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<mods:title id="711C5331930535083DD53AD88CC838A4">Species delimitation of colour polymorphic Cladophorus (Coleoptera: Lycidae) from New Guinea</mods:title>
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<treatment id="A312190FEC16FF87509F3897B9DBFDE8" ID-DOI="http://doi.org/10.5281/zenodo.6018010" ID-GBIF-Taxon="134090568" ID-Zenodo-Dep="6018010" LSID="urn:lsid:plazi:treatment:A312190FEC16FF87509F3897B9DBFDE8" httpUri="http://treatment.plazi.org/id/A312190FEC16FF87509F3897B9DBFDE8" lastPageId="4" lastPageNumber="509" pageId="3" pageNumber="508">
<subSubSection id="63A1FB92EC16FF80509F3897B93DFA75" box="[151,609,1414,1441]" pageId="3" pageNumber="508" type="nomenclature">
<paragraph id="2B04A819EC16FF80509F3897B93DFA75" blockId="3.[151,609,1414,1441]" box="[151,609,1414,1441]" pageId="3" pageNumber="508">
<heading id="704C1F75EC16FF80509F3897B93DFA75" bold="true" box="[151,609,1414,1441]" fontSize="11" level="1" pageId="3" pageNumber="508" reason="1">
<taxonomicName id="ECBBD39AEC16FF80509F3897B93DFA75" authority="Guerin-Meneville, 1830" authorityName="Guerin-Meneville" authorityYear="1830" box="[151,609,1414,1441]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="19CF740BEC16FF80509F3897B93DFA75" bold="true" box="[151,609,1414,1441]" pageId="3" pageNumber="508">
<emphasis id="19CF740BEC16FF80509F3897BA6FFA74" bold="true" box="[151,307,1414,1440]" italics="true" pageId="3" pageNumber="508">Cladophorus</emphasis>
<bibRefCitation id="4F2AD5E8EC16FF8051323896B93DFA75" author="Guerin-Meneville" box="[314,609,1415,1441]" pageId="3" pageNumber="508" refString="Guerin-Meneville, F. C. (1830) Crustaces, arachnides et insectes. In: Duperrey, L. I. (Ed.), Voyage autour du monde ... sur la Corvette de Sa Majeste, La Coquille, pendant les annes 1822, 1823, 1824 et 1825 ... Zoologie. Vol. 2. Part 2. Bertrand, Paris, pl. II, fig. 9." type="journal article" year="1830">Guérin-Méneville, 1830</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC16FF80509F38DDBE65F9CB" pageId="3" pageNumber="508" type="reference_group">
<paragraph id="2B04A819EC16FF80509F38DDB9E7FA36" blockId="3.[151,1337,1484,1567]" box="[151,699,1484,1506]" pageId="3" pageNumber="508">
<taxonomicName id="ECBBD39AEC16FF80509F38DDB946FA36" authority="Guerin-Meneville, 1830" authorityName="Guerin-Meneville" authorityYear="1830" box="[151,538,1484,1506]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="19CF740BEC16FF80509F38DDBA40FA36" box="[151,284,1484,1506]" italics="true" pageId="3" pageNumber="508">Cladophorus</emphasis>
<bibRefCitation id="4F2AD5E8EC16FF80512938DDB946FA36" author="Guerin-Meneville" box="[289,538,1484,1506]" pageId="3" pageNumber="508" refString="Guerin-Meneville, F. C. (1830) Crustaces, arachnides et insectes. In: Duperrey, L. I. (Ed.), Voyage autour du monde ... sur la Corvette de Sa Majeste, La Coquille, pendant les annes 1822, 1823, 1824 et 1825 ... Zoologie. Vol. 2. Part 2. Bertrand, Paris, pl. II, fig. 9." type="journal article" year="1830">Guérin-Méneville, 1830</bibRefCitation>
</taxonomicName>
: Plate II,
<figureCitation id="B380B49CEC16FF80527238DDB9EBFA36" box="[634,695,1484,1506]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="3" pageNumber="508">Fig. 9</figureCitation>
.
</paragraph>
<paragraph id="2B04A819EC16FF80509F38FABE65F9CB" blockId="3.[151,1337,1484,1567]" pageId="3" pageNumber="508">
<emphasis id="19CF740BEC16FF80509F38FABA77F9D5" box="[151,299,1515,1537]" italics="true" pageId="3" pageNumber="508">=Odontocerus</emphasis>
<bibRefCitation id="4F2AD5E8EC16FF80513938FAB975F9D5" author="Guerin-Meneville" box="[305,553,1515,1537]" pageId="3" pageNumber="508" refString="Guerin-Meneville, F. C. (1838) Crustaces, arachnides et insectes. In: Duperrey, L. I. (Ed.), Voyage autour du monde ... sur la Corvette de Sa Majeste, La Coquille, pendant les annes 1822, 1823, 1824 et 1825 ... Zoologie. Vol. 2. Part 2. Bertrand, Paris, pp. 24 - 76." type="book chapter" year="1838">Guérin-Méneville, 1838</bibRefCitation>
: 72 (a homonym of
<emphasis id="19CF740BEC16FF8052FD38FAB824F9D5" box="[757,888,1515,1537]" italics="true" pageId="3" pageNumber="508">Odontocerus</emphasis>
Stephens, 1829 (Trichoptera)) =
<taxonomicName id="ECBBD39AEC16FF8050AD3B18BAE2F9CB" authority="Strand, 1936: 169" authorityName="Strand" authorityPageNumber="169" authorityYear="1936" box="[165,446,1545,1567]" class="Insecta" family="Lycidae" genus="Spacekia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="19CF740BEC16FF8050AD3B18BA5CF9CB" box="[165,256,1545,1567]" italics="true" pageId="3" pageNumber="508">Spacekia</emphasis>
<bibRefCitation id="4F2AD5E8EC16FF80510F3B18BAD7F9CB" author="Strand" box="[263,395,1545,1567]" pageId="3" pageNumber="508" refString="Strand, E. (1936) Miscellanea nomenclatorica zoologica et paleaontologica. Folia Zoologica et Hydrobiologica, 9, 167 - 170." type="journal article" year="1936">Strand, 1936</bibRefCitation>
: 169
</taxonomicName>
;
<bibRefCitation id="4F2AD5E8EC16FF8051C03B18B919F9CB" author="Bocak" box="[456,581,1545,1567]" pageId="3" pageNumber="508" refString="Bocak, L. (1998) Nomenclatural notes on taxa of the family Lycidae described by Guerin-Meneville (Insecta: Coleoptera). Annales Zoologici, 48, 245 - 251." type="journal article" year="1998">Bocak 1998</bibRefCitation>
: 247 (an objective synonym of
<taxonomicName id="ECBBD39AEC16FF80538C3B18BE72F9CB" authority="Guerin-Meneville, 1830" authorityName="Guerin-Meneville" authorityYear="1830" box="[900,1326,1545,1567]" class="Insecta" family="Lycidae" genus="Metriorrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="19CF740BEC16FF80538C3B18BF73F9CB" box="[900,1071,1545,1567]" italics="true" pageId="3" pageNumber="508">Metriorrhynchus</emphasis>
<bibRefCitation id="4F2AD5E8EC16FF80543D3B18BE72F9CB" author="Guerin-Meneville" box="[1077,1326,1545,1567]" pageId="3" pageNumber="508" refString="Guerin-Meneville, F. C. (1830) Crustaces, arachnides et insectes. In: Duperrey, L. I. (Ed.), Voyage autour du monde ... sur la Corvette de Sa Majeste, La Coquille, pendant les annes 1822, 1823, 1824 et 1825 ... Zoologie. Vol. 2. Part 2. Bertrand, Paris, pl. II, fig. 9." type="journal article" year="1830">Guérin-Méneville, 1830</bibRefCitation>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC16FF80509F3B5DB83FF8E9" pageId="3" pageNumber="508" type="diagnosis">
<paragraph id="2B04A819EC16FF80509F3B5DB83FF8E9" blockId="3.[151,1437,1612,2033]" pageId="3" pageNumber="508">
<emphasis id="19CF740BEC16FF80509F3B5DBA4DF9B1" bold="true" box="[151,273,1612,1637]" pageId="3" pageNumber="508">Diagnosis.</emphasis>
<taxonomicName id="ECBBD39AEC16FF8051123B5DBAF0F9B0" authorityName="Guerin-Meneville" authorityYear="1830" box="[282,428,1612,1636]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="19CF740BEC16FF8051123B5DBAF0F9B0" box="[282,428,1612,1636]" italics="true" pageId="3" pageNumber="508">Cladophorus</emphasis>
</taxonomicName>
is defined by long lamellae in male antennomeres 310, the characteristic shape of the wide, transverse pronotum, and the large, often brightly coloured body. The phallus reminds those of
<emphasis id="19CF740BEC16FF8054F43B61BEC7F953" box="[1276,1435,1648,1671]" italics="true" pageId="3" pageNumber="508">Cautiromimus</emphasis>
<bibRefCitation id="4F2AD5E8EC16FF80509F3B85BA77F979" author="Kleine" box="[151,299,1684,1709]" pageId="3" pageNumber="508" refString="Kleine, R. (1926) Coleoptera Lycidae. Nova Guinea. Resultats des expeditions scientifiques a la Nouvelle Guinee, 15, 91 - 195." type="journal article" year="1926">Kleine, 1926</bibRefCitation>
, it is usually robust and short and, unlike
<emphasis id="19CF740BEC16FF8052F73B84B8C2F978" box="[767,926,1685,1708]" italics="true" pageId="3" pageNumber="508">Cautiromimus</emphasis>
, does not have any clearly sclerotized part of the internal duct. The dissection of male genitalia is usually needed for the reliable identification.
<emphasis id="19CF740BEC16FF8054D73BA9BECDF904" box="[1247,1425,1720,1744]" italics="true" pageId="3" pageNumber="508">Metriorrhynchu</emphasis>
s Gemminger et Harold, 1 869 has prolonged pronotum and internal sac with multiple thorns or spirally coiled sclerotized structure in internal sac (
<bibRefCitation id="4F2AD5E8EC16FF80524B3A11B847F8CD" author="Bocak" box="[579,795,1792,1817]" pageId="3" pageNumber="508" refString="Bocak, L., Matsuda, K. &amp; Yagi, T. (2006) A revision of Metriorrhynchus from the Philippines with molecular evidence of an Australian origin of the Oriental Metriorrhynchus fauna (Coleoptera: Lycidae). European Journal of Entomology, 103, 115 - 126. https: // doi. org / 10.14411 / eje. 2006.017" type="journal article" year="2006">
Bocak
<emphasis id="19CF740BEC16FF8052903A10B991F8CC" box="[664,717,1792,1816]" italics="true" pageId="3" pageNumber="508">et al</emphasis>
. 2006
</bibRefCitation>
,
<bibRefCitation id="4F2AD5E8EC16FF8053233A11B8E7F8CC" author="Bocak" box="[811,955,1792,1817]" pageId="3" pageNumber="508" refString="Bocak, L. (2007) A Revision of Metriorrhynchus (Coleoptera: Lycidae) from the Greater Sunda Islands and Continental Asia. Raffles Bulletin of Zoology, 55, 253 - 260." type="journal article" year="2007">Bocak 2007</bibRefCitation>
).
<emphasis id="19CF740BEC16FF8053DB3A10BF04F8CC" box="[979,1112,1793,1816]" italics="true" pageId="3" pageNumber="508">Porrostoma</emphasis>
Castelnau, 1838 and
<emphasis id="19CF740BEC16FF8055543A10BEC0F8CC" box="[1372,1436,1793,1816]" italics="true" pageId="3" pageNumber="508">Ditua</emphasis>
<bibRefCitation id="4F2AD5E8EC16FF80509F3A35BA3CF8E9" author="Waterhouse" box="[151,352,1828,1853]" pageId="3" pageNumber="508" refString="Waterhouse, C. O. (1879) Illustrations of the typical specimens of Coleoptera in the collection of the British Museum. Part I - Lycidae. British Museum, London, 93 pp." type="book" year="1879">Waterhouse, 1879</bibRefCitation>
differ in serrate male antennae (
<bibRefCitation id="4F2AD5E8EC16FF8052C23A35B80BF8E9" author="Bocak" box="[714,855,1828,1853]" pageId="3" pageNumber="508" refString="Bocak, L. (2002) Generic revision and phylogenetic analysis of Metriorrhynchinae. European Journal of Entomology, 99, 315 - 351. https: // doi. org / 10.14411 / eje. 2002.043" type="journal article" year="2002">Bocak 2002</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC16FF8750CF3A59BF46FE04" lastPageId="4" lastPageNumber="509" pageId="3" pageNumber="508" type="description">
<paragraph id="2B04A819EC16FF8050CF3A59BEC1F825" blockId="3.[151,1437,1612,2033]" pageId="3" pageNumber="508">
<emphasis id="19CF740BEC16FF8050CF3A59BA28F8B5" bold="true" box="[199,372,1864,1889]" pageId="3" pageNumber="508">Redescription.</emphasis>
Body medium-sized to large,
<quantity id="EC4305FCEC16FF8052F13A59B8FAF8B4" box="[761,934,1864,1889]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" metricValueMax="1.61" metricValueMin="1.01" pageId="3" pageNumber="508" unit="mm" value="13.1" valueMax="16.1" valueMin="10.1">10.116.1 mm</quantity>
long, often brightly coloured; pronotum uniformly yellow or black coloured, elytra often bi-coloured; seldom dark coloured body parts with blue metallic shine. Head slightly transverse, cranium without rostrum, eyes relatively small, hemispherically prominent. Male antennomeres 310 flabellate, lamella of antennomere 3 1.3 to 4.7 times longer than its stem, lamellae of antennomeres 58 regularly slightly longer, up to six times longer than stem of corresponding antennomere (
<figureCitation id="B380B49CEC16FF80556C3AC9BEC1F825" box="[1380,1437,2008,2033]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="3" pageNumber="508">Figs.</figureCitation>
</paragraph>
<paragraph id="2B04A819EC11FF87509F3D86BF46FE04" blockId="4.[151,1436,151,573]" pageId="4" pageNumber="509">
619), antennal lamellae attached to basal or apical part of stem; maxillary palpi 4-segmented, stout, apical palpomere securiform, labial palpi 3-segmented, similar in shape to maxillary ones. Pronotum transverse, with clearly marked seven areoles, wide and rounded anteriorly, sometimes narrowed frontally and with prominent anterior angles (
<figureCitation id="B380B49CEC11FF8751463C15BA85FEC8" box="[334,473,260,285]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="4" pageNumber="509">Figs. 2032</figureCitation>
). Scutellum flat, emarginate at apex. Elytra slender, wider and longer than abdomen, with complete four primary and five secondary longitudinal costae (
<figureCitation id="B380B49CEC11FF8753DD3C36BF3AFE94" box="[981,1126,295,320]" captionStart="FIGURES 33 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 52. Elytron. 33 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="4" pageNumber="509">Figs. 3349</figureCitation>
), transverse costae dense, reticulate cells tiny, slightly to strongly transverse. Phallus usually stout, only
<taxonomicName id="ECBBD39AEC11FF87543D3C5DBF98FEB0" authorityName="Bocak" authorityYear="2017" box="[1077,1220,332,356]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="haiaensis">
<emphasis id="19CF740BEC11FF87543D3C5DBF98FEB0" box="[1077,1220,332,356]" italics="true" pageId="4" pageNumber="509">C. haiaensis</emphasis>
</taxonomicName>
with very slender phallus, gradually acuminate apically, internal sac partly sclerotized, without thorns, phallobasal membrane pigmented and only partly sclerotized (
<figureCitation id="B380B49CEC11FF87525F3C85B9BFFE79" box="[599,739,404,429]" captionStart="FIGURES 53 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 78. Male genitalia ventrally and laterally. 53 54, Cladophorus bicolor Kleine; 55 56, C. boceki sp. nov.; 57 58, C. craterensis sp. nov.; 59 60, C. haiaensis sp. nov.; 61 62, C. humeralis Kleine; 63 64, C. kailakiensis sp. nov.; 65 66, C. manokwarensis sp. nov.; 67 68, C. mindikensis sp. nov.; 69 70, C. motykai sp. nov.; 71 72, C. pallescens sp. nov.; 73 74, C. praecipuus Kleine; 75 76, C. riedeli sp. nov.; 77 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="4" pageNumber="509">Figs. 5378</figureCitation>
). Ovipositor with moderately wide coxites; valvifers slender; vagina membranous with short spermaduct and gradually widened spermatheca.
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC11FF8750CF3CCDB9DBFDE8" pageId="4" pageNumber="509" type="distribution">
<paragraph id="2B04A819EC11FF8750CF3CCDB9DBFDE8" blockId="4.[151,1436,151,573]" pageId="4" pageNumber="509">
<emphasis id="19CF740BEC11FF8750CF3CCDBA3CFE21" bold="true" box="[199,352,476,501]" pageId="4" pageNumber="509">Distribution.</emphasis>
Australian Region: New
<collectingCountry id="53ACE889EC11FF87528A3CCDB98AFE21" box="[642,726,476,501]" name="Guinea-Bissau" pageId="4" pageNumber="509">Guinea</collectingCountry>
, Yapen,
<collectingCountry id="53ACE889EC11FF87533F3CCDB8FDFE21" box="[823,929,476,501]" name="Australia" pageId="4" pageNumber="509">Australia</collectingCountry>
. The identity of the Australian species needs further investigation (see Introduction for details). All previous records outside these regions refer to other genera (
<bibRefCitation id="4F2AD5E8EC11FF8750963F35BA76FDE8" author="Bocak" box="[158,298,548,573]" pageId="4" pageNumber="509" refString="Bocak, L. (2002) Generic revision and phylogenetic analysis of Metriorrhynchinae. European Journal of Entomology, 99, 315 - 351. https: // doi. org / 10.14411 / eje. 2002.043" type="journal article" year="2002">Bocak 2002</bibRefCitation>
,
<bibRefCitation id="4F2AD5E8EC11FF87513D3F35B968FDE8" author="Sklenarova" box="[309,564,548,573]" pageId="4" pageNumber="509" refString="Sklenarova, K., Chesters, D. &amp; Bocak, L. (2013) Phylogeography of poorly dispersing net-winged beetles: A role of drifting India in the origin of Afrotropical and Oriental fauna. PLoS ONE, 8, e 67957. https: // doi. org / 10.1371 / journal. pone. 0067957" type="journal article" year="2013">
Sklenarova
<emphasis id="19CF740BEC11FF8751B23F34BAB7FDE8" box="[442,491,548,572]" italics="true" pageId="4" pageNumber="509">et al</emphasis>
. 2013
</bibRefCitation>
,
<bibRefCitation id="4F2AD5E8EC11FF8752373F35B920FDE8" author="Sklenarova" box="[575,636,548,572]" pageId="4" pageNumber="509" refString="Sklenarova, K., Kubecek, V. &amp; Bocak, L. (2014) Subtribal classification of Metriorrhynchini (Insecta: Coleoptera: Lycidae): an integrative approach using molecular phylogeny and morphology of adults and larvae. Arthropod Systematics &amp; Phylogeny, 72, 37 - 54." type="journal article" year="2014">2014</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>