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<document id="0883195E889CB1AF2A07FD2A3CD8D392" ID-CLB-Dataset="32076" ID-DOI="10.11646/zootaxa.4320.3.6" ID-GBIF-Dataset="e80fc464-0c2a-4003-a867-ab8b5b3b399e" ID-ISSN="1175-5326" ID-Zenodo-Dep="1051951" ID-ZooBank="Ca50E5A5-B417-4000-B67D-5F6028Da9154" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505712517045" checkinUser="plazi" docAuthor="Bocak, Ladislav" docDate="2017" docId="A312190FEC1CFF89509F3840B89FFEC9" docLanguage="en" docName="zootaxa.4320.3.6.pdf" docOrigin="Zootaxa 4320 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cladophorus pallescens Bocak 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="515" masterDocId="5F2B6177EC15FF8350083D11BB5CFFD4" masterDocTitle="Species delimitation of colour polymorphic Cladophorus (Coleoptera: Lycidae) from New Guinea" masterLastPageNumber="522" masterPageNumber="505" pageNumber="514" updateTime="1698468596363" updateUser="ExternalLinkService">
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<mods:title id="499BDDC6BEF74CE53376902FF4289755">Species delimitation of colour polymorphic Cladophorus (Coleoptera: Lycidae) from New Guinea</mods:title>
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<mods:namePart id="979D78284943532B2525A521D50B0F2C">Bocak, Ladislav</mods:namePart>
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<mods:date id="0871DC71A0D3CE1B957B198616BBECD2">2017</mods:date>
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<mods:number id="E8267206AD040589F856B8076817274D">2017-09-18</mods:number>
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<mods:identifier id="28DD0D583360E20DD2B0E98A28525901" type="DOI">10.11646/zootaxa.4320.3.6</mods:identifier>
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<treatment id="A312190FEC1CFF89509F3840B89FFEC9" ID-DOI="http://doi.org/10.5281/zenodo.6018077" ID-GBIF-Taxon="134090574" ID-Zenodo-Dep="6018077" LSID="urn:lsid:plazi:treatment:A312190FEC1CFF89509F3840B89FFEC9" httpUri="http://treatment.plazi.org/id/A312190FEC1CFF89509F3840B89FFEC9" lastPageId="10" lastPageNumber="515" pageId="9" pageNumber="514">
<subSubSection id="63A1FB92EC1CFF8A509F3840BAF0FA58" pageId="9" pageNumber="514" type="nomenclature">
<paragraph id="2B04A819EC1CFF8A509F3840B947FABF" blockId="9.[151,539,1361,1421]" box="[151,539,1361,1387]" pageId="9" pageNumber="514">
<heading id="704C1F75EC1CFF8A509F3840B947FABF" bold="true" box="[151,539,1361,1387]" fontSize="11" level="1" pageId="9" pageNumber="514" reason="1">
<emphasis id="19CF740BEC1CFF8A509F3840B947FABF" bold="true" box="[151,539,1361,1387]" pageId="9" pageNumber="514">
<taxonomicName id="ECBBD39AEC1CFF8A509F3840BAEAFABF" authority="Bocak, 2017" authorityName="Bocak" authorityYear="2017" box="[151,438,1361,1387]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="pallescens" status="sp. nov.">
<emphasis id="19CF740BEC1CFF8A509F3840BAEAFABF" bold="true" box="[151,438,1361,1387]" italics="true" pageId="9" pageNumber="514">Cladophorus pallescens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02FCC970EC1CFF8A51B53840B947FABF" box="[445,539,1361,1387]" pageId="9" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="2B04A819EC1CFF8A509F3865BAF0FA58" blockId="9.[151,539,1361,1421]" box="[151,428,1396,1421]" pageId="9" pageNumber="514">
(
<figureCitation id="B380B49CEC1CFF8A50973865BA7AFA58" box="[159,294,1396,1421]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="9" pageNumber="514">Figs. 17, 29</figureCitation>
,
<figureCitation id="B380B49CEC1CFF8A513A3865BA0DFA59" box="[306,337,1396,1421]" captionStart="FIGURES 33 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 52. Elytron. 33 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="9" pageNumber="514">46</figureCitation>
,
<figureCitation id="B380B49CEC1CFF8A51553865BAF8FA58" box="[349,420,1396,1420]" captionStart="FIGURES 53 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 78. Male genitalia ventrally and laterally. 53 54, Cladophorus bicolor Kleine; 55 56, C. boceki sp. nov.; 57 58, C. craterensis sp. nov.; 59 60, C. haiaensis sp. nov.; 61 62, C. humeralis Kleine; 63 64, C. kailakiensis sp. nov.; 65 66, C. manokwarensis sp. nov.; 67 68, C. mindikensis sp. nov.; 69 70, C. motykai sp. nov.; 71 72, C. pallescens sp. nov.; 73 74, C. praecipuus Kleine; 75 76, C. riedeli sp. nov.; 77 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="9" pageNumber="514">7172</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC1CFF8A509F38ADB8E2FA2C" pageId="9" pageNumber="514" type="materials_examined">
<paragraph id="2B04A819EC1CFF8A509F38ADB8E2FA2C" blockId="9.[151,1437,1468,2033]" pageId="9" pageNumber="514">
<materialsCitation id="9BD3A244EC1CFF8A509F38ADBA3CFA2C" ID-GBIF-Occurrence="1638540616" collectionCode="LMBC" country="Indonesia" elevation="1825" latitude="-3.6666667" location="Male" longLatPrecision="1307" longitude="138.66667" pageId="9" pageNumber="514" specimenCount="2" stateProvince="Papua" typeStatus="holotype">
<emphasis id="19CF740BEC1CFF8A509F38ADBA1AFA01" bold="true" box="[151,326,1468,1493]" pageId="9" pageNumber="514">
<typeStatus id="F40016BBEC1CFF8A509F38ADBB8EFA01" box="[151,210,1468,1493]" pageId="9" pageNumber="514">Type</typeStatus>
material.
</emphasis>
<typeStatus id="F40016BBEC1CFF8A514638ADBAE6FA01" box="[334,442,1468,1493]" pageId="9" pageNumber="514" type="holotype">Holotype</typeStatus>
.
<location id="2E64FEC2EC1CFF8A51CD38ADB95DFA01" LSID="urn:lsid:plazi:treatment:A312190FEC1CFF89509F3840B89FFEC9:2E64FEC2EC1CFF8A51CD38ADB95DFA01" box="[453,513,1468,1493]" country="Indonesia" latitude="-3.6666667" longLatPrecision="1307" longitude="138.66667" name="Male" pageId="9" pageNumber="514" stateProvince="Papua">Male</location>
,
<collectingCountry id="53ACE889EC1CFF8A520638ADB922FA01" box="[526,638,1468,1493]" name="Indonesia" pageId="9" pageNumber="514">Indonesia</collectingCountry>
,
<collectingRegion id="E97F66FBEC1CFF8A528238ACB993FA00" box="[650,719,1469,1492]" country="Indonesia" name="Papua Barat" pageId="9" pageNumber="514">Papua</collectingRegion>
prov.,
<locationDeviation id="3AB6B13BEC1CFF8A531638ADBF49FA01" box="[798,1045,1468,1493]" pageId="9" pageNumber="514">
<quantity id="EC4305FCEC1CFF8A531638ADB804FA01" box="[798,856,1468,1493]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="514" unit="km" value="3.0">3 km</quantity>
N of Bokondini
</locationDeviation>
,
<geoCoordinate id="4E8FCEDEEC1CFF8A542938ADBF22FA00" box="[1057,1150,1468,1492]" direction="south" orientation="latitude" pageId="9" pageNumber="514" precision="925" value="-3.6666667">03°40´S</geoCoordinate>
<geoCoordinate id="4E8FCEDEEC1CFF8A548D38ADBFA8FA00" box="[1157,1268,1468,1492]" direction="east" orientation="longitude" pageId="9" pageNumber="514" precision="925" value="138.66667">138°40´E</geoCoordinate>
,
<elevation id="A0964F2AEC1CFF8A550838ADBEC0FA00" box="[1280,1436,1468,1493]" metricMagnitude="3" metricUnit="m" metricValue="1.825" metricValueMax="1.9" metricValueMin="1.75" pageId="9" pageNumber="514" unit="m" value="1825.0" valueMax="1900.0" valueMin="1750.0">
<quantity id="EC4305FCEC1CFF8A550838ADBEC0FA00" box="[1280,1436,1468,1493]" metricMagnitude="3" metricUnit="m" metricValue="1.825" metricValueMax="1.9" metricValueMin="1.75" pageId="9" pageNumber="514" unit="m" value="1825.0" valueMax="1900.0" valueMin="1750.0">17501900 m</quantity>
</elevation>
(JB0121,
<collectionCode id="4DAA30DCEC1CFF8A510C38F1BA04FA2C" box="[260,344,1504,1528]" pageId="9" pageNumber="514">LMBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9BD3A244EC1CFF8A516338F1B8E6FA2C" ID-GBIF-Occurrence="1638540608" box="[363,954,1503,1528]" collectionCode="LMBC" country="Indonesia" elevation="1825" latitude="-3.6666667" location="Male" longLatPrecision="1307" longitude="138.66667" pageId="9" pageNumber="514" specimenCount="1" stateProvince="Papua" typeStatus="paratype">
<typeStatus id="F40016BBEC1CFF8A516338F1BA8CFA2C" box="[363,464,1504,1528]" pageId="9" pageNumber="514" type="paratype">Paratype</typeStatus>
, male, same locality data (JB0103,
<collectionCode id="4DAA30DCEC1CFF8A535638F1B8EEFA2C" box="[862,946,1504,1528]" pageId="9" pageNumber="514">LMBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC1CFF8A50CF3B15BB8CF979" pageId="9" pageNumber="514" type="diagnosis">
<paragraph id="2B04A819EC1CFF8A50CF3B15BB8CF979" blockId="9.[151,1437,1468,2033]" pageId="9" pageNumber="514">
<emphasis id="19CF740BEC1CFF8A50CF3B15BA1DF9C9" bold="true" box="[199,321,1540,1565]" pageId="9" pageNumber="514">Diagnosis.</emphasis>
<taxonomicName id="ECBBD39AEC1CFF8A514F3B15BA80F9C8" authorityName="Bocak" authorityYear="2017" box="[327,476,1540,1564]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="pallescens">
<emphasis id="19CF740BEC1CFF8A514F3B15BA80F9C8" box="[327,476,1540,1564]" italics="true" pageId="9" pageNumber="514">C. pallescens</emphasis>
</taxonomicName>
is the only species with black, metallically shining pronotum and yellow elytra except for a humeral patch (
<figureCitation id="B380B49CEC1CFF8A51853B36BAB2F994" box="[397,494,1575,1600]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="9" pageNumber="514">Figs. 29</figureCitation>
,
<figureCitation id="B380B49CEC1CFF8A51F53B36B941F994" box="[509,541,1575,1600]" captionStart="FIGURES 33 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 52. Elytron. 33 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="9" pageNumber="514">46</figureCitation>
). The yellow form of closely related
<taxonomicName id="ECBBD39AEC1CFF8A53D93B39BF1AF994" box="[977,1094,1576,1600]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="19CF740BEC1CFF8A53D93B39BF1AF994" box="[977,1094,1576,1600]" italics="true" pageId="9" pageNumber="514">C. bicolor</emphasis>
</taxonomicName>
can be similar in the general appearance, but all available
<taxonomicName id="ECBBD39AEC1CFF8A51ED3B5DB905F9B0" box="[485,601,1612,1636]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="19CF740BEC1CFF8A51ED3B5DB905F9B0" box="[485,601,1612,1636]" italics="true" pageId="9" pageNumber="514">C. bicolor</emphasis>
</taxonomicName>
have at least a small apical part of elytra infuscate (
<figureCitation id="B380B49CEC1CFF8A54BD3B5DBE57F9B0" box="[1205,1291,1612,1637]" captionStart="FIGURES 33 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 52. Elytron. 33 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="9" pageNumber="514">Fig. 33</figureCitation>
). Further,
<taxonomicName id="ECBBD39AEC1CFF8A558A3B5DBA50F95C" authorityName="Bocak" authorityYear="2017" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="pallescens">
<emphasis id="19CF740BEC1CFF8A558A3B5DBA50F95C" italics="true" pageId="9" pageNumber="514">C. pallescens</emphasis>
</taxonomicName>
has a robust phallus with the short apex (
<figureCitation id="B380B49CEC1CFF8A52E93B7EB836F95C" box="[737,874,1647,1672]" captionStart="FIGURES 53 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 78. Male genitalia ventrally and laterally. 53 54, Cladophorus bicolor Kleine; 55 56, C. boceki sp. nov.; 57 58, C. craterensis sp. nov.; 59 60, C. haiaensis sp. nov.; 61 62, C. humeralis Kleine; 63 64, C. kailakiensis sp. nov.; 65 66, C. manokwarensis sp. nov.; 67 68, C. mindikensis sp. nov.; 69 70, C. motykai sp. nov.; 71 72, C. pallescens sp. nov.; 73 74, C. praecipuus Kleine; 75 76, C. riedeli sp. nov.; 77 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="9" pageNumber="514">Figs. 7172</figureCitation>
), long antennal lamellae (
<figureCitation id="B380B49CEC1CFF8A54853B7EBFBCF95C" box="[1165,1248,1647,1672]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="9" pageNumber="514">Fig. 15</figureCitation>
), and quite large eyes.
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC1CFF8950CF3BA6BF7DFF00" lastPageId="10" lastPageNumber="515" pageId="9" pageNumber="514" type="description">
<paragraph id="2B04A819EC1CFF8A50CF3BA6BE5BF824" blockId="9.[151,1437,1468,2033]" pageId="9" pageNumber="514">
<emphasis id="19CF740BEC1CFF8A50CF3BA6BA05F904" bold="true" box="[199,345,1719,1744]" pageId="9" pageNumber="514">Description.</emphasis>
Male. Body medium-sized,
<quantity id="EC4305FCEC1CFF8A529F3BA9B866F91B" box="[663,826,1720,1744]" metricMagnitude="-2" metricUnit="m" metricValue="1.045" metricValueMax="1.0699999999999998" metricValueMin="1.02" pageId="9" pageNumber="514" unit="mm" value="10.45" valueMax="10.7" valueMin="10.2">10.210.7 mm</quantity>
long, dark brown to black, pronotum and very narrow humeral margin of elytra dark coloured, rest of elytra light yellow (
<figureCitation id="B380B49CEC1CFF8A53A43BCDBF51F920" box="[940,1037,1756,1781]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="9" pageNumber="514">Figs. 29</figureCitation>
,
<figureCitation id="B380B49CEC1CFF8A54143BCDBF60F921" box="[1052,1084,1756,1781]" captionStart="FIGURES 33 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 52. Elytron. 33 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="9" pageNumber="514">46</figureCitation>
). Head small, with quite large hemispherically prominent eyes, frontal interocular distance 1.7 times maximum diameter; antennae flabellate, long, reaching to three quarters of elytral length if inclined, antennomere 3 with lamella 3.0 times longer than stem of antennomere, lamella attached to apex of antennomere (
<figureCitation id="B380B49CEC1CFF8A532D3A56B824F8B4" box="[805,888,1863,1888]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="9" pageNumber="514">Fig. 17</figureCitation>
). Pronotum transverse, 1.4 times wider than long at midline, lateral margins slightly elevated, frontal margin moderately projected, frontal angles sharply marked, lateral margins straight, with small bump in posterior third, posterior angles projected, all carinae well-developed (
<figureCitation id="B380B49CEC1CFF8A50973AA5BBAAF818" box="[159,246,1972,1997]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="9" pageNumber="514">Fig. 29</figureCitation>
). Elytra flat, parallel-sided, transverse costae well-developed, regular, elytral areoles slightly transverse (
<figureCitation id="B380B49CEC1CFF8A50973AC9BBA8F825" box="[159,244,2008,2033]" captionStart="FIGURES 33 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 52. Elytron. 33 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="9" pageNumber="514">Fig. 45</figureCitation>
). Phallus robust, parallel-sided in apical half, with short slender apical process (
<figureCitation id="B380B49CEC1CFF8A54793AC9BFA7F824" box="[1137,1275,2008,2033]" captionStart="FIGURES 53 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 78. Male genitalia ventrally and laterally. 53 54, Cladophorus bicolor Kleine; 55 56, C. boceki sp. nov.; 57 58, C. craterensis sp. nov.; 59 60, C. haiaensis sp. nov.; 61 62, C. humeralis Kleine; 63 64, C. kailakiensis sp. nov.; 65 66, C. manokwarensis sp. nov.; 67 68, C. mindikensis sp. nov.; 69 70, C. motykai sp. nov.; 71 72, C. pallescens sp. nov.; 73 74, C. praecipuus Kleine; 75 76, C. riedeli sp. nov.; 77 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="9" pageNumber="514">Figs. 7172</figureCitation>
).
