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<document id="172D4B83BABCE6BBE581EE2CC69A9D2D" ID-DOI="10.11865/zs.2023401" ID-ISSN="2095-6827" ID-Zenodo-Dep="10941626" ID-ZooBank="46AA3E9C-C065-415D-946B-0813AF169B14" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712576771858" checkinUser="felipe" docAuthor="Zhan, Zhihong &amp; Young, Daniel K." docDate="2023" docId="A51987E07F5FFF91FF02A882FC97FB63" docLanguage="en" docName="ZoolSyst.48.4.279-360.pdf" docOrigin="Zoological Systematics 48 (4)" docSource="https://www.mendeley.com/catalogue/cd623f69-5c7c-3c23-9a34-77dc60046e57/" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Lucanus derani subsp. derani Nagai 2000" docType="treatment" docVersion="3" lastPageNumber="292" masterDocId="5920FF987F54FF9CFF8AAE6BFFD3FFEA" masterDocTitle="A taxonomic assessment and redefinition of the Lucanus fortunei species group in China (Coleoptera: Lucanidae: Lucaninae)" masterLastPageNumber="360" masterPageNumber="279" pageNumber="290" updateTime="1712578213860" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F185889D86F8A86D21370E5FA8145E4D">A taxonomic assessment and redefinition of the Lucanus fortunei species group in China (Coleoptera: Lucanidae: Lucaninae)</mods:title>
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<mods:namePart id="0607B40E7CF704F8F4B01306F8261AD2">Zhan, Zhihong</mods:namePart>
<mods:affiliation id="1BED4AAC38A508612E701F7060764BC3">Department of Entomology, College of Plant Protection, Nanjing Agricultural University, Nanjing 210000, China &amp; Department of Entomology, University of Wisconsin, Madison, WI 53706, USA Corresponding author, E-mail: young @ entomology. wisc. edu</mods:affiliation>
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<mods:namePart id="4B88A4E195008B30D581372C7650FEA8">Young, Daniel K.</mods:namePart>
<mods:affiliation id="CFCC87CBE00C11469A1157CB6A22B790">Department of Entomology, University of Wisconsin, Madison, WI 53706, USA Corresponding author, E-mail: young @ entomology. wisc. edu</mods:affiliation>
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<mods:title id="61ED30BE604AE26D57EFA90A8B014630">Zoological Systematics</mods:title>
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<mods:date id="7892CB67E8419FDB0BEC68A22CB6B410">2023</mods:date>
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<mods:number id="AB2A848FF9D6D331D716B4077490FF2C">48</mods:number>
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<mods:url id="482D1DF517B40104F2C65207753E8FDB">https://www.mendeley.com/catalogue/cd623f69-5c7c-3c23-9a34-77dc60046e57/</mods:url>
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<mods:classification id="01AE4AE1F02596DA357A851D770FF509">journal article</mods:classification>
<mods:identifier id="1C0C09F926B8512070B8B8C9C29DA8C2" type="DOI">10.11865/zs.2023401</mods:identifier>
<mods:identifier id="ECB755D01DD2B83DA0108D6826023E65" type="ISSN">2095-6827</mods:identifier>
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<treatment id="A51987E07F5FFF91FF02A882FC97FB63" ID-DOI="http://doi.org/10.5281/zenodo.10941397" ID-Zenodo-Dep="10941397" LSID="urn:lsid:plazi:treatment:A51987E07F5FFF91FF02A882FC97FB63" httpUri="http://treatment.plazi.org/id/A51987E07F5FFF91FF02A882FC97FB63" lastPageId="13" lastPageNumber="292" pageId="11" pageNumber="290">
<subSubSection id="65AA657D7F5FFF97FF02A882FBF6F8EA" box="[136,1061,1768,1793]" pageId="11" pageNumber="290" type="nomenclature">
<paragraph id="2D0F36F67F5FFF97FF02A882FBF6F8EA" blockId="11.[136,1452,1768,2085]" box="[136,1061,1768,1793]" pageId="11" pageNumber="290">
<heading id="7647819A7F5FFF97FF02A882FBF6F8EA" box="[136,1061,1768,1793]" fontSize="10" level="2" pageId="11" pageNumber="290" reason="2">
<taxonomicName id="EAB04D757F5FFF97FF02A882FDC2F8EA" ID-CoL="7ZC5C" authority="Nagai, 2000" authorityName="Nagai" authorityYear="2000" box="[136,529,1768,1792]" class="Insecta" family="Lucanidae" genus="Lucanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="290" phylum="Arthropoda" rank="subSpecies" species="derani" subSpecies="derani">
<emphasis id="1FC4EAE47F5FFF97FF02A882FDC2F8EA" bold="true" box="[136,529,1768,1792]" pageId="11" pageNumber="290">
<emphasis id="1FC4EAE47F5FFF97FF02A882FE57F8EA" bold="true" box="[136,388,1768,1792]" italics="true" pageId="11" pageNumber="290">Lucanus derani derani</emphasis>
<bibRefCitation id="49214B077F5FFF97FE01A883FDC2F8EA" author="Nagai" box="[395,529,1768,1792]" pageId="11" pageNumber="290" pagination="73 - 108" refId="ref50690" refString="Nagai, S. 2000. Twelve new species, three new subspecies, two new status and with the check list of the family Lucanidae of northern Myanmar. Note on Eurasian Insects, 3: 73 - 108." type="journal article" year="2000">Nagai, 2000</bibRefCitation>
</emphasis>
</taxonomicName>
(
<figureCitation id="B58B2A737F5FFF97FDA4A883FD7AF8EA" box="[558,681,1768,1792]" captionStart="Figures 15–21" captionStartId="12.[155,227,1639,1661]" captionTargetBox="[195,1392,244,1600]" captionTargetId="figure-225@12.[189,1399,243,1603]" captionTargetPageId="12" captionText="Figures 15–21. Habitus of Lucanus derani derani. 15. Male, dorsal view. 16. Male, ventral view. 17. Male, lateral view. 18. Median male. 19. Minor male. 20. Female, dorsal view. 21. Female, ventral view. Scale bar =10mm." figureDoi="http://doi.org/10.5281/zenodo.10941634" httpUri="https://zenodo.org/record/10941634/files/figure.png" pageId="11" pageNumber="290">Figs 15–21</figureCitation>
,
<figureCitation id="B58B2A737F5FFF97FD3EA882FD0CF8EB" box="[692,735,1769,1793]" captionStart="Figures 244–259" captionStartId="67.[155,227,1984,2006]" captionTargetBox="[193,1395,268,1930]" captionTargetId="figure-83@67.[189,1399,266,1936]" captionTargetPageId="67" captionText="Figures 244–259. Head characters of Lucanus spp., male. 244. L. chengyuani. 245. L. datunensis. 246. L. delavayi. 247. L. derani derani. 248. L. derani fukinukiae. 249. L. fortunei. 250. L. fujianensis. 251. L. kirchneri. 252. L. klapperichi. 253. L. moae. 254. L. swinhoei. 255. L. wuyishanensis. 256. L. brivioi. 257. L. cenwanglaoshanus. 258. L. deuveinaus. 259. L. fairmairei. Scale bar=10mm." figureDoi="http://doi.org/10.5281/zenodo.10941723" httpUri="https://zenodo.org/record/10941723/files/figure.png" pageId="11" pageNumber="290">247</figureCitation>
,
<figureCitation id="B58B2A737F5FFF97FD63A882FCC7F8EB" box="[745,788,1769,1793]" captionStart="Figures 276–307" captionStartId="69.[155,227,1902,1924]" captionTargetBox="[191,1398,252,1869]" captionTargetId="figure-151@69.[189,1398,247,1874]" captionTargetPageId="69" captionText="Figures 276–307. Prothoracic tibial characters of Lucanus spp., male. 276. L. chengyuani. 277. L. datunensis. 278. L. delavayi. 279. L. derani derani. 280. L. derani fukinukiae. 281. L. fortunei. 282. L. fujianensis. 283. L. kirchneri. 284. L. klapperichi. 285. L. moae. 286. L. swinhoei. 287. L. wuyishanensis. 288. L. brivioi. 289. L. cenwanglaoshanus. 290. L. deuveinaus. 291. L. fairmairei. 292. L. fonti. 293. L. formosus. 294. L. fujitai. 295. L. hewenjiae. 296. L. liuweii. 297. L. miwai. 298. L. szetschuanicus. 299. L. yulaoensis. 300. L. zhanbishengi. 301. L. zhuxiangi. 302. L. dirki. 303. L. kraatzi kraatzi. 304. L. liupengyui. 305. L. nobilis. 306. L. shulini. 307. L. parryi. Scale bar=10mm." figureDoi="http://doi.org/10.5281/zenodo.10941729" httpUri="https://zenodo.org/record/10941729/files/figure.png" pageId="11" pageNumber="290">279</figureCitation>
,
<figureCitation id="B58B2A737F5FFF97FC95A882FC99F8EB" box="[799,842,1769,1793]" captionStart="Figures 308–323" captionStartId="70.[155,227,1993,2015]" captionTargetBox="[192,1396,270,1938]" captionTargetId="figure-92@70.[189,1399,267,1953]" captionTargetPageId="70" captionText="Figures 308–323. Head characters of Lucanus spp., female. 308. L. delavayi. 309. L. derani derani. 310. L. derani fukinukiae. 311. L. fortunei. 312. L. fujianensis. 313. L. kirchneri. 314. L. klapperichi. 315. L. moae. 316. L. swinhoei. 317. L. wuyishanensis. 318. L. brivioi. 319. L. cenwanglaoshanus. 320. L. deuveinaus. 321. L. fairmairei. 322. L. fonti. 323. L. formosus. Scale bar=10mm." figureDoi="http://doi.org/10.5281/zenodo.10941731" httpUri="https://zenodo.org/record/10941731/files/figure.png" pageId="11" pageNumber="290">309</figureCitation>
,
<figureCitation id="B58B2A737F5FFF97FCDEA882FCACF8EB" box="[852,895,1769,1793]" captionStart="Figures 336–363" captionStartId="72.[155,227,1929,1951]" captionTargetBox="[190,1395,375,1882]" captionTargetId="figure-194@72.[189,1399,373,1885]" captionTargetPageId="72" captionText="Figures 336–363. Prothoracic tibial characters of Lucanus spp., female. 336. L. delavayi. 337. L. derani derani. 338. L. derani fukinukiae. 339. L. fortunei. 340. L. fujianensis. 341. L. kirchneri. 342. L. klapperichi. 343. L. moae. 344. L. swinhoei. 345. L. wuyishanensis. 346. L. brivioi. 347. L. cenwanglaoshanus. 348. L. deuveinaus. 349. L. fairmairei. 350. L. fonti. 351. L. formosus. 352. L. hewenjiae. 353. L. liuweii. 354. L. miwai. 355. L. szetschuanicus. 356. L. zhanbishengi. 357. L. zhuxiangi. 358. L. dirki. 359. L. kraatzi kraatzi. 360. L. liupengyui. 361. L. nobilis. 362. L. shulini. 363. L. parryi. Scale bar=10mm." figureDoi="http://doi.org/10.5281/zenodo.10941737" httpUri="https://zenodo.org/record/10941737/files/figure.png" pageId="11" pageNumber="290">337</figureCitation>
,
<figureCitation id="B58B2A737F5FFF97FC03A882FC39F8EB" box="[905,1002,1769,1793]" captionStart="Figures 364–387" captionStartId="73.[155,227,1862,1884]" captionTargetBox="[191,1396,386,1822]" captionTargetId="figure-202@73.[189,1399,384,1822]" captionTargetPageId="73" captionText="Figures 364–387. Male genitalia of Lucanus spp., dorsal view. 364. L. datunensis, dorsal view. 365. L. datunensis, ventral view. 366. L. datunensis, lateral view. 367. L. delavayi, dorsal view. 368. L. delavayi, ventral view. 369. L. delavayi, lateral view. 370. L. derani derani, dorsal view. 371. L. derani derani, ventral view. 372. L. derani derani, lateral view. 373. L. derani fukinukiae, dorsal view. 374. L. derani fukinukiae, ventral view. 375. L. derani fukinukiae, lateral view. 376. L. fortunei, dorsal view. 377. L. fortunei, ventral view. 378. L. fortunei, lateral view. 379. L. fujianensis, dorsal view. 380. L. fujianensis, ventral view. 381. L. fujianensis, lateral view. 382. L. kirchneri, dorsal view. 383. L. kirchneri, ventral view. 384. L. kirchneri, lateral view. 385. L. klapperichi, dorsal view. 386. L. klapperichi, ventral view. 387. L. klapperichi, lateral view. Scale bar=10mm." figureDoi="http://doi.org/10.5281/zenodo.10941741" httpUri="https://zenodo.org/record/10941741/files/figure.png" pageId="11" pageNumber="290">370–372</figureCitation>
,
<figureCitation id="B58B2A737F5FFF97FC7EA883FBCDF8EA" box="[1012,1054,1768,1792]" captionStart="Figures 457–466" captionStartId="77.[155,227,1951,1973]" captionTargetBox="[189,1399,239,1939]" captionTargetId="figure-98@77.[189,1399,239,1940]" captionTargetPageId="77" captionText="Figures 457–466. Lucanus spp. 457. Male of L. datunensis flying from its grass habitat from Taiwan, photoed by JingZhi Lin. 458. Male of L. delavayi from Sichuan. 459. Male of L. derani derani from Yunnan. 460. Male of L. derani fukinukiae from Yunnan. 461. Male of L. fortunei from Fujian. 462. Male of L. kirchneri collected from host plant in Fujian. 463. Male of L. klapperichi on host plant in Fujian. 464. A pair of L. moae collected in Sichuan. 465. Male of L. swinhoei in Fujian. 466. Male of L. wuyishanensis in Hunan." figureDoi="http://doi.org/10.5281/zenodo.10941751" httpUri="https://zenodo.org/record/10941751/files/figure.png" pageId="11" pageNumber="290">459</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="65AA657D7F5FFF97FF02A967FBCEF8AB" pageId="11" pageNumber="290" type="reference_group">
<paragraph id="2D0F36F67F5FFF97FF02A967FDC3F8CA" blockId="11.[136,1452,1768,2085]" box="[136,528,1803,1825]" pageId="11" pageNumber="290">
<treatmentCitationGroup id="0DA011D87F5FFF97FF02A967FDC3F8CA" box="[136,528,1803,1825]" pageId="11" pageNumber="290">
<treatmentCitation id="AC1110E77F5FFF97FF02A967FE13F8CA" author="Nagai" box="[136,448,1803,1825]" page="91" pageId="11" pageNumber="290" year="2000">
<taxonomicName id="EAB04D757F5FFF97FF02A967FE13F8CA" ID-CoL="72P4R" authority="Nagai, 2000: 91" authorityName="Nagai" authorityPageNumber="91" authorityYear="2000" box="[136,448,1803,1825]" class="Insecta" family="Lucanidae" genus="Lucanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="290" phylum="Arthropoda" rank="species" species="derani">
<emphasis id="1FC4EAE47F5FFF97FF02A967FECCF8CA" box="[136,287,1803,1825]" italics="true" pageId="11" pageNumber="290">Lucanus derani</emphasis>
<bibRefCitation id="49214B077F5FFF97FEAEA960FE13F8CA" author="Nagai" box="[292,448,1803,1825]" pageId="11" pageNumber="290" pagination="73 - 108" refId="ref50690" refString="Nagai, S. 2000. Twelve new species, three new subspecies, two new status and with the check list of the family Lucanidae of northern Myanmar. Note on Eurasian Insects, 3: 73 - 108." type="journal article" year="2000">Nagai, 2000: 91</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 17.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2D0F36F67F5FFF97FF02A947FBCEF8AB" blockId="11.[136,1452,1768,2085]" box="[136,1053,1835,1857]" pageId="11" pageNumber="290">
<taxonomicName id="EAB04D757F5FFF97FF02A947FDEEF8AA" ID-CoL="6QP5T" authority="Huang &amp; Chen, 2010: 89" authorityName="Huang &amp; Chen" authorityPageNumber="89" authorityYear="2010" box="[136,573,1835,1857]" class="Insecta" family="Lucanidae" genus="Lucanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="290" phylum="Arthropoda" rank="species" species="fukinukiae">
<emphasis id="1FC4EAE47F5FFF97FF02A947FE97F8AA" box="[136,324,1835,1857]" italics="true" pageId="11" pageNumber="290">Lucanus fukinukiae</emphasis>
Huang &amp; Chen, 2010: 89
</taxonomicName>
, plate 37, figs 10–11 (Possibly misidentification).
</paragraph>
</subSubSection>
<subSubSection id="65AA657D7F5FFF91FF34A90BFC97FB63" lastPageId="13" lastPageNumber="292" pageId="11" pageNumber="290" type="materials_examined">
<paragraph id="2D0F36F67F5FFF97FF34A90BFF15F7CF" blockId="11.[136,1452,1768,2085]" pageId="11" pageNumber="290">
<materialsCitation id="9DD83CAB7F5FFF97FF34A90BFC62F857" collectingDate="2018-05-26" collectorName="Zhihong Zhan &amp; Adult &amp; Light-trap &amp; Nagai" country="China" county="Yingjiang County" determinerName="Z. Zhan" elevation="1350" latitude="24.6944" location="Mang City" longLatPrecision="7" longitude="97.9433" municipality="Material" pageId="11" pageNumber="290" specimenCount="6" specimenCount-male="6" stateProvince="Yunnan">
<collectingMunicipality id="CD6BAC8C7F5FFF97FF34A90BFECBF892" box="[190,280,1888,1912]" pageId="11" pageNumber="290">Material</collectingMunicipality>
examined. [
<specimenCount id="3BB6FD7F7F5FFF97FE16A90BFE6CF892" box="[412,447,1888,1912]" count="5" pageId="11" pageNumber="290" type="male">5♂</specimenCount>
]:
<collectingCountry id="55A776667F5FFF97FE5CA90BFDFCF892" box="[470,559,1888,1912]" name="China" pageId="11" pageNumber="290">CHINA</collectingCountry>
:
<collectingRegion id="EF74F8147F5FFF97FDB2A90AFD21F892" box="[568,754,1888,1912]" country="China" name="Yunnan" pageId="11" pageNumber="290">Yunnan Province</collectingRegion>
,
<location id="286F602D7F5FFF97FD77A90AFCA3F892" LSID="urn:lsid:plazi:treatment:A51987E07F5FFF91FF02A882FC97FB63:286F602D7F5FFF97FD77A90AFCA3F892" box="[765,880,1888,1913]" country="China" county="Yingjiang County" latitude="24.6944" longLatPrecision="7" longitude="97.9433" municipality="Material" name="Mang City" pageId="11" pageNumber="290" stateProvince="Yunnan">Mang City</location>
//
<location id="286F602D7F5FFF97FC07A90BFBEBF892" LSID="urn:lsid:plazi:treatment:A51987E07F5FFF91FF02A882FC97FB63:286F602D7F5FFF97FC07A90BFBEBF892" box="[909,1080,1888,1912]" country="China" county="Yingjiang County" latitude="24.6944" longLatPrecision="7" longitude="97.9433" municipality="Material" name="Dehong Region" pageId="11" pageNumber="290" stateProvince="Yunnan">Dehong Region</location>
,
<collectingCounty id="C46E4E7A7F5FFF97FBC8A90BFAD2F892" box="[1090,1281,1888,1912]" pageId="11" pageNumber="290">Yingjiang County</collectingCounty>
// [
<emphasis id="1FC4EAE47F5FFF97FAACA930FF6EF873" bold="true" pageId="11" pageNumber="290">德±傣Dzss 治州</emphasis>
,
<emphasis id="1FC4EAE47F5FFF97FF58A915FEF2F873" bold="true" box="[210,289,1918,1945]" pageId="11" pageNumber="290">Ω江县</emphasis>
]; Alt.
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</quantity>
;
<date id="590E10367F5FFF97FE48A9E8FD8BF871" box="[450,600,1923,1947]" pageId="11" pageNumber="290" value="2018-05-26">
<collectingDate id="494AE9DE7F5FFF97FE48A9E8FD8BF871" box="[450,600,1923,1947]" pageId="11" pageNumber="290" value="2018-05-26">26 May. 2018</collectingDate>
</date>
//
<collectorName id="804553207F5FFF97FDFCA9E8FCDCF871" box="[630,783,1923,1947]" pageId="11" pageNumber="290">Zhihong Zhan</collectorName>
leg.; [2nd label]:
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;
<geoCoordinate id="488450317F5FFF97FBDCA9E8FB1BF871" box="[1110,1224,1923,1947]" degrees="97.9433" direction="east" orientation="longitude" pageId="11" pageNumber="290" precision="5" value="97.9433">97.9433°E</geoCoordinate>
<collectorName id="804553207F5FFF97FB47A9E8FAD9F871" box="[1229,1290,1923,1947]" pageId="11" pageNumber="290">Adult</collectorName>
// collected by
<collectorName id="804553207F5FFF97FF02A9CEFF25F857" box="[136,246,1957,1981]" pageId="11" pageNumber="290">Light-trap</collectorName>
//; [3rd label]:
<taxonomicName id="EAB04D757F5FFF97FE05A9CDFE39F857" authorityName="Scopoli" authorityYear="1763" box="[399,490,1958,1981]" class="Insecta" family="Lucanidae" genus="Lucanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="290" phylum="Arthropoda" rank="genus">
<emphasis id="1FC4EAE47F5FFF97FE05A9CDFE39F857" box="[399,490,1958,1981]" italics="true" pageId="11" pageNumber="290">Lucanus</emphasis>
</taxonomicName>
//
<taxonomicName id="EAB04D757F5FFF97FD8CA9CEFD4BF857" authorityName="Nagai" authorityYear="2000" box="[518,664,1957,1981]" class="Insecta" family="Lucanidae" genus="Lucanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="290" phylum="Arthropoda" rank="species" species="derani">
<emphasis id="1FC4EAE47F5FFF97FD8CA9CEFD4BF857" box="[518,664,1957,1981]" italics="true" pageId="11" pageNumber="290">derani derani</emphasis>
</taxonomicName>
//
<collectorName id="804553207F5FFF97FD3EA9CEFD27F857" box="[692,756,1957,1981]" pageId="11" pageNumber="290">Nagai</collectorName>
<specimenCount id="3BB6FD7F7F5FFF97FD71A9CDFCDCF857" box="[763,783,1958,1981]" count="1" pageId="11" pageNumber="290" type="male">♂</specimenCount>
// det.
<determinerName id="D8D3DA327F5FFF97FCD3A9CDFC62F857" box="[857,945,1957,1981]" pageId="11" pageNumber="290">Z. Zhan</determinerName>
</materialsCitation>
;
<materialsCitation id="9DD83CAB7F5FFF97FC36A9CEFF15F7CF" collectingDate="2018-05-26" collectorName="Zhihong Zhan &amp; Adult &amp; Light-trap &amp; Nagai" country="China" county="Yingjiang County" determinerName="Z. Zhan." elevation="1350" latitude="24.6944" location="Dehong Region" longLatPrecision="7" longitude="97.9433" municipality="Mang City" pageId="11" pageNumber="290" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan">
[
<specimenCount id="3BB6FD7F7F5FFF97FC4FA9CEFC35F857" box="[965,998,1957,1981]" count="1" pageId="11" pageNumber="290" type="female">1♀</specimenCount>
]:
<collectingCountry id="55A776667F5FFF97FC77A9CEFB85F857" box="[1021,1110,1957,1981]" name="China" pageId="11" pageNumber="290">CHINA</collectingCountry>
:
<collectingRegion id="EF74F8147F5FFF97FBD5A9CDFACAF857" box="[1119,1305,1957,1981]" country="China" name="Yunnan" pageId="11" pageNumber="290">Yunnan Province</collectingRegion>
,
<collectingMunicipality id="CD6BAC8C7F5FFF97FAA9A9CDFA45F857" box="[1315,1430,1957,1982]" pageId="11" pageNumber="290">Mang City</collectingMunicipality>
//
<location id="286F602D7F5FFF97FF02A9A3FEE6F80A" LSID="urn:lsid:plazi:treatment:A51987E07F5FFF91FF02A882FC97FB63:286F602D7F5FFF97FF02A9A3FEE6F80A" box="[136,309,1992,2016]" country="China" county="Yingjiang County" latitude="24.6944" longLatPrecision="7" longitude="97.9433" municipality="Mang City" name="Dehong Region" pageId="11" pageNumber="290" stateProvince="Yunnan">Dehong Region</location>
,
<collectingCounty id="C46E4E7A7F5FFF97FECAA9A3FDD2F80A" box="[320,513,1992,2016]" pageId="11" pageNumber="290">Yingjiang County</collectingCounty>
// [
<emphasis id="1FC4EAE47F5FFF97FDA3A9A8FD37F834" bold="true" box="[553,740,1987,2014]" pageId="11" pageNumber="290">德±傣Dzss治州</emphasis>
,
<emphasis id="1FC4EAE47F5FFF97FD72A9A8FC9BF834" bold="true" box="[760,840,1987,2014]" pageId="11" pageNumber="290">Ω江县</emphasis>
]; Alt.
<quantity id="EA489B137F5FFF97FC1AA9A3FC0EF80A" box="[912,989,1992,2016]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="11" pageNumber="290" unit="m" value="1350.0">
<elevation id="A69DD1C57F5FFF97FC1AA9A3FC0EF80A" box="[912,989,1992,2016]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="11" pageNumber="290" unit="m" value="1350.0">1350m</elevation>
</quantity>
;
<date id="590E10367F5FFF97FC60A9A3FB52F80A" box="[1002,1153,1992,2017]" pageId="11" pageNumber="290" value="2018-05-26">
<collectingDate id="494AE9DE7F5FFF97FC60A9A3FB52F80A" box="[1002,1153,1992,2017]" pageId="11" pageNumber="290" value="2018-05-26">26 May. 2018</collectingDate>
</date>
//
<collectorName id="804553207F5FFF97FB2BA9A3FAE8F80A" box="[1185,1339,1992,2016]" pageId="11" pageNumber="290">Zhihong Zhan</collectorName>
leg.; [2nd label]:
<geoCoordinate id="488450317F5FFF97FF5EA980FE9FF7E9" box="[212,332,2027,2051]" degrees="24.6944" direction="north" orientation="latitude" pageId="11" pageNumber="290" precision="5" value="24.6944">24.6944°N</geoCoordinate>
;
<geoCoordinate id="488450317F5FFF97FED2A980FE19F7E9" box="[344,458,2027,2051]" degrees="97.9433" direction="east" orientation="longitude" pageId="11" pageNumber="290" precision="5" value="97.9433">97.9433°E</geoCoordinate>
<collectorName id="804553207F5FFF97FE5AA980FDDEF7E9" box="[464,525,2027,2051]" pageId="11" pageNumber="290">Adult</collectorName>
// collected by
<collectorName id="804553207F5FFF97FD3CA980FCF7F7E9" box="[694,804,2027,2051]" pageId="11" pageNumber="290">Light-trap</collectorName>
//; [3rd label]:
<taxonomicName id="EAB04D757F5FFF97FC4AA980FBC8F7E8" authorityName="Scopoli" authorityYear="1763" box="[960,1051,2027,2050]" class="Insecta" family="Lucanidae" genus="Lucanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="290" phylum="Arthropoda" rank="genus">
<emphasis id="1FC4EAE47F5FFF97FC4AA980FBC8F7E8" box="[960,1051,2027,2050]" italics="true" pageId="11" pageNumber="290">Lucanus</emphasis>
</taxonomicName>
//
<taxonomicName id="EAB04D757F5FFF97FBB3A980FB1EF7E9" authorityName="Nagai" authorityYear="2000" box="[1081,1229,2027,2051]" class="Insecta" family="Lucanidae" genus="Lucanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="290" phylum="Arthropoda" rank="species" species="derani">
<emphasis id="1FC4EAE47F5FFF97FBB3A980FB1EF7E9" box="[1081,1229,2027,2051]" italics="true" pageId="11" pageNumber="290">derani derani</emphasis>
</taxonomicName>
//
<collectorName id="804553207F5FFF97FB61A980FAF8F7E9" box="[1259,1323,2027,2051]" pageId="11" pageNumber="290">Nagai</collectorName>
<specimenCount id="3BB6FD7F7F5FFF97FAB9A980FA94F7E8" box="[1331,1351,2027,2050]" count="1" pageId="11" pageNumber="290" type="female">♀</specimenCount>
// det.
<determinerName id="D8D3DA327F5FFF97FA1EA980FF15F7CF" pageId="11" pageNumber="290">Z. Zhan.</determinerName>
</materialsCitation>
</paragraph>
<caption id="79CF667E7F58FF90FF11A80CFBC0F976" ID-DOI="http://doi.org/10.5281/zenodo.10941634" ID-Zenodo-Dep="10941634" httpUri="https://zenodo.org/record/10941634/files/figure.png" pageId="12" pageNumber="291" startId="12.[155,227,1639,1661]" targetBox="[195,1392,244,1600]" targetPageId="12" targetType="figure">
<paragraph id="2D0F36F67F58FF90FF11A80CFBC0F976" blockId="12.[155,1432,1639,1693]" pageId="12" pageNumber="291">
Figures 15–21. Habitus of
<taxonomicName id="EAB04D757F58FF90FE15A803FDA8F996" authorityName="Nagai" authorityYear="2000" box="[415,635,1639,1661]" class="Insecta" family="Lucanidae" genus="Lucanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="291" phylum="Arthropoda" rank="subSpecies" species="derani" subSpecies="derani">
<emphasis id="1FC4EAE47F58FF90FE15A803FDA8F996" box="[415,635,1639,1661]" italics="true" pageId="12" pageNumber="291">Lucanus derani derani</emphasis>
</taxonomicName>
. 15. Male, dorsal view. 16. Male, ventral view. 17. Male, lateral view. 18. Median male. 19. Minor male. 20. Female, dorsal view. 21. Female, ventral view. Scale bar =10 mm.
</paragraph>
</caption>
<paragraph id="2D0F36F67F58FF90FF37A8B1FD7CF918" blockId="12.[136,1452,1754,2090]" box="[189,687,1754,1778]" pageId="12" pageNumber="291">
Body length 33.3–48.
<quantity id="EA489B137F58FF90FE22A8B0FE31F918" box="[424,482,1755,1778]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="291" unit="mm" value="7.0">7 mm</quantity>
(
<specimenCount id="3BB6FD7F7F58FF90FE7BA8B0FDD6F918" box="[497,517,1755,1778]" count="1" pageId="12" pageNumber="291" type="male">♂</specimenCount>
);
<quantity id="EA489B137F58FF90FD97A8B1FDA8F918" box="[541,635,1754,1778]" metricMagnitude="-2" metricUnit="m" metricValue="2.59" pageId="12" pageNumber="291" unit="mm" value="25.9">25. 9 mm</quantity>
(
<specimenCount id="3BB6FD7F7F58FF90FD01A8B0FD4CF918" box="[651,671,1755,1778]" count="1" pageId="12" pageNumber="291" type="female">♀</specimenCount>
).
</paragraph>
<paragraph id="2D0F36F67F58FF91FF37A896FC1EFD4E" blockId="12.[136,1452,1754,2090]" lastBlockId="13.[136,1452,236,1161]" lastPageId="13" lastPageNumber="292" pageId="12" pageNumber="291">Description. Major male. Head and pronotum reddish to brownish, covered with a thick vestiture of long, yellowish-amber setae; elytra covered with vestiture of short, yellowish-amber setae; elytra brownish, lighter than color of head and pronotum, having black stripes along sutural and lateral margins; head, including mandibles, 1.2× width of pronotum and elytra. Head reddish to brownish; mandibles weakly incurved along basal 1/3, straight at middle and weakly incurved along apical 1/3; major inner mandibular tooth located 1/3 distance to apex, triangular protruding forward and inflated on either side, 3–5 separated, small inner mandibular teeth located below major inner mandibular tooth; 2–3 sharp small inner mandibular teeth located between major tooth and mandibular apex; apical mandibular fork strongly developed, widely open from front, upper branch of apical fork 2× length of lower apical branch fork; lateral ridges of head subparallel, weakly rounded, evenly convex at end of head; clypeolabrum weakly developed, flattened, single-pointed along midlength. Pronotum brownish with metallic luster, usually darker than head; straight along apex and basal 1/3, weakly rounded at midlength; length 0.8× of head and mandibles; prothoracic tibiae yellowish-amber, bearing five well developed spines from apex to midlength; mesothoracic tibiae bright yellowish, each bearing two spines along midlength; metathoracic tibiae bright yellowish, black along apex, smooth and without any spines along basal 1/3. Elytra brownish but less mat than head and pronotum, covered with a thin vestiture of short, yellowish-amber setae, less setose than head and pronotum; lateral elytral margins and suture blackish; oval, widest along apical 1/3, strongly narrow along basal 1/3. Median male as major male except: head with major inner mandibular tooth single-pointed, weakly triangular pointing forward; &lt;4 small inner mandibular teeth located below major tooth, 1–2 sharp small inner mandibular teeth located between major tooth and mandibular apex. Minor male as major and median males except: head with major inner mandibular tooth weakly developed to absent, if present, single-pointed and not triangular; &lt;4 small inner mandibular teeth located within mandibular margins. Genitalia. Ventral plate of 9th abdominal segment abruptly widened behind cephalic base and constricted before caudal expansion; basal piece usually well defined, 0.4–0.5× length of entire aedeagus; apical flagellum enlarged, flagellum 1.1× length of aedeagus; aedeagus slender, incurved at basal 1/3, triangular distally.</paragraph>
<paragraph id="2D0F36F67F59FF91FF37ACC4FE32FC15" blockId="13.[136,1452,236,1161]" pageId="13" pageNumber="292">Female. Body dark brownish, covered with vestiture of long yellowish-amber setae; usually smaller than major and median males, subequal in length or larger than minor male. Head brownish; surface clearly punctate; mandibles incurved along basal 1/3 and apical 1/3; mandibular tooth weakly developed, asymmetrical; right mandibular tooth triangular; left mandibular tooth smooth; preocular and postocular margins weakly developed, not forming acute angle or protruding apically; clypeolabrum weakly developed, triangular, single-pointed at middle. Pronotum brownish, with metallic luster; obscurely punctate; prothoracic tibiae reddish, punctate, each bearing four spines from apex to midlength; distal two spines strongly developed; prothoracic tibiae 1.2–1.4× width of mesothoracic and metathoracic tibiae; mesothoracic tibiae reddish to yellowish-amber, each bearing a single spine along midlength; metathoracic tibiae reddish to yellowish-amber, bearing a single spine along basal 1/3. Elytra with apices brownish to black, disc with metallic luster; oval, widest at apical 1/3, strongly narrow along basal 1/3.</paragraph>
<paragraph id="2D0F36F67F59FF91FF37AA62FE3CFBC8" blockId="13.[136,1452,236,1161]" box="[189,495,1033,1058]" pageId="13" pageNumber="292">
Distribution.
<collectingCountry id="55A776667F59FF91FEC6AA62FE5FFBCB" box="[332,396,1033,1057]" name="China" pageId="13" pageNumber="292">China</collectingCountry>
(Yunan).
</paragraph>
<paragraph id="2D0F36F67F59FF91FF37AA47FBB8FBAF" blockId="13.[136,1452,236,1161]" box="[189,1131,1068,1093]" pageId="13" pageNumber="292">
Phenology. Adult specimens of
<taxonomicName id="EAB04D757F59FF91FD9AAA46FD6DFBAE" authorityName="Nagai" authorityYear="2000" box="[528,702,1068,1092]" class="Insecta" family="Lucanidae" genus="Lucanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="292" phylum="Arthropoda" rank="species" species="derani">
<emphasis id="1FC4EAE47F59FF91FD9AAA46FD6DFBAE" box="[528,702,1068,1092]" italics="true" pageId="13" pageNumber="292">L. derani derani</emphasis>
</taxonomicName>
were collected during mid- to late May.
</paragraph>
<paragraph id="2D0F36F67F59FF91FF37AA24FB6EFB8D" blockId="13.[136,1452,236,1161]" box="[189,1213,1103,1127]" pageId="13" pageNumber="292">
Natural history. Host plant data for
<taxonomicName id="EAB04D757F59FF91FDB2AA24FD35FB8D" authorityName="Nagai" authorityYear="2000" box="[568,742,1103,1127]" class="Insecta" family="Lucanidae" genus="Lucanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="292" phylum="Arthropoda" rank="species" species="derani">
<emphasis id="1FC4EAE47F59FF91FDB2AA24FD35FB8D" box="[568,742,1103,1127]" italics="true" pageId="13" pageNumber="292">L. derani derani</emphasis>
</taxonomicName>
is unknown. Adult longevity is 10–13 days.
</paragraph>
<paragraph id="2D0F36F67F59FF91FF37AA1AFC97FB63" blockId="13.[136,1452,236,1161]" box="[189,836,1137,1161]" pageId="13" pageNumber="292">Collecting methods. Light-trap, hand, and net-trap collected.</paragraph>
</subSubSection>
</treatment>
</document>
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