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<document id="43FA032EFA70FD3F2355FAE3FEF5E1D9" ID-CLB-Dataset="30877" ID-DOI="10.11646/zootaxa.4377.1.6" ID-GBIF-Dataset="bf31986f-33c6-4a7b-8b58-736c352e54e4" ID-ISSN="1175-5326" ID-Zenodo-Dep="1163298" ID-ZooBank="CD3ED409-1877-4141-9AE3-ADAC1BE329BE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1517396338004" checkinUser="plazi" docAuthor="Zatwarnicki, Tadeusz &amp; Irwin, Anthony G." docDate="2018" docId="A26487EBFFE65B74FF0C0E002A6AFE6A" docLanguage="en" docName="zootaxa.4377.1.6.pdf" docOrigin="Zootaxa 4377 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scatella (Neoscatella) australis Zatwarnicki &amp; Irwin 2018, new combination" docType="treatment" docVersion="7" lastPageNumber="94" masterDocId="5E5DFF93FFE45B77FF9B0C3C295FFFC5" masterDocTitle="Taxonomic notes on the genera Scatella and Scatophila (Diptera: Ephydridae) with a remark on Trixoscelis chilensis (Trixoscelididae)" masterLastPageNumber="109" masterPageNumber="91" pageNumber="93" updateTime="1698517930452" updateUser="ExternalLinkService">
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<mods:title id="F6F823B005F41976D3F848CDDAD5A6DE">Taxonomic notes on the genera Scatella and Scatophila (Diptera: Ephydridae) with a remark on Trixoscelis chilensis (Trixoscelididae)</mods:title>
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<mods:namePart id="987CC0F26E9AB0562A82ACE474E09C62">Zatwarnicki, Tadeusz</mods:namePart>
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<mods:namePart id="6465518C7E186E5A7EF2EB1F4FD1E24F">Irwin, Anthony G.</mods:namePart>
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<mods:title id="93463AE79B81623C5BB5CE2ED652E217">Zootaxa</mods:title>
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<mods:date id="B4531A9062A1026A06E46E330F46327F">2018</mods:date>
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<treatment id="A26487EBFFE65B74FF0C0E002A6AFE6A" ID-DOI="http://doi.org/10.5281/zenodo.5967093" ID-GBIF-Taxon="140561292" ID-Zenodo-Dep="5967093" LSID="urn:lsid:plazi:treatment:A26487EBFFE65B74FF0C0E002A6AFE6A" httpUri="http://treatment.plazi.org/id/A26487EBFFE65B74FF0C0E002A6AFE6A" lastPageId="3" lastPageNumber="94" pageId="2" pageNumber="93">
<subSubSection id="62D76576FFE65B75FF0C0E0029B4FDB2" pageId="2" pageNumber="93" type="nomenclature">
<paragraph id="2A7236FDFFE65B75FF0C0E002A2EFD93" blockId="2.[151,881,572,631]" box="[151,881,572,598]" pageId="2" pageNumber="93">
<heading id="713A8191FFE65B75FF0C0E002A2EFD93" bold="true" box="[151,881,572,598]" fontSize="11" level="1" pageId="2" pageNumber="93" reason="1">
<emphasis id="18B9EAEFFFE65B75FF0C0E002A2EFD93" bold="true" box="[151,881,572,598]" pageId="2" pageNumber="93">
<taxonomicName id="EDCD4D7EFFE65B75FF0C0E002BD9FD93" authority="(Walker)" authorityName="Zatwarnicki &amp; Irwin" authorityYear="2018" baseAuthorityName="Zatwarnicki &amp; Irwin" baseAuthorityYear="1853" box="[151,646,572,598]" class="Insecta" family="Ephydridae" genus="Scatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="australis" status="comb. nov." subGenus="Neoscatella">
<emphasis id="18B9EAEFFFE65B75FF0C0E0029A8FD93" bold="true" box="[151,247,572,598]" italics="true" pageId="2" pageNumber="93">Scatella</emphasis>
(
<emphasis id="18B9EAEFFFE65B75FE930E0028CBFD93" bold="true" box="[264,404,572,598]" italics="true" pageId="2" pageNumber="93">Neoscatella</emphasis>
)
<emphasis id="18B9EAEFFFE65B75FE3E0E002B51FD93" bold="true" box="[421,526,572,598]" italics="true" pageId="2" pageNumber="93">australis</emphasis>
(Walker)
</taxonomicName>
,
<taxonomicNameLabel id="038A5794FFE65B75FD0F0E002A33FD93" box="[660,876,572,598]" pageId="2" pageNumber="93">new combination</taxonomicNameLabel>
.
</emphasis>
</heading>
</paragraph>
<paragraph id="2A7236FDFFE65B75FF0C0E6229B4FDB2" blockId="2.[151,881,572,631]" box="[151,235,606,631]" pageId="2" pageNumber="93">
(
<figureCitation id="B2F62A78FFE65B75FF040E6229BFFDB2" box="[159,224,606,631]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[151,1436,1033,1837]" captionTargetId="figure@2.[151,1436,1033,1837]" captionTargetPageId="2" captionText="FIGURE 1. Lectotype male of Scatella australis (Walker, 1853), lateral view." httpUri="https://zenodo.org/record/1163300/files/figure.png" pageId="2" pageNumber="93">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="62D76576FFE65B75FF0C0E9A2AD6FD1E" pageId="2" pageNumber="93" type="reference_group">
<paragraph id="2A7236FDFFE65B75FF0C0E9A2B4BFD79" blockId="2.[151,905,678,731]" box="[151,532,678,700]" pageId="2" pageNumber="93">
<treatmentCitationGroup id="0ADD11D3FFE65B75FF0C0E9A2B4BFD79" box="[151,532,678,700]" pageId="2" pageNumber="93">
<taxonomicName id="EDCD4D7EFFE65B75FF0C0E9A2B4FFD79" ID-CoL="8C2CW" authority="Walker, 1853: 409" authorityName="Walker" authorityPageNumber="409" authorityYear="1853" box="[151,528,678,700]" class="Insecta" family="Ephydridae" genus="Ephydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="18B9EAEFFFE65B75FF0C0E9A2812FD79" box="[151,333,678,700]" italics="true" pageId="2" pageNumber="93">Ephydra australis</emphasis>
<treatmentCitation id="AB6C10ECFFE65B75FEC80E9A2B4FFD79" author="Walker" box="[339,528,678,700]" page="409" pageId="2" pageNumber="93" year="1853">
<bibRefCitation id="4E5C4B0CFFE65B75FEC80E9A2882FD79" author="Walker" box="[339,477,678,700]" pageId="2" pageNumber="93" refString="Walker, F. (1853) Diptera. In: Saunders, W. W. (Ed.), Insecta Saundersiana or characters of undescribed Insects in the collection of William Wilson Saunders. Esq., F. R. S., F. L. S. &amp; c., London, pp. 253 - 414." type="book chapter" year="1853">Walker, 1853</bibRefCitation>
: 409
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2A7236FDFFE65B75FF0C0EF92AD6FD1E" blockId="2.[151,905,678,731]" box="[151,905,709,731]" pageId="2" pageNumber="93">
<treatmentCitationGroup id="0ADD11D3FFE65B75FF0C0EF92AD6FD1E" box="[151,905,709,731]" pageId="2" pageNumber="93">
<taxonomicName id="EDCD4D7EFFE65B75FF0C0EF92BB9FD1E" ID-CoL="8DSQ6" authority="Mathis &amp; Wirth, 1981: 24" authorityName="Mathis &amp; Wirth" authorityPageNumber="24" authorityYear="1981" box="[151,742,709,731]" class="Insecta" family="Ephydridae" genus="Scatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="insularis" subGenus="Neoscatella">
<emphasis id="18B9EAEFFFE65B75FF0C0EF92889FD1E" box="[151,470,709,731]" italics="true" pageId="2" pageNumber="93">Scatella (Neoscatella) insularis</emphasis>
<treatmentCitation id="AB6C10ECFFE65B75FE470EF92BB9FD1E" author="Mathis" box="[476,742,709,731]" page="24" pageId="2" pageNumber="93" year="1981">
<bibRefCitation id="4E5C4B0CFFE65B75FE470EF92B9FFD1E" author="Mathis" box="[476,704,709,731]" pageId="2" pageNumber="93" refString="Mathis, W. N. &amp; Wirth, W. W. (1981) Studies of Ephydrinae (Diptera: Ephydridae), IV: Revision of the Australian species of subgenus Neoscatella Malloch. Smithsonian Contributions to Zoology, 325, iii + 27 pp." type="book" year="1981">Mathis &amp; Wirth, 1981</bibRefCitation>
: 24
</treatmentCitation>
</taxonomicName>
,
<taxonomicNameLabel id="038A5794FFE65B75FD6B0EF92ADCFD1F" box="[752,899,709,730]" pageId="2" pageNumber="93">
<emphasis id="18B9EAEFFFE65B75FD6B0EF92ADCFD1F" bold="true" box="[752,899,709,730]" pageId="2" pageNumber="93">new synonym</emphasis>
</taxonomicNameLabel>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="62D76576FFE65B75FF0C0F3B2CCCFC10" pageId="2" pageNumber="93" type="materials_examined">
<paragraph id="2A7236FDFFE65B75FF0C0F3B2857FC49" blockId="2.[151,1436,775,981]" pageId="2" pageNumber="93">
<emphasis id="18B9EAEFFFE65B75FF0C0F3B2817FCE5" bold="true" box="[151,328,775,800]" pageId="2" pageNumber="93">Type material.</emphasis>
The lectotype female of
<taxonomicName id="EDCD4D7EFFE65B75FDF50F342BB1FCE5" authorityName="Walker" authorityYear="1853" box="[622,750,775,800]" class="Insecta" family="Ephydridae" genus="Ephydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="18B9EAEFFFE65B75FDF50F342BB1FCE5" box="[622,750,775,800]" italics="true" pageId="2" pageNumber="93">S. australis</emphasis>
</taxonomicName>
(herein designated) is in poor condition, nearly completely (apart from abdomen and wings) covered by fungal mycelium (
<figureCitation id="B2F62A78FFE65B75FCE40F102A9AFC81" box="[895,965,812,837]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[151,1436,1033,1837]" captionTargetId="figure@2.[151,1436,1033,1837]" captionTargetPageId="2" captionText="FIGURE 1. Lectotype male of Scatella australis (Walker, 1853), lateral view." httpUri="https://zenodo.org/record/1163300/files/figure.png" pageId="2" pageNumber="93">Fig. 1</figureCitation>
), labelled: &quot;68 4&quot;, &quot;
<taxonomicName id="EDCD4D7EFFE65B75FB290F102C48FC80" authorityName="Walker" authorityYear="1853" box="[1202,1303,812,837]" class="Insecta" family="Ephydridae" genus="Ephydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="australis">australis</taxonomicName>
&quot;, &quot;V D L&quot;, &quot;Tasmania. W. W. Saunders B. M. 1868 - 4.&quot;, &quot;
<taxonomicName id="EDCD4D7EFFE65B75FD2B0F732AACFCAD" authority="Walker" authorityName="Walker" box="[688,1011,847,872]" class="Insecta" family="Ephydridae" genus="Neoscatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="australis">Neoscatella australis Walker</taxonomicName>
det. B. H. Cogan 1975&quot; preserved in NHMUK.
</paragraph>
<paragraph id="2A7236FDFFE65B75FF5C0FA42CCCFC10" blockId="2.[151,1436,775,981]" pageId="2" pageNumber="93">
<typeStatus id="F576885FFFE65B75FF5C0FA429A0FC75" box="[199,255,920,944]" pageId="2" pageNumber="93">Type</typeStatus>
locality of the senior synonym is &quot;
<collectingRegion id="E809F81FFFE65B75FD100FA42BE6FC75" box="[651,697,920,944]" country="Turkey" name="Van" pageId="2" pageNumber="93">Van</collectingRegion>
Dieman's Land&quot; (=
<collectingRegion id="E809F81FFFE65B75FC3A0FAB2D4EFC75" box="[929,1041,919,944]" country="Australia" name="Tasmania" pageId="2" pageNumber="93">Tasmania</collectingRegion>
); localities of junior synonym
<typeStatus id="F576885FFFE65B75FAF70FA42CC3FC75" box="[1388,1436,920,944]" pageId="2" pageNumber="93">type</typeStatus>
series are two islands (Little Dog and Fisher) between the northeast coast of
<collectingRegion id="E809F81FFFE65B75FC6A0F802D03FC10" box="[1009,1116,956,981]" country="Australia" name="Tasmania" pageId="2" pageNumber="93">Tasmania</collectingRegion>
and southeastern
<collectingCountry id="52DA766DFFE65B75FABE0F802CD0FC10" box="[1317,1423,956,981]" name="Australia" pageId="2" pageNumber="93">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="7EB26675FFE65B75FF0C0B7F2AF0F89C" box="[151,943,1859,1881]" httpUri="https://zenodo.org/record/1163300/files/figure.png" pageId="2" pageNumber="93" targetBox="[151,1436,1033,1837]" targetPageId="2">
<paragraph id="2A7236FDFFE65B75FF0C0B7F2AF0F89C" blockId="2.[151,943,1859,1881]" box="[151,943,1859,1881]" pageId="2" pageNumber="93">
<emphasis id="18B9EAEFFFE65B75FF0C0B7F284CF89D" bold="true" box="[151,275,1859,1881]" pageId="2" pageNumber="93">FIGURE 1.</emphasis>
Lectotype male of
<taxonomicName id="EDCD4D7EFFE65B75FE4C0B7F2A77F89C" authority="(Walker, 1853)" authorityName="Zatwarnicki &amp; Irwin" authorityYear="2018" baseAuthorityName="Zatwarnicki &amp; Irwin" baseAuthorityYear="1853" box="[471,808,1859,1881]" class="Insecta" family="Ephydridae" genus="Scatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="18B9EAEFFFE65B75FE4C0B7F2BD7F89C" box="[471,648,1859,1881]" italics="true" pageId="2" pageNumber="93">Scatella australis</emphasis>
(Walker, 1853)
</taxonomicName>
, lateral view.
</paragraph>
</caption>
<subSubSection id="62D76576FFE65B75FF5C0BBC2D34F821" pageId="2" pageNumber="93" type="diagnosis">
<paragraph id="2A7236FDFFE65B75FF5C0BBC2D34F821" blockId="2.[151,1436,1920,2020]" pageId="2" pageNumber="93">
<emphasis id="18B9EAEFFFE65B75FF5C0BBC281EF85C" bold="true" box="[199,321,1920,1945]" pageId="2" pageNumber="93">Diagnosis.</emphasis>
The species is closely related to
<taxonomicName id="EDCD4D7EFFE65B75FD2B0BBD2ABCF85C" authority="Mathis &amp; Wirth" authorityName="Mathis &amp; Wirth" authorityYear="1981" box="[688,995,1920,1945]" class="Insecta" family="Ephydridae" genus="Scatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="austrina">
<emphasis id="18B9EAEFFFE65B75FD2B0BBD2A76F85D" box="[688,809,1921,1944]" italics="true" pageId="2" pageNumber="93">S. austrina</emphasis>
Mathis &amp; Wirth
</taxonomicName>
, but is distinguished from other species by the costal vein of male thickened; wing with two subapical spots in cell R5 apicad of posterior crossvein, if connected to form one white area, aligned almost perpendicularly to longitudinal veins.
</paragraph>
</subSubSection>
<subSubSection id="62D76576FFE75B74FF5C0CAB2C2AFF75" box="[199,1397,151,176]" pageId="3" pageNumber="94" type="distribution">
<paragraph id="2A7236FDFFE75B74FF5C0CAB2C2AFF75" blockId="3.[151,1437,151,431]" box="[199,1397,151,176]" pageId="3" pageNumber="94">
<emphasis id="18B9EAEFFFE75B74FF5C0CAB283FFF75" bold="true" box="[199,352,151,176]" pageId="3" pageNumber="94">Distribution.</emphasis>
Australasian/Oceanian:
<collectingCountry id="52DA766DFFE75B74FDE80CAB2B85FF75" box="[627,730,151,176]" name="Australia" pageId="3" pageNumber="94">Australia</collectingCountry>
(
<collectingRegion id="E809F81FFFE75B74FD720CAB2A06FF75" box="[745,857,151,176]" country="Australia" name="Tasmania" pageId="3" pageNumber="94">Tasmania</collectingRegion>
, and two nearby islands: Little Dog and Fisher).
</paragraph>
</subSubSection>
<subSubSection id="62D76576FFE75B74FF5C0C802A6AFE6A" pageId="3" pageNumber="94" type="discussion">
<paragraph id="2A7236FDFFE75B74FF5C0C802A6AFE6A" blockId="3.[151,1437,151,431]" pageId="3" pageNumber="94">
<emphasis id="18B9EAEFFFE75B74FF5C0C802864FF10" bold="true" box="[199,315,188,213]" pageId="3" pageNumber="94">Remarks.</emphasis>
<bibRefCitation id="4E5C4B0CFFE75B74FEDF0C802BC9FF10" author="Mathis" box="[324,662,188,213]" pageId="3" pageNumber="94" refString="Mathis, W. N. &amp; Zatwarnicki, T. (1995) World catalog of shore flies (Diptera: Ephydridae). International Memoirs of Entomology, 4, i - vi + 1 - 423." type="journal article" year="1995">Mathis &amp; Zatwarnicki (1995)</bibRefCitation>
reported that the
<typeStatus id="F576885FFFE75B74FCFD0C812AC9FF10" box="[870,918,189,213]" pageId="3" pageNumber="94">type</typeStatus>
of
<taxonomicName id="EDCD4D7EFFE75B74FC5A0C802DB8FF10" authority="Walker" authorityName="Walker" authorityYear="1853" box="[961,1255,188,213]" class="Insecta" family="Ephydridae" genus="Ephydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="18B9EAEFFFE75B74FC5A0C802DD2FF10" box="[961,1165,188,213]" italics="true" pageId="3" pageNumber="94">Ephydra australis</emphasis>
Walker
</taxonomicName>
was apparently lost (note that they inadvertently referred to it as
<taxonomicName id="EDCD4D7EFFE75B74FD2E0CE32AD8FF3D" authorityName="Loew" authorityYear="1860" box="[693,903,223,248]" class="Insecta" family="Ephydridae" genus="Notiphila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="18B9EAEFFFE75B74FD2E0CE32AD8FF3D" box="[693,903,223,248]" italics="true" pageId="3" pageNumber="94">Notiphila australis</emphasis>
</taxonomicName>
), but it has been rediscovered in NHMUK (
<figureCitation id="B2F62A78FFE75B74FAF40CE329FBFED9" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[151,1436,1033,1837]" captionTargetId="figure@2.[151,1436,1033,1837]" captionTargetPageId="2" captionText="FIGURE 1. Lectotype male of Scatella australis (Walker, 1853), lateral view." httpUri="https://zenodo.org/record/1163300/files/figure.png" pageId="3" pageNumber="94">Fig. 1</figureCitation>
). Detailed examination shows it is the same species as
<taxonomicName id="EDCD4D7EFFE75B74FCAF0D392DCEFED8" authority="Mathis and Wirth" authorityName="Mathis and Wirth" box="[820,1169,260,285]" class="Insecta" family="Ephydridae" genus="Scatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="18B9EAEFFFE75B74FCAF0D392AE9FED8" box="[820,950,260,285]" italics="true" pageId="3" pageNumber="94">S. insularis</emphasis>
Mathis and Wirth
</taxonomicName>
: wing with a pale spot towards the base of cell R1, which features in some females of
<taxonomicName id="EDCD4D7EFFE75B74FCFF0D142ABCFE85" box="[868,995,295,320]" class="Insecta" family="Ephydridae" genus="Scatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="18B9EAEFFFE75B74FCFF0D142ABCFE85" box="[868,995,295,320]" italics="true" pageId="3" pageNumber="94">S. insularis</emphasis>
</taxonomicName>
, but not in other species of this group. Also other spots are identical as on the photograph in
<bibRefCitation id="4E5C4B0CFFE75B74FD680D722D0FFEA2" author="Mathis" box="[755,1104,334,359]" pageId="3" pageNumber="94" refString="Mathis, W. N. &amp; Wirth, W. W. (1981) Studies of Ephydrinae (Diptera: Ephydridae), IV: Revision of the Australian species of subgenus Neoscatella Malloch. Smithsonian Contributions to Zoology, 325, iii + 27 pp." type="book" year="1981">Mathis &amp; Wirth (1981, fig. 44)</bibRefCitation>
, anterior acrostichal setae are short as on the illustration nr.
<quantity id="ED359B18FFE75B74FE7C0D4E2B7EFE4E" box="[487,545,370,395]" metricMagnitude="0" metricUnit="m" metricValue="1.1684" pageId="3" pageNumber="94" unit="in" value="46.0">46 in</quantity>
above paper. The
<typeStatus id="F576885FFFE75B74FD6A0D4E2A0CFE4E" box="[753,851,370,395]" pageId="3" pageNumber="94" type="holotype">holotype</typeStatus>
of the junior synonym is a male and is preserved in Australian National Insect Collection (Canberra,
<collectingCountry id="52DA766DFFE75B74FD260DAA2A76FE6A" box="[701,809,406,431]" name="Australia" pageId="3" pageNumber="94">Australia</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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