</paragraph>
<paragraph id="2B04A819EC1FFF8950CF3D86BF7DFF00" blockId="10.[151,1436,151,285]" pageId="10" pageNumber="515">
<emphasis id="19CF740BEC1FFF8950CF3D86BA21FF64" bold="true" box="[199,381,151,176]" pageId="10" pageNumber="515">Measurements.</emphasis>
BL
<quantity id="EC4305FCEC1FFF8951A63D89B90FFF7B" box="[430,595,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.045" metricValueMax="1.0699999999999998" metricValueMin="1.02" pageId="10" pageNumber="515" unit="mm" value="10.45" valueMax="10.7" valueMin="10.2">10.210.7 mm</quantity>
, PL
<quantity id="EC4305FCEC1FFF89528E3D89B870FF7B" box="[646,812,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="10" pageNumber="515" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.301.40 mm</quantity>
, PW
<quantity id="EC4305FCEC1FFF8953603D86BF51FF7B" box="[872,1037,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.9249999999999998" metricValueMax="1.95" metricValueMin="1.9" pageId="10" pageNumber="515" unit="mm" value="1.9249999999999998" valueMax="1.95" valueMin="1.9">1.901.95 mm</quantity>
, WH
<quantity id="EC4305FCEC1FFF8954463D86BFAFFF7B" box="[1102,1267,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.1749999999999998" metricValueMax="1.2" metricValueMin="1.15" pageId="10" pageNumber="515" unit="mm" value="1.1749999999999998" valueMax="1.2" valueMin="1.15">1.151.20 mm</quantity>
, Edist
<quantity id="EC4305FCEC1FFF8955373D89BEC4FF7B" box="[1343,1432,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="515" unit="mm" value="0.8">0.8 mm</quantity>
, Ediam
<quantity id="EC4305FCEC1FFF8950EF3DADBA11FF00" box="[231,333,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="10" pageNumber="515" unit="mm" value="0.47">0.47 mm</quantity>
, LE
<quantity id="EC4305FCEC1FFF89518A3DADB957FF00" box="[386,523,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" metricValueMax="8.9" metricValueMin="8.5" pageId="10" pageNumber="515" unit="mm" value="8.7" valueMax="8.9" valueMin="8.5">8.58.9 mm</quantity>
, antennomere 3: stem
<quantity id="EC4305FCEC1FFF89530F3DADB803FF00" box="[775,863,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="515" unit="mm" value="0.8">0.8 mm</quantity>
, lamella
<quantity id="EC4305FCEC1FFF8953CC3DADBF41FF00" box="[964,1053,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="515" unit="mm" value="2.4">2.4 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC1FFF8950CF3DCEB82EFF2C" box="[199,882,223,248]" pageId="10" pageNumber="515" type="distribution">
<paragraph id="2B04A819EC1FFF8950CF3DCEB82EFF2C" blockId="10.[151,1436,151,285]" box="[199,882,223,248]" pageId="10" pageNumber="515">
<emphasis id="19CF740BEC1FFF8950CF3DCEBA3CFF2C" bold="true" box="[199,352,223,248]" pageId="10" pageNumber="515">Distribution.</emphasis>
<collectingRegion id="E97F66FBEC1FFF89516F3DCEBAE5FF2C" box="[359,441,223,248]" country="Papua New Guinea" name="Central" pageId="10" pageNumber="515">Central</collectingRegion>
New
<collectingCountry id="53ACE889EC1FFF8951F43DCEB90CFF2C" box="[508,592,223,248]" name="Guinea-Bissau" pageId="10" pageNumber="515">Guinea</collectingCountry>
, Tolikara regency (
<figureCitation id="B380B49CEC1FFF89532F3DCEB835FF2C" box="[807,873,223,248]" captionStart="FIGURES 1 3. 1 2" captionStartId="2.[151,264,1396,1419]" captionTargetBox="[151,1436,339,1375]" captionTargetId="figure@2.[151,1436,339,1375]" captionTargetPageId="2" captionText="FIGURES 1 3. 1 2, General appearance of Cladophorus bicolor in the Central mountains of New Guinea. 3, Sampled localities and the distribution of Cladophorus species" httpUri="https://zenodo.org/record/1051953/files/figure.png" pageId="10" pageNumber="515">Fig. 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC1FFF8950CF3C15B89FFEC9" box="[199,963,260,285]" pageId="10" pageNumber="515" type="etymology">
<paragraph id="2B04A819EC1FFF8950CF3C15B89FFEC9" blockId="10.[151,1436,151,285]" box="[199,963,260,285]" pageId="10" pageNumber="515">
<emphasis id="19CF740BEC1FFF8950CF3C15BA12FEC9" bold="true" box="[199,334,260,285]" pageId="10" pageNumber="515">Etymology.</emphasis>
The species name refers to the pale coloration of elytra.
</paragraph>
</subSubSection>
</treatment>
</document>