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<document id="170042D40231521550C8015D2EB1F2FC" ID-CLB-Dataset="299708" ID-DOI="10.5281/zenodo.11512680" ID-GBIF-Dataset="ccd739a4-10c3-4700-b27e-459bd82f1676" ID-ISSN="0068-547X" ID-Zenodo-Dep="11512680" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1717720566437" checkinUser="felipe" docAuthor="Weinell, Jeffrey L., Hooper, Errol, Leviton, Alan E. &amp; Brown, Rafe M." docDate="2019" docId="A20AE119FFB6FFA6FD980BCE4C9A5AB2" docLanguage="en" docName="ProcCalifAcadSci.66.1.1-49.pdf" docOrigin="Proceedings of the California Academy of Sciences 66 (1)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Colubridae Oppel 1811" docType="key" docVersion="5" lastPageNumber="12" masterDocId="5E339961FFBDFFADFFD00A21480B5C14" masterDocTitle="Illustrated Key to the Snakes of the Philippines" masterLastPageNumber="49" masterPageNumber="1" pageNumber="12" updateTime="1720732226891" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C8F03C7141E854EE2D1266C413ADF4EE">Illustrated Key to the Snakes of the Philippines</mods:title>
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<mods:namePart id="E53EC457939B02B26AB2D6A0C2DFB253">Weinell, Jeffrey L.</mods:namePart>
<mods:affiliation id="7D36811F956319B905FF402576ED5096">Department of Ecology and Evolutionary Biology and Biodiversity Institute, University of Kansas, Lawrence, Kansas 66045;</mods:affiliation>
<mods:nameIdentifier id="F017CEDBAFE20929D9DCE1497751AE6F" type="email">jweine2@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F4C4AF0081CD680276663162F79F7057">Hooper, Errol</mods:namePart>
<mods:affiliation id="1A405E00F5918A83C8B008D10955AB8A">Department of Ecology and Evolutionary Biology and Biodiversity Institute, University of Kansas, Lawrence, Kansas 66045;</mods:affiliation>
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<mods:namePart id="83845869018725530BCB15CF9622C602">Leviton, Alan E.</mods:namePart>
<mods:affiliation id="FBCEA4AAD911417668CEF4E1FDA55234">Herpetology Division, Institute of Biodiversity Science &amp; Sustainability, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, California 94118. * Corresponding author; Email: jweine 2 @ gmail. com</mods:affiliation>
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<mods:namePart id="2EB912C23DC4BF771B34531BD426BAF7">Brown, Rafe M.</mods:namePart>
<mods:affiliation id="497C0FE7C63A4F84B49ACA77E1E1969A">Department of Ecology and Evolutionary Biology and Biodiversity Institute, University of Kansas, Lawrence, Kansas 66045;</mods:affiliation>
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<mods:date id="B7BD35F885F22B0B38D9CE777A51E247">2019</mods:date>
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<treatment id="A20AE119FFB6FFA6FD980BCE4C9A5AB2" ID-DOI="http://doi.org/10.5281/zenodo.12728056" ID-GBIF-Taxon="235040639" ID-Zenodo-Dep="12728056" LSID="urn:lsid:plazi:treatment:A20AE119FFB6FFA6FD980BCE4C9A5AB2" httpUri="http://treatment.plazi.org/id/A20AE119FFB6FFA6FD980BCE4C9A5AB2" lastPageNumber="12" pageId="11" pageNumber="12">
<subSubSection id="62B90384FFB6FFA6FD980BCE4AC25E12" box="[584,713,495,518]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="2A1C500FFFB6FFA6FD980BCE4AC25E12" blockId="11.[485,812,495,550]" box="[584,713,495,518]" pageId="11" pageNumber="12">
<taxonomicName id="EDA32B8CFFB6FFA6FD980BCE4AC25E12" ID-CoL="8G9" authority="Oppel, 1811" authorityName="Oppel" authorityYear="1811" box="[584,713,495,518]" class="Reptilia" family="Colubridae" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="family">
<emphasis id="18D78C1DFFB6FFA6FD980BCE4AC25E12" bold="true" box="[584,713,495,518]" pageId="11" pageNumber="12">Colubridae</emphasis>
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<subSubSection id="62B90384FFB6FFA6FE35082E4B275E32" box="[485,812,527,550]" pageId="11" pageNumber="12" type="multiple">
<paragraph id="2A1C500FFFB6FFA6FE35082E4B275E32" blockId="11.[485,812,495,550]" box="[485,812,527,550]" pageId="11" pageNumber="12">Key to Philippine subfamilies.</paragraph>
</subSubSection>
<subSubSection id="62B90384FFB6FFA6FFAB081B4C9A5AB2" pageId="11" pageNumber="12" type="key">
<key id="5E32073EFFB6FFA6FFAB081B4C9A5AB2" pageId="11" pageNumber="12">
<keyStep id="915748AAFFB6FFA6FFAB081B4C9A5E85" pageId="11" pageNumber="12">
<paragraph id="2A1C500FFFB6FFA6FFAB081B4C9A5E65" blockId="11.[123,1169,570,657]" pageId="11" pageNumber="12">
<keyLead id="9152F33AFFB6FFA6FFAB081B4C9A5E65" pageId="11" pageNumber="12">
1a. Temporal scales absent; parietal scales in contact with supralabial scales (
<figureCitation id="B2984C8AFFB6FFA6FC69081B4C105E45" box="[953,1051,570,593]" captionStart="FIGURE" captionStartId="10.[157,169,993,1012]" captionTargetBox="[294,1004,793,980]" captionTargetId="figure-213@10.[294,1004,792,980]" captionTargetPageId="10" captionText="FIGURE 12. Lateral view of head of (A) Calamaria gervaisii (KU 322329), (B) Lycodon muelleri (KU 327575). Temporal scales (green), supralabial scales (purplish-gray), parietal scale (blue)." figureDoi="http://doi.org/10.5281/zenodo.12725257" httpUri="https://zenodo.org/record/12725257/files/figure.png" pageId="11" pageNumber="12">Fig. 12A</figureCitation>
)................................................................... Calamariinae (p. 18)
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFB6FFA6FFAB085B4C9A5E85" blockId="11.[123,1169,570,657]" box="[123,1169,634,657]" pageId="11" pageNumber="12">
<keyLead id="9152F33AFFB6FFA6FFAB085B4C9A5E85" box="[123,1169,634,657]" pageId="11" pageNumber="12">
1b. Temporal scales present; parietal scales not in contact with supralabial scales (
<figureCitation id="B2984C8AFFB6FFA6FC3D085B4C465E85" box="[1005,1101,634,657]" captionStart="FIGURE" captionStartId="10.[157,169,993,1012]" captionTargetBox="[294,1004,793,980]" captionTargetId="figure-213@10.[294,1004,792,980]" captionTargetPageId="10" captionText="FIGURE 12. Lateral view of head of (A) Calamaria gervaisii (KU 322329), (B) Lycodon muelleri (KU 327575). Temporal scales (green), supralabial scales (purplish-gray), parietal scale (blue)." figureDoi="http://doi.org/10.5281/zenodo.12725257" httpUri="https://zenodo.org/record/12725257/files/figure.png" pageId="11" pageNumber="12">Fig. 12B</figureCitation>
)... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFB6FFA6FFAB08854C9A5F6F" pageId="11" pageNumber="12">
<paragraph id="2A1C500FFFB6FFA6FFAB08854C9A5F0F" blockId="11.[123,1174,676,891]" pageId="11" pageNumber="12">
<keyLead id="9152F33AFFB6FFA6FFAB08854C9A5F0F" pageId="11" pageNumber="12">
2a. Supralabial scales either do not border eye, or supralabial scales 46 border eye; if supralabial scales 46 border eye, then dorsal body scales reduce from 19→17 or from 17→15 longitudinal rows posteriorly, and lateral body scales (except possibly first row) keeled (
<figureCitation id="B2984C8AFFB6FFA6FC1508C54C1C5EEF" box="[965,1047,740,763]" captionStart="FIGURE" captionStartId="7.[155,167,853,872]" captionTargetBox="[119,1176,587,845]" captionTargetId="figure-735@7.[119,1176,587,845]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal body scales: (A) spinose (Acrochordus granulatus; KU 302951), (B) keeled (Opisthotropis typica; KU 327424), (C) smooth (Oligodon maculatus; KU 321699)." figureDoi="http://doi.org/10.5281/zenodo.11512690" httpUri="https://zenodo.org/record/11512690/files/figure.png" pageId="11" pageNumber="12">Fig. 3B</figureCitation>
)...................................................................... Natricinae (p. 19)
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFB6FFA6FFAB09054C9A5F6F" blockId="11.[123,1174,676,891]" pageId="11" pageNumber="12">
<keyLead id="9152F33AFFB6FFA6FFAB09054C9A5F6F" pageId="11" pageNumber="12">
2b. One or more supralabial scales border eye; if supralabial scales 46 border eye and dorsal body scales reduce from 19→17 or 17→15 longitudinal rows posteriorly, then lateral body scales are smooth (
<figureCitation id="B2984C8AFFB6FFA6FE71094549F85F6F" box="[417,499,868,891]" captionStart="FIGURE" captionStartId="7.[155,167,853,872]" captionTargetBox="[119,1176,587,845]" captionTargetId="figure-735@7.[119,1176,587,845]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal body scales: (A) spinose (Acrochordus granulatus; KU 302951), (B) keeled (Opisthotropis typica; KU 327424), (C) smooth (Oligodon maculatus; KU 321699)." figureDoi="http://doi.org/10.5281/zenodo.11512690" httpUri="https://zenodo.org/record/11512690/files/figure.png" pageId="11" pageNumber="12">Fig. 3C</figureCitation>
)................................................ 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="915748AAFFB6FFA6FFAB09AE4C9A58B2" pageId="11" pageNumber="12">
<paragraph id="2A1C500FFFB6FFA6FFAB09AE4C9A5FF2" blockId="11.[123,1174,911,1190]" pageId="11" pageNumber="12">
<keyLead id="9152F33AFFB6FFA6FFAB09AE4C9A5FF2" pageId="11" pageNumber="12">
3a. Lateral body scales in oblique rows (
<figureCitation id="B2984C8AFFB6FFA6FD8009AE4AB35FB2" box="[592,696,911,934]" captionStart="FIGURE" captionStartId="11.[155,167,1419,1438]" captionTargetBox="[320,990,1197,1400]" captionTargetId="figure-898@11.[293,1003,1197,1408]" captionTargetPageId="11" captionText="FIGURE 15. Lateral body scales of (A) Dendrelaphis marenae (KU 314131): oblique rows, (B) Oligodon maculatus (KU 321699): horizontal rows." figureDoi="http://doi.org/10.5281/zenodo.11512708" httpUri="https://zenodo.org/record/11512708/files/figure.png" pageId="11" pageNumber="12">Fig. 15A</figureCitation>
); dorsal body scales reduce from 17→13, 15→13, 15→11, or 13→11 longitudinal scale rows posteriorly; third supralabial scale does not border eye................................................... Ahaetulliinae (p. 17)
</keyLead>
</paragraph>
<paragraph id="2A1C500FFFB6FFA6FFAB09CE4C9A58B2" blockId="11.[123,1174,911,1190]" pageId="11" pageNumber="12">
<keyLead id="9152F33AFFB6FFA6FFAB09CE4C9A58B2" pageId="11" pageNumber="12">
3b. One of the following combinations of characters true: (1) lateral body scales in horizontal, not oblique rows (
<figureCitation id="B2984C8AFFB6FFA6FEE20E2E49985832" box="[306,403,1039,1062]" captionStart="FIGURE" captionStartId="10.[156,168,1348,1367]" captionTargetBox="[313,1004,1049,1338]" captionTargetId="figure-241@10.[294,1004,1049,1338]" captionText="FIGURE 13. Lateral view of head of (A) Psammodynastes pulverulentus (KU 329688), (B) Cyclocorus lineatus alcalai (KU 324539). Nasal scale (green) undivided (A) or divided (B)." figureDoi="http://doi.org/10.5281/zenodo.12725259" httpUri="https://zenodo.org/record/12725259/files/figure.png" pageId="11" pageNumber="12">Fig. 13B</figureCitation>
), and dorsal body scales in 13 longitudinal rows at midbody; (2) lateral body scales in oblique rows (
<figureCitation id="B2984C8AFFB6FFA6FE0B0E0E4A355852" box="[475,574,1071,1094]" captionStart="FIGURE" captionStartId="11.[155,167,1419,1438]" captionTargetBox="[320,990,1197,1400]" captionTargetId="figure-898@11.[293,1003,1197,1408]" captionTargetPageId="11" captionText="FIGURE 15. Lateral body scales of (A) Dendrelaphis marenae (KU 314131): oblique rows, (B) Oligodon maculatus (KU 321699): horizontal rows." figureDoi="http://doi.org/10.5281/zenodo.11512708" httpUri="https://zenodo.org/record/11512708/files/figure.png" pageId="11" pageNumber="12">Fig. 15A</figureCitation>
) and dorsal body scales in 1517 longitudinal rows at midbody, not reducing to fewer than 15 rows before vent; (3) lateral body scales in oblique or horizontal rows (compare
<figureCitation id="B2984C8AFFB6FFA6FE670E4E4A375892" box="[439,572,1135,1158]" captionStart="FIGURE" captionStartId="11.[155,167,1419,1438]" captionTargetBox="[320,990,1197,1400]" captionTargetId="figure-898@11.[293,1003,1197,1408]" captionTargetPageId="11" captionText="FIGURE 15. Lateral body scales of (A) Dendrelaphis marenae (KU 314131): oblique rows, (B) Oligodon maculatus (KU 321699): horizontal rows." figureDoi="http://doi.org/10.5281/zenodo.11512708" httpUri="https://zenodo.org/record/11512708/files/figure.png" pageId="11" pageNumber="12">Fig. 15A, B</figureCitation>
), and dorsal body scales in 1923 longitudinal rows at midbody.................................................................... 4
</keyLead>
</paragraph>
</keyStep>
<caption id="7EDC0087FFB6FFA6FF4B0FAA498D59A3" ID-DOI="http://doi.org/10.5281/zenodo.11512708" ID-Zenodo-Dep="11512708" httpUri="https://zenodo.org/record/11512708/files/figure.png" pageId="11" pageNumber="12" startId="11.[155,167,1419,1438]" targetBox="[320,990,1197,1400]" targetPageId="11" targetType="figure">
<paragraph id="2A1C500FFFB6FFA6FF4B0FAA498D59A3" blockId="11.[123,1174,1419,1463]" pageId="11" pageNumber="12">
<smallCapsWord id="2CFAC6D3FFB6FFA6FF4B0FAA48D35988" baselines="1433,1432" box="[155,216,1419,1438]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="11" pageNumber="12">FIGURE</smallCapsWord>
15. Lateral body scales of (A)
<taxonomicName id="EDA32B8CFFB6FFA6FE230FAA4AB3598A" authorityName="Vogel &amp; Van Rooijen" authorityYear="2008" box="[499,696,1419,1438]" class="Reptilia" family="Colubridae" genus="Dendrelaphis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="marenae">
<emphasis id="18D78C1DFFB6FFA6FE230FAA4AB3598A" box="[499,696,1419,1438]" italics="true" pageId="11" pageNumber="12">Dendrelaphis marenae</emphasis>
</taxonomicName>
(KU 314131): oblique rows, (B)
<taxonomicName id="EDA32B8CFFB6FFA6FC350FAA4C9D598A" baseAuthorityName="Taylor" baseAuthorityYear="1918" box="[997,1174,1419,1438]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="18D78C1DFFB6FFA6FC350FAA4C9D598A" box="[997,1174,1419,1438]" italics="true" pageId="11" pageNumber="12">Oligodon maculatus</emphasis>
</taxonomicName>
(KU 321699): horizontal rows.
</paragraph>
</caption>
<keyStep id="915748AAFFB6FFA6FFAB0FEE4C9A5AB2" pageId="11" pageNumber="12">
<paragraph id="2A1C500FFFB6FFA6FFAB0FEE4C9A5A32" blockId="11.[123,1173,1487,1702]" pageId="11" pageNumber="12">
<keyLead id="9152F33AFFB6FFA6FFAB0FEE4C9A5A32" pageId="11" pageNumber="12">4a. Dorsal body scales in 17 smooth longitudinal rows throughout length of body; number of ventral body scales 144185; subcaudal scales arranged in 89145 pairs; numerous small teeth, 3050, on each of maxillary and dentary bones.................... Sibynophiinae (p. 19)</keyLead>
</paragraph>
<paragraph id="2A1C500FFFB6FFA6FFAB0C0E4C9A5AB2" blockId="11.[123,1173,1487,1702]" pageId="11" pageNumber="12">
<keyLead id="9152F33AFFB6FFA6FFAB0C0E4C9A5AB2" pageId="11" pageNumber="12">4b. Dorsal body scales in 1523 smooth or keeled longitudinal rows at midbody; if dorsal body scales in 17 smooth longitudinal rows throughout length of body, then subcaudal scales arranged in &lt;80 or&gt; 190 pairs; fewer than 30 teeth on each of maxillary and dentary bones..................................................................... Colubrinae (p. 18)</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
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