210 lines
32 KiB
XML
210 lines
32 KiB
XML
<document id="D90594E54A0C6AE5E556F6C073A8EFA8" ID-DOI="10.1111/j.1096-3642.2012.00844.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5408589" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630349144448" checkinUser="marcus" docAuthor="Adnet, Sylvain, Cappetta, Henri, Guinot, Guillaume & Sciara, Giuseppe Notarbartolo Di" docDate="2012" docId="A260879AFFCDA218FF3988D432480353" docLanguage="en" docName="j.1096-3642.2012.00844.x.pdf" docOrigin="Zoological Journal of the Linnean Society 166 (1)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2012.00844.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Argoubia barbei Adnet & Cappetta & Guinot & Sciara 2012, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="144" masterDocId="5E59FFE2FFC6A214FFD68A4B313F043C" masterDocTitle="Evolutionary history of the devilrays (Chondrichthyes: Myliobatiformes) from fossil and morphological inference" masterLastPageNumber="159" masterPageNumber="132" pageNumber="143" updateTime="1699240823188" updateUser="plazi">
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<mods:title id="117F637E8E72AFECAB7DD76BB79C2793">Evolutionary history of the devilrays (Chondrichthyes: Myliobatiformes) from fossil and morphological inference</mods:title>
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<mods:namePart id="CF31D0A0D356BDB02C7C53E7764F1DD6">Adnet, Sylvain</mods:namePart>
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<mods:namePart id="EA7E629878D4F5F5DCF3C49ED1BA62A0">Cappetta, Henri</mods:namePart>
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<mods:namePart id="C06711C41E0EE3EA8E7071A7F7D1DCFB">Guinot, Guillaume</mods:namePart>
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<mods:roleTerm id="223FE78D0B64746A211449FC4280C14F">Author</mods:roleTerm>
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<mods:namePart id="0CE3D346751EC383FFFB52D0B36C1FC9">Sciara, Giuseppe Notarbartolo Di</mods:namePart>
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<mods:title id="39E108D6BEE8E5E6DB37631793386AD4">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="9CFF205B94D38861A1779FC0CF39D0B6">2012</mods:date>
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<mods:number id="574CE5A397EA92CDEA82392371A8F8D6">2015-07-06</mods:number>
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<mods:number id="ABBE9EAB7C919F47B19C2ADE1B82064E">166</mods:number>
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<mods:url id="20283C620E7A008715D662742FFFDB58">http://dx.doi.org/10.1111/j.1096-3642.2012.00844.x</mods:url>
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<treatment id="A260879AFFCDA218FF3988D432480353" ID-DOI="http://doi.org/10.5281/zenodo.5479986" ID-Zenodo-Dep="5479986" LSID="urn:lsid:plazi:treatment:A260879AFFCDA218FF3988D432480353" httpUri="http://treatment.plazi.org/id/A260879AFFCDA218FF3988D432480353" lastPageId="12" lastPageNumber="144" pageId="11" pageNumber="143">
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<subSubSection id="62D36507FFCDA21FFF3988D433A80689" box="[239,663,671,693]" pageId="11" pageNumber="143" type="nomenclature">
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<paragraph id="2A76368CFFCDA21FFF3988D433A80689" blockId="11.[239,663,671,693]" box="[239,663,671,693]" pageId="11" pageNumber="143">
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<heading id="713E81E0FFCDA21FFF3988D433A80689" box="[239,663,671,693]" centered="true" fontSize="9" level="2" pageId="11" pageNumber="143" reason="4">
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<taxonomicName id="EDC94D0FFFCDA21FFF3988D430F60688" authority="Adnet & Cappetta & Guinot & Sciara, 2012" authorityName="Adnet & Cappetta & Guinot & Sciara" authorityYear="2012" box="[239,457,671,692]" class="Elasmobranchii" family="Myliobatidae" genus="Argoubia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="11" pageNumber="159" phylum="Chordata" rank="species" species="barbei" status="gen. et sp. nov.">ARGOUBIA BARBEI</taxonomicName>
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<emphasis id="18BDEA9EFFCDA21FFE0588E833070689" bold="true" box="[467,568,672,693]" pageId="11" pageNumber="143">
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<taxonomicNameLabel id="038E57E5FFCDA21FFE0588E833070689" box="[467,568,672,693]" pageId="11" pageNumber="143" rank="species">SP. NOV.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="B2F22A09FFCDA21FFD9C88EB33B00689" box="[586,655,671,693]" captionStart="Figure 5" captionStartId="12.[164,243,1598,1617]" captionTargetBox="[172,1437,202,1563]" captionTargetId="figure-70@12.[164,1445,194,1568]" captionTargetPageId="12" captionText="Figure 5. Argoubia barbei gen. nov., sp. nov. A-E, Antero-lateral tooth, holotype, DAK3-1. A, lingual view; B, profile; C, labial view; D, occlusal view; E, basal view. F-J, Antero-lateral tooth, paratype, DAK3-2. F, lingual view; G, profile; H, labial view; I, occlusal view; J, basal view." figureDoi="http://doi.org/10.5281/zenodo.5408599" httpUri="https://zenodo.org/record/5408599/files/figure.png" pageId="11" pageNumber="143">FIG. 5</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="62D36507FFCDA218FF46888D32480353" lastPageId="12" lastPageNumber="144" pageId="11" pageNumber="143" type="materials_examined">
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<paragraph id="2A76368CFFCDA21FFF46888D33C806E0" blockId="11.[144,759,710,732]" box="[144,759,710,732]" pageId="11" pageNumber="143">
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<emphasis id="18BDEA9EFFCDA21FFF46888D31C606E7" box="[144,249,710,731]" italics="true" pageId="11" pageNumber="143">Material:</emphasis>
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Two complete teeth and several fragments.
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</paragraph>
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<paragraph id="2A76368CFFCDA21FFF46894E30E30764" blockId="11.[144,758,773,856]" pageId="11" pageNumber="143">
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<emphasis id="18BDEA9EFFCDA21FFF46894E301A0726" box="[144,293,773,794]" italics="true" pageId="11" pageNumber="143">Type locality:</emphasis>
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Locality DAK3. Unit 2 level B1, close to El Argoub, south of Ad Dakhla, south-western
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<collectingCountry id="52DE761CFFCDA21FFF46890931D2076B" box="[144,237,834,855]" name="Morocco" pageId="11" pageNumber="143">Morocco</collectingCountry>
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(
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<bibRefCitation id="4E584B7DFFCDA21FFF28890930EF0764" author="Adnet S & Cappetta H & Tabuce R" box="[254,464,834,856]" pageId="11" pageNumber="143" pagination="860 - 870" refId="ref16589" refString="Adnet S, Cappetta H, Tabuce R. 2010. A Middle-Late Eocene vertebrate fauna (marine fish and mammals) from southwestern Morocco; preliminary report: age and palaeobiogeographical implications. Geological Magazine 147: 860 - 870." type="journal article" year="2010">
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Adnet
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<emphasis id="18BDEA9EFFCDA21FFE9B89093041076B" box="[333,382,834,855]" italics="true" pageId="11" pageNumber="143">et al</emphasis>
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., 2010
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</bibRefCitation>
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).
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</paragraph>
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<paragraph id="2A76368CFFCDA21FFF4689CA33C80788" blockId="11.[144,759,896,949]" pageId="11" pageNumber="143">
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<emphasis id="18BDEA9EFFCDA21FFF4689CA300E07A9" box="[144,305,896,918]" italics="true" pageId="11" pageNumber="143">Type horizon:</emphasis>
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Samlat Formation, Gerran member (
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<bibRefCitation id="4E584B7DFFCDA21FFF4189D430660789" author="Ratschiller LK" box="[151,345,927,949]" pageId="11" pageNumber="143" pagination="55 - 190" refId="ref19042" refString="Ratschiller LK. 1967. Sahara, correlazioni geologicolithostratigrafiche fra Sahara Centrale ed Occidentale (con note gologiche generali e brevi cenni sulle possibilita dell'Africa Nord-Occidentale). Memorie del Museo Tridentino di Scienze Naturali 16: 55 - 190." type="journal article" year="1967">Ratschiller, 1967</bibRefCitation>
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) – late Middle Eocene/Late Eocene.
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</paragraph>
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<paragraph id="2A76368CFFCDA21FFF46899633EB002E" blockId="11.[144,758,989,1042]" pageId="11" pageNumber="143">
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<emphasis id="18BDEA9EFFCDA21FFF468996303207CE" box="[144,269,989,1010]" italics="true" pageId="11" pageNumber="143">Etymology:</emphasis>
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Species dedicated to Mr Gérard Barbe for his help in the field and for donation of material.
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</paragraph>
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<paragraph id="2A76368CFFCDA21FFF468E7030520053" blockId="11.[144,758,1082,1135]" pageId="11" pageNumber="143">
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<materialsCitation id="9AA13CD1FFCDA21FFF468E7030520053" location="University of Montpellier." pageId="11" pageNumber="143" specimenCount="1" typeStatus="holotype">
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<emphasis id="18BDEA9EFFCDA21FFF468E7031C5006C" box="[144,250,1083,1104]" italics="true" pageId="11" pageNumber="143">
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<typeStatus id="F572882EFFCDA21FFF468E7031CB006C" box="[144,244,1083,1104]" pageId="11" pageNumber="143" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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DAK3-1 (
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<figureCitation id="B2F22A09FFCDA21FFEA78E7030DB006C" box="[369,484,1082,1105]" captionStart="Figure 5" captionStartId="12.[164,243,1598,1617]" captionTargetBox="[172,1437,202,1563]" captionTargetId="figure-70@12.[164,1445,194,1568]" captionTargetPageId="12" captionText="Figure 5. Argoubia barbei gen. nov., sp. nov. A-E, Antero-lateral tooth, holotype, DAK3-1. A, lingual view; B, profile; C, labial view; D, occlusal view; E, basal view. F-J, Antero-lateral tooth, paratype, DAK3-2. F, lingual view; G, profile; H, labial view; I, occlusal view; J, basal view." figureDoi="http://doi.org/10.5281/zenodo.5408599" httpUri="https://zenodo.org/record/5408599/files/figure.png" pageId="11" pageNumber="143">Fig. 5A–E</figureCitation>
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) housed at the
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<location id="2F166057FFCDA21FFD498E7030520053" LSID="urn:lsid:plazi:treatment:A260879AFFCDA218FF3988D432480353:2F166057FFCDA21FFD498E7030520053" name="University of Montpellier." pageId="11" pageNumber="143">University of Montpellier.</location>
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="2A76368CFFCDA21FFF468ED330C90091" blockId="11.[144,502,1176,1198]" box="[144,502,1176,1198]" pageId="11" pageNumber="143">
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<emphasis id="18BDEA9EFFCDA21FFF468ED330390091" box="[144,262,1176,1197]" italics="true" pageId="11" pageNumber="143">Diagnosis:</emphasis>
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Same as the genus.
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</paragraph>
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<paragraph id="2A76368CFFCDA21FFF468E9C325905AF" blockId="11.[144,759,1238,1903]" lastBlockId="11.[806,1422,198,1413]" pageId="11" pageNumber="143">
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<emphasis id="18BDEA9EFFCDA21FFF468E9C302700D0" box="[144,280,1239,1260]" italics="true" pageId="11" pageNumber="143">Description:</emphasis>
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The teeth of this species are rather large, reaching between 3 and
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<quantity id="ED319B69FFCDA21FFE7E8EBE30D60136" box="[424,489,1269,1290]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="143" unit="mm" value="4.0">4 mm</quantity>
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width for anterolateral teeth. The
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<typeStatus id="F572882EFFCDA21FFEC68F58304E0115" box="[272,369,1299,1321]" pageId="11" pageNumber="143" type="holotype">holotype</typeStatus>
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||
(
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||
<figureCitation id="B2F22A09FFCDA21FFE548F5830CA0115" box="[386,501,1299,1321]" captionStart="Figure 5" captionStartId="12.[164,243,1598,1617]" captionTargetBox="[172,1437,202,1563]" captionTargetId="figure-70@12.[164,1445,194,1568]" captionTargetPageId="12" captionText="Figure 5. Argoubia barbei gen. nov., sp. nov. A-E, Antero-lateral tooth, holotype, DAK3-1. A, lingual view; B, profile; C, labial view; D, occlusal view; E, basal view. F-J, Antero-lateral tooth, paratype, DAK3-2. F, lingual view; G, profile; H, labial view; I, occlusal view; J, basal view." figureDoi="http://doi.org/10.5281/zenodo.5408599" httpUri="https://zenodo.org/record/5408599/files/figure.png" pageId="11" pageNumber="143">Fig. 5A–E</figureCitation>
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) is broader than high in labial view and rather thick in profile (
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<quantity id="ED319B69FFCDA21FFD778F7933C8017B" box="[673,759,1330,1351]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="11" pageNumber="143" unit="mm" value="3.2">3.2 mm</quantity>
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wide,
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<quantity id="ED319B69FFCDA21FFF018F1A3013015A" box="[215,300,1361,1382]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="11" pageNumber="143" unit="mm" value="2.4">2.4 mm</quantity>
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high). The maximum width is located above the middle of the tooth and the base of the crown is practically as broad as the occlusal face. In labial view (
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<figureCitation id="B2F22A09FFCDA21FFE9E8FE730A201FE" box="[328,413,1452,1474]" captionStart="Figure 5" captionStartId="12.[164,243,1598,1617]" captionTargetBox="[172,1437,202,1563]" captionTargetId="figure-70@12.[164,1445,194,1568]" captionTargetPageId="12" captionText="Figure 5. Argoubia barbei gen. nov., sp. nov. A-E, Antero-lateral tooth, holotype, DAK3-1. A, lingual view; B, profile; C, labial view; D, occlusal view; E, basal view. F-J, Antero-lateral tooth, paratype, DAK3-2. F, lingual view; G, profile; H, labial view; I, occlusal view; J, basal view." figureDoi="http://doi.org/10.5281/zenodo.5408599" httpUri="https://zenodo.org/record/5408599/files/figure.png" pageId="11" pageNumber="143">Fig. 5C</figureCitation>
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), the edges of the crown are generally convex, and distinctly and irregularly cut off above the limit of the visor. In occlusal view (
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<figureCitation id="B2F22A09FFCDA21FFF4E8C4331D10222" box="[152,238,1544,1566]" captionStart="Figure 5" captionStartId="12.[164,243,1598,1617]" captionTargetBox="[172,1437,202,1563]" captionTargetId="figure-70@12.[164,1445,194,1568]" captionTargetPageId="12" captionText="Figure 5. Argoubia barbei gen. nov., sp. nov. A-E, Antero-lateral tooth, holotype, DAK3-1. A, lingual view; B, profile; C, labial view; D, occlusal view; E, basal view. F-J, Antero-lateral tooth, paratype, DAK3-2. F, lingual view; G, profile; H, labial view; I, occlusal view; J, basal view." figureDoi="http://doi.org/10.5281/zenodo.5408599" httpUri="https://zenodo.org/record/5408599/files/figure.png" pageId="11" pageNumber="143">Fig. 5D</figureCitation>
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), the occlusal face, very slightly depressed, is less developed transversely than the rest of the crown. The enameloid of this face is completely smooth and bright. The labial margin of the face is irregular. Its lingual edge is more salient and cut off by well-developed irregular cusps with rounded extremities and separated by weak to deep notches. The labial face (
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<figureCitation id="B2F22A09FFCDA21FFE808C95309602C8" box="[342,425,1758,1780]" captionStart="Figure 5" captionStartId="12.[164,243,1598,1617]" captionTargetBox="[172,1437,202,1563]" captionTargetId="figure-70@12.[164,1445,194,1568]" captionTargetPageId="12" captionText="Figure 5. Argoubia barbei gen. nov., sp. nov. A-E, Antero-lateral tooth, holotype, DAK3-1. A, lingual view; B, profile; C, labial view; D, occlusal view; E, basal view. F-J, Antero-lateral tooth, paratype, DAK3-2. F, lingual view; G, profile; H, labial view; I, occlusal view; J, basal view." figureDoi="http://doi.org/10.5281/zenodo.5408599" httpUri="https://zenodo.org/record/5408599/files/figure.png" pageId="11" pageNumber="143">Fig. 5C</figureCitation>
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) of the crown bears a very strong ornamentation of more or less vertical ridges separated by deep vertical hollows. These ridges are very salient, irregular, and bear villi directed upwards. These villi have oblique directions when the tooth is observed in labial view. The basal part of this labial face is limited by a distinct crest that corresponds to a labial visor. The lower part of this visor is well developed and oblique in profile (
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<figureCitation id="B2F22A09FFCDA21FFAFD8B6934BE050B" box="[1323,1409,289,312]" captionStart="Figure 5" captionStartId="12.[164,243,1598,1617]" captionTargetBox="[172,1437,202,1563]" captionTargetId="figure-70@12.[164,1445,194,1568]" captionTargetPageId="12" captionText="Figure 5. Argoubia barbei gen. nov., sp. nov. A-E, Antero-lateral tooth, holotype, DAK3-1. A, lingual view; B, profile; C, labial view; D, occlusal view; E, basal view. F-J, Antero-lateral tooth, paratype, DAK3-2. F, lingual view; G, profile; H, labial view; I, occlusal view; J, basal view." figureDoi="http://doi.org/10.5281/zenodo.5408599" httpUri="https://zenodo.org/record/5408599/files/figure.png" pageId="11" pageNumber="143">Fig. 5B</figureCitation>
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). It is smooth and, in labial view (
|
||
<figureCitation id="B2F22A09FFCDA21FFAFD8B0B34BF056A" box="[1323,1408,320,342]" captionStart="Figure 5" captionStartId="12.[164,243,1598,1617]" captionTargetBox="[172,1437,202,1563]" captionTargetId="figure-70@12.[164,1445,194,1568]" captionTargetPageId="12" captionText="Figure 5. Argoubia barbei gen. nov., sp. nov. A-E, Antero-lateral tooth, holotype, DAK3-1. A, lingual view; B, profile; C, labial view; D, occlusal view; E, basal view. F-J, Antero-lateral tooth, paratype, DAK3-2. F, lingual view; G, profile; H, labial view; I, occlusal view; J, basal view." figureDoi="http://doi.org/10.5281/zenodo.5408599" httpUri="https://zenodo.org/record/5408599/files/figure.png" pageId="11" pageNumber="143">Fig. 5C</figureCitation>
|
||
), it is higher and slightly depressed marginally on both sides.
|
||
</paragraph>
|
||
<paragraph id="2A76368CFFCDA21FFCE98BD7329006B4" blockId="11.[806,1422,198,1413]" pageId="11" pageNumber="143">
|
||
The lingual face is oblique in profile view (
|
||
<figureCitation id="B2F22A09FFCDA21FFAFD8BD73440058E" box="[1323,1407,412,434]" captionStart="Figure 5" captionStartId="12.[164,243,1598,1617]" captionTargetBox="[172,1437,202,1563]" captionTargetId="figure-70@12.[164,1445,194,1568]" captionTargetPageId="12" captionText="Figure 5. Argoubia barbei gen. nov., sp. nov. A-E, Antero-lateral tooth, holotype, DAK3-1. A, lingual view; B, profile; C, labial view; D, occlusal view; E, basal view. F-J, Antero-lateral tooth, paratype, DAK3-2. F, lingual view; G, profile; H, labial view; I, occlusal view; J, basal view." figureDoi="http://doi.org/10.5281/zenodo.5408599" httpUri="https://zenodo.org/record/5408599/files/figure.png" pageId="11" pageNumber="143">Fig. 5B</figureCitation>
|
||
), parallel to the labial face. It is restricted at its base by a distinct bulge, rounded in profile. This bulge overhangs the root in occlusal view. The lingual face is covered by vertical ridges that are shorter, less developed, and less irregular than those of the labial face. They stop above the bulge. Marginally, some ridges are oblique.
|
||
</paragraph>
|
||
<paragraph id="2A76368CFFCDA21FFCE988DA342807E4" blockId="11.[806,1422,198,1413]" pageId="11" pageNumber="143">
|
||
The root is not very high, about a quarter of the tooth in profile view (
|
||
<figureCitation id="B2F22A09FFCDA21FFBCE88E4355306F9" box="[1048,1132,687,709]" captionStart="Figure 5" captionStartId="12.[164,243,1598,1617]" captionTargetBox="[172,1437,202,1563]" captionTargetId="figure-70@12.[164,1445,194,1568]" captionTargetPageId="12" captionText="Figure 5. Argoubia barbei gen. nov., sp. nov. A-E, Antero-lateral tooth, holotype, DAK3-1. A, lingual view; B, profile; C, labial view; D, occlusal view; E, basal view. F-J, Antero-lateral tooth, paratype, DAK3-2. F, lingual view; G, profile; H, labial view; I, occlusal view; J, basal view." figureDoi="http://doi.org/10.5281/zenodo.5408599" httpUri="https://zenodo.org/record/5408599/files/figure.png" pageId="11" pageNumber="143">Fig. 5B</figureCitation>
|
||
), and distinctly narrower than the crown. In lingual view, the width of the root itself is greater just below the crown than basally. The root is higher medially than on the margins. There are three lobes (
|
||
<figureCitation id="B2F22A09FFCDA21FFC388961357C077C" box="[1006,1091,810,832]" captionStart="Figure 5" captionStartId="12.[164,243,1598,1617]" captionTargetBox="[172,1437,202,1563]" captionTargetId="figure-70@12.[164,1445,194,1568]" captionTargetPageId="12" captionText="Figure 5. Argoubia barbei gen. nov., sp. nov. A-E, Antero-lateral tooth, holotype, DAK3-1. A, lingual view; B, profile; C, labial view; D, occlusal view; E, basal view. F-J, Antero-lateral tooth, paratype, DAK3-2. F, lingual view; G, profile; H, labial view; I, occlusal view; J, basal view." figureDoi="http://doi.org/10.5281/zenodo.5408599" httpUri="https://zenodo.org/record/5408599/files/figure.png" pageId="11" pageNumber="143">Fig. 5E</figureCitation>
|
||
): two marginal, with a flat basal face of triangular outline, and one median that is narrow and also has a flat basal face. The lobes are separated by broad and rather deep and W- shaped furrows. Irregularly placed foramina open on the labial, lingual, and basal faces of the root.
|
||
</paragraph>
|
||
<paragraph id="2A76368CFFCDA21FFCE989AA358301B9" blockId="11.[806,1422,198,1413]" pageId="11" pageNumber="143">
|
||
The
|
||
<typeStatus id="F572882EFFCDA21FFCAE89A932E107CB" box="[888,990,994,1015]" pageId="11" pageNumber="143" type="paratype">paratype</typeStatus>
|
||
(
|
||
<figureCitation id="B2F22A09FFCDA21FFC2589AA355F07CB" box="[1011,1120,993,1015]" captionStart="Figure 5" captionStartId="12.[164,243,1598,1617]" captionTargetBox="[172,1437,202,1563]" captionTargetId="figure-70@12.[164,1445,194,1568]" captionTargetPageId="12" captionText="Figure 5. Argoubia barbei gen. nov., sp. nov. A-E, Antero-lateral tooth, holotype, DAK3-1. A, lingual view; B, profile; C, labial view; D, occlusal view; E, basal view. F-J, Antero-lateral tooth, paratype, DAK3-2. F, lingual view; G, profile; H, labial view; I, occlusal view; J, basal view." figureDoi="http://doi.org/10.5281/zenodo.5408599" httpUri="https://zenodo.org/record/5408599/files/figure.png" pageId="11" pageNumber="143">Fig. 5F–J</figureCitation>
|
||
) is smaller and exhibits some differences. The occlusal face (
|
||
<figureCitation id="B2F22A09FFCDA21FFB378E4B3414002A" box="[1249,1323,1024,1046]" captionStart="Figure 5" captionStartId="12.[164,243,1598,1617]" captionTargetBox="[172,1437,202,1563]" captionTargetId="figure-70@12.[164,1445,194,1568]" captionTargetPageId="12" captionText="Figure 5. Argoubia barbei gen. nov., sp. nov. A-E, Antero-lateral tooth, holotype, DAK3-1. A, lingual view; B, profile; C, labial view; D, occlusal view; E, basal view. F-J, Antero-lateral tooth, paratype, DAK3-2. F, lingual view; G, profile; H, labial view; I, occlusal view; J, basal view." figureDoi="http://doi.org/10.5281/zenodo.5408599" httpUri="https://zenodo.org/record/5408599/files/figure.png" pageId="11" pageNumber="143">Fig. 5I</figureCitation>
|
||
) is less distinctly separated from the labial face, with a lingual edge more strongly cut off. In profile view (
|
||
<figureCitation id="B2F22A09FFCDA21FFCF88E1732BA004E" box="[814,901,1116,1138]" captionStart="Figure 5" captionStartId="12.[164,243,1598,1617]" captionTargetBox="[172,1437,202,1563]" captionTargetId="figure-70@12.[164,1445,194,1568]" captionTargetPageId="12" captionText="Figure 5. Argoubia barbei gen. nov., sp. nov. A-E, Antero-lateral tooth, holotype, DAK3-1. A, lingual view; B, profile; C, labial view; D, occlusal view; E, basal view. F-J, Antero-lateral tooth, paratype, DAK3-2. F, lingual view; G, profile; H, labial view; I, occlusal view; J, basal view." figureDoi="http://doi.org/10.5281/zenodo.5408599" httpUri="https://zenodo.org/record/5408599/files/figure.png" pageId="11" pageNumber="143">Fig. 5G</figureCitation>
|
||
), the tooth is less high than the
|
||
<typeStatus id="F572882EFFCDA21FFAF38E1734B7004E" box="[1317,1416,1116,1138]" pageId="11" pageNumber="143" type="holotype">holotype</typeStatus>
|
||
, and, in occlusal (or basal) view (
|
||
<figureCitation id="B2F22A09FFCDA21FFB458E3135C400AC" box="[1171,1275,1146,1168]" captionStart="Figure 5" captionStartId="12.[164,243,1598,1617]" captionTargetBox="[172,1437,202,1563]" captionTargetId="figure-70@12.[164,1445,194,1568]" captionTargetPageId="12" captionText="Figure 5. Argoubia barbei gen. nov., sp. nov. A-E, Antero-lateral tooth, holotype, DAK3-1. A, lingual view; B, profile; C, labial view; D, occlusal view; E, basal view. F-J, Antero-lateral tooth, paratype, DAK3-2. F, lingual view; G, profile; H, labial view; I, occlusal view; J, basal view." figureDoi="http://doi.org/10.5281/zenodo.5408599" httpUri="https://zenodo.org/record/5408599/files/figure.png" pageId="11" pageNumber="143">Fig. 5I–J</figureCitation>
|
||
), an angular labial outline can be observed medially. The lingual face of the crown is less ornamented, with only three smooth vertical ridges. The labial face bears strong vertical crests but less irregular than in the
|
||
<typeStatus id="F572882EFFCDA21FFAF08EBE34B50137" box="[1318,1418,1269,1291]" pageId="11" pageNumber="143" type="holotype">holotype</typeStatus>
|
||
. The marginal outlines of the crown are not cut off in labial view. The root shows four lobes and three furrows (
|
||
<figureCitation id="B2F22A09FFCDA21FFC478F1A32DF015A" box="[913,992,1360,1383]" captionStart="Figure 5" captionStartId="12.[164,243,1598,1617]" captionTargetBox="[172,1437,202,1563]" captionTargetId="figure-70@12.[164,1445,194,1568]" captionTargetPageId="12" captionText="Figure 5. Argoubia barbei gen. nov., sp. nov. A-E, Antero-lateral tooth, holotype, DAK3-1. A, lingual view; B, profile; C, labial view; D, occlusal view; E, basal view. F-J, Antero-lateral tooth, paratype, DAK3-2. F, lingual view; G, profile; H, labial view; I, occlusal view; J, basal view." figureDoi="http://doi.org/10.5281/zenodo.5408599" httpUri="https://zenodo.org/record/5408599/files/figure.png" pageId="11" pageNumber="143">Fig. 5J</figureCitation>
|
||
). It is broader just below the crown than at the level of the basal face.
|
||
</paragraph>
|
||
<paragraph id="2A76368CFFCDA21FFCF08FE635460352" blockId="11.[806,1422,1452,1903]" pageId="11" pageNumber="143">
|
||
<emphasis id="18BDEA9EFFCDA21FFCF08FE6329801FE" box="[806,935,1453,1474]" italics="true" pageId="11" pageNumber="143">Discussion:</emphasis>
|
||
Despite the morphological differences observed between the
|
||
<typeStatus id="F572882EFFCDA21FFBF08F8035B801DD" box="[1062,1159,1483,1505]" pageId="11" pageNumber="143" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="F572882EFFCDA21FFB178F80341601DC" box="[1217,1321,1483,1504]" pageId="11" pageNumber="143" type="paratype">paratype</typeStatus>
|
||
, one can consider that they fall within the range of intraspecific variation. It is possible that the more salient and more cut off edge of the lingual face observed on the
|
||
<typeStatus id="F572882EFFCDA21FFCF08C0D32B30267" box="[806,908,1606,1627]" pageId="11" pageNumber="143" type="paratype">paratype</typeStatus>
|
||
corresponds to a dental sexual dimorphism. In this case, the
|
||
<typeStatus id="F572882EFFCDA21FFC2F8C2F35600245" box="[1017,1119,1636,1657]" pageId="11" pageNumber="143" type="paratype">paratype</typeStatus>
|
||
may have belonged to a male. Several other fragmentary teeth of this species have been collected. However, they were not illustrated because of their poor state of preservation. Some teeth are more transversally developed with a distinctly polyaulacorhize root, indicating that the teeth described above would be anterior or anterolateral teeth. These more lateral teeth are also characterized by a less high crown.
|
||
</paragraph>
|
||
<caption id="7EB66604FFCAA218FF728C75335202B7" ID-DOI="http://doi.org/10.5281/zenodo.5408599" ID-Zenodo-Dep="5408599" httpUri="https://zenodo.org/record/5408599/files/figure.png" pageId="12" pageNumber="144" startId="12.[164,243,1598,1617]" targetBox="[172,1437,202,1563]" targetPageId="12">
|
||
<paragraph id="2A76368CFFCAA218FF728C75335202B7" blockId="12.[164,1441,1598,1676]" pageId="12" pageNumber="144">
|
||
<emphasis id="18BDEA9EFFCAA218FF728C753394026D" bold="true" box="[164,683,1598,1617]" pageId="12" pageNumber="144">
|
||
Figure 5.
|
||
<taxonomicName id="EDC94D0FFFCAA218FECF8C7530E7026D" authority="Adnet & Cappetta & Guinot & Sciara, 2012" authorityName="Adnet & Cappetta & Guinot & Sciara" authorityYear="2012" box="[281,472,1598,1617]" class="Elasmobranchii" family="Myliobatidae" genus="Argoubia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="12" pageNumber="144" phylum="Chordata" rank="species" species="barbei" status="gen. nov., sp. nov.">
|
||
<emphasis id="18BDEA9EFFCAA218FECF8C7530E7026D" bold="true" box="[281,472,1598,1617]" italics="true" pageId="12" pageNumber="144">Argoubia barbei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="038E57E5FFCAA218FE098C753394026D" box="[479,683,1598,1617]" pageId="12" pageNumber="144" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A-E, Antero-lateral tooth, holotype, DAK3-1. A, lingual view; B, profile; C, labial view; D, occlusal view; E, basal view. F-J, Antero-lateral tooth, paratype, DAK3-2. F, lingual view; G, profile; H, labial view; I, occlusal view; J, basal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2A76368CFFCAA218FF6A8CB632480353" blockId="12.[164,779,1789,1903]" lastBlockId="12.[826,1441,1789,1903]" pageId="12" pageNumber="144">
|
||
<taxonomicName id="EDC94D0FFFCAA218FF6A8CB63049032E" authority="Adnet & Cappetta & Guinot & Sciara, 2012" authorityName="Adnet & Cappetta & Guinot & Sciara" authorityYear="2012" box="[188,374,1789,1810]" class="Elasmobranchii" family="Myliobatidae" genus="Argoubia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="12" pageNumber="144" phylum="Chordata" rank="species" species="barbei" status="gen. et sp. nov.">
|
||
<emphasis id="18BDEA9EFFCAA218FF6A8CB63049032E" box="[188,374,1789,1810]" italics="true" pageId="12" pageNumber="144">Argoubia barbei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="038E57E5FFCAA218FEA88CB63315032E" box="[382,554,1789,1810]" pageId="12" pageNumber="144" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
can be easily separated from
|
||
<taxonomicName id="EDC94D0FFFCAA218FEF08D57330A030D" authority="Adnet & Cappetta & Guinot & Sciara, 2012" authorityName="Adnet & Cappetta & Guinot & Sciara" authorityYear="2012" box="[294,565,1820,1841]" class="Elasmobranchii" family="Myliobatidae" genus="Oromobula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="12" pageNumber="144" phylum="Chordata" rank="species" species="dakhlaensis" status="gen. et sp. nov.">
|
||
<emphasis id="18BDEA9EFFCAA218FEF08D57330A030D" box="[294,565,1820,1841]" italics="true" pageId="12" pageNumber="144">Oromobula dakhlaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="038E57E5FFCAA218FDEB8D5733D7030D" box="[573,744,1820,1841]" pageId="12" pageNumber="144" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
by its relatively less high and much thicker teeth. The ornamentation of the labial and lingual faces of the crown is also stronger, and the root is more massive with less high lobes in the former. It is also important to note that these taxa do not co-exist in the same level.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |