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<document id="D2FEFC0670D46316243D10EE6BADB98E" ID-CLB-Dataset="54548" ID-DOI="10.11646/zootaxa.5222.5.1" ID-GBIF-Dataset="a2aef7c8-09e1-48f1-983e-00132055a5a2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7471736" ID-ZooBank="F4983B58-7301-4A3F-AC08-3E5120450EA1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1671694447742" checkinUser="plazi" docAuthor="Evenhuis, Neal L. &amp; Gharali, Babak" docDate="2022" docId="A1541B07261EFFA0FF60F8D01BB9FCC6" docLanguage="en" docName="zootaxa.5222.5.1.pdf" docOrigin="Zootaxa 5222 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Empidideicus (Anomaloptilus) Hesse" docType="key" docVersion="7" lastPageNumber="404" masterDocId="5D6D637F261FFFA3FFF7FFEB1E24FFF4" masterDocTitle="The subgenus Empidideicus (Anomaloptilus) (Diptera: Mythicomyiidae) in Madagascar" masterLastPageNumber="426" masterPageNumber="401" pageNumber="402" updateTime="1699413895303" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="10F8CAE225B3D0791A8D43F8C83213F4">
<mods:title id="9F4D7BBCF6E1DFBB772C8E0AAE1A1014">The subgenus Empidideicus (Anomaloptilus) (Diptera: Mythicomyiidae) in Madagascar</mods:title>
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<mods:name id="36D06F3D6F5A438588E3784BE053E57B" type="personal">
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<mods:namePart id="2510F79CEE894A6CB75D15B2F95F87FB">Evenhuis, Neal L.</mods:namePart>
<mods:nameIdentifier id="37D075764A0330D0A9D7AC57A4289B1A" type="ORCID">0000-0002-1314-755X</mods:nameIdentifier>
<mods:affiliation id="24A6A2A47BA73A377C0C52B3A6264B08">J. Linsley Gressitt Center for Research in Entomology, Bishop Museum, 1525 Bernice Street, Honolulu, Hawaii 96817 - 2704, USA neale @ bishopmuseum. org; https: // orcid. org / 0000 - 0002 - 1314 - 755 X</mods:affiliation>
<mods:nameIdentifier id="B4CE2EA08F6FF9FDF3CCB48D60B0FAF1" type="email">neale@bishopmuseum.org</mods:nameIdentifier>
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<mods:namePart id="749E2E3273F605E93A19F3501523AB0F">Gharali, Babak</mods:namePart>
<mods:nameIdentifier id="F79E2B74DFF52AB36E7A3F334948E0CF" type="ORCID">0000-0003-2490-156X</mods:nameIdentifier>
<mods:affiliation id="42D1F1E01E08AC8F5BC7DE6E4F27A538">Plant Protection Research Department, Qazvin Agricultural and Natural Resources Research and Education Center, AREEO, Qazvin, Iran. bgaharaei @ yahoo. com; https: // orcid. org / 0000 - 0003 - 2490 - 156 X</mods:affiliation>
<mods:nameIdentifier id="AC83882A9A3B5A088614965308C648C2" type="email">bgaharaei@yahoo.com</mods:nameIdentifier>
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<mods:title id="42CCB3E0DAAEA3798CE959B57913078F">Zootaxa</mods:title>
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<mods:date id="BACEAE4B541CDFF5ED384BB06FA497D6">2022</mods:date>
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<mods:number id="3C8299E38514C7C506631FCA5B9C598E">2022-12-22</mods:number>
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<treatment id="A1541B07261EFFA0FF60F8D01BB9FCC6" ID-DOI="http://doi.org/10.5281/zenodo.7473304" ID-GBIF-Taxon="207927022" ID-Zenodo-Dep="7473304" LSID="urn:lsid:plazi:treatment:A1541B07261EFFA0FF60F8D01BB9FCC6" httpUri="http://treatment.plazi.org/id/A1541B07261EFFA0FF60F8D01BB9FCC6" lastPageId="3" lastPageNumber="404" pageId="1" pageNumber="402">
<subSubSection id="61E7F99A261EFFA2FF60F8D01DAAF8A1" box="[151,910,1851,1878]" pageId="1" pageNumber="402" type="nomenclature">
<paragraph id="2942AA11261EFFA2FF60F8D01DAAF8A1" blockId="1.[151,910,1851,1878]" box="[151,910,1851,1878]" pageId="1" pageNumber="402">
<heading id="720A1D7D261EFFA2FF60F8D01DAAF8A1" bold="true" box="[151,910,1851,1878]" fontSize="11" level="1" pageId="1" pageNumber="402" reason="1">
<emphasis id="1B897603261EFFA2FF60F8D01DAAF8A1" bold="true" box="[151,910,1851,1878]" pageId="1" pageNumber="402">
Key to the Malagasy species of
<taxonomicName id="EEFDD192261EFFA2FDEAF8D01DAAF8A1" authorityName="Hesse" box="[541,910,1851,1878]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="402" phylum="Arthropoda" rank="subGenus" subGenus="Anomaloptilus">
<emphasis id="1B897603261EFFA2FDEAF8D01CE6F8A2" bold="true" box="[541,706,1851,1878]" italics="true" pageId="1" pageNumber="402">Empidideicus</emphasis>
(
<emphasis id="1B897603261EFFA2FD25F8D01DA0F8A2" bold="true" box="[722,900,1851,1878]" italics="true" pageId="1" pageNumber="402">Anomaloptilus</emphasis>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61E7F99A261EFFA0FF60F86B1BB9FCC6" lastPageId="3" lastPageNumber="404" pageId="1" pageNumber="402" type="key">
<keyStep id="9209B2B4261EFFA2FF60F86B1BB8F824" pageId="1" pageNumber="402">
<paragraph id="2942AA11261EFFA2FF60F86B1BB8F847" blockId="1.[151,1437,1920,2030]" pageId="1" pageNumber="402">
<keyLead id="920C0924261EFFA2FF60F86B1BB8F847" pageId="1" pageNumber="402">
1. Clypeus elongate, extending well beyond length of oral margin, forcing proboscis to be reflexed downward and backward (
<figureCitation id="B1C6B694261EFFA2FA82F86B1ECBF847" captionStart="FIGURES 3–4" captionStartId="2.[151,264,1920,1944]" captionTargetBox="[157,1423,240,1880]" captionTargetId="figure-140@2.[151,1436,227,1896]" captionTargetPageId="2" captionText="FIGURES 3–4. Schematic illustration showing differences in clypeus lengths of Empidideicus (Anomaloptilus) species. 3, E. (A.) celluliferus Hesse; 4, E. (A.) rhinoclypeatus Evenhuis. Arrows point to clypeus. (Redrawn from Evenhuis 2007; figs 2–3)." figureDoi="http://doi.org/10.5281/zenodo.7471742" httpUri="https://zenodo.org/record/7471742/files/figure.png" pageId="1" pageNumber="402">Fig. 4</figureCitation>
).......................................................................
<taxonomicName id="EEFDD192261EFFA2FB95F8751BB8F847" authority="Evenhuis" authorityName="Evenhuis" box="[1122,1436,1949,1971]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="403" phylum="Arthropoda" rank="species" species="rhinoclypeatus" subGenus="Anomaloptilus">
<emphasis id="1B897603261EFFA2FB95F8751B19F847" bold="true" box="[1122,1341,1949,1971]" pageId="1" pageNumber="402">
<emphasis id="1B897603261EFFA2FB95F8751A56F847" bold="true" box="[1122,1138,1950,1971]" italics="true" pageId="1" pageNumber="402">E</emphasis>
. (
<emphasis id="1B897603261EFFA2FB71F8761AB2F846" bold="true" box="[1158,1174,1949,1970]" italics="true" pageId="1" pageNumber="402">A</emphasis>
.)
<emphasis id="1B897603261EFFA2FB5DF8761B19F847" bold="true" box="[1194,1341,1949,1971]" italics="true" pageId="1" pageNumber="402">rhinoclypeatus</emphasis>
</emphasis>
Evenhuis
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2942AA11261EFFA2FF60F8501BB8F824" blockId="1.[151,1437,1920,2030]" box="[151,1436,1979,2001]" pageId="1" pageNumber="402">
<keyLead id="920C0924261EFFA2FF60F8501BB8F824" box="[151,1436,1979,2001]" pageId="1" pageNumber="402">
–. Clypeus not elongate, never extending beyond length of oral margin (
<figureCitation id="B1C6B694261EFFA2FC8AF8501D92F824" box="[893,950,1979,2001]" captionStart="FIGURES 3–4" captionStartId="2.[151,264,1920,1944]" captionTargetBox="[157,1423,240,1880]" captionTargetId="figure-140@2.[151,1436,227,1896]" captionTargetPageId="2" captionText="FIGURES 3–4. Schematic illustration showing differences in clypeus lengths of Empidideicus (Anomaloptilus) species. 3, E. (A.) celluliferus Hesse; 4, E. (A.) rhinoclypeatus Evenhuis. Arrows point to clypeus. (Redrawn from Evenhuis 2007; figs 2–3)." figureDoi="http://doi.org/10.5281/zenodo.7471742" httpUri="https://zenodo.org/record/7471742/files/figure.png" pageId="1" pageNumber="402">Fig. 3</figureCitation>
)...................................... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9209B2B4261EFFA1FF60F8331BB8FF3E" lastPageId="2" lastPageNumber="403" pageId="1" pageNumber="402">
<paragraph id="2942AA11261EFFA2FF60F8331BB8F819" blockId="1.[151,1437,1920,2030]" box="[151,1436,2008,2030]" pageId="1" pageNumber="402">
<keyLead id="920C0924261EFFA2FF60F8331BB8F819" box="[151,1436,2008,2030]" pageId="1" pageNumber="402">
2. Proboscis extremely short, barely projecting beyond oral margin (
<figureCitation id="B1C6B694261EFFA2FCAEF8331D85F819" box="[857,929,2008,2030]" captionStart="FIGURES 20–22" captionStartId="8.[151,264,1965,1989]" captionTargetBox="[151,1363,1475,1942]" captionTargetId="figure-302@8.[151,1435,1475,1942]" captionTargetPageId="8" captionText="FIGURES 20–22. Malagasy Empidideicus (Anomaloptilus), heads. 20, E. (A.) contrasticus, n. sp., male, frontal view; 21, E. (A.) irwini, n. sp., male, oblique view; 22, E. (A.) rhinoclypeatus Evenhuis, male, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7471752" httpUri="https://zenodo.org/record/7471752/files/figure.png" pageId="1" pageNumber="402">Fig. 21</figureCitation>
); head wider than long in dorsal view (
<figureCitation id="B1C6B694261EFFA2FAEFF8331B79F819" box="[1304,1373,2008,2030]" captionStart="FIGURES 17–19" captionStartId="8.[151,264,1390,1414]" captionTargetBox="[151,1435,741,1367]" captionTargetId="figure-245@8.[151,1435,741,1367]" captionTargetPageId="8" captionText="FIGURES 17–19. Malagasy Empidideicus (Anomaloptilus), heads. 17, E. (A.) andriambahomananii, n. sp., male, oblique view; 18, E. (A.) irwini, n. sp., female, dorsal view; 19, E. (A.) rinhai, n. sp., male, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7471750" httpUri="https://zenodo.org/record/7471750/files/figure.png" pageId="1" pageNumber="402">Fig. 18</figureCitation>
)... 3
</keyLead>
</paragraph>
<paragraph id="2942AA11261DFFA1FF60FF731BB8FF3E" blockId="2.[151,1436,151,202]" pageId="2" pageNumber="403">
<keyLead id="920C0924261DFFA1FF60FF731BB8FF3E" pageId="2" pageNumber="403">
–. Proboscis clearly projecting beyond oral margin for at least 1/2 head height; head longer than wide in dorsal view (
<figureCitation id="B1C6B694261DFFA1FAC6FF7C1B53FF59" box="[1329,1399,151,173]" captionStart="FIGURES 17–19" captionStartId="8.[151,264,1390,1414]" captionTargetBox="[151,1435,741,1367]" captionTargetId="figure-245@8.[151,1435,741,1367]" captionTargetPageId="8" captionText="FIGURES 17–19. Malagasy Empidideicus (Anomaloptilus), heads. 17, E. (A.) andriambahomananii, n. sp., male, oblique view; 18, E. (A.) irwini, n. sp., female, dorsal view; 19, E. (A.) rinhai, n. sp., male, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7471750" httpUri="https://zenodo.org/record/7471750/files/figure.png" pageId="2" pageNumber="403">Fig. 17</figureCitation>
).................................................................................................... 4
</keyLead>
</paragraph>
</keyStep>
<caption id="7D82FA99261DFFA1FF60F86B1BBCF848" ID-DOI="http://doi.org/10.5281/zenodo.7471742" ID-Zenodo-Dep="7471742" httpUri="https://zenodo.org/record/7471742/files/figure.png" pageId="2" pageNumber="403" startId="2.[151,264,1920,1944]" targetBox="[157,1423,240,1880]" targetPageId="2">
<paragraph id="2942AA11261DFFA1FF60F86B1BBCF848" blockId="2.[151,1437,1920,1980]" pageId="2" pageNumber="403">
<emphasis id="1B897603261DFFA1FF60F86B1F18F86C" bold="true" box="[151,316,1920,1944]" pageId="2" pageNumber="403">FIGURES 3–4.</emphasis>
Schematic illustration showing differences in clypeus lengths of
<taxonomicName id="EEFDD192261DFFA1FC2EF86B1B2AF86C" box="[985,1294,1920,1944]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="subGenus" subGenus="Anomaloptilus">
<emphasis id="1B897603261DFFA1FC2EF86B1A47F86C" box="[985,1123,1920,1944]" italics="true" pageId="2" pageNumber="403">Empidideicus</emphasis>
(
<emphasis id="1B897603261DFFA1FB85F86B1B22F86C" box="[1138,1286,1920,1944]" italics="true" pageId="2" pageNumber="403">Anomaloptilus</emphasis>
)
</taxonomicName>
species.
<emphasis id="1B897603261DFFA1FA9BF86B1B5DF86C" bold="true" box="[1388,1401,1920,1944]" pageId="2" pageNumber="403">3</emphasis>
,
<taxonomicName id="EEFDD192261DFFA1FA71F86A1F59F84F" authority="Hesse" authorityName="Hesse" authorityYear="1938" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="celluliferus" subGenus="Anomaloptilus">
<emphasis id="1B897603261DFFA1FA71F86A1BB9F86C" box="[1414,1437,1921,1944]" italics="true" pageId="2" pageNumber="403">E.</emphasis>
(
<emphasis id="1B897603261DFFA1FF57F84F1E92F84F" box="[160,182,1956,1979]" italics="true" pageId="2" pageNumber="403">A.</emphasis>
)
<emphasis id="1B897603261DFFA1FF33F84F1F12F848" box="[196,310,1956,1980]" italics="true" pageId="2" pageNumber="403">celluliferus</emphasis>
Hesse
</taxonomicName>
;
<emphasis id="1B897603261DFFA1FE71F84F1FB7F848" bold="true" box="[390,403,1956,1980]" pageId="2" pageNumber="403">4</emphasis>
,
<taxonomicName id="EEFDD192261DFFA1FE57F84E1D1FF848" authority="Evenhuis. Arrows" authorityName="Evenhuis. Arrows" box="[416,827,1956,1980]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="rhinoclypeatus" subGenus="Anomaloptilus">
<emphasis id="1B897603261DFFA1FE57F84E1F92F848" box="[416,438,1957,1980]" italics="true" pageId="2" pageNumber="403">E.</emphasis>
(
<emphasis id="1B897603261DFFA1FE33F84F1FFFF84F" box="[452,475,1956,1979]" italics="true" pageId="2" pageNumber="403">A.</emphasis>
)
<emphasis id="1B897603261DFFA1FE1EF84F1C5BF848" box="[489,639,1956,1980]" italics="true" pageId="2" pageNumber="403">rhinoclypeatus</emphasis>
Evenhuis. Arrows
</taxonomicName>
point to clypeus. (Redrawn from
<bibRefCitation id="4D6CD7E0261DFFA1FB66F84F1B08F848" author="Evenhuis, N. L." box="[1169,1324,1956,1980]" pageId="2" pageNumber="403" pagination="55 - 62" refId="ref15043" refString="Evenhuis, N. L. (2007) A remarkable new species of Empidideicus (Diptera: Mythicomyiidae) from Madagascar. Zootaxa, 1474 (1), 55 - 62. https: // doi. org / 10.11646 / zootaxa. 1474.1.2" type="journal article" year="2007">Evenhuis 2007</bibRefCitation>
; figs 2–3).
</paragraph>
</caption>
<keyStep id="9209B2B4261CFFA0FF60FF731BB9FECB" pageId="3" pageNumber="404">
<paragraph id="2942AA11261CFFA0FF60FF731BB9FF13" blockId="3.[151,1437,151,819]" pageId="3" pageNumber="404">
<keyLead id="920C0924261CFFA0FF60FF731BB9FF13" pageId="3" pageNumber="404">
3. Wing vein R 4+5 ending in wing margin before level of end of M 2 (
<figureCitation id="B1C6B694261CFFA0FCA6FF7C1DBDFF59" box="[849,921,151,173]" captionStart="FIGURES 40–47" captionStartId="15.[151,264,1607,1631]" captionTargetBox="[170,1416,309,1582]" captionTargetId="figure-81@15.[170,1416,309,1584]" captionTargetPageId="15" captionText="FIGURES 40–47. Malagasy Empidideicus (Anomaloptilus), wings. 40, E. (A.) andriambahomananii, n. sp.; 41, E. (A.) contrasticus, n. sp.; 42, E. (A.) hauseri, n. sp.; 43, E. (A.) irwini, n. sp.; 44, E. (A.) nanus, n. sp.; 45, E. (A.) regius, n. sp.; 46, E. (A.) rhinoclypeatus Evenhuis; 47, E. (A.) rinhai, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471764" httpUri="https://zenodo.org/record/7471764/files/figure.png" pageId="3" pageNumber="404">Fig. 41</figureCitation>
); cell r4+5 at wing margin as wide as cell m2; eyes in males clearly dichoptic below level of antennae; prescutellar area in females with yellow marks............................................................... (northwest
<collectingCountry id="51EAEA81261CFFA0FCCDFF391D94FF1C" box="[826,944,210,232]" name="Madagascar" pageId="3" pageNumber="404">Madagascar</collectingCountry>
) ...
<emphasis id="1B897603261CFFA0FC23FF391DC0FF13" bold="true" box="[980,996,210,231]" italics="true" pageId="3" pageNumber="404">E</emphasis>
.
<taxonomicName id="EEFDD192261CFFA0FC07FF391B78FF1C" authority="Evenhuis &amp; Gharali" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[1008,1372,210,232]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="contrasticus" status="sp. nov." subGenus="Anomaloptilus">
<emphasis id="1B897603261CFFA0FC07FF391AB0FF13" bold="true" box="[1008,1172,210,232]" pageId="3" pageNumber="404">
(
<emphasis id="1B897603261CFFA0FC0FFF391A2CFF13" bold="true" box="[1016,1032,210,231]" italics="true" pageId="3" pageNumber="404">A</emphasis>
.)
<emphasis id="1B897603261CFFA0FBEBFF391AB0FF13" bold="true" box="[1052,1172,210,231]" italics="true" pageId="3" pageNumber="404">contrasticus</emphasis>
</emphasis>
Evenhuis &amp; Gharali
</taxonomicName>
,
<emphasis id="1B897603261CFFA0FA91FF391BB9FF13" bold="true" box="[1382,1437,210,231]" pageId="3" pageNumber="404">
<taxonomicNameLabel id="00BACB78261CFFA0FA91FF391BB9FF13" box="[1382,1437,210,231]" pageId="3" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="2942AA11261CFFA0FF60FF1B1BB9FECB" blockId="3.[151,1437,151,819]" pageId="3" pageNumber="404">
<keyLead id="920C0924261CFFA0FF60FF1B1BB9FECB" pageId="3" pageNumber="404">
–. Wing vein R 4+5 ending in wing margin well beyond level of end of M 2 (
<figureCitation id="B1C6B694261CFFA0FC6EFF041DC6FEF1" box="[921,994,239,261]" captionStart="FIGURES 40–47" captionStartId="15.[151,264,1607,1631]" captionTargetBox="[170,1416,309,1582]" captionTargetId="figure-81@15.[170,1416,309,1584]" captionTargetPageId="15" captionText="FIGURES 40–47. Malagasy Empidideicus (Anomaloptilus), wings. 40, E. (A.) andriambahomananii, n. sp.; 41, E. (A.) contrasticus, n. sp.; 42, E. (A.) hauseri, n. sp.; 43, E. (A.) irwini, n. sp.; 44, E. (A.) nanus, n. sp.; 45, E. (A.) regius, n. sp.; 46, E. (A.) rhinoclypeatus Evenhuis; 47, E. (A.) rinhai, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471764" httpUri="https://zenodo.org/record/7471764/files/figure.png" pageId="3" pageNumber="404">Fig. 43</figureCitation>
); cell r4+5 at wing margin narrower than cell m2; eyes in males holoptic or nearly so below level of antennae; prescutellar area in females black, without yellow marks................................................ (southwest
<collectingCountry id="51EAEA81261CFFA0FC8FFEC11DCAFEB4" box="[888,1006,298,320]" name="Madagascar" pageId="3" pageNumber="404">Madagascar</collectingCountry>
) ...
<emphasis id="1B897603261CFFA0FBE5FEC11A06FECB" bold="true" box="[1042,1058,298,319]" italics="true" pageId="3" pageNumber="404">E</emphasis>
.
<taxonomicName id="EEFDD192261CFFA0FBD9FEC11B78FEB4" authority="Evenhuis &amp; Gharali" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[1070,1372,298,320]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="irwini" status="sp. nov." subGenus="Anomaloptilus">
<emphasis id="1B897603261CFFA0FBD9FEC11AB1FECB" bold="true" box="[1070,1173,298,320]" pageId="3" pageNumber="404">
(
<emphasis id="1B897603261CFFA0FBC1FEC11A62FECB" bold="true" box="[1078,1094,298,319]" italics="true" pageId="3" pageNumber="404">A</emphasis>
.)
<emphasis id="1B897603261CFFA0FBADFEC11AB1FECB" bold="true" box="[1114,1173,298,319]" italics="true" pageId="3" pageNumber="404">irwini</emphasis>
</emphasis>
Evenhuis &amp; Gharali
</taxonomicName>
,
<emphasis id="1B897603261CFFA0FA91FEC11BB9FECB" bold="true" box="[1382,1437,298,319]" pageId="3" pageNumber="404">
<taxonomicNameLabel id="00BACB78261CFFA0FA91FEC11BB9FECB" box="[1382,1437,298,319]" pageId="3" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9209B2B4261CFFA0FF60FEA31BB9FE63" pageId="3" pageNumber="404">
<paragraph id="2942AA11261CFFA0FF60FEA31BB8FEA9" blockId="3.[151,1437,151,819]" box="[151,1436,327,349]" pageId="3" pageNumber="404">
<keyLead id="920C0924261CFFA0FF60FEA31BB8FEA9" box="[151,1436,327,349]" pageId="3" pageNumber="404">
4. Second antennal flagellomere 1/3 or more length of first flagellomere (
<figureCitation id="B1C6B694261CFFA0FC7DFEAC1DF4FEA9" box="[906,976,327,349]" captionStart="FIGURES 23–24" captionStartId="10.[151,264,1309,1333]" captionTargetBox="[151,1435,181,1284]" captionTargetId="figure-17@10.[151,1435,181,1285]" captionTargetPageId="10" captionText="FIGURES 23–24. Malagasy Empidideicus (Anomaloptilus), antennae. 23, E. (A.) andriambahomananii, n. sp.; 24, E. (A.) nanus, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471754" httpUri="https://zenodo.org/record/7471754/files/figure.png" pageId="3" pageNumber="404">Fig. 23</figureCitation>
).................................... 5
</keyLead>
</paragraph>
<paragraph id="2942AA11261CFFA0FF60FE8E1BB9FE63" blockId="3.[151,1437,151,819]" pageId="3" pageNumber="404">
<keyLead id="920C0924261CFFA0FF60FE8E1BB9FE63" pageId="3" pageNumber="404">
–. Second antennal flagellomere reduced in size, less than 1/4 length of first flagellomere (
<figureCitation id="B1C6B694261CFFA0FBD9FE8E1A50FE8E" box="[1070,1140,357,379]" captionStart="FIGURES 23–24" captionStartId="10.[151,264,1309,1333]" captionTargetBox="[151,1435,181,1284]" captionTargetId="figure-17@10.[151,1435,181,1285]" captionTargetPageId="10" captionText="FIGURES 23–24. