239 lines
37 KiB
XML
239 lines
37 KiB
XML
<document id="5E2AE495BA4B9E4F3B3B1B111DD96DBA" ID-DOI="10.1111/j.1096-3642.2012.00844.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5408589" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630349144448" checkinUser="marcus" docAuthor="Adnet, Sylvain, Cappetta, Henri, Guinot, Guillaume & Sciara, Giuseppe Notarbartolo Di" docDate="2012" docId="A260879AFFC9A204FC908F7F32FE00F1" docLanguage="en" docName="j.1096-3642.2012.00844.x.pdf" docOrigin="Zoological Journal of the Linnean Society 166 (1)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2012.00844.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Eoplinthicus underwoodi Adnet & Cappetta & Guinot & Sciara 2012, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="148" masterDocId="5E59FFE2FFC6A214FFD68A4B313F043C" masterDocTitle="Evolutionary history of the devilrays (Chondrichthyes: Myliobatiformes) from fossil and morphological inference" masterLastPageNumber="159" masterPageNumber="132" pageNumber="147" updateTime="1699240823188" updateUser="plazi">
|
||
<mods:mods id="2F4DC7DBBDFB8A6CED427254A12E67DB" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="2266A5C70E673A460EE991D385FD567B">
|
||
<mods:title id="2FBAF1BD7F15910BC29537BBF8F1AF5E">Evolutionary history of the devilrays (Chondrichthyes: Myliobatiformes) from fossil and morphological inference</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="5CD3AF6826A30998A649E63B063914A0" type="personal">
|
||
<mods:role id="F0DF33C30276FB296626128A03D272CC">
|
||
<mods:roleTerm id="CC340F51CB7709DC04C02BDE94C6AB7F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="DAD78649C8AEA45E0F5C12B5CD4E1AB2">Adnet, Sylvain</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="D86AFCD674D37382D6826E444C053BED" type="personal">
|
||
<mods:role id="61EEA6CB9C24CB94AD868AF8B0A153EE">
|
||
<mods:roleTerm id="A0269B5C300E82A625DE123DB4E94A90">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="AB9026BF6E860A343578F91C50CFCFFC">Cappetta, Henri</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="294BD4C723174DCE3968784BBEE87F67" type="personal">
|
||
<mods:role id="865438D4B801A3BAD08BBDC6EAB6F0F5">
|
||
<mods:roleTerm id="A4F0E8BB85F30FA19C124D1CCE69937F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C6367976513B32CD0E03B0E38A1FB647">Guinot, Guillaume</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="F8058DB32B75888F902135238ABA61EA" type="personal">
|
||
<mods:role id="55E92FB00112F8BB77D6DF9EEAD97F18">
|
||
<mods:roleTerm id="F99FC0E5F2435D6E6E06C038548C50D7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8E58C362371F54742BDE00F70ED17765">Sciara, Giuseppe Notarbartolo Di</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="17E2A202B715980EB01FD2568DEE87F3">text</mods:typeOfResource>
|
||
<mods:relatedItem id="A54465BFABFA25F35A636DE2D234C4D6" type="host">
|
||
<mods:titleInfo id="5497F7C8A143AEBBAC5BEE621CCEA8EA">
|
||
<mods:title id="44082932E0482320CB17A9FE0691C83A">Zoological Journal of the Linnean Society</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="CA613A63E28FB7B93CC01017D103FF2B">
|
||
<mods:date id="97810585B3886322C969C9E5DE0408C6">2012</mods:date>
|
||
<mods:detail id="2B9740A94336ADB1D7D301F396D25BD8" type="pubDate">
|
||
<mods:number id="6A1985698BA6A5271948440AAF5A9E00">2015-07-06</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="EBC0ED261342EA0ED9EE5C439EFD12EE" type="volume">
|
||
<mods:number id="18FB1A48C64F179DD53EF92C5C492D5F">166</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="04B977FC5A283A5A006C9A46E554CE9E" type="issue">
|
||
<mods:number id="756353746D10A4E456FA815CE2A0D09B">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="887732985DA1358157C4F257BECD2DD9" unit="page">
|
||
<mods:start id="E31875A565320A6F8B360A9FE17D79BF">132</mods:start>
|
||
<mods:end id="7077E23CD8C8B60F1D84DE38581C631A">159</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="45062FC9945B3F5E520756161B8AA9DA">
|
||
<mods:url id="5BC20E597C869E65572E80694DCD4851">http://dx.doi.org/10.1111/j.1096-3642.2012.00844.x</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="89026A30B2F9EF88CC8BA0CA3F3F8F79">journal article</mods:classification>
|
||
<mods:identifier id="2CAADFC692334BDD1EFD539AC6EE9D19" type="DOI">10.1111/j.1096-3642.2012.00844.x</mods:identifier>
|
||
<mods:identifier id="E3AA6D00C5670FF9B8FF84FB7E20E1C5" type="ISSN">0024-4082</mods:identifier>
|
||
<mods:identifier id="B71A6DBCD9885DDEE78CD126BB2B640E" type="Zenodo-Dep">5408589</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="A260879AFFC9A204FC908F7F32FE00F1" LSID="urn:lsid:plazi:treatment:A260879AFFC9A204FC908F7F32FE00F1" httpUri="http://treatment.plazi.org/id/A260879AFFC9A204FC908F7F32FE00F1" lastPageId="16" lastPageNumber="148" pageId="15" pageNumber="147">
|
||
<subSubSection id="62D36507FFC9A21BFC908F7F34520175" box="[838,1389,1331,1353]" pageId="15" pageNumber="147" type="nomenclature">
|
||
<paragraph id="2A76368CFFC9A21BFC908F7F34520175" blockId="15.[810,1417,1300,1353]" box="[838,1389,1331,1353]" pageId="15" pageNumber="147">
|
||
<heading id="713E81E0FFC9A21BFC908F7F34520175" box="[838,1389,1331,1353]" centered="true" fontSize="9" level="2" pageId="15" pageNumber="147" reason="2">
|
||
<taxonomicName id="EDC94D0FFFC9A21BFC908F7F35A10174" authority="Adnet & Cappetta & Guinot & Sciara, 2012" authorityName="Adnet & Cappetta & Guinot & Sciara" authorityYear="2012" box="[838,1182,1332,1353]" class="Elasmobranchii" family="Mobulidae" genus="Eoplinthicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="15" pageNumber="147" phylum="Chordata" rank="species" species="underwoodi" status="sp. nov.">EOPLINTHICUS UNDERWOODI</taxonomicName>
|
||
<emphasis id="18BDEA9EFFC9A21BFB7F8F7C34310174" bold="true" box="[1193,1294,1331,1352]" pageId="15" pageNumber="147">
|
||
<taxonomicNameLabel id="038E57E5FFC9A21BFB7F8F7C34310174" box="[1193,1294,1331,1352]" pageId="15" pageNumber="147" rank="species">SP. NOV.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="B2F22A09FFC9A21BFAF68F78345A0175" box="[1312,1381,1331,1353]" captionStart="Figure 7" captionStartId="15.[144,223,1058,1077]" captionTargetBox="[144,1421,204,1018]" captionTargetId="figure-276@15.[144,1425,197,1028]" captionTargetPageId="15" captionText="Figure 7. Eoplinthicus underwoodi sp. nov. A-E, DAK2A-2, holotype: tooth position unknown. A, lingual view; B, labial view; C, occlusal view; D, basal view; E, profile. F-G, DAK2A-3, paratype, tooth position unknown. F, lingual view; G, occlusal view. H-J, DAK2B-2, tooth position unknown. H, lingual view; I, labial view; J, occlusal view. K-O, DAK2B-3: young element, tooth position unknown. K, lingual view; L, labial view; M, occlusal view; N, basal view; O, profile." figureDoi="http://doi.org/10.5281/zenodo.5408603" httpUri="https://zenodo.org/record/5408603/files/figure.png" pageId="15" pageNumber="147">FIG. 7</figureCitation>
|
||
)
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62D36507FFC9A204FCF08F1132FE00F1" lastPageId="16" lastPageNumber="148" pageId="15" pageNumber="147" type="materials_examined">
|
||
<paragraph id="2A76368CFFC9A21BFCF08F1134B8014C" blockId="15.[806,1415,1370,1392]" box="[806,1415,1370,1392]" pageId="15" pageNumber="147">
|
||
<emphasis id="18BDEA9EFFC9A21BFCF08F1132B00153" box="[806,911,1370,1391]" italics="true" pageId="15" pageNumber="147">Material:</emphasis>
|
||
Four complete teeth and many fragments.
|
||
</paragraph>
|
||
<paragraph id="2A76368CFFC9A21BFCF08FD7353301D3" blockId="15.[806,1421,1435,1519]" pageId="15" pageNumber="147">
|
||
<emphasis id="18BDEA9EFFC9A21BFCF08FD73280018C" box="[806,959,1435,1457]" italics="true" pageId="15" pageNumber="147">Type locality:</emphasis>
|
||
Locality DAK2. Unit 2, levels B1 and B2, south of Ad Dakhla, south-western
|
||
<collectingCountry id="52DE761CFFC9A21BFAE68FF134B201F3" box="[1328,1421,1466,1487]" name="Morocco" pageId="15" pageNumber="147">Morocco</collectingCountry>
|
||
(
|
||
<bibRefCitation id="4E584B7DFFC9A21BFCF88F92353F01D3" author="Adnet S & Cappetta H & Tabuce R" box="[814,1024,1497,1519]" pageId="15" pageNumber="147" pagination="860 - 870" refId="ref16589" refString="Adnet S, Cappetta H, Tabuce R. 2010. A Middle-Late Eocene vertebrate fauna (marine fish and mammals) from southwestern Morocco; preliminary report: age and palaeobiogeographical implications. Geological Magazine 147: 860 - 870." type="journal article" year="2010">
|
||
Adnet
|
||
<emphasis id="18BDEA9EFFC9A21BFCAB8F92329201D2" box="[893,941,1497,1518]" italics="true" pageId="15" pageNumber="147">et al</emphasis>
|
||
., 2010
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2A76368CFFC9A21BFCF08C5034B10272" blockId="15.[806,1422,1562,1615]" pageId="15" pageNumber="147">
|
||
<emphasis id="18BDEA9EFFC9A21BFCF08C5032F70213" box="[806,968,1562,1584]" italics="true" pageId="15" pageNumber="147">Type horizon:</emphasis>
|
||
Samlat Formation, Gerran member (
|
||
<bibRefCitation id="4E584B7DFFC9A21BFCF88C7232CF0273" author="Ratschiller LK" box="[814,1008,1593,1615]" pageId="15" pageNumber="147" pagination="55 - 190" refId="ref19042" refString="Ratschiller LK. 1967. Sahara, correlazioni geologicolithostratigrafiche fra Sahara Centrale ed Occidentale (con note gologiche generali e brevi cenni sulle possibilita dell'Africa Nord-Occidentale). Memorie del Museo Tridentino di Scienze Naturali 16: 55 - 190." type="journal article" year="1967">Ratschiller, 1967</bibRefCitation>
|
||
) – late Middle Eocene/Late Eocene.
|
||
</paragraph>
|
||
<paragraph id="2A76368CFFC9A21BFCF08C31357C0292" blockId="15.[806,1421,1658,1711]" pageId="15" pageNumber="147">
|
||
<emphasis id="18BDEA9EFFC9A21BFCF08C31329B02B3" box="[806,932,1658,1679]" italics="true" pageId="15" pageNumber="147">Etymology:</emphasis>
|
||
After C. J. Underwood, for his work on the Fayum faunas in
|
||
<collectingCountry id="52DE761CFFC9A21BFC2E8CD235000292" box="[1016,1087,1689,1710]" name="Egypt" pageId="15" pageNumber="147">Egypt</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="2A76368CFFC9A21BFCF08C9135A402CC" blockId="15.[806,1179,1754,1776]" box="[806,1179,1754,1776]" pageId="15" pageNumber="147">
|
||
<emphasis id="18BDEA9EFFC9A21BFCF08C9132AF02D3" box="[806,912,1754,1775]" italics="true" pageId="15" pageNumber="147">
|
||
<typeStatus id="F572882EFFC9A21BFCF08C9132B502D3" box="[806,906,1754,1775]" pageId="15" pageNumber="147" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
DAK2A-2 (
|
||
<figureCitation id="B2F22A09FFC9A21BFBCC8C9135AF02CC" box="[1050,1168,1754,1776]" captionStart="Figure 7" captionStartId="15.[144,223,1058,1077]" captionTargetBox="[144,1421,204,1018]" captionTargetId="figure-276@15.[144,1425,197,1028]" captionTargetPageId="15" captionText="Figure 7. Eoplinthicus underwoodi sp. nov. A-E, DAK2A-2, holotype: tooth position unknown. A, lingual view; B, labial view; C, occlusal view; D, basal view; E, profile. F-G, DAK2A-3, paratype, tooth position unknown. F, lingual view; G, occlusal view. H-J, DAK2B-2, tooth position unknown. H, lingual view; I, labial view; J, occlusal view. K-O, DAK2B-3: young element, tooth position unknown. K, lingual view; L, labial view; M, occlusal view; N, basal view; O, profile." figureDoi="http://doi.org/10.5281/zenodo.5408603" httpUri="https://zenodo.org/record/5408603/files/figure.png" pageId="15" pageNumber="147">Fig. 7A–E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2A76368CFFC9A204FCF08D57301E0632" blockId="15.[806,1421,1820,1903]" lastBlockId="16.[164,779,198,526]" lastPageId="16" lastPageNumber="148" pageId="15" pageNumber="147">
|
||
<emphasis id="18BDEA9EFFC9A21BFCF08D5732A2030D" box="[806,925,1820,1841]" italics="true" pageId="15" pageNumber="147">Diagnosis:</emphasis>
|
||
The teeth have a small size, less than
|
||
<quantity id="ED319B69FFC9A21BFCF08D713258036C" box="[806,871,1850,1872]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="147" unit="mm" value="5.0">5 mm</quantity>
|
||
broad, with a roughly hexagonal outline in occlusal view. Marginolabial and marginolingual angles very obtuse and rounded. Occlusal face as developed or less developed than the complete width of the crown, and very hollowed transversally. Oblique and well-developed labial face covered by corrugated enameloid. Sharp and salient labial visor and well-marked lingual bulge at the base of the crown. Root as high as the crown, with about six lobes separated by rather deep and broad W- shaped furrows. Sharp basal face of the lobes. The two marginal lobes may be shorter and diverging more than the others.
|
||
</paragraph>
|
||
<paragraph id="2A76368CFFD6A204FF72886D30400232" blockId="16.[164,779,550,1796]" pageId="16" pageNumber="148">
|
||
<emphasis id="18BDEA9EFFD6A204FF72886D30130607" box="[164,300,550,571]" italics="true" pageId="16" pageNumber="148">Description:</emphasis>
|
||
The
|
||
<typeStatus id="F572882EFFD6A204FEA0886D30E80600" box="[374,471,550,572]" pageId="16" pageNumber="148" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="B2F22A09FFD6A204FE38886D335E0600" box="[494,609,550,572]" captionStart="Figure 7" captionStartId="15.[144,223,1058,1077]" captionTargetBox="[144,1421,204,1018]" captionTargetId="figure-276@15.[144,1425,197,1028]" captionTargetPageId="15" captionText="Figure 7. Eoplinthicus underwoodi sp. nov. A-E, DAK2A-2, holotype: tooth position unknown. A, lingual view; B, labial view; C, occlusal view; D, basal view; E, profile. F-G, DAK2A-3, paratype, tooth position unknown. F, lingual view; G, occlusal view. H-J, DAK2B-2, tooth position unknown. H, lingual view; I, labial view; J, occlusal view. K-O, DAK2B-3: young element, tooth position unknown. K, lingual view; L, labial view; M, occlusal view; N, basal view; O, profile." figureDoi="http://doi.org/10.5281/zenodo.5408603" httpUri="https://zenodo.org/record/5408603/files/figure.png" pageId="16" pageNumber="148">Fig. 7A–E</figureCitation>
|
||
) is about 2.4 times broader (
|
||
<quantity id="ED319B69FFD6A204FE87880F30970666" box="[337,424,580,602]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="16" pageNumber="148" unit="mm" value="4.8">4.8 mm</quantity>
|
||
) than long (
|
||
<quantity id="ED319B69FFD6A204FDE5880F334A0666" box="[563,629,580,602]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="148" unit="mm" value="2.0">2 mm</quantity>
|
||
) and
|
||
<quantity id="ED319B69FFD6A204FD60880F32340666" box="[694,779,580,602]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="16" pageNumber="148" unit="mm" value="2.3">2.3 mm</quantity>
|
||
high. In occlusal view (
|
||
<figureCitation id="B2F22A09FFD6A204FE7D8828333F0645" box="[427,512,611,633]" captionStart="Figure 7" captionStartId="15.[144,223,1058,1077]" captionTargetBox="[144,1421,204,1018]" captionTargetId="figure-276@15.[144,1425,197,1028]" captionTargetPageId="15" captionText="Figure 7. Eoplinthicus underwoodi sp. nov. A-E, DAK2A-2, holotype: tooth position unknown. A, lingual view; B, labial view; C, occlusal view; D, basal view; E, profile. F-G, DAK2A-3, paratype, tooth position unknown. F, lingual view; G, occlusal view. H-J, DAK2B-2, tooth position unknown. H, lingual view; I, labial view; J, occlusal view. K-O, DAK2B-3: young element, tooth position unknown. K, lingual view; L, labial view; M, occlusal view; N, basal view; O, profile." figureDoi="http://doi.org/10.5281/zenodo.5408603" httpUri="https://zenodo.org/record/5408603/files/figure.png" pageId="16" pageNumber="148">Fig. 7C</figureCitation>
|
||
), the outline is roughly hexagonal, with marginal angles less than 90°. The occlusal face is narrower than the crown at the level of marginal angles and its shape is elliptical on the whole. This face has a deep transverse hollow with an angular bottom. This hollow is asymmetrical, with a labial wall more abrupt than the lingual one. Its surface is sometimes irregular, with a shiny enameloid. Its lingual margin is more irregular than its labial margin, which is rather convex. The labial face (
|
||
<figureCitation id="B2F22A09FFD6A204FF7A89DE31C00796" box="[172,255,917,939]" captionStart="Figure 7" captionStartId="15.[144,223,1058,1077]" captionTargetBox="[144,1421,204,1018]" captionTargetId="figure-276@15.[144,1425,197,1028]" captionTargetPageId="15" captionText="Figure 7. Eoplinthicus underwoodi sp. nov. A-E, DAK2A-2, holotype: tooth position unknown. A, lingual view; B, labial view; C, occlusal view; D, basal view; E, profile. F-G, DAK2A-3, paratype, tooth position unknown. F, lingual view; G, occlusal view. H-J, DAK2B-2, tooth position unknown. H, lingual view; I, labial view; J, occlusal view. K-O, DAK2B-3: young element, tooth position unknown. K, lingual view; L, labial view; M, occlusal view; N, basal view; O, profile." figureDoi="http://doi.org/10.5281/zenodo.5408603" httpUri="https://zenodo.org/record/5408603/files/figure.png" pageId="16" pageNumber="148">Fig. 7B</figureCitation>
|
||
) is oblique in profile (
|
||
<figureCitation id="B2F22A09FFD6A204FDC389DE33570796" box="[533,616,917,939]" captionStart="Figure 7" captionStartId="15.[144,223,1058,1077]" captionTargetBox="[144,1421,204,1018]" captionTargetId="figure-276@15.[144,1425,197,1028]" captionTargetPageId="15" captionText="Figure 7. Eoplinthicus underwoodi sp. nov. A-E, DAK2A-2, holotype: tooth position unknown. A, lingual view; B, labial view; C, occlusal view; D, basal view; E, profile. F-G, DAK2A-3, paratype, tooth position unknown. F, lingual view; G, occlusal view. H-J, DAK2B-2, tooth position unknown. H, lingual view; I, labial view; J, occlusal view. K-O, DAK2B-3: young element, tooth position unknown. K, lingual view; L, labial view; M, occlusal view; N, basal view; O, profile." figureDoi="http://doi.org/10.5281/zenodo.5408603" httpUri="https://zenodo.org/record/5408603/files/figure.png" pageId="16" pageNumber="148">Fig. 7E</figureCitation>
|
||
) and slightly convex. The labial visor is sharp and distinctly overhangs the root. The lower part of the visor is rather flat and well developed labiolingually. In occlusal view, the visor outline consists of a labial transversal segment slightly convex and of two shorter and straighter marginal segments, with strongly marked and rounded angles. The labial face is strongly corrugated. The lingual face is less developed than the labial one and more vertical in profile (
|
||
<figureCitation id="B2F22A09FFD6A204FDB98EE333FA0082" box="[623,709,1192,1214]" captionStart="Figure 7" captionStartId="15.[144,223,1058,1077]" captionTargetBox="[144,1421,204,1018]" captionTargetId="figure-276@15.[144,1425,197,1028]" captionTargetPageId="15" captionText="Figure 7. Eoplinthicus underwoodi sp. nov. A-E, DAK2A-2, holotype: tooth position unknown. A, lingual view; B, labial view; C, occlusal view; D, basal view; E, profile. F-G, DAK2A-3, paratype, tooth position unknown. F, lingual view; G, occlusal view. H-J, DAK2B-2, tooth position unknown. H, lingual view; I, labial view; J, occlusal view. K-O, DAK2B-3: young element, tooth position unknown. K, lingual view; L, labial view; M, occlusal view; N, basal view; O, profile." figureDoi="http://doi.org/10.5281/zenodo.5408603" httpUri="https://zenodo.org/record/5408603/files/figure.png" pageId="16" pageNumber="148">Fig. 7E</figureCitation>
|
||
). It is also very ornamented, with smooth vertical folds mainly in the median part of the face. In lingual view (
|
||
<figureCitation id="B2F22A09FFD6A204FF7A8F4F303E0126" box="[172,257,1284,1306]" captionStart="Figure 7" captionStartId="15.[144,223,1058,1077]" captionTargetBox="[144,1421,204,1018]" captionTargetId="figure-276@15.[144,1425,197,1028]" captionTargetPageId="15" captionText="Figure 7. Eoplinthicus underwoodi sp. nov. A-E, DAK2A-2, holotype: tooth position unknown. A, lingual view; B, labial view; C, occlusal view; D, basal view; E, profile. F-G, DAK2A-3, paratype, tooth position unknown. F, lingual view; G, occlusal view. H-J, DAK2B-2, tooth position unknown. H, lingual view; I, labial view; J, occlusal view. K-O, DAK2B-3: young element, tooth position unknown. K, lingual view; L, labial view; M, occlusal view; N, basal view; O, profile." figureDoi="http://doi.org/10.5281/zenodo.5408603" httpUri="https://zenodo.org/record/5408603/files/figure.png" pageId="16" pageNumber="148">Fig. 7A</figureCitation>
|
||
), oblique segments unite the marginal angles with the extremities of the occlusal face of the crown. The transversal bulge of the base of the lingual face is well developed, rounded, and salient. The root is practically as high as the crown. It consists of six lobes separated by broad and deep W- shaped furrows. The marginal lobes are shorter than the median ones and even diverging. Their basal face is sharp and not flattened (
|
||
<figureCitation id="B2F22A09FFD6A204FECD8FB2304C0232" box="[283,371,1529,1551]" captionStart="Figure 7" captionStartId="15.[144,223,1058,1077]" captionTargetBox="[144,1421,204,1018]" captionTargetId="figure-276@15.[144,1425,197,1028]" captionTargetPageId="15" captionText="Figure 7. Eoplinthicus underwoodi sp. nov. A-E, DAK2A-2, holotype: tooth position unknown. A, lingual view; B, labial view; C, occlusal view; D, basal view; E, profile. F-G, DAK2A-3, paratype, tooth position unknown. F, lingual view; G, occlusal view. H-J, DAK2B-2, tooth position unknown. H, lingual view; I, labial view; J, occlusal view. K-O, DAK2B-3: young element, tooth position unknown. K, lingual view; L, labial view; M, occlusal view; N, basal view; O, profile." figureDoi="http://doi.org/10.5281/zenodo.5408603" httpUri="https://zenodo.org/record/5408603/files/figure.png" pageId="16" pageNumber="148">Fig. 7D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2A76368CFFD6A204FF6A8C5C33940338" blockId="16.[164,779,550,1796]" pageId="16" pageNumber="148">
|
||
The other specimens (
|
||
<figureCitation id="B2F22A09FFD6A204FE188C5C337E0212" box="[462,577,1559,1582]" captionStart="Figure 7" captionStartId="15.[144,223,1058,1077]" captionTargetBox="[144,1421,204,1018]" captionTargetId="figure-276@15.[144,1425,197,1028]" captionTargetPageId="15" captionText="Figure 7. Eoplinthicus underwoodi sp. nov. A-E, DAK2A-2, holotype: tooth position unknown. A, lingual view; B, labial view; C, occlusal view; D, basal view; E, profile. F-G, DAK2A-3, paratype, tooth position unknown. F, lingual view; G, occlusal view. H-J, DAK2B-2, tooth position unknown. H, lingual view; I, labial view; J, occlusal view. K-O, DAK2B-3: young element, tooth position unknown. K, lingual view; L, labial view; M, occlusal view; N, basal view; O, profile." figureDoi="http://doi.org/10.5281/zenodo.5408603" httpUri="https://zenodo.org/record/5408603/files/figure.png" pageId="16" pageNumber="148">Fig. 7F–O</figureCitation>
|
||
) have a similar morphology. Their occlusal face is sometimes more transversally developed, with vertical segments joining their extremities and the marginal angles in lingual view (
|
||
<figureCitation id="B2F22A09FFD6A204FE518CD930E5029B" box="[391,474,1682,1704]" captionStart="Figure 7" captionStartId="15.[144,223,1058,1077]" captionTargetBox="[144,1421,204,1018]" captionTargetId="figure-276@15.[144,1425,197,1028]" captionTargetPageId="15" captionText="Figure 7. Eoplinthicus underwoodi sp. nov. A-E, DAK2A-2, holotype: tooth position unknown. A, lingual view; B, labial view; C, occlusal view; D, basal view; E, profile. F-G, DAK2A-3, paratype, tooth position unknown. F, lingual view; G, occlusal view. H-J, DAK2B-2, tooth position unknown. H, lingual view; I, labial view; J, occlusal view. K-O, DAK2B-3: young element, tooth position unknown. K, lingual view; L, labial view; M, occlusal view; N, basal view; O, profile." figureDoi="http://doi.org/10.5281/zenodo.5408603" httpUri="https://zenodo.org/record/5408603/files/figure.png" pageId="16" pageNumber="148">Fig. 7F</figureCitation>
|
||
). The largest specimen (
|
||
<figureCitation id="B2F22A09FFD6A204FF7A8CFB302202FA" box="[172,285,1712,1734]" captionStart="Figure 7" captionStartId="15.[144,223,1058,1077]" captionTargetBox="[144,1421,204,1018]" captionTargetId="figure-276@15.[144,1425,197,1028]" captionTargetPageId="15" captionText="Figure 7. Eoplinthicus underwoodi sp. nov. A-E, DAK2A-2, holotype: tooth position unknown. A, lingual view; B, labial view; C, occlusal view; D, basal view; E, profile. F-G, DAK2A-3, paratype, tooth position unknown. F, lingual view; G, occlusal view. H-J, DAK2B-2, tooth position unknown. H, lingual view; I, labial view; J, occlusal view. K-O, DAK2B-3: young element, tooth position unknown. K, lingual view; L, labial view; M, occlusal view; N, basal view; O, profile." figureDoi="http://doi.org/10.5281/zenodo.5408603" httpUri="https://zenodo.org/record/5408603/files/figure.png" pageId="16" pageNumber="148">Fig. 7H–J</figureCitation>
|
||
) has a polyaulacorhize root that consists of nine lobes, irregularly spaced, whereas the smallest (
|
||
<figureCitation id="B2F22A09FFD6A204FF7A8CA5301E0338" box="[172,289,1774,1796]" captionStart="Figure 7" captionStartId="15.[144,223,1058,1077]" captionTargetBox="[144,1421,204,1018]" captionTargetId="figure-276@15.[144,1425,197,1028]" captionTargetPageId="15" captionText="Figure 7. Eoplinthicus underwoodi sp. nov. A-E, DAK2A-2, holotype: tooth position unknown. A, lingual view; B, labial view; C, occlusal view; D, basal view; E, profile. F-G, DAK2A-3, paratype, tooth position unknown. F, lingual view; G, occlusal view. H-J, DAK2B-2, tooth position unknown. H, lingual view; I, labial view; J, occlusal view. K-O, DAK2B-3: young element, tooth position unknown. K, lingual view; L, labial view; M, occlusal view; N, basal view; O, profile." figureDoi="http://doi.org/10.5281/zenodo.5408603" httpUri="https://zenodo.org/record/5408603/files/figure.png" pageId="16" pageNumber="148">Fig. 7K–O</figureCitation>
|
||
) has a less concave occlusal face.
|
||
</paragraph>
|
||
<paragraph id="2A76368CFFD6A204FF728D5732FE00F1" blockId="16.[164,779,1820,1903]" lastBlockId="16.[826,1442,198,1229]" pageId="16" pageNumber="148">
|
||
<emphasis id="18BDEA9EFFD6A204FF728D57301B030D" box="[164,292,1820,1841]" italics="true" pageId="16" pageNumber="148">Discussion:</emphasis>
|
||
Compared with the
|
||
<typeStatus id="F572882EFFD6A204FDC18D573377030D" box="[535,584,1820,1841]" pageId="16" pageNumber="148">type</typeStatus>
|
||
species †
|
||
<taxonomicName id="EDC94D0FFFD6A204FD618D5733A20373" authority="Cappetta & Stringer, 2002" authorityName="Cappetta & Stringer" authorityYear="2002" class="Elasmobranchii" family="Mobulidae" genus="Eoplinthicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="148" phylum="Chordata" rank="species" species="yazooensis">
|
||
<emphasis id="18BDEA9EFFD6A204FD618D573059036C" italics="true" pageId="16" pageNumber="148">Eoplinthicus yazooensis</emphasis>
|
||
<bibRefCitation id="4E584B7DFFD6A204FEBB8D7133A20373" author="Cappetta H & Stringer GL" box="[365,669,1850,1872]" pageId="16" pageNumber="148" pagination="51 - 56" refId="ref17057" refString="Cappetta H, Stringer GL. 2002. A new batoid genus (Neoselachii: Myliobatiformes) from the Yazoo Clay (Upper Eocene) of Louisiana, U. S. A. Tertiary Research 21: 51 - 56." type="journal article" year="2002">Cappetta & Stringer, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
, from the Priabonian of
|
||
<collectingRegion id="E80DF86EFFD6A204FE8E8D1230F30353" box="[344,460,1881,1903]" country="United States of America" name="Louisiana" pageId="16" pageNumber="148">Louisiana</collectingRegion>
|
||
,
|
||
<collectingCountry id="52DE761CFFD6A204FE378D1233260352" box="[481,537,1881,1902]" name="United States of America" pageId="16" pageNumber="148">USA</collectingCountry>
|
||
, several differences can be noted. The teeth of †
|
||
<taxonomicName id="EDC94D0FFFD6A204FBA28A8D342504E7" authority="Adnet & Cappetta & Guinot & Sciara, 2012" authorityName="Adnet & Cappetta & Guinot & Sciara" authorityYear="2012" box="[1140,1306,198,219]" class="Elasmobranchii" family="Mobulidae" genus="Eoplinthicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="148" phylum="Chordata" rank="species" species="underwoodi" status="sp. nov.">
|
||
<emphasis id="18BDEA9EFFD6A204FBA28A8D342504E7" box="[1140,1306,198,219]" italics="true" pageId="16" pageNumber="148">E. underwoodi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="038E57E5FFD6A204FAF78A8D344904E7" box="[1313,1398,198,219]" pageId="16" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
|
||
are smaller and much less transversally elongated. Their occlusal face is more lingually developed, with a deeper transverse hollow that does not show the ‘lip-like’ design of the
|
||
<typeStatus id="F572882EFFD6A204FB938B0A3549056A" box="[1093,1142,321,342]" pageId="16" pageNumber="148">type</typeStatus>
|
||
species. The ornamentation of the labial face is less corrugated. The differences are more marked at the level of the root. In the
|
||
<typeStatus id="F572882EFFD6A204FCEC8BD73254058D" box="[826,875,412,433]" pageId="16" pageNumber="148">type</typeStatus>
|
||
species, the lobes are more numerous (six versus 11/12), with a flat basal face, and the base of the root is parallel to the base of the crown in labial or lingual view. Such differences seem to indicate a less derived morphology in †
|
||
<taxonomicName id="EDC94D0FFFD6A204FC2C885C359F0617" authority="Adnet & Cappetta & Guinot & Sciara, 2012" authorityName="Adnet & Cappetta & Guinot & Sciara" authorityYear="2012" box="[1018,1184,534,556]" class="Elasmobranchii" family="Mobulidae" genus="Eoplinthicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="148" phylum="Chordata" rank="species" species="underwoodi" status="sp. nov.">
|
||
<emphasis id="18BDEA9EFFD6A204FC2C885C359F0617" box="[1018,1184,534,556]" italics="true" pageId="16" pageNumber="148">E. underwoodi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="038E57E5FFD6A204FB7A885C34390610" box="[1196,1286,535,556]" pageId="16" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
|
||
compared to †
|
||
<taxonomicName id="EDC94D0FFFD6A204FC90887D32E30677" authorityName="Cappetta & Stringer" authorityYear="2002" box="[838,988,566,587]" class="Elasmobranchii" family="Mobulidae" genus="Eoplinthicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="148" phylum="Chordata" rank="species" species="yazooensis">
|
||
<emphasis id="18BDEA9EFFD6A204FC90887D32E30677" box="[838,988,566,587]" italics="true" pageId="16" pageNumber="148">E. yazooensis</emphasis>
|
||
</taxonomicName>
|
||
and species of †
|
||
<taxonomicName id="EDC94D0FFFD6A204FB48887E342C0676" authorityName="Cope" authorityYear="1869" box="[1182,1299,565,586]" class="Elasmobranchii" family="Myliobatidae" genus="Plinthicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="148" phylum="Chordata" rank="genus">
|
||
<emphasis id="18BDEA9EFFD6A204FB48887E342C0676" box="[1182,1299,565,586]" italics="true" pageId="16" pageNumber="148">Plinthicus</emphasis>
|
||
</taxonomicName>
|
||
, resembling the morphology of some species of †
|
||
<taxonomicName id="EDC94D0FFFD6A204FAC0881F34A50655" authorityName="CAPPETTA" authorityYear="1976" box="[1302,1434,596,617]" class="Elasmobranchii" family="Mobulidae" genus="Burnhamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="148" phylum="Chordata" rank="genus">
|
||
<emphasis id="18BDEA9EFFD6A204FAC0881F34A50655" box="[1302,1434,596,617]" italics="true" pageId="16" pageNumber="148">Burnhamia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="4E584B7DFFD6A204FCEC8839354906B4" author="Cappetta H & Stringer GL" box="[826,1142,626,648]" pageId="16" pageNumber="148" pagination="51 - 56" refId="ref17057" refString="Cappetta H, Stringer GL. 2002. A new batoid genus (Neoselachii: Myliobatiformes) from the Yazoo Clay (Upper Eocene) of Louisiana, U. S. A. Tertiary Research 21: 51 - 56." type="journal article" year="2002">Cappetta & Stringer (2002)</bibRefCitation>
|
||
suggested that the crown was little marked by functional wear as in mobulids and contrary to the rhinopterids and myliobatids. The same functional assumption could be seriously made with the concave occlusal surface observed in material from Dakhla. At present, the new species †
|
||
<taxonomicName id="EDC94D0FFFD6A204FC90896132D30703" authority="Adnet & Cappetta & Guinot & Sciara, 2012" authorityName="Adnet & Cappetta & Guinot & Sciara" authorityYear="2012" box="[838,1004,810,831]" class="Elasmobranchii" family="Mobulidae" genus="Eoplinthicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="148" phylum="Chordata" rank="species" species="underwoodi" status="sp. nov.">
|
||
<emphasis id="18BDEA9EFFD6A204FC90896132D30703" box="[838,1004,810,831]" italics="true" pageId="16" pageNumber="148">E. underwoodi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="038E57E5FFD6A204FC28896135610703" box="[1022,1118,810,831]" pageId="16" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is known only from the Bartonian–Priabonian deposits of south-western
|
||
<collectingCountry id="52DE761CFFD6A204FCEC892C32A80740" box="[826,919,871,892]" name="Morocco" pageId="16" pageNumber="148">Morocco</collectingCountry>
|
||
although its presence may be spatially extended to the contemporaneous deposits from Fayum,
|
||
<collectingCountry id="52DE761CFFD6A204FC4C89EE32E10786" box="[922,990,933,954]" name="Egypt" pageId="16" pageNumber="148">Egypt</collectingCountry>
|
||
(
|
||
<bibRefCitation id="4E584B7DFFD6A204FC3889EF35C90785" author="Underwood CJ & Ward DJ & King C & Antar SM & Zalmout IS & Gingerich PD" box="[1006,1270,932,954]" pageId="16" pageNumber="148" pagination="47 - 66" refId="ref19560" refString="Underwood CJ, Ward DJ, King C, Antar SM, Zalmout IS, Gingerich PD. 2011. Shark and ray faunas in the Middle and Late Eocene of the Fayum Area, Egypt. Proceedings of the Geologists' Association of London 122: 47 - 66." type="journal article" year="2011">
|
||
Underwood
|
||
<emphasis id="18BDEA9EFFD6A204FBAC89EE35950785" box="[1146,1194,932,954]" italics="true" pageId="16" pageNumber="148">et al</emphasis>
|
||
., 2011
|
||
</bibRefCitation>
|
||
as †
|
||
<taxonomicName id="EDC94D0FFFD6A204FAFB89EF32B307E4" class="Elasmobranchii" family="Mobulidae" genus="Eoplinthicus" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="148" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="18BDEA9EFFD6A204FAFB89EF326007E4" italics="true" pageId="16" pageNumber="148">Eoplinthicus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
). The occurrence of this species elsewhere is suspected but currently remains debatable.
|
||
<bibRefCitation id="4E584B7DFFD6A204FA9F89AA32F9002A" author="Case GR & West RM" pageId="16" pageNumber="148" pagination="105 - 120" refId="ref17281" refString="Case GR, West RM. 1991. Geology and paleontology of the Eocene Drazinda Shale Member of the Kirthar Formation, central Western Pakistan. Part II: Late Eocene fishes. Tertiary Research 12: 105 - 120." type="journal article" year="1991">Case & West (1991)</bibRefCitation>
|
||
reported a single broken tooth as
|
||
<taxonomicName id="EDC94D0FFFD6A204FAB98E4B32810008" class="Elasmobranchii" family="Myliobatidae" genus="Rhinoptera" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="148" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="18BDEA9EFFD6A204FAB98E4B32AF0008" italics="true" pageId="16" pageNumber="148">Rhinoptera</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
from the Late Eocene of
|
||
<collectingCountry id="52DE761CFFD6A204FB288E54345B0009" box="[1278,1380,1055,1077]" name="Pakistan" pageId="16" pageNumber="148">Pakistan</collectingCountry>
|
||
that shows a deflected extremity of the crown in labial view (
|
||
<bibRefCitation id="4E584B7DFFD6A204FC538E173548004D" author="Case GR & West RM" box="[901,1143,1116,1137]" pageId="16" pageNumber="148" pagination="105 - 120" refId="ref17281" refString="Case GR, West RM. 1991. Geology and paleontology of the Eocene Drazinda Shale Member of the Kirthar Formation, central Western Pakistan. Part II: Late Eocene fishes. Tertiary Research 12: 105 - 120." type="journal article" year="1991">Case & West, 1991</bibRefCitation>
|
||
: pl. 4, fig. 1), indicating that the occlusal face is straighter than the base of the crown, as in †
|
||
<taxonomicName id="EDC94D0FFFD6A204FB928ED235EB0092" authorityName="Cappetta & Stringer" authorityYear="2002" box="[1092,1236,1177,1198]" class="Elasmobranchii" family="Mobulidae" genus="Eoplinthicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="148" phylum="Chordata" rank="genus">
|
||
<emphasis id="18BDEA9EFFD6A204FB928ED235EB0092" box="[1092,1236,1177,1198]" italics="true" pageId="16" pageNumber="148">Eoplinthicus</emphasis>
|
||
</taxonomicName>
|
||
and contrary to
|
||
<taxonomicName id="EDC94D0FFFD6A204FCEC8EFC328500F0" box="[826,954,1207,1228]" class="Elasmobranchii" family="Myliobatidae" genus="Rhinoptera" kingdom="Animalia" order="Myliobatiformes" pageId="16" pageNumber="148" phylum="Chordata" rank="genus">
|
||
<emphasis id="18BDEA9EFFD6A204FCEC8EFC328500F0" box="[826,954,1207,1228]" italics="true" pageId="16" pageNumber="148">Rhinoptera</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |