig" captionStartId="22.[189,232,1831,1857]" captionTargetBox="[236,1394,275,1759]" captionTargetId="figure-17@22.[189,1399,265,1759]" captionTargetPageId="22" captionText="Fig. 12. Cyparium achardi sp. nov., paratype, ♂ (CELC): aedeagi. A. Lateral view. B. Frontal view. C. Dorsal view. D. Internal sac, frontal view. E. Internal sac, dorsal view. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: hk = hook; op = opening; pa = parameres. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7033424" httpUri="https://zenodo.org/record/7033424/files/figure.png" pageId="17" pageNumber="18">Fig. 12A</figureCitation>
).
</paragraph>
<caption id="7997BFC1F408FFC29A9DF8DDFD2FF87D" ID-DOI="http://doi.org/10.5281/zenodo.7033416" ID-Zenodo-Dep="7033416" httpUri="https://zenodo.org/record/7033416/files/figure.png" pageId="18" pageNumber="19" startId="18.[189,232,1831,1857]" targetBox="[192,1393,273,1749]" targetPageId="18">
<paragraph id="2D57EF49F408FFC29A9DF8DDFD2FF87D" blockId="18.[189,1399,1831,1999]" pageId="18" pageNumber="19">
<emphasis id="1F9C335BF408FFC29A9DF8DDFEC0F8F3" bold="true" box="[189,268,1831,1857]" pageId="18" pageNumber="19">Fig. 8.</emphasis>
<taxonomicName id="EAE894CAF408FFC29B35F8D2FE21F8F0" authority="Groll &amp; Lopes-Andrade, 2022" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[277,493,1832,1858]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="achardi" status="sp. nov.">
<emphasis id="1F9C335BF408FFC29B35F8D2FE21F8F0" box="[277,493,1832,1858]" italics="true" pageId="18" pageNumber="19">Cyparium achardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AF8E20F408FFC29BD6F8D2FD9EF8F0" box="[502,594,1832,1858]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="1F9C335BF408FFC2987BF8DDFD57F8F3" bold="true" box="[603,667,1831,1857]" pageId="18" pageNumber="19">A–C.</emphasis>
Paratype, ♂ (CELC).
<emphasis id="1F9C335BF408FFC29987F8DDFC70F8F3" bold="true" box="[935,956,1831,1857]" pageId="18" pageNumber="19">A</emphasis>
. Labium.
<emphasis id="1F9C335BF408FFC29E19F8D2FB87F8F0" bold="true" box="[1081,1099,1832,1858]" pageId="18" pageNumber="19">B</emphasis>
. Hypopharynx.
<emphasis id="1F9C335BF408FFC29F2DF8DDFAEDF8F3" bold="true" box="[1293,1313,1831,1857]" pageId="18" pageNumber="19">C</emphasis>
. Head, ventral view.
<emphasis id="1F9C335BF408FFC29B42F8B1FEB9F8D7" bold="true" box="[354,373,1867,1893]" pageId="18" pageNumber="19">D</emphasis>
. Holotype, ♂ (CELC), pronotum, dorsal view.
<emphasis id="1F9C335BF408FFC2998DF8B0FC2AF8D6" bold="true" box="[941,998,1866,1892]" pageId="18" pageNumber="19">E–G</emphasis>
. Paratype, ♂ (CELC), prothorax.
<emphasis id="1F9C335BF408FFC29A9DF894FF1DF83A" bold="true" box="[189,209,1902,1928]" pageId="18" pageNumber="19">E</emphasis>
. Frontal view.
<emphasis id="1F9C335BF408FFC29BB2F894FE68F83A" bold="true" box="[402,420,1902,1928]" pageId="18" pageNumber="19">F</emphasis>
. Ventral view.
<emphasis id="1F9C335BF408FFC29844F894FDB7F83A" bold="true" box="[612,635,1902,1928]" pageId="18" pageNumber="19">G</emphasis>
. Inner view. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: ns = notosternal suture; pr = profurca; st = sclerotized plate. Scale bars: A–B = 0.05 mm; C = 0.1 mm; D–G = 0.5 mm.
</paragraph>
</caption>
<caption id="7997BFC1F409FFC39A9DF917FEBFF869" ID-DOI="http://doi.org/10.5281/zenodo.7033418" ID-Zenodo-Dep="7033418" httpUri="https://zenodo.org/record/7033418/files/figure.png" pageId="19" pageNumber="20" startId="19.[189,232,1773,1799]" targetBox="[189,1398,285,1696]" targetPageId="19">
<paragraph id="2D57EF49F409FFC39A9DF917FEBFF869" blockId="19.[187,1399,1773,2011]" pageId="19" pageNumber="20">
<emphasis id="1F9C335BF409FFC39A9DF917FEC3F8B5" bold="true" box="[189,271,1773,1799]" pageId="19" pageNumber="20">Fig. 9.</emphasis>
<taxonomicName id="EAE894CAF409FFC39B3AF917FE38F8B5" authority="Groll &amp; Lopes-Andrade, 2022" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[282,500,1773,1799]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="achardi" status="sp. nov.">
<emphasis id="1F9C335BF409FFC39B3AF917FE38F8B5" box="[282,500,1773,1799]" italics="true" pageId="19" pageNumber="20">Cyparium achardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AF8E20F409FFC39820F914FDACF8BA" box="[512,608,1774,1800]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♂ (CELC).
<emphasis id="1F9C335BF409FFC39959F917FC42F8B5" bold="true" box="[889,910,1773,1799]" pageId="19" pageNumber="20">A</emphasis>
. Prothorax, lateral view.
<emphasis id="1F9C335BF409FFC39EE4F917FB30F8B5" bold="true" box="[1220,1276,1773,1799]" pageId="19" pageNumber="20">B–C</emphasis>
. Scutellar plate.
<emphasis id="1F9C335BF409FFC39B24F8EAFED4F898" bold="true" box="[260,280,1808,1834]" pageId="19" pageNumber="20">B</emphasis>
. Dorsal view.
<emphasis id="1F9C335BF409FFC39B9DF8EAFE1EF898" bold="true" box="[445,466,1808,1834]" pageId="19" pageNumber="20">C</emphasis>
. Apically slanted view.
<emphasis id="1F9C335BF409FFC398C7F8EAFCD2F898" bold="true" box="[743,798,1808,1834]" pageId="19" pageNumber="20">D–E</emphasis>
. Meso- and metathorax.
<emphasis id="1F9C335BF409FFC39E1FF8EAFB98F898" bold="true" box="[1087,1108,1808,1834]" pageId="19" pageNumber="20">D</emphasis>
. Ventral view.
<emphasis id="1F9C335BF409FFC39F21F8EAFAD9F898" bold="true" box="[1281,1301,1808,1834]" pageId="19" pageNumber="20">E</emphasis>
. Lateral view.
<emphasis id="1F9C335BF409FFC39B28F8CEFED6F8FC" bold="true" box="[264,282,1844,1870]" pageId="19" pageNumber="20">F</emphasis>
. Metanotum. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: al = alacrista; ap = anterior phragma; ip = intercoxal plate; ml = median line; mma = median membranous area; mp = mesoventral process; mvl = mesoventral line; mxl = mesocoxal line; pp = prosternal process; ps = prescutellar line; px = procoxal rest; sss = scutoscutellar suture. Scale bars: A, D–F = 0.2 mm; B–C = 0.1 mm.
</paragraph>
</caption>
<caption id="7997BFC1F40EFFC49A9DF911FABBF820" ID-DOI="http://doi.org/10.5281/zenodo.7033420" ID-Zenodo-Dep="7033420" httpUri="https://zenodo.org/record/7033420/files/figure.png" pageId="20" pageNumber="21" startId="20.[189,232,1771,1797]" targetBox="[189,1392,273,1719]" targetPageId="20">
<paragraph id="2D57EF49F40EFFC49A9DF911FABBF820" blockId="20.[188,1399,1771,1939]" pageId="20" pageNumber="21">
<emphasis id="1F9C335BF40EFFC49A9DF911FED3F8B7" bold="true" box="[189,287,1771,1797]" pageId="20" pageNumber="21">Fig. 10.</emphasis>
<taxonomicName id="EAE894CAF40EFFC49B0AF916FDCAF8B7" authority="Groll &amp; Lopes-Andrade, 2022" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[298,518,1771,1798]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="achardi" status="sp. nov.">
<emphasis id="1F9C335BF40EFFC49B0AF916FDCAF8B7" box="[298,518,1771,1798]" italics="true" pageId="20" pageNumber="21">Cyparium achardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AF8E20F40EFFC49832F916FDBFF8B4" box="[530,627,1772,1798]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♂ (CELC).
<emphasis id="1F9C335BF40EFFC499AFF911FC0BF8B7" bold="true" box="[911,967,1771,1797]" pageId="20" pageNumber="21">A–B</emphasis>
. Metendosternite.
<emphasis id="1F9C335BF40EFFC49E90F911FB09F8B7" bold="true" box="[1200,1221,1771,1797]" pageId="20" pageNumber="21">A</emphasis>
. Dorsal view.
<emphasis id="1F9C335BF40EFFC49A9DF8F4FF1DF89A" bold="true" box="[189,209,1806,1832]" pageId="20" pageNumber="21">B</emphasis>
. Lateral view.
<emphasis id="1F9C335BF40EFFC49BABF8F4FE08F89A" bold="true" box="[395,452,1806,1832]" pageId="20" pageNumber="21">C–D</emphasis>
. Elytron.
<emphasis id="1F9C335BF40EFFC49861F8F4FD9AF89A" bold="true" box="[577,598,1806,1832]" pageId="20" pageNumber="21">C</emphasis>
. Entire.
<emphasis id="1F9C335BF40EFFC498E2F8F4FD1BF89A" bold="true" box="[706,727,1806,1832]" pageId="20" pageNumber="21">D</emphasis>
. Apex.
<emphasis id="1F9C335BF40EFFC4991BF8F4FC81F89A" bold="true" box="[827,845,1806,1832]" pageId="20" pageNumber="21">E</emphasis>
. Hind wing.
<emphasis id="1F9C335BF40EFFC499D1F8F4FBE5F89A" bold="true" box="[1009,1065,1806,1832]" pageId="20" pageNumber="21">F–H</emphasis>
. Legs.
<emphasis id="1F9C335BF40EFFC49EA6F8F4FB54F89A" bold="true" box="[1158,1176,1806,1832]" pageId="20" pageNumber="21">F</emphasis>
. Fore.
<emphasis id="1F9C335BF40EFFC49ED1F8F4FAC4F89A" bold="true" box="[1265,1288,1806,1832]" pageId="20" pageNumber="21">G</emphasis>
. Middle.
<emphasis id="1F9C335BF40EFFC49A9DF8C8FF18F8FE" bold="true" box="[189,212,1842,1868]" pageId="20" pageNumber="21">H</emphasis>
. Hind. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: CAS = cubitoanal strut; fa = furcal arms; MP 1+2 = medial field; MSP = medial spur; RA 4 = radius anterior; RP 3+4 = radius posterior; sr = ‘stalk ridge’; vlf = ventral longitudinal flange. Scale bars:A–D, F–H = 0.2 mm; E = 0.5 mm.
</paragraph>
</caption>
<caption id="7997BFC1F40FFFC59A9DF8DDFAA2F81E" ID-DOI="http://doi.org/10.5281/zenodo.7033422" ID-Zenodo-Dep="7033422" httpUri="https://zenodo.org/record/7033422/files/figure.png" pageId="21" pageNumber="22" startId="21.[189,232,1831,1857]" targetBox="[189,1392,266,1756]" targetPageId="21">
<paragraph id="2D57EF49F40FFFC59A9DF8DDFAA2F81E" blockId="21.[189,1399,1831,1964]" pageId="21" pageNumber="22">
<emphasis id="1F9C335BF40FFFC59A9DF8DDFEECF8F3" bold="true" box="[189,288,1831,1857]" pageId="21" pageNumber="22">Fig. 11.</emphasis>
<taxonomicName id="EAE894CAF40FFFC59B0EF8D2FDC0F8F0" authority="Groll &amp; Lopes-Andrade, 2022" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[302,524,1832,1858]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="achardi" status="sp. nov.">
<emphasis id="1F9C335BF40FFFC59B0EF8D2FDC0F8F0" box="[302,524,1832,1858]" italics="true" pageId="21" pageNumber="22">Cyparium achardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AF8E20F40FFFC5983BF8D2FDB1F8F0" box="[539,637,1832,1858]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="1F9C335BF40FFFC598ABF8DDFD07F8F3" bold="true" box="[651,715,1831,1857]" pageId="21" pageNumber="22">A, C</emphasis>
. Holotype, ♂ (CELC).
<emphasis id="1F9C335BF40FFFC599DDF8D2FBADF8F3" bold="true" box="[1021,1121,1831,1858]" pageId="21" pageNumber="22">B, D–G</emphasis>
. Paratype ♂ (CELC).
<emphasis id="1F9C335BF40FFFC59A9DF8B0FF39F8D6" bold="true" box="[189,245,1866,1892]" pageId="21" pageNumber="22">A–B</emphasis>
. Abdomen, dorsal view (A) and ventral view (B).
<emphasis id="1F9C335BF40FFFC5997BF8B0FCBCF8D6" bold="true" box="[859,880,1866,1892]" pageId="21" pageNumber="22">C</emphasis>
. Ventrite 1.
<emphasis id="1F9C335BF40FFFC59E29F8B1FBD0F8D7" bold="true" box="[1033,1052,1867,1893]" pageId="21" pageNumber="22">D</emphasis>
. Tergite VIII.
<emphasis id="1F9C335BF40FFFC59EEEF8B1FB2CF8D7" bold="true" box="[1230,1248,1867,1893]" pageId="21" pageNumber="22">E</emphasis>
. Terminalia.
<emphasis id="1F9C335BF40FFFC59A9FF894FF03F83A" bold="true" box="[191,207,1902,1928]" pageId="21" pageNumber="22">F</emphasis>
. Tergite IX.
<emphasis id="1F9C335BF40FFFC59B5CF894FE5FF83A" bold="true" box="[380,403,1902,1928]" pageId="21" pageNumber="22">G</emphasis>
. Sternite VIII. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: st = sternite; tg = tergite; vs = ventral strut. Scale bars: A–B = 0.4 mm; C–G = 0.1 mm.
</paragraph>
</caption>
<caption id="7997BFC1F40CFFC69A9DF8DDFED2F81E" ID-DOI="http://doi.org/10.5281/zenodo.7033424" ID-Zenodo-Dep="7033424" httpUri="https://zenodo.org/record/7033424/files/figure.png" pageId="22" pageNumber="23" startId="22.[189,232,1831,1857]" targetBox="[236,1394,275,1759]" targetPageId="22">
<paragraph id="2D57EF49F40CFFC69A9DF8DDFED2F81E" blockId="22.[189,1400,1831,1964]" pageId="22" pageNumber="23">
<emphasis id="1F9C335BF40CFFC69A9DF8DDFED1F8F3" bold="true" box="[189,285,1831,1857]" pageId="22" pageNumber="23">Fig. 12.</emphasis>
<taxonomicName id="EAE894CAF40CFFC69B07F8D2FDCCF8F0" authority="Groll &amp; Lopes-Andrade, 2022" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[295,512,1832,1858]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="achardi" status="sp. nov.">
<emphasis id="1F9C335BF40CFFC69B07F8D2FDCCF8F0" box="[295,512,1832,1858]" italics="true" pageId="22" pageNumber="23">Cyparium achardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AF8E20F40CFFC6982AF8D2FDA5F8F0" box="[522,617,1832,1858]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♂ (CELC): aedeagi.
<emphasis id="1F9C335BF40CFFC699C9F8DDFC32F8F3" bold="true" box="[1001,1022,1831,1857]" pageId="22" pageNumber="23">A</emphasis>
. Lateral view.
<emphasis id="1F9C335BF40CFFC69E95F8D2FB05F8F0" bold="true" box="[1205,1225,1832,1858]" pageId="22" pageNumber="23">B</emphasis>
. Frontal view.
<emphasis id="1F9C335BF40CFFC69A9DF8B0FF1EF8D6" bold="true" box="[189,210,1866,1892]" pageId="22" pageNumber="23">C</emphasis>
. Dorsal view.
<emphasis id="1F9C335BF40CFFC69BA1F8B1FE59F8D7" bold="true" box="[385,405,1867,1893]" pageId="22" pageNumber="23">D</emphasis>
. Internal sac, frontal view.
<emphasis id="1F9C335BF40CFFC698FCF8B1FD22F8D7" bold="true" box="[732,750,1867,1893]" pageId="22" pageNumber="23">E</emphasis>
. Internal sac, dorsal view. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: hk = hook; op = opening; pa = parameres. Scale bars = 0.2 mm.
</paragraph>
</caption>
<caption id="7997BFC1F40DFFC79A9DF8EAFC5FF826" ID-DOI="http://doi.org/10.5281/zenodo.7033426" ID-Zenodo-Dep="7033426" httpUri="https://zenodo.org/record/7033426/files/figure.png" pageId="23" pageNumber="24" startId="23.[189,232,1808,1834]" targetBox="[189,1398,266,1752]" targetPageId="23">
<paragraph id="2D57EF49F40DFFC79A9DF8EAFC5FF826" blockId="23.[187,1399,1808,1940]" pageId="23" pageNumber="24">
<emphasis id="1F9C335BF40DFFC79A9DF8EAFED4F898" bold="true" box="[189,280,1808,1834]" pageId="23" pageNumber="24">Fig. 13.</emphasis>
<taxonomicName id="EAE894CAF40DFFC79B3EF8EBFE3FF898" authority="Groll &amp; Lopes-Andrade, 2022" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[286,499,1808,1835]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="achardi" status="sp. nov.">
<emphasis id="1F9C335BF40DFFC79B3EF8EBFE3FF898" box="[286,499,1808,1835]" italics="true" pageId="23" pageNumber="24">Cyparium achardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AF8E20F40DFFC79BD9F8EBFD98F899" box="[505,596,1809,1835]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♀ (CELC).
<emphasis id="1F9C335BF40DFFC79977F8EAFCA0F898" bold="true" box="[855,876,1808,1834]" pageId="23" pageNumber="24">A</emphasis>
. Tergite VIII.
<emphasis id="1F9C335BF40DFFC79E2DF8EAFBEDF898" bold="true" box="[1037,1057,1808,1834]" pageId="23" pageNumber="24">B</emphasis>
. Sternite VIII.
<emphasis id="1F9C335BF40DFFC79EEDF8EAFB2EF898" bold="true" box="[1229,1250,1808,1834]" pageId="23" pageNumber="24">C</emphasis>
. Terminalia.
<emphasis id="1F9C335BF40DFFC79A9BF8CEFF1CF8FC" bold="true" box="[187,208,1844,1870]" pageId="23" pageNumber="24">D</emphasis>
. Sclerite of vaginal plate.
<emphasis id="1F9C335BF40DFFC79806F8CEFDF6F8FC" bold="true" box="[550,570,1844,1870]" pageId="23" pageNumber="24">E</emphasis>
. Ovipositor. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: bc = bursa copulatrix; dg = distal gonocoxites; gs = gonostyli; sc = sclerite; sp = spermatheca. Scale bars: A–C = 0.2 mm; D–E = 0.1 mm.
</paragraph>
</caption>
<paragraph id="2D57EF49F402FFC89ACAFEF3FE9DFE91" blockId="24.[188,1399,265,573]" box="[234,337,265,291]" pageId="24" pageNumber="25">
<emphasis id="1F9C335BF402FFC89ACAFEF3FE9DFE91" bold="true" box="[234,337,265,291]" pageId="24" pageNumber="25">Females</emphasis>
</paragraph>
<paragraph id="2D57EF49F402FFC89A9DFED6FE92FD8E" blockId="24.[188,1399,265,573]" pageId="24" pageNumber="25">
MEASUREMENTS (n = 1,
<typeStatus id="F25351EBF402FFC89BCDFED6FD9BFEF4" box="[493,599,300,326]" pageId="24" pageNumber="25" type="paratype">paratype</typeStatus>
; in mm). Antennomeres (length(width)): 0.20(0.06), 0.13(0.05), 0.11(0.04), 0.12(0.05), 0.11(0.06), 0.08(0.07), 0.11(0.11), 0.10(0.13), 0.11(0.14), 0.10(0.16), 0.16(0.15); (n = 4,
<typeStatus id="F25351EBF402FFC89B02FE89FE47FE3F" box="[290,395,371,397]" pageId="24" pageNumber="25" type="paratype">paratype</typeStatus>
; in mm): TL 3.32–3.40 (mean = 3.37, standard deviation ± 0.04), PL 1.23–1.39 (1.33 ± 0.07), PA 0.97–1.05 (1.01 ± 0.04), PB 2.05–2.17 (2.10 ± 0.05), SL 0.18–0.21 (0.19 ± 0.01), SW 0.17–0.20 (0.19 ± 0.01), EI 1.90–2.00 (1.94 ± 0.05), EL 2.27–2.40 (2.35 ± 0.05), EW 1.25–1.27 (1.25 ± 0.01), EH 0.80–1.00 (0.88 ± 0.09), HW 0.81–0.84 (0.84 ±0.02), IS 0.16–0.25 (0.20 ±0.04), WA 0.15–0.21 (0.18 ± 0.02), MC 0.87–0.95 (0.90 ± 0.03), MB 0.42–0.52 (0.46 ± 0.04), VL 0.67–0.75 (0.71 ± 0.03).
</paragraph>
<paragraph id="2D57EF49F402FFC89A9DFD9EFB9AFCB8" blockId="24.[185,1400,611,779]" pageId="24" pageNumber="25">
Antennae longer than in males, surpassing mesanepisternum (
<figureCitation id="B5D3F3CCF402FFC899B5FD99FC22FDCC" box="[917,1006,611,638]" captionStart="Fig" captionStartId="16.[189,232,1814,1840]" captionTargetBox="[189,1398,273,1759]" captionTargetId="figure-16@16.[189,1398,265,1759]" captionTargetPageId="16" captionText="Fig. 7. Cyparium achardi sp. nov. A–D. Holotype, ♂ (CELC). A. Dorsal view. B. Ventral view. C. Lateral view. D. Frontal view. E–F. Antennae, paratype, ♂ (CELC) (E) and paratype, ♀ (CELC) (F). G–J. Paratype, ♂ (CELC). G. Labrum. H–I. Mandible. H. Left. I. Right. J. Maxilla. Specimens collected at Mata do Paraíso, Viçosa (MG, Brazil). Scale bars: A–C = 1.0 mm; D–F = 0.2 mm; G–J = 0.1." figureDoi="http://doi.org/10.5281/zenodo.7033414" httpUri="https://zenodo.org/record/7033414/files/figure.png" pageId="24" pageNumber="25">Fig. 7F</figureCitation>
). Tergite VIII triangular, rounded posteriorly; punctation dense, coarse (inconspicuous in some specimens); subglabrous (
<figureCitation id="B5D3F3CCF402FFC89E81FD7DFAC7FD13" box="[1185,1291,647,673]" captionStart="Fig" captionStartId="23.[189,232,1808,1834]" captionTargetBox="[189,1398,266,1752]" captionTargetId="figure-12@23.[189,1398,264,1758]" captionTargetPageId="23" captionText="Fig. 13. Cyparium achardi sp. nov., paratype, ♀ (CELC). A. Tergite VIII.B. Sternite VIII.C. Terminalia. D. Sclerite of vaginal plate. E. Ovipositor. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: bc = bursa copulatrix; dg = distal gonocoxites; gs = gonostyli; sc = sclerite; sp = spermatheca. Scale bars: A–C = 0.2 mm; D–E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033426" httpUri="https://zenodo.org/record/7033426/files/figure.png" pageId="24" pageNumber="25">Fig. 13A</figureCitation>
).Sternite VIII rectangular with strigulate microsculpture (
<figureCitation id="B5D3F3CCF402FFC898D8FD50FCAAFD76" box="[760,870,682,708]" captionStart="Fig" captionStartId="23.[189,232,1808,1834]" captionTargetBox="[189,1398,266,1752]" captionTargetId="figure-12@23.[189,1398,264,1758]" captionTargetPageId="23" captionText="Fig. 13. Cyparium achardi sp. nov., paratype, ♀ (CELC). A. Tergite VIII.B. Sternite VIII.C. Terminalia. D. Sclerite of vaginal plate. E. Ovipositor. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: bc = bursa copulatrix; dg = distal gonocoxites; gs = gonostyli; sc = sclerite; sp = spermatheca. Scale bars: A–C = 0.2 mm; D–E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033426" httpUri="https://zenodo.org/record/7033426/files/figure.png" pageId="24" pageNumber="25">Fig. 13B</figureCitation>
). Vagina and bursa copulatrix membranous; vaginal plate with an apical sclerite and a pair of spermatheca at base of oviduct (
<figureCitation id="B5D3F3CCF402FFC89EACFD37FAD1FD55" box="[1164,1309,717,743]" captionStart="Fig" captionStartId="23.[189,232,1808,1834]" captionTargetBox="[189,1398,266,1752]" captionTargetId="figure-12@23.[189,1398,264,1758]" captionTargetPageId="23" captionText="Fig. 13. Cyparium achardi sp. nov., paratype, ♀ (CELC). A. Tergite VIII.B. Sternite VIII.C. Terminalia. D. Sclerite of vaginal plate. E. Ovipositor. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: bc = bursa copulatrix; dg = distal gonocoxites; gs = gonostyli; sc = sclerite; sp = spermatheca. Scale bars: A–C = 0.2 mm; D–E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033426" httpUri="https://zenodo.org/record/7033426/files/figure.png" pageId="24" pageNumber="25">Fig. 13C–D</figureCitation>
). Distal gonocoxites parallel, straight and thick; gonostyli larger at apex (
<figureCitation id="B5D3F3CCF402FFC89998FD0AFB85FCB8" box="[952,1097,752,778]" captionStart="Fig" captionStartId="23.[189,232,1808,1834]" captionTargetBox="[189,1398,266,1752]" captionTargetId="figure-12@23.[189,1398,264,1758]" captionTargetPageId="23" captionText="Fig. 13. Cyparium achardi sp. nov., paratype, ♀ (CELC). A. Tergite VIII.B. Sternite VIII.C. Terminalia. D. Sclerite of vaginal plate. E. Ovipositor. Specimen collected at Mata do Paraíso, Viçosa (MG, Brazil). Abbreviations: bc = bursa copulatrix; dg = distal gonocoxites; gs = gonostyli; sc = sclerite; sp = spermatheca. Scale bars: A–C = 0.2 mm; D–E = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7033426" httpUri="https://zenodo.org/record/7033426/files/figure.png" pageId="24" pageNumber="25">Figs 13C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="65F2BCC2F402FFC89A9DFCC8FCAAFBB5" pageId="24" pageNumber="25" type="biology_ecology">
<paragraph id="2D57EF49F402FFC89A9DFCC8FE8CFCF9" blockId="24.[189,320,817,844]" box="[189,320,817,844]" pageId="24" pageNumber="25">
<heading id="761F5825F402FFC89A9DFCC8FE8CFCF9" bold="true" box="[189,320,817,844]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="1F9C335BF402FFC89A9DFCC8FE8CFCF9" bold="true" box="[189,320,817,844]" pageId="24" pageNumber="25">Host fungi</emphasis>
</heading>
</paragraph>
<paragraph id="2D57EF49F402FFC89A9DFC9AFCAAFBB5" blockId="24.[188,1399,864,1032]" pageId="24" pageNumber="25">
Adults were collected from
<taxonomicName id="EAE894CAF402FFC8983DFC9AFC44FCC8" authority="Singer" authorityName="Singer" box="[541,904,864,890]" class="Agaricomycetes" family="Omphalotaceae" genus="Marasmiellus" kingdom="Fungi" order="Agaricales" pageId="24" pageNumber="25" phylum="Basidiomycota" rank="species" species="volvatus">
<emphasis id="1F9C335BF402FFC8983DFC9AFCE2FCC8" box="[541,814,864,890]" italics="true" pageId="24" pageNumber="25">Marasmiellus volvatus</emphasis>
Singer
</taxonomicName>
(
<taxonomicName id="EAE894CAF402FFC899BDFC9AFBD3FCC8" box="[925,1055,864,890]" class="Agaricomycetes" kingdom="Fungi" order="Agaricales" pageId="24" pageNumber="25" phylum="Basidiomycota" rank="order">Agaricales</taxonomicName>
,
<taxonomicName id="EAE894CAF402FFC89E11FC9AFB26FCC8" authorityName="Bresinsky" authorityYear="1985" box="[1073,1258,864,890]" class="Agaricomycetes" family="Omphalotaceae" kingdom="Fungi" order="Agaricales" pageId="24" pageNumber="25" phylum="Basidiomycota" rank="family">Omphalotaceae</taxonomicName>
) (1 record, 4 individuals),
<taxonomicName id="EAE894CAF402FFC89B51FC79FDF7FC2C" box="[369,571,899,926]" class="Agaricomycetes" family="Omphalotaceae" genus="Marasmiellus" kingdom="Fungi" order="Agaricales" pageId="24" pageNumber="25" phylum="Basidiomycota" rank="species" species="undetermined">
<emphasis id="1F9C335BF402FFC89B51FC79FDDEFC2F" box="[369,530,899,925]" italics="true" pageId="24" pageNumber="25">Marasmiellus</emphasis>
sp.
</taxonomicName>
Murrill (1, 2),
<taxonomicName id="EAE894CAF402FFC898D3FC7EFB15FC2C" authority="(Mont.) Fr." authorityName="Fr." baseAuthorityName="Mont." box="[755,1241,899,926]" class="Agaricomycetes" family="Marasmiaceae" genus="Marasmius" kingdom="Fungi" order="Agaricales" pageId="24" pageNumber="25" phylum="Basidiomycota" rank="species" species="haematocephalus">
<emphasis id="1F9C335BF402FFC898D3FC7EFB81FC2F" box="[755,1101,899,925]" italics="true" pageId="24" pageNumber="25">Marasmius haematocephalus</emphasis>
(Mont.) Fr.
</taxonomicName>
(
<taxonomicName id="EAE894CAF402FFC89ED1FC79FABFFC2F" box="[1265,1395,899,925]" class="Agaricomycetes" kingdom="Fungi" order="Agaricales" pageId="24" pageNumber="25" phylum="Basidiomycota" rank="order">Agaricales</taxonomicName>
,
<taxonomicName id="EAE894CAF402FFC89A9EFC5DFEA5FC73" authorityName="Roze ex Kuhner" authorityYear="1980" box="[190,361,935,961]" class="Agaricomycetes" family="Marasmiaceae" kingdom="Fungi" order="Agaricales" pageId="24" pageNumber="25" phylum="Basidiomycota" rank="family">Marasmiaceae</taxonomicName>
) (1, 1),
<taxonomicName id="EAE894CAF402FFC89BFFFC5DFC1CFC73" authority="(Sacc.) P. Mohr" authorityName="P. Mohr" baseAuthorityName="Sacc." box="[479,976,935,961]" class="Agaricomycetes" family="Agaricaceae" genus="Leucocoprinus" kingdom="Fungi" order="Agaricales" pageId="24" pageNumber="25" phylum="Basidiomycota" rank="species" species="ianthinus">
<emphasis id="1F9C335BF402FFC89BFFFC5DFCC6FC73" box="[479,778,935,961]" italics="true" pageId="24" pageNumber="25">Leucocoprinus ianthinus</emphasis>
(Sacc.) P.Mohr
</taxonomicName>
(
<taxonomicName id="EAE894CAF402FFC899C8FC5DFBA6FC73" box="[1000,1130,935,961]" class="Agaricomycetes" kingdom="Fungi" order="Agaricales" pageId="24" pageNumber="25" phylum="Basidiomycota" rank="order">Agaricales</taxonomicName>
,
<taxonomicName id="EAE894CAF402FFC89E5DFC5DFADCFC73" box="[1149,1296,935,961]" class="Agaricomycetes" family="Agaricaceae" kingdom="Fungi" order="Agaricales" pageId="24" pageNumber="25" phylum="Basidiomycota" rank="family">Agaricaceae</taxonomicName>
) (1, 1),
<taxonomicName id="EAE894CAF402FFC89A9DFC31FDC8FC56" authority="(Sowerby) Pat." authorityName="Pat." baseAuthorityName="Sowerby" box="[189,516,970,996]" class="Agaricomycetes" family="Agaricaceae" genus="Leucocoprinus" kingdom="Fungi" order="Agaricales" pageId="24" pageNumber="25" phylum="Basidiomycota" rank="species" species="cepistipes">
<emphasis id="1F9C335BF402FFC89A9DFC31FE83FC56" box="[189,335,971,996]" italics="true" pageId="24" pageNumber="25">L. cepistipes</emphasis>
(Sowerby) Pat.
</taxonomicName>
(1, 1),
<emphasis id="1F9C335BF402FFC8987BFC31FD22FC56" box="[603,750,971,996]" italics="true" pageId="24" pageNumber="25">Leucoprinus</emphasis>
sp. (1, 1),
<taxonomicName id="EAE894CAF402FFC89948FC30FBDCFC56" box="[872,1040,970,996]" class="Agaricomycetes" family="Hygrophoraceae" genus="Hygrocybe" kingdom="Fungi" order="Agaricales" pageId="24" pageNumber="25" phylum="Basidiomycota" rank="species" species="undetermined">
<emphasis id="1F9C335BF402FFC89948FC30FC24FC56" box="[872,1000,970,996]" italics="true" pageId="24" pageNumber="25">Hygrocybe</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="EAE894CAF402FFC89E3EFC30FB6CFC56" box="[1054,1184,970,996]" class="Agaricomycetes" kingdom="Fungi" order="Agaricales" pageId="24" pageNumber="25" phylum="Basidiomycota" rank="order">Agaricales</taxonomicName>
,
<taxonomicName id="EAE894CAF402FFC89E8BFC30FAA3FC56" authorityName="Lotsy" authorityYear="1907" box="[1195,1391,970,996]" class="Agaricomycetes" family="Hygrophoraceae" kingdom="Fungi" order="Agaricales" pageId="24" pageNumber="25" phylum="Basidiomycota" rank="family">Hygrophoraceae</taxonomicName>
) (1, 1),
<taxonomicName id="EAE894CAF402FFC89B2BFC14FE2EFBBA" box="[267,482,1006,1032]" class="Agaricomycetes" family="Agaricaceae" genus="Leucoagaricus" kingdom="Fungi" order="Agaricales" pageId="24" pageNumber="25" phylum="Basidiomycota" rank="species" species="undetermined">
<emphasis id="1F9C335BF402FFC89B2BFC14FE75FBB5" box="[267,441,1006,1031]" italics="true" pageId="24" pageNumber="25">Leucoagaricus</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="EAE894CAF402FFC89BD2FC17FDB8FBB5" box="[498,628,1005,1031]" class="Agaricomycetes" kingdom="Fungi" order="Agaricales" pageId="24" pageNumber="25" phylum="Basidiomycota" rank="order">Agaricales</taxonomicName>
,
<taxonomicName id="EAE894CAF402FFC8985EFC17FCDDFBB5" box="[638,785,1005,1031]" class="Agaricomycetes" family="Agaricaceae" kingdom="Fungi" order="Agaricales" pageId="24" pageNumber="25" phylum="Basidiomycota" rank="family">Agaricaceae</taxonomicName>
) (1, 1).
</paragraph>
</subSubSection>
<subSubSection id="65F2BCC2F402FFC89A9DFBD5FBFCFAB6" pageId="24" pageNumber="25" type="discussion">
<paragraph id="2D57EF49F402FFC89A9DFBD5FEE3FBFB" blockId="24.[189,303,1071,1097]" box="[189,303,1071,1097]" pageId="24" pageNumber="25">
<heading id="761F5825F402FFC89A9DFBD5FEE3FBFB" bold="true" box="[189,303,1071,1097]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="1F9C335BF402FFC89A9DFBD5FEE3FBFB" bold="true" box="[189,303,1071,1097]" pageId="24" pageNumber="25">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="2D57EF49F402FFC89A9DFBA7FBFCFAB6" blockId="24.[189,1399,1117,1284]" pageId="24" pageNumber="25">
Similar to
<taxonomicName id="EAE894CAF402FFC89B63FBA7FE1BFBC5" authorityName="Achard" authorityYear="1922" box="[323,471,1117,1143]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pygidiale">
<emphasis id="1F9C335BF402FFC89B63FBA7FE1BFBC5" box="[323,471,1117,1143]" italics="true" pageId="24" pageNumber="25">C. pygidiale</emphasis>
</taxonomicName>
, which occurs in Jataí,
<collectingRegion id="EF2C21ABF402FFC89925FBA7FC80FBC5" box="[773,844,1117,1143]" country="Brazil" name="Goias" pageId="24" pageNumber="25">Goiás</collectingRegion>
,
<collectingCountry id="55FFAFD9F402FFC8997DFBA7FC69FBC5" box="[861,933,1117,1143]" name="Brazil" pageId="24" pageNumber="25">Brazil</collectingCountry>
(
<quantity id="EA1042ACF402FFC8999AFBA7FBEAFBC5" box="[954,1062,1117,1143]" metricMagnitude="6" metricUnit="m" metricValue="1.174" pageId="24" pageNumber="25" unit="km" value="1174.0">1174 km</quantity>
from the collection site of
<taxonomicName id="EAE894CAF402FFC89A9DFB7BFEF6FB28" authority="Groll &amp; Lopes-Andrade, 2022" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[189,314,1152,1179]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="achardi" status="sp. nov.">
<emphasis id="1F9C335BF402FFC89A9DFB7BFEF6FB28" box="[189,314,1152,1179]" italics="true" pageId="24" pageNumber="25">C. achardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04AF8E20F402FFC89B64FB7BFE68FB28" box="[324,420,1152,1179]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
), but they differ in the humeral and apical region of the elytra, and in the scutellar shield (blackish in
<taxonomicName id="EAE894CAF402FFC89B84FB5EFDE8FB0F" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[420,548,1187,1214]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="achardi">
<emphasis id="1F9C335BF402FFC89B84FB5EFDE8FB0F" box="[420,548,1187,1214]" italics="true" pageId="24" pageNumber="25">C. achardi</emphasis>
</taxonomicName>
and reddish in
<taxonomicName id="EAE894CAF402FFC898C5FB5EFCBBFB0F" authorityName="Achard" authorityYear="1922" box="[741,887,1187,1214]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pygidiale">
<emphasis id="1F9C335BF402FFC898C5FB5EFCBBFB0F" box="[741,887,1187,1214]" italics="true" pageId="24" pageNumber="25">C. pygidiale</emphasis>
</taxonomicName>
); elytral punctation is also different: rows 1–3 are coarser in
<taxonomicName id="EAE894CAF402FFC89BB9FB3DFDE5FB53" authorityName="Achard" authorityYear="1922" box="[409,553,1223,1249]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pygidiale">
<emphasis id="1F9C335BF402FFC89BB9FB3DFDE5FB53" box="[409,553,1223,1249]" italics="true" pageId="24" pageNumber="25">C. pygidiale</emphasis>
</taxonomicName>
and the remainder are coarser in
<taxonomicName id="EAE894CAF402FFC89996FB3DFBFDFB53" authorityName="von Groll &amp; Lopes-Andrade" authorityYear="2022" box="[950,1073,1223,1249]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="achardi">
<emphasis id="1F9C335BF402FFC89996FB3DFBFDFB53" box="[950,1073,1223,1249]" italics="true" pageId="24" pageNumber="25">C. achardi</emphasis>
</taxonomicName>
; furthermore, rows 1–3 are somewhat outward directed, while they are inward directed in
<taxonomicName id="EAE894CAF402FFC899BAFB10FBE5FAB6" authorityName="Achard" authorityYear="1922" box="[922,1065,1258,1284]" class="Insecta" family="Staphylinidae" genus="Cyparium" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pygidiale">
<emphasis id="1F9C335BF402FFC899BAFB10FBE5FAB6" box="[922,1065,1258,1284]" italics="true" pageId="24" pageNumber="25">C. pygidiale</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="65F2BCC2F402FFC89A9DFAD1FD6EFA25" pageId="24" pageNumber="25" type="distribution">
<paragraph id="2D57EF49F402FFC89A9DFAD1FE9AFAF7" blockId="24.[189,342,1323,1349]" box="[189,342,1323,1349]" pageId="24" pageNumber="25">
<heading id="761F5825F402FFC89A9DFAD1FE9AFAF7" bold="true" box="[189,342,1323,1349]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="1F9C335BF402FFC89A9DFAD1FE9AFAF7" bold="true" box="[189,342,1323,1349]" pageId="24" pageNumber="25">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="2D57EF49F402FFC89A9DFAA0FD6EFA25" blockId="24.[189,1398,1370,1431]" pageId="24" pageNumber="25">
Known only from Mata da Biologia, campus of the Universidade Federal de Viçosa, Viçosa, state of
<collectingRegion id="EF2C21ABF402FFC89A9FFA87FEACFA25" box="[191,352,1405,1431]" country="Brazil" name="Minas Gerais" pageId="24" pageNumber="25">Minas Gerais</collectingRegion>
, Southeast
<collectingCountry id="55FFAFD9F402FFC89BC5FA87FDE1FA25" box="[485,557,1405,1431]" name="Brazil" pageId="24" pageNumber="25">Brazil</collectingCountry>
(
<figureCitation id="B5D3F3CCF402FFC8981CFA87FD59FA25" box="[572,661,1405,1431]" captionStart="Fig" captionStartId="65.[189,232,1952,1978]" captionTargetBox="[189,1398,1082,1910]" captionTargetId="figure-297@65.[189,1398,1082,1910]" captionTargetPageId="65" captionText="Fig. 46. A. Distribution map of species of Cyparium Erichson, 1845, South America. B. Mata da Biologia. C. Mata do Paraíso." figureDoi="http://doi.org/10.5281/zenodo.7033501" httpUri="https://zenodo.org/record/7033501/files/figure.png" pageId="24" pageNumber="25">Fig. 46</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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