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<mods:title id="D0A385D465A1D0F31854ED1C21B8184D">New taxonomic and faunistic data on Liocranidae (Arachnida: Araneae) of West Palaearctic), with nine new species of Mesiotelus Simon, 1897</mods:title>
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<mods:namePart id="6C0EF855A0B57D6A67392340AE5605BC">Zamani, Alireza</mods:namePart>
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<mods:namePart id="1B7DFF1AD5E0DE1249D7AC919D8858CC">Fomichev, Alexander A.</mods:namePart>
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<mods:affiliation id="9E9FE71DE8D7E54C8718137B99CFE222">Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia.</mods:affiliation>
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<mods:namePart id="C50BC031F02D5A38727E2A0C39B3AED0">Naumova, Maria</mods:namePart>
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<mods:affiliation id="3FB6E6B726270B0246A43E509AF7AAE3">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd., 1000 Sofia, Bulgaria.</mods:affiliation>
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<mods:namePart id="406C855429F5F62C7E0139CF49F201B7">Kaya, Rahşen S.</mods:namePart>
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<mods:affiliation id="D9EB27C908B0644B98857A471B9BACE5">Department of Biology, Faculty of Arts and Science, Bursa Uludağ University, TR- 16059, Bursa, Türkiye.</mods:affiliation>
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<mods:affiliation id="F1400FBA4CDD3CB733B16100E35B7931">Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa. &amp; Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia. &amp; Institute for Biological Problems of the North, Portovaya Str. 18, Magadan 685000, Russia.</mods:affiliation>
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<taxonomicName id="EDC74D31FFB9FFC209F095BCD1D848D8" ID-CoL="98YST" authority="Zamani &amp; Marusik, 2021" authorityName="Zamani &amp; Marusik" authorityYear="2021" box="[151,707,1195,1222]" class="Arachnida" family="Liocranidae" genus="Mesiotelus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="202" phylum="Arthropoda" rank="species" species="patricki">
<emphasis id="18B3EAA0FFB9FFC209F095BCD1D848D8" bold="true" box="[151,707,1195,1222]" pageId="12" pageNumber="202">
<emphasis id="18B3EAA0FFB9FFC209F095BCD26548DB" bold="true" box="[151,382,1195,1222]" italics="true" pageId="12" pageNumber="202">Mesiotelus patricki</emphasis>
Zamani &amp; Marusik, 2021
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<figureCitation id="B2FC2A37FFB9FFC209F095D8D3F048F4" box="[151,235,1231,1257]" captionStart="FIGURES 712" captionStartId="3.[151,264,1969,1993]" captionTargetBox="[152,1436,635,1936]" captionTargetId="figure-223@3.[151,1436,633,1946]" captionTargetPageId="3" captionText="FIGURES 712. Dorsal habitus of Mesiotelus cyprius (7), M. damavandicus sp. nov. (8), M. kravetsi sp. nov. (9), M. mediterraneus sp. nov. (10), M. patricki (11) and M. sistanus sp. nov. (12). 78, 1012—female; 9—male. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13915747" httpUri="https://zenodo.org/record/13915747/files/figure.png" pageId="12" pageNumber="202">Figs 11</figureCitation>
,
<figureCitation id="B2FC2A37FFB9FFC2099095D8D22648F7" box="[247,317,1231,1258]" captionStart="FIGURES 5260" captionStartId="18.[151,264,1898,1922]" captionTargetBox="[151,1436,181,1875]" captionTargetId="figure-18@18.[151,1436,181,1875]" captionTargetPageId="18" captionText="FIGURES 5260. Epigyne (5253, 5560) and receptacles (54) of Mesiotelus kulczynskii (5254), M. mediterraneus sp. nov. (5557) and M. patricki (5860). 55, 58—intact, ventral; 52, 56, 59—macerated, ventral; 5354, 57, 60—macerated, dorsal. Scale bars: 0.1 mm. Abbreviations: Ah—anterior hood, Hl—lateral margin of anterior hood." figureDoi="http://doi.org/10.5281/zenodo.13915763" httpUri="https://zenodo.org/record/13915763/files/figure.png" pageId="12" pageNumber="202">5860</figureCitation>
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<taxonomicName id="EDC74D31FFB9FFC209F09403D2774936" ID-CoL="3ZZPF" authorityName="Drensky" authorityYear="1935" box="[151,364,1300,1324]" class="Arachnida" family="Liocranidae" genus="Mesiotelus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="202" phylum="Arthropoda" rank="species" species="scopensis">
<emphasis id="18B3EAA0FFB9FFC209F09403D2774936" box="[151,364,1300,1324]" italics="true" pageId="12" pageNumber="202">Mesiotelus scopensis</emphasis>
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:
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<bibRefCitation id="4E564B43FFB9FFC2081D9403D1774936" author="Zamani, A. &amp; Dimitrov, D. &amp; Weiss, I. &amp; Alimohammadi, S. &amp; Rafiei-Jahed, R. &amp; Esyunin, S. L. &amp; Moradmand, M. &amp; Chatzaki, M. &amp; Marusik, Y. M." box="[378,620,1299,1324]" pageId="12" pageNumber="202" pagination="569 - 591" refId="ref14711" refString="Zamani, A., Dimitrov, D., Weiss, I., Alimohammadi, S., Rafiei-Jahed, R., Esyunin, S. L., Moradmand, M., Chatzaki, M. &amp; Marusik, Y. M. (2020) New data on the spider fauna of Iran (Arachnida: Araneae), Part VII. Arachnology, 18 (6), 569 - 591. https: // doi. org / 10.13156 / arac. 2020.18.6.569" type="journal article" year="2020">
Zamani
<emphasis id="18B3EAA0FFB9FFC208AA9403D2E14931" box="[461,506,1300,1324]" italics="true" pageId="12" pageNumber="202">et al</emphasis>
. 2020: 584
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(in part).
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<treatmentCitation id="AB6610A3FFB9FFC209F09425D1BC4957" author="Zamani, A. &amp; Marusik, Y. M." box="[151,679,1330,1354]" page="110" pageId="12" pageNumber="202" year="2021">
<taxonomicName id="EDC74D31FFB9FFC209F09425D1BC4957" ID-CoL="98YST" authority="Zamani &amp; Marusik 2021 a: 110" authorityName="Zamani &amp; Marusik" authorityPageNumber="110" authorityYear="2021" box="[151,679,1330,1354]" class="Arachnida" family="Liocranidae" genus="Mesiotelus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="202" phylum="Arthropoda" rank="species" species="patricki">
<emphasis id="18B3EAA0FFB9FFC209F09425D2474957" box="[151,348,1330,1354]" italics="true" pageId="12" pageNumber="202">Mesiotelus patricki</emphasis>
<bibRefCitation id="4E564B43FFB9FFC208039425D1BC4957" author="Zamani, A. &amp; Marusik, Y. M." box="[356,679,1330,1354]" pageId="12" pageNumber="202" pagination="95 - 126" refId="ref14802" refString="Zamani, A. &amp; Marusik, Y. M. (2021 a) A new genus and ten new species of spiders (Arachnida, Araneae) from Iran. ZooKeys, 1054, 95 - 126. https: // doi. org / 10.3897 / zookeys. 1054.70408" type="journal article" year="2021">Zamani &amp; Marusik 2021a: 110</bibRefCitation>
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, figs 4EF, 11AE, 12AC (
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);
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<bibRefCitation id="4E564B43FFB9FFC20D679425D65D4957" author="Zamani, A. &amp; Marusik, Y. M." box="[1024,1350,1330,1354]" pageId="12" pageNumber="202" pagination="557 - 564" refId="ref14854" refString="Zamani, A. &amp; Marusik, Y. M. (2021 b) Two new species of Liocranidae (Arachnida: Aranei) from the Caucasus and northern Iran. Arthropoda Selecta, 30 (4), 557 - 564. https: // doi. org / 10.15298 / arthsel. 30.4.12" type="journal article" year="2021">Zamani &amp; Marusik 2021b: 560</bibRefCitation>
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, fig. 4H (
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).
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<emphasis id="18B3EAA0FFB9FFC209F09480D21D49AC" bold="true" box="[151,262,1431,1457]" pageId="12" pageNumber="202">Material.</emphasis>
<materialsCitation id="9AAF3CEFFFB9FFC208699480D67D49AC" box="[270,1382,1431,1458]" collectingDate="1974-07-13" collectionCode="MHNG" collectorName="A. Senglet" country="Iran" latitude="37.383335" location="Golestan Prov." longLatPrecision="1181" longitude="55.833332" pageId="12" pageNumber="202" specimenCount="1" specimenCount-female="1" stateProvince="Golestan">
<collectingCountry id="52D07622FFB9FFC208699480D24E49AC" box="[270,341,1431,1457]" name="Iran" pageId="12" pageNumber="202">IRAN</collectingCountry>
:
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<collectingRegion id="E803F850FFB9FFC208029480D11749AC" box="[357,524,1431,1457]" country="Iran" name="Golestan" pageId="12" pageNumber="202">Golestan Prov.</collectingRegion>
</emphasis>
:
<specimenCount id="3CC1FD3BFFB9FFC20B7D9480D12649AC" box="[538,573,1431,1457]" count="1" pageId="12" pageNumber="202" type="female">1♀</specimenCount>
(
<collectionCode id="4CD6AE77FFB9FFC20B2C9480D1BD49AC" box="[587,678,1431,1457]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="12" pageNumber="202" type="Museum">MHNG</collectionCode>
), Tang-e Rah,
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,
<geoCoordinate id="4FF35075FFB9FFC20AD49480D71649AF" box="[947,1037,1431,1458]" degrees="55" direction="east" minutes="50" orientation="longitude" pageId="12" pageNumber="202" precision="925" value="55.833332">55°50'E</geoCoordinate>
,
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<collectingDate id="4E3DE99AFFB9FFC20D7C9480D7B849AC" box="[1051,1187,1431,1457]" pageId="12" pageNumber="202" value="1974-07-13">13.VII.1974</collectingDate>
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(leg.
<collectorName id="87325364FFB9FFC20D859480D64549AC" box="[1250,1374,1431,1458]" pageId="12" pageNumber="202">A. Senglet</collectorName>
)
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.
</paragraph>
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<emphasis id="18B3EAA0FFB9FFC209A094ACD25949C8" bold="true" box="[199,322,1467,1493]" pageId="12" pageNumber="202">Diagnosis.</emphasis>
The epigyne of this species is somewhat similar to that of
<taxonomicName id="EDC74D31FFB9FFC20AB994ABD79B49C8" authorityName="Charitonov" authorityYear="1946" box="[990,1152,1467,1493]" class="Arachnida" family="Liocranidae" genus="Mesiotelus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="202" phylum="Arthropoda" rank="species" species="kulczynskii">
<emphasis id="18B3EAA0FFB9FFC20AB994ABD79B49C8" box="[990,1152,1467,1493]" italics="true" pageId="12" pageNumber="202">M. kulczynskii</emphasis>
</taxonomicName>
(
<figureCitation id="B2FC2A37FFB9FFC20DE894ACD60849CB" box="[1167,1299,1467,1494]" captionStart="FIGURES 5260" captionStartId="18.[151,264,1898,1922]" captionTargetBox="[151,1436,181,1875]" captionTargetId="figure-18@18.[151,1436,181,1875]" captionTargetPageId="18" captionText="FIGURES 5260. Epigyne (5253, 5560) and receptacles (54) of Mesiotelus kulczynskii (5254), M. mediterraneus sp. nov. (5557) and M. patricki (5860). 55, 58—intact, ventral; 52, 56, 59—macerated, ventral; 5354, 57, 60—macerated, dorsal. Scale bars: 0.1 mm. Abbreviations: Ah—anterior hood, Hl—lateral margin of anterior hood." figureDoi="http://doi.org/10.5281/zenodo.13915763" httpUri="https://zenodo.org/record/13915763/files/figure.png" pageId="12" pageNumber="202">Figs 5254</figureCitation>
), but can be separated from it by the relatively wider anterior hood with distinct lateral margins (
<emphasis id="18B3EAA0FFB9FFC20D3A94F7D76F49E4" box="[1117,1140,1504,1529]" italics="true" pageId="12" pageNumber="202">vs</emphasis>
. lacking), and subparallel posterior lateral margins (
<emphasis id="18B3EAA0FFB9FFC208DC9713D2C94A00" box="[443,466,1540,1565]" italics="true" pageId="12" pageNumber="202">vs</emphasis>
. diverging), and receptacles spaced by their length (
<emphasis id="18B3EAA0FFB9FFC20D459713D7224A00" box="[1058,1081,1540,1565]" italics="true" pageId="12" pageNumber="202">vs</emphasis>
. less). For male, see Zamani &amp; Marusik (2021).
</paragraph>
</subSubSection>
<subSubSection id="62DD6539FFB9FFC209A0975CD1754BEC" pageId="12" pageNumber="202" type="description">
<paragraph id="2A7836B2FFB9FFC209A0975CD15B4B04" blockId="12.[151,1437,1431,2033]" pageId="12" pageNumber="202">
<emphasis id="18B3EAA0FFB9FFC209A0975CD2424A78" bold="true" box="[199,345,1611,1637]" pageId="12" pageNumber="202">Description.</emphasis>
Female. Habitus as in
<figureCitation id="B2FC2A37FFB9FFC20B3F975CD1B24A78" box="[600,681,1611,1637]" captionStart="FIGURES 712" captionStartId="3.[151,264,1969,1993]" captionTargetBox="[152,1436,635,1936]" captionTargetId="figure-223@3.[151,1436,633,1946]" captionTargetPageId="3" captionText="FIGURES 712. Dorsal habitus of Mesiotelus cyprius (7), M. damavandicus sp. nov. (8), M. kravetsi sp. nov. (9), M. mediterraneus sp. nov. (10), M. patricki (11) and M. sistanus sp. nov. (12). 78, 1012—female; 9—male. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13915747" httpUri="https://zenodo.org/record/13915747/files/figure.png" pageId="12" pageNumber="202">Fig. 11</figureCitation>
. Total length 5.50. Carapace 2.40 long, 2.00 wide. Eye sizes: AME: 0.13, ALE: 0.15, PME: 0.12, PLE: 0.14. Carapace, chelicerae, maxillae, labium and sternum light yellowish-brown. Legs slightly lighter than carapace, without annulations. Abdomen beige, lighter ventrally, without patterns. Spinnerets uniformly light yellowish-brown. Ventral paired tibial spines: I, II: 2p, III, IV:
<specimenCount id="3CC1FD3BFFB9FFC20D1097A0D7814ACC" box="[1143,1178,1719,1745]" count="3" pageId="12" pageNumber="202" type="generic">3p.</specimenCount>
Measurements of legs: I: 8.31 (2.31, 1.23, 2.11, 1.66, 1.00), II: 7.32 (2.00, 1.08, 1.80, 1.50, 0.94), III: 6.91 (1.87, 0.93, 1.51, 1.75, 0.85), IV: 9.80 (2.56, 1.12, 2.37, 2.72, 1.03).
</paragraph>
<paragraph id="2A7836B2FFB9FFC209A09634D1C74BD0" blockId="12.[151,1437,1431,2033]" pageId="12" pageNumber="202">
Epigyne as in
<figureCitation id="B2FC2A37FFB9FFC208149634D2EC4B23" box="[371,503,1827,1854]" captionStart="FIGURES 5260" captionStartId="18.[151,264,1898,1922]" captionTargetBox="[151,1436,181,1875]" captionTargetId="figure-18@18.[151,1436,181,1875]" captionTargetPageId="18" captionText="FIGURES 5260. Epigyne (5253, 5560) and receptacles (54) of Mesiotelus kulczynskii (5254), M. mediterraneus sp. nov. (5557) and M. patricki (5860). 55, 58—intact, ventral; 52, 56, 59—macerated, ventral; 5354, 57, 60—macerated, dorsal. Scale bars: 0.1 mm. Abbreviations: Ah—anterior hood, Hl—lateral margin of anterior hood." figureDoi="http://doi.org/10.5281/zenodo.13915763" httpUri="https://zenodo.org/record/13915763/files/figure.png" pageId="12" pageNumber="202">Figs 5860</figureCitation>
; epigynal plate 1.4 times longer than wide; fovea ca. 2. time longer than wide; anterior hood (
<emphasis id="18B3EAA0FFB9FFC208269650D27B4B7C" box="[321,352,1863,1889]" italics="true" pageId="12" pageNumber="202">Ah</emphasis>
) with lateral margins (
<emphasis id="18B3EAA0FFB9FFC20B019650D1994B7C" box="[614,642,1863,1889]" italics="true" pageId="12" pageNumber="202">Hl</emphasis>
) longer that hoods width, hood ca. 2 times thinner than fovea; fovea lacking distinct transversal wrinkles, posterior part of fovea V-shaped, anterior part of lateral margins subparallel; receptacles oval, ca. 1.6 times longer than wide, inclined, spaced by about their length; copulatory ducts almost of receptacles length, spaced by 1.5 times their width.
</paragraph>
<paragraph id="2A7836B2FFB9FFC209A096C0D1754BEC" blockId="12.[151,1437,1431,2033]" box="[199,622,2007,2033]" pageId="12" pageNumber="202">Male. See Zamani &amp; Marusik (2021).</paragraph>
</subSubSection>
<subSubSection id="62DD6539FFB8FFC309A09181D70D4CE4" pageId="13" pageNumber="203" type="discussion">
<paragraph id="2A7836B2FFB8FFC309A09181D70D4CE4" blockId="13.[151,1436,150,249]" pageId="13" pageNumber="203">
<emphasis id="18B3EAA0FFB8FFC309A09181D25F4CAD" bold="true" box="[199,324,150,176]" pageId="13" pageNumber="203">Comment.</emphasis>
<materialsCitation id="9AAF3CEFFFB8FFC308289181D7094CE4" country="Iran" location="Golestan Province" pageId="13" pageNumber="203" specimenCount="1" stateProvince="Golestan">
Although the female described here was not collected with a male, we are quite certain of our identification as the locality reported here is only approximately
<quantity id="ED3F9B57FFB8FFC30A0A91ADD0BD4CC9" box="[877,934,186,212]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="203" unit="km" value="9.0">9 km</quantity>
of aerial distance from the
<typeStatus id="F57C8810FFB8FFC30DBE91ACD6124CC8" box="[1241,1289,187,213]" pageId="13" pageNumber="203">type</typeStatus>
locality.
<emphasis id="18B3EAA0FFB8FFC309A091C9D27A4CE5" bold="true" box="[199,353,222,248]" pageId="13" pageNumber="203">Distribution.</emphasis>
Known only from
<collectingRegion id="E803F850FFB8FFC30B5091C9D0134CE5" box="[567,776,222,248]" country="Iran" name="Golestan" pageId="13" pageNumber="203">Golestan Province</collectingRegion>
, northern
<collectingCountry id="52D07622FFB8FFC30A1E91C8D0BD4CE4" box="[889,934,223,249]" name="Iran" pageId="13" pageNumber="203">Iran</collectingCountry>
(
<figureCitation id="B2FC2A37FFB8FFC30AD291C9D7114CE4" box="[949,1034,222,249]" captionStart="FIGURE 75" captionStartId="21.[151,250,1840,1864]" captionTargetBox="[151,1436,853,1817]" captionTargetId="figure-284@21.[151,1436,853,1817]" captionTargetPageId="21" captionText="FIGURE 75. Collection localities of the studied liocranid species. 1—Agroeca angirasu; 2—A. cuprea; 3—A. inopina; 4—A. proxima; 5—Mesiotelus alanyensis sp. nov. (in part, specimens from Antalya); 6—M. bosmansi sp. nov.; 7—M. confusus sp. nov.; 8—M. cyprius; 9—M. damavandicus sp. nov.; 10—M. scopensis; 11—M. kravetsi sp. nov.; 12—M. mediterraneus sp. nov. and M. alanyensis sp. nov. (in part, specimens from Mersin); 13—M. patricki; 14—M. sistanus sp. nov.; 15—M. tadzhikus sp. nov.; 16—M. turanicus sp. nov.; 17—M. kulczynskii; 18—Scotina celans; 19—Sestakovaia annulipes. Some symbols could refer to more than one closely situated locality." figureDoi="http://doi.org/10.5281/zenodo.13915769" httpUri="https://zenodo.org/record/13915769/files/figure.png" pageId="13" pageNumber="203">Fig. 75</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>