235 lines
33 KiB
XML
235 lines
33 KiB
XML
<document id="4A7AB722BCC7100025F82F20AEA229BD" ID-DOI="10.5281/zenodo.5172580" ID-ISSN="1942-1354" ID-Zenodo-Dep="5172580" ID-ZooBank="F8317A32-DFD7-42EA-8BC6-DCB5A3659F6C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628270718788" checkinUser="felipe" docAuthor="Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa" docDate="2017" docId="A24087A739680427FF7FFEB1FA06FB2C" docLanguage="en" docName="InsectaMundi.0547.1-36.pdf" docOrigin="Insecta Mundi 2017 (547)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Tomarus rosettae" docType="treatment" docVersion="4" lastPageNumber="10" masterDocId="5E79FFDF3963042CFFBFFF93FF92FF96" masterDocTitle="The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay" masterLastPageNumber="36" masterPageNumber="1" pageNumber="10" updateTime="1699038600670" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
|
||
<mods:mods id="C74BF68AC44E3C88E6717B139DFA31C3" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="D1AED7EB6D78D638B7B714235B29A573">
|
||
<mods:title id="D980C80BE615C2553425A87EE169C6DA">The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="720200BD460AA3093ADCBDA3DAD75797" type="personal">
|
||
<mods:role id="3D3FB5615A941CAB2A0F4FFCB0DBE40E">
|
||
<mods:roleTerm id="EE6284D2B0A910988998526F24D766F9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2E72D685BE225E41DFF30DF470560D07">Neita-Moreno, Jhon César</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="5644FEE8A27B7B3CC70DBB24D94A6478" type="personal">
|
||
<mods:role id="BFB7C52F794DBB61D993124BFBF510FC">
|
||
<mods:roleTerm id="E7692B26A1D27251BCA5F886C9913023">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B79825B6B0172FF55D1FE2D25FE3F73B">Alexander, Recursos Biológicos</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="1BCC4B2CF5AC39C472D902406B30B767" type="personal">
|
||
<mods:role id="6C795A86B0FB0AE37F92CE92B963613A">
|
||
<mods:roleTerm id="58CBE53CFB8B1571CA690F02023DD53F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B434534D80D8379C4C0C12FEADEDCB21">Agustín, San</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="F47EEF97E09D57371B126706906BDD66" type="personal">
|
||
<mods:role id="3C70A8B6609B5F01AB19F40E256353E9">
|
||
<mods:roleTerm id="864BB8EC3B4268DA1CA921FDD5A49B91">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E0AC75D969D23880EEED48393451F5ED">de, Villa</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="B7A11EFD41D54AA59FBB7156EF3DCFA9" type="personal">
|
||
<mods:role id="D2F5A04B9F00A47629973952634D590E">
|
||
<mods:roleTerm id="09F7E4569BDC3FD1DA8F57F562B47008">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="AC5C7FC90CD468DBB2263B4BC061A751">Leyva</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="46ED363B0A7CA1AD4FAFF043A10A3AC4" type="personal">
|
||
<mods:role id="BA191696D92BA9317DC579E1678E0D05">
|
||
<mods:roleTerm id="8FB1B3A5654B129092D3A1AB4FB25EED">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1C5F4B6F5CEA72AD31C23E206F63CD6D">Boyacá</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="408F8679A874898C714DE674E40C46E0" type="personal">
|
||
<mods:role id="8BEFD62BA5A9F5B313940CD766A414F0">
|
||
<mods:roleTerm id="1DB75FCF112AEFDE42E483055E7CCC94">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="DB54CFFEFC3645893BD17EEAD009F1C5">Ratcliffe, Brett C.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="7E38686C8575493168214F88321160EC" type="personal">
|
||
<mods:role id="88D818614CE8B757B114148649121C2E">
|
||
<mods:roleTerm id="6A2D7658C4549F973B15F4F4A5BFEF12">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F62EFF958ECA39674FDC844248E9772E">Collections, Systematics Research</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="E12318991B272C4FF78D3E783C3284CE" type="personal">
|
||
<mods:role id="1ACBE7D886963BAAA45E4719A45F72C8">
|
||
<mods:roleTerm id="BDAA4A149D8276485FC4E050EF3008EB">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C5FACE368285027D1B49624F43FAB8C2">of, University</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="EB23DC931C3A5F6F2277D4F74B8F8896" type="personal">
|
||
<mods:role id="C59FEBFAEBF77CBA2E7FB038B912CCE7">
|
||
<mods:roleTerm id="E2BDECEA58E259BA4785B1C0BAC6E4C7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="86ACE1D0CBE9F60872C528E29F0D89D4">Museum, Nebraska State</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="A4A9F14332813D0027EE93D48708E8AF" type="personal">
|
||
<mods:role id="04541D7C9F423E6BAB62EBA7F5A5A343">
|
||
<mods:roleTerm id="A27D3292C8B37688B64F869666FA1224">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1AE66B4017F5202248B40A9CE14696DF">Hall, Nebraska</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="4FD2C225D20D8B6C1CD584609DF99514" type="personal">
|
||
<mods:role id="9362E3DFA7F73CC3F6BA58EB7E83B497">
|
||
<mods:roleTerm id="4E0F3C5EA2DE26AFF59F72B6FDF76302">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="7CC4A84DF304AA982092BD0DB5982779">of, University</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="686EE98DA1442C27A292BACC57EE3F7D" type="personal">
|
||
<mods:role id="A7FD1BF5843EEB458DEAC7EB3B052811">
|
||
<mods:roleTerm id="D5300DF3238F891F89A85F7827DE2AA0">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="5BA68EB364A014B37F6491DB2A3DC148">Nebraska</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="710A79E6F5FFBBC43AB0B18A8967AA96" type="personal">
|
||
<mods:role id="946187616F3F453AEEF21E90E767D0C9">
|
||
<mods:roleTerm id="E9C0AB1504D14A51C37D0384C1F7BD22">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="66F7201D8063BC12D921C8DD35312BED">Lincoln</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="0CFC0FF8E7A2A304B791FDB5ED887A2C" type="personal">
|
||
<mods:role id="591F251ADC782F09B459776AABC11229">
|
||
<mods:roleTerm id="B04E3E5F4B271DD4B03773BC9AF2B04E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C5FEE72C6BCB36797B189B43FDAC55FE">Ne</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="E76C2E27763518EB4A12DC6C64D90466" type="personal">
|
||
<mods:role id="A361F2E9A81FE887EE143950B809E56C">
|
||
<mods:roleTerm id="7D9EC90F672568E5E522CD17D41F5D34">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="3C9F1D775918FCA1F35E7FB46A9B9AF2">Usa</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="E75E60B6468195D1AC38193818FEA3EF">text</mods:typeOfResource>
|
||
<mods:relatedItem id="B83A97F47C7510473771A80465058004" type="host">
|
||
<mods:titleInfo id="2134DBBE2C0E9DFAA8E5778767A522E8">
|
||
<mods:title id="7D2BFA8EF724E60F75372772761CEA15">Insecta Mundi</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="30BCAB0973253770E8D8CD6E05720B2D">
|
||
<mods:date id="CCEC7836EC23B7F20A046DD05A471B15">2017</mods:date>
|
||
<mods:detail id="3CE3D8291B67C959F9AF81DD53C0EA30" type="pubDate">
|
||
<mods:number id="272BABFAD63DF5E0BA7FB47DC5F54E7D">2017-04-28</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="725EBDE7F2B795FE4043788F65401873" type="volume">
|
||
<mods:number id="4C36B106C1930991CD8C9E3139295BE0">2017</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="294E3BF38598CEA9F05E96A5EAF76A36" type="issue">
|
||
<mods:number id="F32276424200F89FE216A8B3091457F7">547</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="011976BB12D68A38A9F0F1AC85BDAB2F" unit="page">
|
||
<mods:start id="88997E9B034CFDE424E2B16F4CC8DD3B">1</mods:start>
|
||
<mods:end id="E77B7461FF82D78200AC0A0AC95DAE3C">36</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="8B627EEB10357318CB470DF199FFE4FB">journal article</mods:classification>
|
||
<mods:identifier id="3FC427D8884A64767324B1C77E42FA22" type="DOI">10.5281/zenodo.5172580</mods:identifier>
|
||
<mods:identifier id="5BA48A882D4AA3B6F7E0F945BEF759FE" type="ISSN">1942-1354</mods:identifier>
|
||
<mods:identifier id="9F56BB23AD1B7302965480BEEEC15C4B" type="Zenodo-Dep">5172580</mods:identifier>
|
||
<mods:identifier id="91CEBF79E4A262256E345739E659CDA0" type="ZooBank">F8317A32-DFD7-42EA-8BC6-DCB5A3659F6C</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="A24087A739680427FF7FFEB1FA06FB2C" ID-DOI="http://doi.org/10.5281/zenodo.5185988" ID-Zenodo-Dep="5185988" LSID="urn:lsid:plazi:treatment:A24087A739680427FF7FFEB1FA06FB2C" httpUri="http://treatment.plazi.org/id/A24087A739680427FF7FFEB1FA06FB2C" lastPageNumber="10" pageId="11" pageNumber="10">
|
||
<subSubSection id="62F3653A39680427FF7FFEB1FD18FEAF" box="[192,650,289,313]" pageId="11" pageNumber="10" type="nomenclature">
|
||
<paragraph id="2A5636B139680427FF7FFEB1FD18FEAF" blockId="11.[192,1010,289,346]" box="[192,650,289,313]" pageId="11" pageNumber="10">
|
||
<heading id="711E81DD39680427FF7FFEB1FD18FEAF" bold="true" box="[192,650,289,313]" fontSize="10" level="2" pageId="11" pageNumber="10" reason="3">
|
||
<taxonomicName id="EDE94D3239680427FF7FFEB1FD18FEAF" authority="(Endrodi, 1968)" baseAuthorityName="Endrodi" baseAuthorityYear="1968" box="[192,650,289,313]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="rosettae">
|
||
<emphasis id="189DEAA339680427FF7FFEB1FD18FEAF" bold="true" box="[192,650,289,313]" pageId="11" pageNumber="10">
|
||
<emphasis id="189DEAA339680427FF7FFEB1FE3DFEAF" bold="true" box="[192,431,290,313]" italics="true" pageId="11" pageNumber="10">Tomarus rosettae</emphasis>
|
||
(
|
||
<bibRefCitation id="4E784B4039680427FE7FFEB2FD13FEAF" author="Endrodi, S." box="[448,641,289,313]" pageId="11" pageNumber="10" pagination="161 - 177" refId="ref14056" refString="Endrodi, S. 1968. Neue Arten der Pentodontini (Col. Dynastinae). Folia Entomologica Hungarica (N. S.) 21: 161 - 177." type="journal article" year="1968">Endrödi, 1968</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62F3653A39680427FF7FFED1FC60FECC" box="[192,1010,321,346]" pageId="11" pageNumber="10" type="description">
|
||
<paragraph id="2A5636B139680427FF7FFED1FC60FECC" blockId="11.[192,1010,289,346]" box="[192,1010,321,346]" pageId="11" pageNumber="10">
|
||
(
|
||
<figureCitation id="B2D22A3439680427FF76FED1FEB2FECC" box="[201,288,322,346]" captionStart="Figure 1" captionStartId="19.[192,276,1846,1868]" captionTargetBox="[207,1426,292,1810]" captionTargetId="figure-64@19.[192,1441,192,1828]" captionTargetPageId="19" captionText="Figure 1. Habitus of Tomarus spp. a) Tomarus bidentulus (Fairmaire) dorsal view. b) Tomarus burmeisteri (Steinheil) dorsal view. c) Tomarus gyas Erichson dorsal view. d) Tomarus rosettae (Endrödi) dorsal view. e) Tomarus roigjunenti n. sp. dorsal view, holotype. f) Tomarus roigjunenti n. sp. dorsal view, allotype." figureDoi="http://doi.org/10.5281/zenodo.5172584" httpUri="https://zenodo.org/record/5172584/files/figure.png" pageId="11" pageNumber="10">Fig. 1d</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439680427FE92FED1FEDFFECC" box="[301,333,322,346]" captionStart="Figure 2" captionStartId="20.[192,276,1817,1839]" captionTargetBox="[229,1331,230,1795]" captionTargetId="figure-18@20.[192,1440,192,1807]" captionTargetPageId="20" captionText="Figure 2. Habitus of Tomarus spp. a) Tomarus rosettae (Endrödi) dorsal view. b) Tomarus spinipenis n. sp. dorsal view, holotype. c) Tomarus spinipenis dorsal view, allotype. d) Tomarus burmeisteri (Steinheil) dorsal view. e) Tomarus villosus (Burmeister) ventral view. f) Tomarus gyas ventral view. g) Tomarus bidentulus (Fairmaire) abdomen segments, ventral view (female). h) Tomarus bidentulus abdomen segments, ventral view (male)." figureDoi="http://doi.org/10.5281/zenodo.5172588" httpUri="https://zenodo.org/record/5172588/files/figure.png" pageId="11" pageNumber="10">2a</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439680427FEE5FED1FEE1FECC" box="[346,371,322,346]" captionStart="Figure 3" captionStartId="21.[192,276,1727,1749]" captionTargetBox="[197,1361,257,1710]" captionTargetId="figure-124@21.[197,1375,192,1717]" captionTargetPageId="21" captionText="Figure 3. Types of Tomarus spp. a) T. bidentulus female holotype, dorsal view. b) T. burmeisteri male holotype, dorsal view. c) T. burmeisteri male holotype, lateral view. d) T. burmeisteri male holotype parameres, caudal view. e) T. burmeisteri male holotype parameres (lateral view). f) Label for T. burmeisteri holotype. g) T. gyas male holotype, dorsal view and labels and parameres, caudal view). h) T. maimon holotype, dorsal view. i) T. rosettae male holotype, dorsal view. j) T. villosus male holotype, dorsal view. k) T. villosus male holotype, dorsal view of head. l) T. villosus male holotype, parameres, caudal view. m) Label for T. villosus holotype." figureDoi="http://doi.org/10.5281/zenodo.5172591" httpUri="https://zenodo.org/record/5172591/files/figure.png" pageId="11" pageNumber="10">3i</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439680427FE3FFED1FE09FECC" box="[384,411,322,346]" captionStart="Figure 4" captionStartId="22.[192,276,1848,1870]" captionTargetBox="[216,1440,216,1825]" captionTargetId="figure-63@22.[192,1442,192,1842]" captionTargetPageId="22" captionText="Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172595" httpUri="https://zenodo.org/record/5172595/files/figure.png" pageId="11" pageNumber="10">4f</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439680427FE19FED2FE52FECF" box="[422,448,321,345]" captionStart="Figure 5" captionStartId="23.[192,276,1853,1875]" captionTargetBox="[215,1440,192,1823]" captionTargetId="figure-63@23.[192,1443,192,1843]" captionTargetPageId="23" captionText="Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172599" httpUri="https://zenodo.org/record/5172599/files/figure.png" pageId="11" pageNumber="10">5f</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439680427FE72FED1FD97FECC" box="[461,517,322,346]" captionStart="Figure 6" captionStartId="24.[192,276,1739,1761]" captionTargetBox="[211,1436,218,1720]" captionTargetId="figure-63@24.[192,1440,192,1730]" captionTargetPageId="24" captionText="Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172601" httpUri="https://zenodo.org/record/5172601/files/figure.png" pageId="11" pageNumber="10">6k–l</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439680427FDAEFED1FDBEFECC" box="[529,556,322,346]" captionStart="Figure 7" captionStartId="25.[192,276,1683,1705]" captionTargetBox="[231,1392,238,1664]" captionTargetId="figure-63@25.[192,1440,192,1671]" captionTargetPageId="25" captionText="Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172605" httpUri="https://zenodo.org/record/5172605/files/figure.png" pageId="11" pageNumber="10">7f</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439680427FD87FED1FDC1FECC" box="[568,595,322,346]" captionStart="Figure 8" captionStartId="26.[192,276,1841,1863]" captionTargetBox="[275,1367,208,1829]" captionTargetId="figure-18@26.[269,1376,192,1840]" captionTargetPageId="26" captionText="Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172607" httpUri="https://zenodo.org/record/5172607/files/figure.png" pageId="11" pageNumber="10">8f</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439680427FDE0FED1FD34FECC" box="[607,678,322,346]" captionStart="Figure 10" captionStartId="28.[192,275,1624,1646]" captionTargetBox="[220,1411,210,1601]" captionTargetId="figure-65@28.[192,1440,192,1613]" captionTargetPageId="28" captionText="Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172614" httpUri="https://zenodo.org/record/5172614/files/figure.png" pageId="11" pageNumber="10">10k–l</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439680427FD0DFED1FD49FECC" box="[690,731,322,346]" captionStart="Figure 11" captionStartId="29.[192,276,1835,1857]" captionTargetBox="[244,1411,241,1815]" captionTargetId="figure-63@29.[192,1442,192,1821]" captionTargetPageId="29" captionText="Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172618" httpUri="https://zenodo.org/record/5172618/files/figure.png" pageId="11" pageNumber="10">11f</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439680427FD58FED1FC82FECC" box="[743,784,322,346]" captionStart="Figure 12" captionStartId="30.[192,274,1776,1798]" captionTargetBox="[210,1434,220,1735]" captionTargetId="figure-61@30.[192,1440,192,1769]" captionTargetPageId="30" captionText="Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172620" httpUri="https://zenodo.org/record/5172620/files/figure.png" pageId="11" pageNumber="10">12f</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439680427FCA2FED1FCD4FECC" box="[797,838,322,346]" captionStart="Figure 13" captionStartId="31.[192,275,1823,1845]" captionTargetBox="[238,1403,236,1798]" captionTargetId="figure-63@31.[192,1440,192,1807]" captionTargetPageId="31" captionText="Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172622" httpUri="https://zenodo.org/record/5172622/files/figure.png" pageId="11" pageNumber="10">13f</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439680427FCEDFED1FCE9FECC" box="[850,891,322,346]" captionStart="Figure 14" captionStartId="32.[192,276,1832,1854]" captionTargetBox="[358,1267,208,1798]" captionTargetId="figure-70@32.[349,1298,192,1821]" captionTargetPageId="32" captionText="Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite." figureDoi="http://doi.org/10.5281/zenodo.5172626" httpUri="https://zenodo.org/record/5172626/files/figure.png" pageId="11" pageNumber="10">14f</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439680427FC38FED2FC23FECF" box="[903,945,321,345]" captionStart="Figure 15" captionStartId="33.[192,276,1817,1839]" captionTargetBox="[238,1409,229,1784]" captionTargetId="figure-88@33.[192,1440,192,1807]" captionTargetPageId="33" captionText="Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness." figureDoi="http://doi.org/10.5281/zenodo.5172630" httpUri="https://zenodo.org/record/5172630/files/figure.png" pageId="11" pageNumber="10">15f</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439680427FC02FED1FC78FECC" box="[957,1002,322,346]" captionStart="Figure 17" captionStartId="35.[192,276,1818,1840]" captionTargetBox="[235,1415,214,1804]" captionTargetId="figure-44@35.[192,1440,192,1807]" captionTargetPageId="35" captionText="Figure 17. Distribution of Tomarus spp. a) T. roigjunenti n. sp. b) T. rosettae. c) T. spinipenis n. sp. d) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172634" httpUri="https://zenodo.org/record/5172634/files/figure.png" pageId="11" pageNumber="10">17b</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62F3653A39680427FF7FFE11FC1CFE0C" box="[192,910,386,410]" pageId="11" pageNumber="10" type="reference_group">
|
||
<paragraph id="2A5636B139680427FF7FFE11FC1CFE0C" blockId="11.[192,910,386,410]" box="[192,910,386,410]" pageId="11" pageNumber="10">
|
||
<treatmentCitationGroup id="0AF9119F39680427FF7FFE11FC1CFE0C" box="[192,910,386,410]" pageId="11" pageNumber="10">
|
||
<treatmentCitation id="AB4810A039680427FF7FFE11FDFFFE0C" author="Endrodi, S." box="[192,621,386,410]" page="168" pageId="11" pageNumber="10" year="1968">
|
||
<taxonomicName id="EDE94D3239680427FF7FFE11FDFFFE0C" authority="Endrodi 1968: 168" authorityName="Endrodi" authorityPageNumber="168" authorityYear="1968" box="[192,621,386,410]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="rosettae">
|
||
<emphasis id="189DEAA339680427FF7FFE11FE17FE0F" box="[192,389,386,409]" italics="true" pageId="11" pageNumber="10">Ligyrus rosettae</emphasis>
|
||
<bibRefCitation id="4E784B4039680427FE33FE11FDFFFE0C" author="Endrodi, S." box="[396,621,386,410]" pageId="11" pageNumber="10" pagination="161 - 177" refId="ref14056" refString="Endrodi, S. 1968. Neue Arten der Pentodontini (Col. Dynastinae). Folia Entomologica Hungarica (N. S.) 21: 161 - 177." type="journal article" year="1968">Endrödi 1968: 168</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
(original combination).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62F3653A39680427FF7FFE51FA06FB2C" pageId="11" pageNumber="10" type="description">
|
||
<paragraph id="2A5636B139680427FF7FFE51FB9BFBCF" blockId="11.[192,1441,449,1114]" pageId="11" pageNumber="10">
|
||
Redescription. Male. Length
|
||
<quantity id="ED119B5439680427FD95FE52FD4EFE4F" box="[554,732,449,473]" metricMagnitude="-2" metricUnit="m" metricValue="2.045" metricValueMax="2.15" metricValueMin="1.94" pageId="11" pageNumber="10" unit="mm" value="20.45" valueMax="21.5" valueMin="19.4">19.4–21.5 mm</quantity>
|
||
; width
|
||
<quantity id="ED119B5439680427FC85FE51FC4CFE4F" box="[826,990,450,474]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" metricValueMax="1.08" metricValueMin="0.94" pageId="11" pageNumber="10" unit="mm" value="10.1" valueMax="10.8" valueMin="9.4">9.4–10.8 mm</quantity>
|
||
. Color dark reddish brown to black. Head: Frons and clypeus coarsely rugose. Frontoclypeal region with 2 distinctly transverse tubercles; tubercles relatively narrowly separated, usually about 2.5 tubercle diameters apart (
|
||
<figureCitation id="B2D22A3439680427FB6AFD91FABAFD8C" box="[1237,1320,514,538]" captionStart="Figure 4" captionStartId="22.[192,276,1848,1870]" captionTargetBox="[216,1440,216,1825]" captionTargetId="figure-63@22.[192,1442,192,1842]" captionTargetPageId="22" captionText="Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172595" httpUri="https://zenodo.org/record/5172595/files/figure.png" pageId="11" pageNumber="10">Fig. 4f</figureCitation>
|
||
). Clypeus strongly rugopunctate to transversely rugose, tapering to broad, slightly emarginate apex with 2 reflexed teeth. Mandibles with 2 teeth, apical tooth rounded, subapical notch shallow (
|
||
<figureCitation id="B2D22A3439680427FB79FDD1FAA7FDCC" box="[1222,1333,578,602]" captionStart="Figure 6" captionStartId="24.[192,276,1739,1761]" captionTargetBox="[211,1436,218,1720]" captionTargetId="figure-63@24.[192,1440,192,1730]" captionTargetPageId="24" captionText="Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172601" httpUri="https://zenodo.org/record/5172601/files/figure.png" pageId="11" pageNumber="10">Fig. 6k–l</figureCitation>
|
||
); labium densely setose on sides, paraglossa developed, apex with small tooth at center (
|
||
<figureCitation id="B2D22A3439680427FB25FDF1FB7FFDEC" box="[1178,1261,610,634]" captionStart="Figure 7" captionStartId="25.[192,276,1683,1705]" captionTargetBox="[231,1392,238,1664]" captionTargetId="figure-63@25.[192,1440,192,1671]" captionTargetPageId="25" captionText="Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172605" httpUri="https://zenodo.org/record/5172605/files/figure.png" pageId="11" pageNumber="10">Fig. 7f</figureCitation>
|
||
); maxilla with galea developed and with 6 teeth (
|
||
<figureCitation id="B2D22A3439680427FDCAFD11FD58FD0F" box="[629,714,641,666]" captionStart="Figure 5" captionStartId="23.[192,276,1853,1875]" captionTargetBox="[215,1440,192,1823]" captionTargetId="figure-63@23.[192,1443,192,1843]" captionTargetPageId="23" captionText="Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172599" httpUri="https://zenodo.org/record/5172599/files/figure.png" pageId="11" pageNumber="10">Fig. 5f</figureCitation>
|
||
). Pronotum: Surface with disc moderately punctate, sides and anterior angles densely punctate to rugopunctate; punctures moderate in size on disc, becoming larger laterally; fovea rugose within. Anterior margin with strong, median tubercle; fovea small, as wide as width between eyes (
|
||
<figureCitation id="B2D22A3439680427FD8DFD71FD15FD6C" box="[562,647,738,762]" captionStart="Figure 8" captionStartId="26.[192,276,1841,1863]" captionTargetBox="[275,1367,208,1829]" captionTargetId="figure-18@26.[269,1376,192,1840]" captionTargetPageId="26" captionText="Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172607" httpUri="https://zenodo.org/record/5172607/files/figure.png" pageId="11" pageNumber="10">Fig. 8f</figureCitation>
|
||
). Elytra: Surface densely punctate; punctures moderate in size with sparser, minute punctures between larger punctures; larger punctures usually ocellate, and with 3 pairs of double rows. Pygidium: Surface densely punctate, punctures becoming denser in basal and lateral angles; punctures small to moderate in size, weakly ocellate. In lateral view, male with convex surface, surface weakly convex in females. Legs: Protibia tridentate and with weak, basal convexity suggestive of 4
|
||
<superScript id="DD9C9BF939680427FECEFC13FE13FC19" attach="left" box="[369,385,896,911]" fontSize="6" pageId="11" pageNumber="10">th</superScript>
|
||
tooth, teeth subequally spaced from each other. Protarsus in male simple, not enlarged. Mesotibia relatively broad, transverse carinae arcuate, spinules long, stout (
|
||
<figureCitation id="B2D22A3439680427FBE5FC31FB4AFC2C" box="[1114,1240,930,954]" captionStart="Figure 10" captionStartId="28.[192,275,1624,1646]" captionTargetBox="[220,1411,210,1601]" captionTargetId="figure-65@28.[192,1440,192,1613]" captionTargetPageId="28" captionText="Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172614" httpUri="https://zenodo.org/record/5172614/files/figure.png" pageId="11" pageNumber="10">Fig. 10k–l</figureCitation>
|
||
). Metatibia with 1 basal, transverses carina interrupted at middle (carina with spine-like setae) and 1 medial, transverse carina; apex truncate with about 21 spinules (
|
||
<figureCitation id="B2D22A3439680427FCBBFC71FCF5FC6C" box="[772,871,994,1018]" captionStart="Figure 10" captionStartId="28.[192,275,1624,1646]" captionTargetBox="[220,1411,210,1601]" captionTargetId="figure-65@28.[192,1440,192,1613]" captionTargetPageId="28" captionText="Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172614" httpUri="https://zenodo.org/record/5172614/files/figure.png" pageId="11" pageNumber="10">Fig. 10l</figureCitation>
|
||
). Venter: Prosternal process long, thick, apex longitudinally oval. Mesosternal surface glabrous, slightly concave at middle. Metasternal surface with sparse, ocellate punctures. Male Genitalia: Parameres (
|
||
<figureCitation id="B2D22A3439680427FCCDFBB1FC41FBAC" box="[882,979,1058,1082]" captionStart="Figure 11" captionStartId="29.[192,276,1835,1857]" captionTargetBox="[244,1411,241,1815]" captionTargetId="figure-63@29.[192,1442,192,1821]" captionTargetPageId="29" captionText="Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172618" httpUri="https://zenodo.org/record/5172618/files/figure.png" pageId="11" pageNumber="10">Fig. 11f</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439680427FC5FFBB1FB9BFBAC" box="[992,1033,1058,1082]" captionStart="Figure 12" captionStartId="30.[192,274,1776,1798]" captionTargetBox="[210,1434,220,1735]" captionTargetId="figure-61@30.[192,1440,192,1769]" captionTargetPageId="30" captionText="Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172620" httpUri="https://zenodo.org/record/5172620/files/figure.png" pageId="11" pageNumber="10">12f</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439680427FBA8FBB1FBACFBAC" box="[1047,1086,1058,1082]" captionStart="Figure 13" captionStartId="31.[192,275,1823,1845]" captionTargetBox="[238,1403,236,1798]" captionTargetId="figure-63@31.[192,1440,192,1807]" captionTargetPageId="31" captionText="Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172622" httpUri="https://zenodo.org/record/5172622/files/figure.png" pageId="11" pageNumber="10">13f</figureCitation>
|
||
) subtriangular, acute apices curving outwards. Internal sac with 3 copulatory lamellae (
|
||
<figureCitation id="B2D22A3439680427FC24FBD1FC6EFBCF" box="[923,1020,1089,1114]" captionStart="Figure 15" captionStartId="33.[192,276,1817,1839]" captionTargetBox="[238,1409,229,1784]" captionTargetId="figure-88@33.[192,1440,192,1807]" captionTargetPageId="33" captionText="Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness." figureDoi="http://doi.org/10.5281/zenodo.5172630" httpUri="https://zenodo.org/record/5172630/files/figure.png" pageId="11" pageNumber="10">Fig. 15f</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2A5636B139680427FF7FFB11FA06FB2C" blockId="11.[192,1440,1154,1210]" pageId="11" pageNumber="10">
|
||
<emphasis id="189DEAA339680427FF7FFB11FEB1FB0C" bold="true" box="[192,291,1154,1178]" pageId="11" pageNumber="10">Female</emphasis>
|
||
. Length 20.0–
|
||
<quantity id="ED119B5439680427FE71FB11FDA5FB0F" box="[462,567,1154,1178]" metricMagnitude="-2" metricUnit="m" metricValue="2.16" pageId="11" pageNumber="10" unit="mm" value="21.6">21.6 mm</quantity>
|
||
; width 9.8–10.0 mm. Similar to male, but pronotal surface with punctures slightly less dense. Gonocoxite larger than gonocoxal sternite, with 2 semicircular furrows (
|
||
<figureCitation id="B2D22A3439680427FA9AFB31FA15FB2C" box="[1317,1415,1186,1210]" captionStart="Figure 14" captionStartId="32.[192,276,1832,1854]" captionTargetBox="[358,1267,208,1798]" captionTargetId="figure-70@32.[349,1298,192,1821]" captionTargetPageId="32" captionText="Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite." figureDoi="http://doi.org/10.5281/zenodo.5172626" httpUri="https://zenodo.org/record/5172626/files/figure.png" pageId="11" pageNumber="10">Fig. 14f</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |