205 lines
35 KiB
XML
205 lines
35 KiB
XML
<document id="C8A82757A07349E4ED73E8BE5E10BD74" ID-CLB-Dataset="32076" ID-DOI="10.11646/zootaxa.4320.3.6" ID-GBIF-Dataset="e80fc464-0c2a-4003-a867-ab8b5b3b399e" ID-ISSN="1175-5326" ID-Zenodo-Dep="1051951" ID-ZooBank="Ca50E5A5-B417-4000-B67D-5F6028Da9154" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505712517045" checkinUser="plazi" docAuthor="Bocak, Ladislav" docDate="2017" docId="A312190FEC12FF8B509F38DBBFEBFE21" docLanguage="en" docName="zootaxa.4320.3.6.pdf" docOrigin="Zootaxa 4320 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cladophorus kailakiensis Bocak 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="513" masterDocId="5F2B6177EC15FF8350083D11BB5CFFD4" masterDocTitle="Species delimitation of colour polymorphic Cladophorus (Coleoptera: Lycidae) from New Guinea" masterLastPageNumber="522" masterPageNumber="505" pageNumber="512" updateTime="1698468596363" updateUser="ExternalLinkService">
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<mods:title id="186A2E8D32697F33DB11E61179CC9C35">Species delimitation of colour polymorphic Cladophorus (Coleoptera: Lycidae) from New Guinea</mods:title>
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<mods:namePart id="AA53037FB871F193512B66CEDBAD1D86">Bocak, Ladislav</mods:namePart>
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<mods:title id="C2A54B2158F1D94735D7EF5EE64CCEEA">Zootaxa</mods:title>
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<mods:part id="637E4827076D6DC8FA5916FF95FEBA30">
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<mods:date id="E806F84E35D6DA90E138683A538C265D">2017</mods:date>
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<mods:number id="820F1CF987A741C9AC23B6946C397F56">2017-09-18</mods:number>
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<treatment id="A312190FEC12FF8B509F38DBBFEBFE21" ID-DOI="http://doi.org/10.5281/zenodo.6018049" ID-GBIF-Taxon="134090579" ID-Zenodo-Dep="6018049" LSID="urn:lsid:plazi:treatment:A312190FEC12FF8B509F38DBBFEBFE21" httpUri="http://treatment.plazi.org/id/A312190FEC12FF8B509F38DBBFEBFE21" lastPageId="8" lastPageNumber="513" pageId="7" pageNumber="512">
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<subSubSection id="63A1FB92EC12FF84509F38DBBAF7F9D2" pageId="7" pageNumber="512" type="nomenclature">
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<paragraph id="2B04A819EC12FF84509F38DBB973FA31" blockId="7.[151,559,1482,1542]" box="[151,559,1482,1509]" pageId="7" pageNumber="512">
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<heading id="704C1F75EC12FF84509F38DBB973FA31" bold="true" box="[151,559,1482,1509]" fontSize="11" level="1" pageId="7" pageNumber="512" reason="1">
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<emphasis id="19CF740BEC12FF84509F38DBB973FA31" bold="true" box="[151,559,1482,1509]" pageId="7" pageNumber="512">
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<taxonomicName id="ECBBD39AEC12FF84509F38DBBA96FA30" authority="Bocak, 2017" authorityName="Bocak" authorityYear="2017" box="[151,458,1482,1508]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="kailakiensis" status="sp. nov.">
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<emphasis id="19CF740BEC12FF84509F38DBBA96FA30" bold="true" box="[151,458,1482,1508]" italics="true" pageId="7" pageNumber="512">Cladophorus kailakiensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="02FCC970EC12FF8451D938DAB973FA31" box="[465,559,1483,1509]" pageId="7" pageNumber="512" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="2B04A819EC12FF84509F38FCBAF7F9D2" blockId="7.[151,559,1482,1542]" box="[151,427,1517,1542]" pageId="7" pageNumber="512">
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(
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<figureCitation id="B380B49CEC12FF84509738FCBA79F9D2" box="[159,293,1517,1542]" captionStart="FIGURES 6 – 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 – 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="7" pageNumber="512">Figs. 11, 25</figureCitation>
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,
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<figureCitation id="B380B49CEC12FF84513938FFBA13F9D2" box="[305,335,1518,1542]" captionStart="FIGURES 33 – 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 – 52. Elytron. 33 – 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 – 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="7" pageNumber="512">41</figureCitation>
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,
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<figureCitation id="B380B49CEC12FF84515338FCBAFFF9D2" box="[347,419,1517,1542]" captionStart="FIGURES 53 – 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 – 78. Male genitalia ventrally and laterally. 53 – 54, Cladophorus bicolor Kleine; 55 – 56, C. boceki sp. nov.; 57 – 58, C. craterensis sp. nov.; 59 – 60, C. haiaensis sp. nov.; 61 – 62, C. humeralis Kleine; 63 – 64, C. kailakiensis sp. nov.; 65 – 66, C. manokwarensis sp. nov.; 67 – 68, C. mindikensis sp. nov.; 69 – 70, C. motykai sp. nov.; 71 – 72, C. pallescens sp. nov.; 73 – 74, C. praecipuus Kleine; 75 – 76, C. riedeli sp. nov.; 77 – 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="7" pageNumber="512">63–64</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="63A1FB92EC12FF84509F3B24BA35F942" pageId="7" pageNumber="512" type="materials_examined">
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<paragraph id="2B04A819EC12FF84509F3B24BA35F942" blockId="7.[151,1437,1589,2010]" pageId="7" pageNumber="512">
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<materialsCitation id="9BD3A244EC12FF84509F3B24BA39F9A6" ID-GBIF-Occurrence="1638540599" collectionCode="LMBC" collectorName="Pt. Moresby" country="Papua New Guinea" elevation="735" latitude="-9.433333" location="Papua New Guinea" longLatPrecision="1299" longitude="147.55" pageId="7" pageNumber="512" specimenCount="2" typeStatus="holotype">
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<emphasis id="19CF740BEC12FF84509F3B24BA19F99A" bold="true" box="[151,325,1589,1614]" pageId="7" pageNumber="512">
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<typeStatus id="F40016BBEC12FF84509F3B24BB8EF99A" box="[151,210,1589,1614]" pageId="7" pageNumber="512">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="F40016BBEC12FF8451443B24BAE4F99A" box="[332,440,1589,1614]" pageId="7" pageNumber="512" type="holotype">Holotype</typeStatus>
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. Male,
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<collectingCountry id="53ACE889EC12FF8452023B27B9B9F99A" box="[522,741,1589,1614]" name="Papua New Guinea" pageId="7" pageNumber="512">Papua New Guinea</collectingCountry>
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,
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<collectorName id="864ECDCFEC12FF8452F93B27B827F99A" box="[753,891,1589,1614]" pageId="7" pageNumber="512">Pt. Moresby</collectorName>
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, Kailaki, Berem,
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||
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,
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<elevation id="A0964F2AEC12FF8455153B24BEC0F999" box="[1309,1436,1589,1614]" metricMagnitude="2" metricUnit="m" metricValue="7.35" metricValueMax="9.4" metricValueMin="5.3" pageId="7" pageNumber="512" unit="m" value="735.0" valueMax="940.0" valueMin="530.0">
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</elevation>
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(A00122,
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<collectionCode id="4DAA30DCEC12FF8451003B4BBA00F9A6" box="[264,348,1626,1650]" pageId="7" pageNumber="512">LMBC</collectionCode>
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)
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="9BD3A244EC12FF8451673B4BBA39F942" ID-GBIF-Occurrence="1638540613" collectionCode="LMBC" collectorName="Pt. Moresby & Varirata N. P." country="Papua New Guinea" elevation="750" latitude="-9.433333" location="Papua New Guinea" longLatPrecision="1299" longitude="147.36667" pageId="7" pageNumber="512" specimenCount="1" typeStatus="paratype">
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<typeStatus id="F40016BBEC12FF8451673B4BBA88F9A6" box="[367,468,1626,1650]" pageId="7" pageNumber="512" type="paratype">Paratype</typeStatus>
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, male,
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||
<collectingCountry id="53ACE889EC12FF84522C3B4BB9A3F9A6" box="[548,767,1625,1650]" name="Papua New Guinea" pageId="7" pageNumber="512">Papua New Guinea</collectingCountry>
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,
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<collectorName id="864ECDCFEC12FF8453013B4BB8C8F9A6" box="[777,916,1625,1650]" pageId="7" pageNumber="512">Pt. Moresby</collectorName>
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,
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<collectorName id="864ECDCFEC12FF8453963B48BF6AF9A5" box="[926,1078,1625,1650]" pageId="7" pageNumber="512">Varirata N. P.</collectorName>
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,
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<geoCoordinate id="4E8FCEDEEC12FF8454493B48BFC2F9A6" box="[1089,1182,1625,1650]" direction="south" orientation="latitude" pageId="7" pageNumber="512" precision="925" value="-9.433333">09°26´S</geoCoordinate>
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<geoCoordinate id="4E8FCEDEEC12FF8454AC3B4BBE4EF9A6" box="[1188,1298,1626,1650]" direction="east" orientation="longitude" pageId="7" pageNumber="512" precision="925" value="147.36667">147°22´E</geoCoordinate>
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,
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<elevation id="A0964F2AEC12FF8455153B4BBEC0F9A5" box="[1309,1436,1626,1650]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="7" pageNumber="512" unit="m" value="750.0" valueMax="800.0" valueMin="700.0">
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<quantity id="EC4305FCEC12FF8455153B4BBEC0F9A5" box="[1309,1436,1626,1650]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="7" pageNumber="512" unit="m" value="750.0" valueMax="800.0" valueMin="700.0">700–800 m</quantity>
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</elevation>
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(
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<accessionNumber id="34E835FAEC12FF8450963B6CBBA1F942" box="[158,253,1661,1686]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A00176" pageId="7" pageNumber="512" type="EnaNcbi">A00176</accessionNumber>
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,
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<collectionCode id="4DAA30DCEC12FF8451003B6FBA00F942" box="[264,348,1662,1686]" pageId="7" pageNumber="512">LMBC</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="63A1FB92EC12FF8450CF3BB0B8E9F89E" pageId="7" pageNumber="512" type="diagnosis">
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<paragraph id="2B04A819EC12FF8450CF3BB0B8E9F89E" blockId="7.[151,1437,1589,2010]" pageId="7" pageNumber="512">
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<emphasis id="19CF740BEC12FF8450CF3BB0BA1DF96E" bold="true" box="[199,321,1697,1722]" pageId="7" pageNumber="512">Diagnosis.</emphasis>
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In contrast with distant relationships,
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<taxonomicName id="ECBBD39AEC12FF84530A3BB3B8F7F96D" authorityName="Bocak" authorityYear="2017" box="[770,939,1697,1721]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="kailakiensis">
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<emphasis id="19CF740BEC12FF84530A3BB3B8F7F96D" box="[770,939,1697,1721]" italics="true" pageId="7" pageNumber="512">C. kailakiensis</emphasis>
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</taxonomicName>
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resembles in the general colour pattern
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<taxonomicName id="ECBBD39AEC12FF84558A3BB3BA18F909" authorityName="Bocak" authorityYear="2017" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="manokwarensis">
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<emphasis id="19CF740BEC12FF84558A3BB3BA18F909" italics="true" pageId="7" pageNumber="512">C. manokwarensis</emphasis>
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</taxonomicName>
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(
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<figureCitation id="B380B49CEC12FF8451523BD4BAF3F90A" box="[346,431,1733,1758]" captionStart="FIGURES 4 – 5. 4" captionStartId="3.[151,264,1220,1243]" captionTargetBox="[159,1417,255,1198]" captionTargetId="figure@3.[151,1436,255,1210]" captionTargetPageId="3" captionText="FIGURES 4 – 5. 4, Phylogenetic hypothesis of Papuan Cladophorus inferred from the cox 1 mtDNA dataset using maximum likelihood criterion. 5, Cladophorus sp., general appearance." httpUri="https://zenodo.org/record/1051955/files/figure.png" pageId="7" pageNumber="512">Figs. 4</figureCitation>
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,
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||
<figureCitation id="B380B49CEC12FF8451C93BD7B950F90A" box="[449,524,1734,1758]" captionStart="FIGURES 33 – 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 – 52. Elytron. 33 – 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 – 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="7" pageNumber="512">37–38</figureCitation>
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). Both species have brightly yellow coloured pronotum, the extensive black coloured humeral part of elytra and brightly coloured rest of elytra (
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<figureCitation id="B380B49CEC12FF8453C23BF8BF06F8D6" box="[970,1114,1769,1794]" captionStart="FIGURES 33 – 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 – 52. Elytron. 33 – 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 – 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="7" pageNumber="512">Figs. 41–42</figureCitation>
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). The molecular phylogeny identified
|
||
<taxonomicName id="ECBBD39AEC12FF8451033A1FBAEDF8F1" authorityName="Bocak" authorityYear="2017" box="[267,433,1805,1829]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="mindikensis">
|
||
<emphasis id="19CF740BEC12FF8451033A1FBAEDF8F1" box="[267,433,1805,1829]" italics="true" pageId="7" pageNumber="512">C. mindikensis</emphasis>
|
||
</taxonomicName>
|
||
as a sister species (
|
||
<figureCitation id="B380B49CEC12FF84529B3A1CB984F8F2" box="[659,728,1805,1830]" captionStart="FIGURES 4 – 5. 4" captionStartId="3.[151,264,1220,1243]" captionTargetBox="[159,1417,255,1198]" captionTargetId="figure@3.[151,1436,255,1210]" captionTargetPageId="3" captionText="FIGURES 4 – 5. 4, Phylogenetic hypothesis of Papuan Cladophorus inferred from the cox 1 mtDNA dataset using maximum likelihood criterion. 5, Cladophorus sp., general appearance." httpUri="https://zenodo.org/record/1051955/files/figure.png" pageId="7" pageNumber="512">Fig. 4</figureCitation>
|
||
) and these species differ in the elytral and pronotal coloration (
|
||
<figureCitation id="B380B49CEC12FF8450973A20BA7AF89E" box="[159,294,1841,1866]" captionStart="FIGURES 6 – 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 – 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="7" pageNumber="512">Figs. 25, 27</figureCitation>
|
||
,
|
||
<figureCitation id="B380B49CEC12FF84513A3A23BA26F89E" box="[306,378,1842,1866]" captionStart="FIGURES 33 – 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 – 52. Elytron. 33 – 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 – 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="7" pageNumber="512">41, 43</figureCitation>
|
||
) and the length of antennal lamellae (
|
||
<figureCitation id="B380B49CEC12FF8453293A20B8F4F89E" box="[801,936,1841,1866]" captionStart="FIGURES 6 – 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 – 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="7" pageNumber="512">Figs. 11, 13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="63A1FB92EC12FF8B50CF3A44B8A2FE78" lastPageId="8" lastPageNumber="513" pageId="7" pageNumber="512" type="description">
|
||
<paragraph id="2B04A819EC12FF8B50CF3A44B82EFEB1" blockId="7.[151,1437,1589,2010]" lastBlockId="8.[151,1437,151,501]" lastPageId="8" lastPageNumber="513" pageId="7" pageNumber="512">
|
||
<emphasis id="19CF740BEC12FF8450CF3A44BA05F8BA" bold="true" box="[199,345,1877,1902]" pageId="7" pageNumber="512">Description.</emphasis>
|
||
Male. Body medium-sized,
|
||
<quantity id="EC4305FCEC12FF8452B83A44B805F8B9" box="[688,857,1877,1902]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" metricValueMax="1.29" metricValueMin="1.05" pageId="7" pageNumber="512" unit="mm" value="11.7" valueMax="12.9" valueMin="10.5">10.5–12.9 mm</quantity>
|
||
long, body dark brown to black, pronotum and substantial part of elytra brightly yellow (
|
||
<figureCitation id="B380B49CEC12FF8452773A68B989F846" box="[639,725,1913,1938]" captionStart="FIGURES 33 – 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 – 52. Elytron. 33 – 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 – 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="7" pageNumber="512">Fig. 41</figureCitation>
|
||
), humeral tenth of elytra black, other parts of elytra brightly yellow (
|
||
<figureCitation id="B380B49CEC12FF8450FC3A8CBA15F862" box="[244,329,1949,1974]" captionStart="FIGURES 33 – 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 – 52. Elytron. 33 – 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 – 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="7" pageNumber="512">Fig. 41</figureCitation>
|
||
). Head small, with small, hemispherically prominent eyes, frontal interocular distance 1.87 times maximum eye diameter; antennae long, reaching beyond two thirds of elytral length, flabellate, antennomere 3 with lamella 2.56 times longer than stem of antennomere, lamella attached close to middle part of antennomere (
|
||
<figureCitation id="B380B49CEC1DFF8B55673D86BBE8FF00" captionStart="FIGURES 6 – 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 – 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="8" pageNumber="513">Fig. 11</figureCitation>
|
||
). Pronotum transverse, 1.3 times wider than long at midline, lateral margins elevated, frontal margin widely rounded, frontal angles inconspicuous, rounded, lateral margins slightly convex, posterior angles moderately prominent, postero-lateral carinae weak (
|
||
<figureCitation id="B380B49CEC1DFF8B526C3C15B9E5FEC9" box="[612,697,260,285]" captionStart="FIGURES 6 – 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 – 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="8" pageNumber="513">Fig. 25</figureCitation>
|
||
). Elytra flat, slightly widened posteriorly, transverse costae weak, regular, elytral areoles slightly transverse (
|
||
<figureCitation id="B380B49CEC1DFF8B529B3C36B9B1FE94" box="[659,749,295,320]" captionStart="FIGURES 33 – 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 – 52. Elytron. 33 – 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 – 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="8" pageNumber="513">Fig. 41</figureCitation>
|
||
). Phallus robust, parallel-sided in middle part, apparently constricted basally, with gradually acuminate apex (
|
||
<figureCitation id="B380B49CEC1DFF8B52D43C5DB83AFEB1" box="[732,870,332,357]" captionStart="FIGURES 53 – 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 – 78. Male genitalia ventrally and laterally. 53 – 54, Cladophorus bicolor Kleine; 55 – 56, C. boceki sp. nov.; 57 – 58, C. craterensis sp. nov.; 59 – 60, C. haiaensis sp. nov.; 61 – 62, C. humeralis Kleine; 63 – 64, C. kailakiensis sp. nov.; 65 – 66, C. manokwarensis sp. nov.; 67 – 68, C. mindikensis sp. nov.; 69 – 70, C. motykai sp. nov.; 71 – 72, C. pallescens sp. nov.; 73 – 74, C. praecipuus Kleine; 75 – 76, C. riedeli sp. nov.; 77 – 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="8" pageNumber="513">Figs. 63–64</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2B04A819EC1DFF8B50CF3C7EB8A2FE78" blockId="8.[151,1437,151,501]" pageId="8" pageNumber="513">
|
||
<emphasis id="19CF740BEC1DFF8B50CF3C7EBA21FE5C" bold="true" box="[199,381,367,392]" pageId="8" pageNumber="513">Measurements.</emphasis>
|
||
BL
|
||
<quantity id="EC4305FCEC1DFF8B51B83C7EB90AFE53" box="[432,598,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" metricValueMax="1.29" metricValueMin="1.05" pageId="8" pageNumber="513" unit="mm" value="11.7" valueMax="12.9" valueMin="10.5">10.5–12.9 mm</quantity>
|
||
, PL
|
||
<quantity id="EC4305FCEC1DFF8B52833C7EB849FE53" box="[651,789,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="8" pageNumber="513" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.5–1.8 mm</quantity>
|
||
, PW
|
||
<quantity id="EC4305FCEC1DFF8B535C3C7EB8A5FE53" box="[852,1017,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="2.425" metricValueMax="2.65" metricValueMin="2.2" pageId="8" pageNumber="513" unit="mm" value="2.425" valueMax="2.65" valueMin="2.2">2.20–2.65 mm</quantity>
|
||
, WH
|
||
<quantity id="EC4305FCEC1DFF8B54343C7EBFBEFE53" box="[1084,1250,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" metricValueMax="1.54" metricValueMin="1.3" pageId="8" pageNumber="513" unit="mm" value="1.42" valueMax="1.54" valueMin="1.3">1.30–1.54 mm</quantity>
|
||
, Edist
|
||
<quantity id="EC4305FCEC1DFF8B55383C61BEC4FE53" box="[1328,1432,368,392]" metricMagnitude="-4" metricUnit="m" metricValue="9.9" pageId="8" pageNumber="513" unit="mm" value="0.99">0.99 mm</quantity>
|
||
, Ediam
|
||
<quantity id="EC4305FCEC1DFF8B50EF3C85BA11FE78" box="[231,333,404,429]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="8" pageNumber="513" unit="mm" value="0.53">0.53 mm</quantity>
|
||
, LE
|
||
<quantity id="EC4305FCEC1DFF8B518A3C85BAB4FE78" box="[386,488,404,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="8" pageNumber="513" unit="mm" value="10.6">10.6 mm</quantity>
|
||
, antennomere 3: stem
|
||
<quantity id="EC4305FCEC1DFF8B52EC3C85B860FE78" box="[740,828,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="513" unit="mm" value="0.9">0.9 mm</quantity>
|
||
, lamella
|
||
<quantity id="EC4305FCEC1DFF8B53A93C85B8A6FE78" box="[929,1018,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="513" unit="mm" value="2.3">2.3 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="63A1FB92EC1DFF8B50CF3CA6BF99FE04" box="[199,1221,439,464]" pageId="8" pageNumber="513" type="distribution">
|
||
<paragraph id="2B04A819EC1DFF8B50CF3CA6BF99FE04" blockId="8.[151,1437,151,501]" box="[199,1221,439,464]" pageId="8" pageNumber="513">
|
||
<emphasis id="19CF740BEC1DFF8B50CF3CA6BA3CFE04" bold="true" box="[199,352,439,464]" pageId="8" pageNumber="513">Distribution.</emphasis>
|
||
Eastern New
|
||
<collectingCountry id="53ACE889EC1DFF8B51F63CA6B90EFE04" box="[510,594,439,464]" name="Guinea-Bissau" pageId="8" pageNumber="513">Guinea</collectingCountry>
|
||
, the southern slope of the Owen Stanley Range (
|
||
<figureCitation id="B380B49CEC1DFF8B547A3CA6BFEBFE04" box="[1138,1207,439,464]" captionStart="FIGURES 1 – 3. 1 – 2" captionStartId="2.[151,264,1396,1419]" captionTargetBox="[151,1436,339,1375]" captionTargetId="figure@2.[151,1436,339,1375]" captionTargetPageId="2" captionText="FIGURES 1 – 3. 1 – 2, General appearance of Cladophorus bicolor in the Central mountains of New Guinea. 3, Sampled localities and the distribution of Cladophorus species" httpUri="https://zenodo.org/record/1051953/files/figure.png" pageId="8" pageNumber="513">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="63A1FB92EC1DFF8B50CF3CCDBFEBFE21" box="[199,1207,476,501]" pageId="8" pageNumber="513" type="etymology">
|
||
<paragraph id="2B04A819EC1DFF8B50CF3CCDBFEBFE21" blockId="8.[151,1437,151,501]" box="[199,1207,476,501]" pageId="8" pageNumber="513">
|
||
<emphasis id="19CF740BEC1DFF8B50CF3CCDBA12FE21" bold="true" box="[199,334,476,501]" pageId="8" pageNumber="513">Etymology.</emphasis>
|
||
The epithet refers to the local name ‘Kailaki’, the
|
||
<typeStatus id="F40016BBEC1DFF8B538C3CCCB8E8FE21" box="[900,948,477,501]" pageId="8" pageNumber="513">type</typeStatus>
|
||
locality of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |