156 lines
17 KiB
XML
156 lines
17 KiB
XML
<document id="B119383A8B55FA07B6796AA9C6933157" ID-CLB-Dataset="172794" ID-DOI="10.11646/zootaxa.5200.1.2" ID-GBIF-Dataset="f95b3c02-51d2-4d9b-8792-fee09a84c2a8" ID-ISSN="1175-5326" ID-Zenodo-Dep="7251836" ID-ZooBank="0C794D44-12BB-42F9-BFCF-EF9B850341EE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666770162308" checkinUser="plazi" docAuthor="Moreno-Juárez, Eric G., Villalobos, José Luis & Álvarez, Fernando" docDate="2022" docId="A2414922F011FF94FF5BF999FD0BFADC" docLanguage="en" docName="zootaxa.5199.1.2.pdf" docOrigin="Zootaxa 5200 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mokayathelphusa Moreno, Villalobos & Alvarez 2022, n. gen." docType="treatment" docVersion="7" lastPageNumber="28" masterDocId="5E78315AF012FF90FFCCFF8BFF96FFE0" masterDocTitle="Two new genera and one new species of freshwater crabs of the subfamily Pseudothelphusinae (Decapoda: Brachyura: Pseudothelphusidae) from southwestern Mexico" masterLastPageNumber="36" masterPageNumber="24" pageNumber="27" updateTime="1699408215080" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="9CB3862B1B4BE02DA5776AE85BED880E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="674EB991B61F2B1588BE6341688ADE70">
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<mods:title id="AE7250157CAD287DA70FDCDB3E0028FA">Two new genera and one new species of freshwater crabs of the subfamily Pseudothelphusinae (Decapoda: Brachyura: Pseudothelphusidae) from southwestern Mexico</mods:title>
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<mods:name id="D22216C5829A80FF0B1D408C8C1F6DD9" type="personal">
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<mods:roleTerm id="9FFE85031B7DCDA1E408BE45BEF2032E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2E6D5F55FB617003629139B037B880EC">Moreno-Juárez, Eric G.</mods:namePart>
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<mods:affiliation id="ACA82CAF0A892F934DDAA24E7CD8FBD0">Posgrado en Ciencias Biológicas, Universidad Nacional Autónoma de México, 04510 Ciudad de México, Mexico. & Colección Nacional de Crustáceos, Instituto de Biología, Universidad Nacional Autónoma de México, 04510 Ciudad de México, Mexico.</mods:affiliation>
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<mods:name id="17E48E35D4C25546C0E5186567D5BB93" type="personal">
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<mods:roleTerm id="ED652A44B3CD30D8295CAFA69AB7E223">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3BC0D7531F48ACCA3FE678FCF67467B6">Villalobos, José Luis</mods:namePart>
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<mods:affiliation id="B91621BE862322CB9910711B22A4B551">Colección Nacional de Crustáceos, Instituto de Biología, Universidad Nacional Autónoma de México, 04510 Ciudad de México, Mexico.</mods:affiliation>
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</mods:name>
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<mods:name id="1905EF422711926234564604676BFB1F" type="personal">
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<mods:roleTerm id="ADA10D52A5139D3E3652D3A36F0D0FB8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7711D5276D9CB2B1D4CF53ADBEE34B1E">Álvarez, Fernando</mods:namePart>
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<mods:affiliation id="632F0067775275042665C0D9BC0B2B0E">Colección Nacional de Crustáceos, Instituto de Biología, Universidad Nacional Autónoma de México, 04510 Ciudad de México, Mexico.</mods:affiliation>
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<mods:typeOfResource id="4C91726334EA2F3D35BD0935C3481E35">text</mods:typeOfResource>
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<mods:title id="05A01C38AC968C6D47A6FBD47A420C58">Zootaxa</mods:title>
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<mods:part id="E63D9EFF2FCACD937E0D32FE68EEA008">
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<mods:date id="506AC5F63C080950C530B4017955EDD5">2022</mods:date>
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<mods:number id="0E365CCA9A2061730FAAA5E1F2B603C2">2022-10-26</mods:number>
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<mods:number id="18104EFD6BEFEFF1A74CC6B1234A71FC">5200</mods:number>
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<mods:number id="03037F80D7C9F517D47620EEC24A7417">1</mods:number>
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<mods:start id="F7FE71D9BBFCDAA3255E603D64ABE268">24</mods:start>
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<treatment id="A2414922F011FF94FF5BF999FD0BFADC" ID-DOI="http://doi.org/10.5281/zenodo.7251746" ID-GBIF-Taxon="203398717" ID-Zenodo-Dep="7251746" LSID="urn:lsid:plazi:treatment:A2414922F011FF94FF5BF999FD0BFADC" httpUri="http://treatment.plazi.org/id/A2414922F011FF94FF5BF999FD0BFADC" lastPageId="4" lastPageNumber="28" pageId="3" pageNumber="27">
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<subSubSection id="62F2ABBFF011FF93FF5BF999FC5FF9B1" pageId="3" pageNumber="27" type="nomenclature">
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<paragraph id="2A57F834F011FF93FF5BF999FCD0F9CD" blockId="3.[151,969,1554,1617]" box="[151,838,1554,1584]" pageId="3" pageNumber="27">
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<heading id="711F4F58F011FF93FF5BF999FCD0F9CD" bold="true" box="[151,838,1554,1584]" fontSize="11" level="1" pageId="3" pageNumber="27" reason="1">
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<emphasis id="189C2426F011FF93FF5BF999FCD0F9CD" bold="true" box="[151,838,1554,1584]" pageId="3" pageNumber="27">
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<taxonomicName id="EDE883B7F011FF93FF5BF999FD7BF9D0" ID-CoL="a92d38db-d4ef-46a5-b1fa-e401275b3495" authority="Moreno, Villalobos & Alvarez" authorityName="Moreno, Villalobos & Alvarez" authorityYear="2022" box="[151,749,1554,1584]" class="Malacostraca" family="Pseudothelphusidae" genus="Mokayathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="27" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="189C2426F011FF93FF5BF999FEE6F9CD" bold="true" box="[151,368,1554,1581]" italics="true" pageId="3" pageNumber="27">Mokayathelphusa</emphasis>
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Moreno, Villalobos & Álvarez
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</taxonomicName>
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<taxonomicNameLabel id="03AF995DF011FF93FD38F998FCD0F9CD" box="[756,838,1555,1581]" pageId="3" pageNumber="27" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="2A57F834F011FF93FF5BF9BDFC5FF9B1" blockId="3.[151,969,1554,1617]" box="[151,969,1590,1617]" pageId="3" pageNumber="27">
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<uri id="5E79F436F011FF93FF5BF9BDFC5FF9B1" box="[151,969,1590,1617]" pageId="3" pageNumber="27">
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urn:lsid:zoobank.org:act:
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<uuid id="5E4EC2E1F011FF93FE60F9BDFC5FF9B1" box="[428,969,1590,1617]" pageId="3" pageNumber="27">A06B6E9E-4953-4B25-9DC9-3F1C592ACF4F</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="62F2ABBFF011FF93FF5BF9F3FDF5F8C2" pageId="3" pageNumber="27" type="diagnosis">
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<paragraph id="2A57F834F011FF93FF5BF9F3FDF5F8C2" blockId="3.[151,1437,1656,2042]" pageId="3" pageNumber="27">
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<emphasis id="189C2426F011FF93FF5BF9F3FE84F972" bold="true" box="[151,274,1656,1682]" pageId="3" pageNumber="27">Diagnosis.</emphasis>
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G1 with the distal portion of principal axis twisted counterclockwise. Apical cavity oriented mesocephalically, compressed and closed cephalically by the internal angle of mesial process. Caudo-marginal projection trilobed distally, middle and distal lobes spiniform of same length, proximal lobe as elongated sheet, curved, extending through distal third of main gonopod axis. Mesial process large, subrectangular, with lateral margin ornamented with several spiniform teeth.
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</paragraph>
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</subSubSection>
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<subSubSection id="62F2ABBFF011FF93FF0BF8A7FB92F8A6" box="[199,1028,1836,1863]" pageId="3" pageNumber="27" type="type_taxon">
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<paragraph id="2A57F834F011FF93FF0BF8A7FB92F8A6" blockId="3.[151,1437,1656,2042]" box="[199,1028,1836,1863]" pageId="3" pageNumber="27">
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<emphasis id="189C2426F011FF93FF0BF8A7FEF4F8A6" bold="true" box="[199,354,1836,1862]" pageId="3" pageNumber="27">
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<typeStatus id="F5534696F011FF93FF0BF8A7FE94F8A6" box="[199,258,1836,1862]" pageId="3" pageNumber="27">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="EDE883B7F011FF93FEA5F8A7FD3AF8A6" ID-CoL="abf9c48e-6380-47ab-bdc6-2a0285498aba" authority="Moreno-Juárez & Villalobos & Álvarez, 2022" authorityName="Moreno-Juárez & Villalobos & Álvarez" authorityYear="2022" box="[361,684,1836,1862]" class="Malacostraca" family="Pseudothelphusidae" genus="Mokayathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="27" phylum="Arthropoda" rank="species" species="angelsotoi" status="sp. nov.">
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<emphasis id="189C2426F011FF93FEA5F8A7FD3AF8A6" box="[361,684,1836,1862]" italics="true" pageId="3" pageNumber="27">Mokayathelphusa angelsotoi</emphasis>
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</taxonomicName>
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<emphasis id="189C2426F011FF93FD7FF8A7FD64F8A6" bold="true" box="[691,754,1836,1862]" pageId="3" pageNumber="27">
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<taxonomicNameLabel id="03AF995DF011FF93FD7FF8A7FD64F8A6" box="[691,754,1836,1862]" pageId="3" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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, by present designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="62F2ABBFF011FF93FF0BF8DBFDE9F852" pageId="3" pageNumber="27" type="etymology">
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<paragraph id="2A57F834F011FF93FF0BF8DBFDE9F852" blockId="3.[151,1437,1656,2042]" pageId="3" pageNumber="27">
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<emphasis id="189C2426F011FF93FF0BF8DBFED8F88A" bold="true" box="[199,334,1872,1898]" pageId="3" pageNumber="27">Etymology.</emphasis>
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The name of the genus is taken from the ancient tribe “Mokaya” that inhabited the Chimalapas region about 3,600 ybp. The word “Mokaya” comes from the earliest Zoque language, that means “people of the corn”. The gender of this name is feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="62F2ABBFF011FF93FF0BF837FF62F81A" pageId="3" pageNumber="27" type="distribution">
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<paragraph id="2A57F834F011FF93FF0BF837FF62F81A" blockId="3.[151,1437,1656,2042]" pageId="3" pageNumber="27">
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<emphasis id="189C2426F011FF93FF0BF837FEF7F836" bold="true" box="[199,353,1980,2006]" pageId="3" pageNumber="27">Distribution.</emphasis>
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Currently known only from the municipality of Santa María Chimalapa, State of
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<collectingRegion id="E82C36D6F011FF93FA8DF837FA0EF836" box="[1345,1432,1980,2006]" country="Mexico" name="Oaxaca" pageId="3" pageNumber="27">Oaxaca</collectingRegion>
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,
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<collectingCountry id="52FFB8A4F011FF93FF5BF86BFF66F81A" box="[151,240,2016,2042]" name="Mexico" pageId="3" pageNumber="27">Mexico</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<caption id="7E97A8BCF016FF94FF5BFC13FD85FC34" ID-DOI="http://doi.org/10.5281/zenodo.7251840" ID-Zenodo-Dep="7251840" httpUri="https://zenodo.org/record/7251840/files/figure.png" pageId="4" pageNumber="28" startId="4.[151,250,920,944]" targetBox="[375,1215,186,895]" targetPageId="4">
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<paragraph id="2A57F834F016FF94FF5BFC13FD85FC34" blockId="4.[151,1436,920,980]" pageId="4" pageNumber="28">
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<emphasis id="189C2426F016FF94FF5BFC13FE84FC50" bold="true" box="[151,274,920,944]" pageId="4" pageNumber="28">FIGURE 2.</emphasis>
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Consensus tree resulting from ML and BI analyzes of the concatenated matrix (COX 1, 16S and H3) from the genus of the
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<taxonomicName id="EDE883B7F016FF94FF14FC37FE37FC34" authorityName="Ortmann" authorityYear="1893" box="[216,417,956,980]" class="Malacostraca" family="Pseudothelphusidae" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="29" phylum="Arthropoda" rank="subFamily" subFamily="Pseudothelphusinae">Pseudothelphusinae</taxonomicName>
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subfamily.
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</paragraph>
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</caption>
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<subSubSection id="62F2ABBFF016FF94FF0BFB89FD0BFADC" pageId="4" pageNumber="28" type="discussion">
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<paragraph id="2A57F834F016FF94FF0BFB89FD0BFADC" blockId="4.[151,1437,1026,1340]" pageId="4" pageNumber="28">
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<emphasis id="189C2426F016FF94FF0BFB89FEADFBFC" bold="true" box="[199,315,1026,1052]" pageId="4" pageNumber="28">Remarks.</emphasis>
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The description of
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<taxonomicName id="EDE883B7F016FF94FDEFFB89FD7CFBFC" authority="Moreno-Juárez & Villalobos & Álvarez, 2022" authorityName="Moreno-Juárez & Villalobos & Álvarez" authorityYear="2022" box="[547,746,1026,1052]" class="Malacostraca" family="Pseudothelphusidae" genus="Mokayathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="28" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="189C2426F016FF94FDEFFB89FD7CFBFC" box="[547,746,1026,1052]" italics="true" pageId="4" pageNumber="28">Mokayathelphusa</emphasis>
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</taxonomicName>
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<emphasis id="189C2426F016FF94FD38FB89FCD3FBFC" bold="true" box="[756,837,1026,1052]" pageId="4" pageNumber="28">
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<taxonomicNameLabel id="03AF995DF016FF94FD38FB89FCD3FBFC" box="[756,837,1026,1052]" pageId="4" pageNumber="28" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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is justified by its unique male G1 morphology and clear genetic separation from related genera. The new genus can be related to other genera of the subfamily
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<taxonomicName id="EDE883B7F016FF94FF5BFBC1FEE3FB84" authorityName="Ortmann" authorityYear="1893" box="[151,373,1098,1124]" class="Malacostraca" family="Pseudothelphusidae" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="28" phylum="Arthropoda" rank="subFamily" subFamily="Pseudothelphusinae">Pseudothelphusinae</taxonomicName>
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by the presence of the caudo-marginal projection, which ends distally in a trilobed distal plate. The phylogenetic analysis (
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<figureCitation id="B2D3E4B1F016FF94FE02FBE5FD84FB68" box="[462,530,1134,1160]" captionStart="FIGURE 2" captionStartId="4.[151,250,920,944]" captionTargetBox="[375,1215,186,895]" captionTargetId="figure-19@4.[371,1216,181,896]" captionTargetPageId="4" captionText="FIGURE 2. Consensus tree resulting from ML and BI analyzes of the concatenated matrix (COX 1, 16S and H3) from the genus of the Pseudothelphusinae subfamily." figureDoi="http://doi.org/10.5281/zenodo.7251840" httpUri="https://zenodo.org/record/7251840/files/figure.png" pageId="4" pageNumber="28">Fig. 2</figureCitation>
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) recovers it as the sister lineage to
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<taxonomicName id="EDE883B7F016FF94FC69FBE5FA0EFB68" authority="Smalley & Adkison, 1984" authorityName="Smalley & Adkison" authorityYear="1984" box="[933,1432,1134,1160]" class="Malacostraca" family="Pseudothelphusidae" genus="Disparithelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="28" phylum="Arthropoda" rank="genus">
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<emphasis id="189C2426F016FF94FC69FBE5FBF1FB68" box="[933,1127,1134,1160]" italics="true" pageId="4" pageNumber="28">Disparithelphusa</emphasis>
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<bibRefCitation id="4E7985C5F016FF94FBA3FBE5FA0EFB68" author="Smalley, A. E. & Adkison, D. L." box="[1135,1432,1134,1160]" pageId="4" pageNumber="28" pagination="127 - 133" refId="ref9071" refString="Smalley, A. E. & Adkison, D. L. (1984) Disparithelphusa pecki, a new genus and species of fresh water crab from Mexico (Brachyura: Pseudothelphusidae). Journal of Crustacean Biology, 4 (1), 127 - 133. https: // doi. org / 10.2307 / 1547901" type="journal article" year="1984">Smalley & Adkison, 1984</bibRefCitation>
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</taxonomicName>
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. Morphologically the G1 of both genera present a torsion that modifies the shape and orientation of the apical cavity and have a trilobed distal plate in the caudo-marginal projection, where the distal lobes are smaller than the proximal one. They can also be differentiated by the size of the mesial process and the ornamentation of its lateral border, while in
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<taxonomicName id="EDE883B7F016FF94FF33FB75FE57FAF8" authorityName="Smalley & Adkison" authorityYear="1984" box="[255,449,1278,1304]" class="Malacostraca" family="Pseudothelphusidae" genus="Disparithelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="28" phylum="Arthropoda" rank="genus">
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<emphasis id="189C2426F016FF94FF33FB75FE57FAF8" box="[255,449,1278,1304]" italics="true" pageId="4" pageNumber="28">Disparithelphusa</emphasis>
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</taxonomicName>
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it is small, oval and without lateral ornamentation; in
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<taxonomicName id="EDE883B7F016FF94FBF1FB75FA92FAF8" authority="Moreno-Juárez & Villalobos & Álvarez, 2022" authorityName="Moreno-Juárez & Villalobos & Álvarez" authorityYear="2022" box="[1085,1284,1278,1304]" class="Malacostraca" family="Pseudothelphusidae" genus="Mokayathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="28" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="189C2426F016FF94FBF1FB75FA92FAF8" box="[1085,1284,1278,1304]" italics="true" pageId="4" pageNumber="28">Mokayathelphusa</emphasis>
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</taxonomicName>
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<emphasis id="189C2426F016FF94FAC2FB75FAC9FAF8" bold="true" box="[1294,1375,1278,1304]" pageId="4" pageNumber="28">
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<taxonomicNameLabel id="03AF995DF016FF94FAC2FB75FAC9FAF8" box="[1294,1375,1278,1304]" pageId="4" pageNumber="28" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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, it is evidently developed and ornamented laterally.
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</paragraph>
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</subSubSection>
|
|
</treatment>
|
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</document> |