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<document ID-CLB-Dataset="55738" ID-DOI="http://dx.doi.org/10.3897/zookeys.1098.73223" ID-GBIF-Dataset="8b26ea49-e662-41f7-a213-47f4b598c91c" ID-PMC="PMC9848896" ID-Pensoft-Pub="1313-2970-1098-1" ID-Pensoft-UUID="B22184E75B1A59C9B985C0DD9F092196" ID-PubMed="36761078" ID-ZooBank="B4F4033F296E43CCBE54B9413BC19268" ModsDocID="1313-2970-1098-1" checkinTime="1651618226429" checkinUser="pensoft" docAuthor="Rakotonirina, Jean Claude &amp; Fisher, Brian L." docDate="2022" docId="A22ED62BF1D95FA98587130050582AEB" docLanguage="en" docName="ZooKeys 1098: 1-180" docOrigin="ZooKeys 1098" docPubDate="2022-05-03" docSource="http://dx.doi.org/10.3897/zookeys.1098.73223" docTitle="Camponotus asara Rakotonirina &amp; Fisher 2022, sp. nov." docType="treatment" docUuid="5C3637B8-D025-44D1-B91D-E49FA3D3CEBC" docUuidSource="ZooBank" docVersion="5" id="B22184E75B1A59C9B985C0DD9F092196" lastPageNumber="1" masterDocId="B22184E75B1A59C9B985C0DD9F092196" masterDocTitle="Revision of the Malagasy Camponotus subgenus Myrmosaga (Hymenoptera, Formicidae) using qualitative and quantitative morphology" masterLastPageNumber="180" masterPageNumber="1" pageNumber="1" updateTime="1732660529870" updateUser="ExternalLinkService">
<mods:mods id="70DB65691F06DB10F14188FF5A8D6DED" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5F71DC06E83DCED1E99C7C5E765DDBBB">
<mods:title id="47215AFAF9CB22E12B426FF77BCB59B2">Revision of the Malagasy Camponotus subgenus Myrmosaga (Hymenoptera, Formicidae) using qualitative and quantitative morphology</mods:title>
</mods:titleInfo>
<mods:name id="E983D50B42AC0133EF8F68F4FAECB4FC" type="personal">
<mods:role id="530152462E041457A9FBA8708E66D60A">
<mods:roleTerm id="925A03C464862D2094D088A3E2251A02">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B586C30AEF2CE584D32DEF0178E0C5A6">Rakotonirina, Jean Claude</mods:namePart>
<mods:affiliation id="963BF3B2B0F7639CDC47C30878BBA2AF">Madagascar Biodiversity Center, BP 6257, Parc Botanique et Zoologique de Tsimbazaza, Antananarivo, Madagascar &amp; Departement d'Entomologie-Culture, Elevage, Sante; Faculte des Sciences, BP 906, Universite d'Antananarivo, Antananarivo, Madagascar</mods:affiliation>
<mods:nameIdentifier id="4D8B22475CC6FE20C4F141DDAAC51B1B" type="email">jcrakoto25@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name id="DB4D679AE395A9D4C5CF3709E43A700B" type="personal">
<mods:role id="AB5E8BE4D6A7C59F0DC91ADC43A8FC02">
<mods:roleTerm id="27E02189606411698997D75D076ACABE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="74F8535045FF59E2CA877CC446233300">Fisher, Brian L.</mods:namePart>
<mods:nameIdentifier id="C7D2EE57DB1BA9D29FCC7EDC7D1F88E4" type="ORCID">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
<mods:affiliation id="87C9442BF0119F9647012CDDBBC471B4">Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="8E0ADDB38A8EDBEB519FF58E2E83DC4E">text</mods:typeOfResource>
<mods:relatedItem id="65179BEFA79C5D2E7EA3B1C4EEEA2384" type="host">
<mods:titleInfo id="5EF7CFAC5D5C00A4D09A9F8849CD6A1D">
<mods:title id="B5F305A57200518819CCF0841F9ED24E">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="B138C0CAE4C643D7114BCE7E9E74CCD0">
<mods:date id="F3C02092F50E7731C50680F358595E02">2022</mods:date>
<mods:detail id="B1497D9BEABD243CE644EF2AA3D2E1A4" type="pubDate">
<mods:number id="824F5AF34293E553FF0D125147C6881E">2022-05-03</mods:number>
</mods:detail>
<mods:detail id="F548A6D5E8252189186E0B6AAF398268" type="volume">
<mods:number id="C89029B6D694FF3CBAAA792F44709408">1098</mods:number>
</mods:detail>
<mods:extent id="EDD12EEBD04A69B179AD5A271E828CAA" unit="page">
<mods:start id="14D3E63CDD3DE39671673F95DF68FD56">1</mods:start>
<mods:end id="80EDBA37E68E96472EDDD6007C09574D">180</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="59A2DB1A15874C3C8413D071AADE84F5">
<mods:url id="7C0E8A989343CB06AE2213B7E121F782">http://dx.doi.org/10.3897/zookeys.1098.73223</mods:url>
</mods:location>
<mods:classification id="102E48E2E58E4FB2061B974419AF162F">journal article</mods:classification>
<mods:identifier id="CA2DDFA36CC0C6802E89D1691BFDF629" type="DOI">http://dx.doi.org/10.3897/zookeys.1098.73223</mods:identifier>
<mods:identifier id="CD4A6BEF75A1B54550B4AD1C9CE544A0" type="Pensoft-Pub">1313-2970-1098-1</mods:identifier>
<mods:identifier id="9AEDF449E9A5B5E985FCF77D5E0A5689" type="ZooBank">B4F4033F296E43CCBE54B9413BC19268</mods:identifier>
<mods:identifier id="5606CC34E6FECCCCFAE0EBC7642E6405" type="Pensoft-UUID">B22184E75B1A59C9B985C0DD9F092196</mods:identifier>
</mods:mods>
<treatment id="A22ED62BF1D95FA98587130050582AEB" ID-GBIF-Taxon="195190286" LSID="urn:lsid:zoobank.org:act:5C3637B8-D025-44D1-B91D-E49FA3D3CEBC" httpUri="http://treatment.plazi.org/id/A22ED62BF1D95FA98587130050582AEB" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="D422569813E60366A833A4042D80C5A2" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="BED8E23F88E53ECE24D3E74A753F549E" pageId="0" pageNumber="1">
<taxonomicName id="A22AD0DF80E0573C0275D36652206956" LSID="http://zoobank.org/5C3637B8-D025-44D1-B91D-E49FA3D3CEBC" authority="Rakotonirina &amp; Fisher, 2022" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus asara" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="asara" status="sp. nov.">Camponotus asara</taxonomicName>
<taxonomicNameLabel id="E48A979C5A933038CE4EEED4CB238306" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="A70D46B8271E215156DF6F6E901B4B34" pageId="0" pageNumber="1" type="description">
<paragraph id="55442E8DC4C2E41A2F6293D1D8558A90" pageId="0" pageNumber="1">
<figureCitation id="4484DF2304674DA089E9E26B541E47D7" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Head in full-face view A C. becki (CASENT 0071380) B C. asara (CASENT 0493252)." figureDoi="10.3897/zookeys.1098.73223.figure27" httpUri="https://binary.pensoft.net/fig/678086" pageId="0" pageNumber="1">Figs 27B</figureCitation>
<figureCitation id="E8F91BF60EABF5BE18C55A90564057B9" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Mesosoma and petiolar node in lateral view A C. asara (CASENT 0493252) B C. bozaka (CASENT 0217309)." figureDoi="10.3897/zookeys.1098.73223.figure28" httpUri="https://binary.pensoft.net/fig/678087" pageId="0" pageNumber="1">, 28A</figureCitation>
<figureCitation id="B88B00D4827212CCAF4DACC77609FD1B" captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Camponotus asara A lateral view B head in full-face view C dorsal view of holotype minor worker CASENT 0493252 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure42" httpUri="https://binary.pensoft.net/fig/678101" pageId="0" pageNumber="1">, 42</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="20A3DA644E77D260A372C3AD976FFB6C" pageId="0" pageNumber="1" type="holotype worker">
<paragraph id="9A9666E0E30289C1938108D5B10886D0" pageId="0" pageNumber="1">Holotype worker.</paragraph>
<paragraph id="5883A9ED0707E3FDFD7027F7475E3F0A" pageId="0" pageNumber="1">
<emphasis id="F254633E7A973C8C0ACEE8C1CB8301C0" bold="true" pageId="0" pageNumber="1">Madagascar</emphasis>
: Province
<emphasis id="2E2FDB94D55FCE8BE6261BFC2E657A21" bold="true" pageId="0" pageNumber="1">Toliara</emphasis>
: PN Zombitse, 19.8 km 84° E Sakaraha, -22.84333, 44.71, 770 m, tropical dry forest, ex rotten log, 05-09 Feb 2003 (Fisher, Griswold et al.) collection code: BLF07523, specimen code: CASENT0493252 (CAS).
</paragraph>
</subSubSection>
<subSubSection id="171C10DE989DCD7712E359D688ADF5CF" pageId="0" pageNumber="1" type="paratypes">
<paragraph id="A948B8DBC5E6492A469A7875E4D3D3CC" pageId="0" pageNumber="1">Paratypes.</paragraph>
<paragraph id="997647B71BD0B6130D42F81E18DBEFDB" pageId="0" pageNumber="1">3 workers, same data as holotype but with the following specimen codes: CASENT0493251, CASENT0837576, CASENT0837577 (NHMUK, CAS).</paragraph>
</subSubSection>
<subSubSection id="699C62AB15A9B6C1548A33B767A33314" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="A609F3E218FEAD22AFE282A07327DF39" pageId="0" pageNumber="1">Additional material examined.</paragraph>
<paragraph id="70F1D10AC7D6771151227979E958A52F" pageId="0" pageNumber="1">
<materialsCitation id="484DBEF586645AAD65FFBC7A08C9816A" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="P. S. Ward, PN Zombitse" country="Madagascar" elevation="760" location="15 km E Sakaraha" specimenCount="1" stateProvince="Toliara">
<emphasis id="803B8EEB4A59A0917E6FEDDA85ED1AEA" bold="true" pageId="0" pageNumber="1">
<collectingCountry id="140D31D389FA824401BB4B6765181DD4" name="Madagascar">Madagascar</collectingCountry>
:
<collectingRegion id="48F23F05F9594F52542F081760F43C95" country="Madagascar" name="Toliara">Toliara</collectingRegion>
</emphasis>
:
<location id="1319FF05F556FAF35E07B1C7B92DFE8B" LSID="urn:lsid:plazi:treatment:A22ED62BF1D95FA98587130050582AEB:1319FF05F556FAF35E07B1C7B92DFE8B" country="Madagascar" name="15 km E Sakaraha" stateProvince="Toliara">
<quantity id="5F2F2726390192E37DB67FECC6D48072" metricMagnitude="4" metricUnit="m" metricValue="1.5" unit="km" value="15.0">15 km</quantity>
E Sakaraha
</location>
, -22.9, 44.68333,
<quantity id="599E62085E39C4ED9C73520E517BD257" metricMagnitude="2" metricUnit="m" metricValue="7.6" unit="m" value="760.0">
<elevation id="3F51387537D9F7582D62665042691419" metricMagnitude="2" metricUnit="m" metricValue="7.6" unit="m" value="760.0">760 m</elevation>
</quantity>
, tropical dry forest (
<collectorName id="4C02B0235E47629E4B35D50F26E3B8B4">P.S. Ward</collectorName>
) (CAS);
<collectorName id="1ABC6089A18135E0377E6C40F9A7F37A">PN Zombitse</collectorName>
</materialsCitation>
,
<materialsCitation id="7B793591C1D77B3C7F4FE248F41AF1C9" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Fisher, Griswold, RS Ambohijanahary, Foret d'Ankazotsihitafototra" country="Madagascar" elevation="770" location="Sakaraha" longitude="84.0" specimenCount="1" stateProvince="Toliara">
<quantity id="53902D5B1E1AA0537AD085BDD462BCE8" metricMagnitude="4" metricUnit="m" metricValue="1.98" unit="km" value="19.8">19.8 km</quantity>
<geoCoordinate id="6AC3C4525E58297B4459330A0B860D90" degrees="84" direction="east" orientation="longitude" precision="55555" value="84.0">84° E</geoCoordinate>
<location id="FDA409AB7FF6006B315C283710621AAD" LSID="urn:lsid:plazi:treatment:A22ED62BF1D95FA98587130050582AEB:FDA409AB7FF6006B315C283710621AAD" country="Madagascar" longitude="84.0" name="Sakaraha" stateProvince="Toliara">Sakaraha</location>
, -22.84333, 44.71,
<quantity id="80FAE1B8F426A0ACD22B8E1032F9D84E" metricMagnitude="2" metricUnit="m" metricValue="7.7" unit="m" value="770.0">
<elevation id="99ADCFCAB6EA4EE1CD37A8F551DA0D98" metricMagnitude="2" metricUnit="m" metricValue="7.7" unit="m" value="770.0">770 m</elevation>
</quantity>
, tropical dry forest (
<collectorName id="763A1404F0B6F98767D03EB6DE44B9DA">Fisher</collectorName>
,
<collectorName id="C37976B2CC469CE2A123F456F1213532">Griswold</collectorName>
et al.) (CAS);
<collectorName id="29AE8AC4E5CC4690C25E2C8161D2D1B4">RS Ambohijanahary</collectorName>
,
<collectorName id="8C4F22EB840D79090C153180BA20AD13">
<normalizedToken id="934B7C288FD11E64A9C52B67106492E7" originalValue="Forêt">Foret</normalizedToken>
<normalizedToken id="42FD7A9FE8FE3F58DEFEDA2CABB380C1" originalValue="d’Ankazotsihitafototra">d'Ankazotsihitafototra</normalizedToken>
</collectorName>
</materialsCitation>
,
<materialsCitation id="E9C1951CFE14770B56561CC19EA525F6" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Fisher, Griswold, RS Ambohijanahary, Foret d'Ankazotsihitafototra" country="Madagascar" elevation="1100" location="34.6 km 314 ° NW Ambaravaranala" specimenCount="1" stateProvince="Toliara">
<location id="C4BDF52A27188FCA6BE143D30EDD4EAB" LSID="urn:lsid:plazi:treatment:A22ED62BF1D95FA98587130050582AEB:C4BDF52A27188FCA6BE143D30EDD4EAB" country="Madagascar" name="34.6 km 314 ° NW Ambaravaranala" stateProvince="Toliara">
<quantity id="5619D175FAFBD91C2DCB438D7842A3B4" metricMagnitude="4" metricUnit="m" metricValue="3.46" unit="km" value="34.6">34.6 km</quantity>
314° NW Ambaravaranala
</location>
, -18.26, 45.41833,
<quantity id="03308A23CD30AE3FAECF995C473C9014" metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">
<elevation id="F5D2B8E4DEE70B98AB28660BC9318503" metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">1100 m</elevation>
</quantity>
, montane rainforest (
<collectorName id="BC8FF6818714BD714C575FB365CE90EF">Fisher</collectorName>
,
<collectorName id="599BC19362AF3D3EC268897286541B91">Griswold</collectorName>
et al.) (CAS);
<collectorName id="A450FC4D0DFC56B28E99B5EB1309159E">RS Ambohijanahary</collectorName>
,
<collectorName id="6709B1AE6D479C48E21911349BADA864">
<normalizedToken id="519DBF4052C2EF1656F36541ACE0C4A5" originalValue="Forêt">Foret</normalizedToken>
<normalizedToken id="5B99C4565DF0B5706F3CC139426F0122" originalValue="d’Ankazotsihitafototra">d'Ankazotsihitafototra</normalizedToken>
</collectorName>
</materialsCitation>
,
<materialsCitation id="9758C8A3F3B17FCBC23BE6B7225CF7B7" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Fisher, Sylvain" country="Madagascar" county="Griswold" elevation="1050" latitude="-22.68333" location="Vohibasia Forest" longLatPrecision="1" longitude="44.83333" municipality="Southern Isoky-Vohimena Forest" specimenCount="1" stateProvince="Toliara">
<locationDeviation id="00A82B0211710971AB800D2D4F95E20C" location="Ambaravaranala">
<quantity id="68E9D3ACE49C1A72D61D9B3E138C9687" metricMagnitude="4" metricUnit="m" metricValue="3.52" unit="km" value="35.2">35.2 km</quantity>
312° NW Ambaravaranala
</locationDeviation>
,
<geoCoordinate id="67450F43ECF1E52E031CE5A79D84C27E" degrees="18.26667" direction="south" orientation="latitude" precision="1" value="-18.26667">-18.26667</geoCoordinate>
,
<geoCoordinate id="BCC723E7416AA42F504955BD9782E6F7" degrees="45.40667" direction="east" orientation="longitude" precision="1" value="45.40667">45.40667</geoCoordinate>
,
<quantity id="0048656DAC94D73944C681E2D59B5EE0" metricMagnitude="3" metricUnit="m" metricValue="1.05" unit="m" value="1050.0">
<elevation id="FB01B0C4CF72A3ACC64242610DB46DE5" metricMagnitude="3" metricUnit="m" metricValue="1.05" unit="m" value="1050.0">1050 m</elevation>
</quantity>
, montane rainforest (
<collectorName id="417B135E77912B5AE1E90C0A53E665D7">Fisher</collectorName>
,
<collectingCounty id="AA0E05D21DDC4CCA93ECEC25FA018306">Griswold</collectingCounty>
et al.) (CAS);
<collectingMunicipality id="974C88999D9F82A02AB02A8E90BA4B8D">Southern Isoky-Vohimena Forest</collectingMunicipality>
,
<geoCoordinate id="74655C0F4DFE5AAC3370AD262BCCAAE9" degrees="22.68333" direction="south" orientation="latitude" precision="1" value="-22.68333">-22.68333</geoCoordinate>
,
<geoCoordinate id="CA2BD0A2D0B34E6D5874C07C5D3AFF96" degrees="44.83333" direction="east" orientation="longitude" precision="1" value="44.83333">44.83333</geoCoordinate>
,
<quantity id="D66C260EC2E19CBC1CE30B5B6890F5C7" metricMagnitude="2" metricUnit="m" metricValue="7.3" unit="m" value="730.0">
<elevation id="0D6FF1C8AFD8A8367B242B75FC07CC53" metricMagnitude="2" metricUnit="m" metricValue="7.3" unit="m" value="730.0">730 m</elevation>
</quantity>
(
<collectorName id="1B96ABF8D5A3818797781C319469A8DB">Sylvain</collectorName>
) (CAS);
<location id="A0B8B89AE8D2EC80056F8AF797284B17" LSID="urn:lsid:plazi:treatment:A22ED62BF1D95FA98587130050582AEB:A0B8B89AE8D2EC80056F8AF797284B17" country="Madagascar" county="Griswold" latitude="-22.68333" longLatPrecision="1" longitude="44.83333" municipality="Southern Isoky-Vohimena Forest" name="Vohibasia Forest" stateProvince="Toliara">Vohibasia Forest</location>
</materialsCitation>
,
<materialsCitation id="31F5890734FE94B34565308A32B015C6" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Sylvain, PN Zombitse" country="Madagascar" elevation="780" location="59 km NE Sakaraha" specimenCount="1" stateProvince="Toliara">
<location id="14D4AA570988DEB4D0D4BA184A03C42F" LSID="urn:lsid:plazi:treatment:A22ED62BF1D95FA98587130050582AEB:14D4AA570988DEB4D0D4BA184A03C42F" country="Madagascar" name="59 km NE Sakaraha" stateProvince="Toliara">
<quantity id="C99CEF03D28906B14DDE6DD363D013A4" metricMagnitude="4" metricUnit="m" metricValue="5.9" unit="km" value="59.0">59 km</quantity>
NE Sakaraha
</location>
, -22.46667, 44.85,
<quantity id="AA4E38E90A9F941DA09A2AD00CEE0011" metricMagnitude="2" metricUnit="m" metricValue="7.8" unit="m" value="780.0">
<elevation id="11C1AE5AE94EC6D401063209C8B39D76" metricMagnitude="2" metricUnit="m" metricValue="7.8" unit="m" value="780.0">780 m</elevation>
</quantity>
(
<collectorName id="D5A0705BD3651D33AB322C55188A4DDF">Sylvain</collectorName>
) (CAS);
<collectorName id="E70E173E362BB0572AEC8C37363A8DBD">PN Zombitse</collectorName>
</materialsCitation>
,
<materialsCitation id="59581604C6CC43D458B92880AE82691F" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="PN Zombitse, Harin'Hala" country="Madagascar" elevation="770" latitude="-22.8405" location="Griswold" longLatPrecision="5" longitude="44.73117" municipality="Fisher" specimenCount="1" stateProvince="Toliara">
<locationDeviation id="9C75DAA4410F128BC387A8F9C9E5EE9C" location="Sakaraha">
<quantity id="C031CA3072BCA5C2C90FC192F2312F9E" metricMagnitude="4" metricUnit="m" metricValue="1.98" unit="km" value="19.8">19.8 km</quantity>
<geoCoordinate id="43C487015FEF70C7D9E653E1A3E74ED4" degrees="84" direction="east" orientation="longitude" precision="55555" value="84.0">84° E</geoCoordinate>
Sakaraha
</locationDeviation>
, -22.84333, 44.71,
<quantity id="78707DC345D5552E1C29BF64D926D41F" metricMagnitude="2" metricUnit="m" metricValue="7.7" unit="m" value="770.0">
<elevation id="C592384F3CDF567958A1760EA1B081EC" metricMagnitude="2" metricUnit="m" metricValue="7.7" unit="m" value="770.0">770 m</elevation>
</quantity>
, tropical dry forest (
<collectingMunicipality id="A9E0539343681CAF72FCFA770C515887">Fisher</collectingMunicipality>
,
<location id="82DA9A379E9FD2DEDF0AB91C982A040F" LSID="urn:lsid:plazi:treatment:A22ED62BF1D95FA98587130050582AEB:82DA9A379E9FD2DEDF0AB91C982A040F" country="Madagascar" latitude="-22.8405" longLatPrecision="5" longitude="44.73117" municipality="Fisher" name="Griswold" stateProvince="Toliara">Griswold</location>
et al.) (CAS); near ANGAP office,
<collectorName id="5C50BB81D38D4DB21849873EF11A022D">PN Zombitse</collectorName>
,
<geoCoordinate id="5D654047A1024767ADCBD5CA9FA7D761" degrees="22.8865" direction="south" orientation="latitude" precision="5" value="-22.8865">-22.8865</geoCoordinate>
,
<geoCoordinate id="B629763E58AFE30DE1B8522601631ADB" degrees="44.69217" direction="east" orientation="longitude" precision="1" value="44.69217">44.69217</geoCoordinate>
,
<quantity id="AF9B7BAA0B887CB4CA13F4F5EFD6B9D3" metricMagnitude="2" metricUnit="m" metricValue="8.4" unit="m" value="840.0">
<elevation id="3F059867587A0D28249E6C7A2313A1CA" metricMagnitude="2" metricUnit="m" metricValue="8.4" unit="m" value="840.0">840 m</elevation>
</quantity>
, deciduous spiny forest (R.
<normalizedToken id="ABF77CE1148249D41F98D07A99055271" originalValue="Harin’Hala">
<collectorName id="A6CDE52E1583727A7EB9C3C6B1E49C41">Harin'Hala</collectorName>
</normalizedToken>
) (CAS); near road,
<collectorName id="8D17593D04BA508AA65171F0B97F0C0E">PN Zombitse</collectorName>
,
<geoCoordinate id="BB6B5325F7A6754A2E48C3C9056E5AE4" degrees="22.8405" direction="south" orientation="latitude" precision="5" value="-22.8405">-22.8405</geoCoordinate>
,
<geoCoordinate id="645364D46D2BB497706496F333639389" degrees="44.73117" direction="east" orientation="longitude" precision="1" value="44.73117">44.73117</geoCoordinate>
,
<quantity id="1F03C9C85F1F4ED3A63FA7FA9A9DF612" metricMagnitude="2" metricUnit="m" metricValue="8.25" unit="m" value="825.0">
<elevation id="AAA6959AB5FEB2B17C5D87EB0C599C77" metricMagnitude="2" metricUnit="m" metricValue="8.25" unit="m" value="825.0">825 m</elevation>
</quantity>
, spiny deciduous forest (R.
<normalizedToken id="D356A01819142773DB0EFC29F456EE76" originalValue="Harin’Hala">
<collectorName id="4802D37CD6A37F3927F43942F35D52DF">Harin'Hala</collectorName>
</normalizedToken>
) (CAS)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9B3CF31E055B29DDF6BE42B9C1600D21" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="92710DC8C0ADDE3FC22A1CF09E0C90CD" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="A1C86470DC6534C9E7147406275CC6E6" pageId="0" pageNumber="1">With head in full-face view, lateral margins of head anterior to eye level, parallel, and lacking erect hairs; lateral cephalic margin rounding to posterior margin; anteromedian clypeal margin continuously forming broad convexity; propodeal dorsum more or less straight and separated from declivity surface by broad angle.</paragraph>
</subSubSection>
<subSubSection id="628BA8C00F82600B1C277F960141C45F" pageId="0" pageNumber="1" type="description">
<paragraph id="B8B6AAE3F80B408BFF095F9AE489B71A" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="071DA58D645E66B389D848B477A09E46" pageId="0" pageNumber="1">
<emphasis id="7369941C7A957C81DDA002872CEF565A" bold="true" pageId="0" pageNumber="1">Minor worker.</emphasis>
In full-face view, head sides anterior to level of eye parallel, converging progressively to posterior margin behind eye level; eyes protruding and large (EL/CL: 0.26
<normalizedToken id="24CC83D410FB2282ED0E20FA734AF6B1" originalValue="±">+/-</normalizedToken>
0.01; 0.23-0.28), breaking lateral cephalic margin, level of its posterior border approximately located at posterior 1/4 of head (PoOc/CS: 0.26
<normalizedToken id="7124CDE16704F4D578A064EE23F8547F" originalValue="±">+/-</normalizedToken>
0.01; 0.24-0.28); frontal carinae posteriorly parallel (FR/CS: 0.26
<normalizedToken id="2E3C90646C0E7E5029065B583774624E" originalValue="±">+/-</normalizedToken>
0.01; 0.23-0.27); clypeus without well-defined anterolateral angle, its anteromedian margin broadly convex; mandible with six teeth, the two apical teeth distantly spaced; antennal scape relatively long (SL/CS: 1.57
<normalizedToken id="7574F9C47A27C3F7746974BB788416AC" originalValue="±">+/-</normalizedToken>
0.08; 1.38-1.68). Promesonotum weakly convex, mesopropodeum almost flat; mesonotum flat immediately anterior to weakly visible metanotal groove; propodeal dorsum approximately straight, rounding progressively towards declivity; propodeal declivity ca. 1/3 length of dorsum. Petiolar node flattened anteroposteriorly or short and high, tapering dorsally; femur of hind leg flattened laterally and twisted near base.
</paragraph>
<paragraph id="E06610CDC9F03343B5CFE334A98E570D" pageId="0" pageNumber="1">First and second gastral tergites with a pair of white spots; erect hairs lacking on lateral margin of head; posterior margin of head with a pair of erect hairs; antennal scape only covered with appressed hairs; pronotum with a pair of erect hairs; posterodorsal angle of propodeum without erect hairs.</paragraph>
<paragraph id="DA54453A09A757DA820D035022292BF8" pageId="0" pageNumber="1">
<emphasis id="AE47CB85A791BE50B504BDDB2F4C62B9" bold="true" pageId="0" pageNumber="1">Major worker.</emphasis>
With characteristics of minor worker, except for the following characters: enlarged head (CS: 3.22
<normalizedToken id="E9689CC905E28EA00A66184BA3BA6048" originalValue="±">+/-</normalizedToken>
0.15; 3.03-3.39; CWb/CL: 0.93
<normalizedToken id="A07979F278F6E880A999ED34E28D644F" originalValue="±">+/-</normalizedToken>
0.01; 0.92-0.93) with noticeable medial excision on posterior margin; straight anterior clypeal margin; apical 1/4 of antennal scape surpassing posterior cephalic margin; propodeum dorsum and declivity the same length.
</paragraph>
</subSubSection>
<subSubSection id="4ABDFAF2508A4BB2500AB011D57963C2" pageId="0" pageNumber="1" type="distribution">
<paragraph id="54EFFEDA4B83838714AB552CE0AF6D2C" pageId="0" pageNumber="1">Distribution and biology.</paragraph>
<paragraph id="13964328D7A0A4500E800A5D3E58B139" pageId="0" pageNumber="1">
<taxonomicName id="3C6E1914E60F5A8E1D7DD9EF2A74A122" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus asara" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="asara">
<emphasis id="6D1C35D5BCDE8CE158E9BF94B3BF6C65" italics="true" pageId="0" pageNumber="1">Camponotus asara</emphasis>
</taxonomicName>
is endemic to Madagascar and geographically restricted to the dry forest of the PN Zombitse in the southern high plateau of the island and the relict montane rainforest of the PN Ambohijanahary (Fig.
<figureCitation id="768170DE89FB23F6A85C49B2B6ABE290" captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Camponotus asara A lateral view B head in full-face view C dorsal view of holotype minor worker CASENT 0493252 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure42" httpUri="https://binary.pensoft.net/fig/678101" pageId="0" pageNumber="1">42D</figureCitation>
). Colony nests can be in rotten logs and in rot pockets above the ground. Workers have been found foraging in leaf litter and on the ground.
</paragraph>
<caption id="188AE458417100CF325A58EDB2CF62EB" doi="10.3897/zookeys.1098.73223.figure42" httpUri="https://binary.pensoft.net/fig/678101" pageId="0" pageNumber="1" start="Figure 42" startId="F42">
<paragraph id="DF79DFF4E8C773749A3F7DEACC420FF4" pageId="0" pageNumber="1">
<emphasis id="93443E5C8701C11E3652FB3E84A42278" bold="true" pageId="0" pageNumber="1">Figure 42.</emphasis>
<taxonomicName id="1EA67F43DB37F62FBECAA45D47CE3689" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus asara" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="asara">
<emphasis id="8F5107ECE78623E301ACD27ADF2A38A5" italics="true" pageId="0" pageNumber="1">Camponotus asara</emphasis>
</taxonomicName>
<emphasis id="F798EEEC8DCEC9C0F1765D7C792C1A59" bold="true" pageId="0" pageNumber="1">A</emphasis>
lateral view
<emphasis id="4F0192E610CF2437EBE83B48893CAE6B" bold="true" pageId="0" pageNumber="1">B</emphasis>
head in full-face view
<emphasis id="C7E8C8B974985C81282CCD8836BA1D63" bold="true" pageId="0" pageNumber="1">C</emphasis>
dorsal view of holotype minor worker CASENT0493252
<emphasis id="096D7BE6067412103EA25E4BEECF0083" bold="true" pageId="0" pageNumber="1">D</emphasis>
distribution map.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="78C9AD1B2E9C6670AA815F5E21E210CB" pageId="0" pageNumber="1" type="discussion">
<paragraph id="0B5A31E976F7B5F7C3CDE25DEB918BBE" pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph id="B16A064ABBCDADA0FBC2828B91E6865F" pageId="0" pageNumber="1">
<taxonomicName id="18FDC8B63C027E9F729CF2B3F12160A6" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus asara" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="asara">
<emphasis id="737B400D57D256A6171661DD37AB8CF9" italics="true" pageId="0" pageNumber="1">Camponotus asara</emphasis>
</taxonomicName>
is morphologically similar to
<taxonomicName id="F328F3B4DC1E13C78470DDC89FD4A145" lsidName="C. bozaka" pageId="0" pageNumber="1" rank="species" species="bozaka">
<emphasis id="E5EBDBF0FB4C0D3F3B57405A52FF45A8" italics="true" pageId="0" pageNumber="1">C. bozaka</emphasis>
</taxonomicName>
and
<taxonomicName id="B456796E89E8962EE662E97C2B628CD3" lsidName="C. becki" pageId="0" pageNumber="1" rank="species" species="becki">
<emphasis id="DDF06B6B6F0AC6B007C34B1A4AD94894" italics="true" pageId="0" pageNumber="1">C. becki</emphasis>
</taxonomicName>
in that its petiolar node is more or less compressed anteroposteriorly in lateral view and its body color is black to dark brown, or the posterior portion of the mesosoma is pale brown to yellow. However,
<taxonomicName id="3DE56D59E2FEFFEDB669090F7619896E" lsidName="C. bozaka" pageId="0" pageNumber="1" rank="species" species="bozaka">
<emphasis id="46083DAB712EEC054F57314BBB5EFD17" italics="true" pageId="0" pageNumber="1">C. bozaka</emphasis>
</taxonomicName>
has a scalelike petiolar node, short and high mesosoma, and propodeal dorsum that joins the declivity in a blunt angle with a length &lt;2
<normalizedToken id="A6D75F1248709F1C2080C9E076655AC2" originalValue="×">x</normalizedToken>
height of the declivity, the junction being angulate. Regarding
<taxonomicName id="ECBD1AB2226898EC6AE8D2F264966B47" lsidName="C. becki" pageId="0" pageNumber="1" rank="species" species="becki">
<emphasis id="37F408A5535E4C470353B87F94F07BBB" italics="true" pageId="0" pageNumber="1">C. becki</emphasis>
</taxonomicName>
, its anteromedian clypeal margin is truncate and a few erect hairs are present on the lateral margin of the head behind the level of the posterior ocular margin.
</paragraph>
<paragraph id="43B8BD88DB4CB338AFCA26433B588F23" pageId="0" pageNumber="1">
The cluster of the samples for
<taxonomicName id="B4A7C5AA690EDB6D1DD956CD29D130F0" lsidName="C. asara" pageId="0" pageNumber="1" rank="species" species="asara">
<emphasis id="058E3535C868E3EB8888CA9CF833EFC7" italics="true" pageId="0" pageNumber="1">C. asara</emphasis>
</taxonomicName>
in the NC-clustering dendrogram is supported by LDA with a classification success of 100%.
</paragraph>
</subSubSection>
<subSubSection id="C6044ACBEF41347F4D13FB5BE66E71B1" pageId="0" pageNumber="1" type="etymology">
<paragraph id="A66D722E7F49880CD2888B12F85EAFB7" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="E6FFD8356C84DB74C900D03FCBC5E910" pageId="0" pageNumber="1">
The species name
<emphasis id="84DD338055A97B6DE7589BF204DD4D80" italics="true" pageId="0" pageNumber="1">asara</emphasis>
is a non-Latin singular noun used in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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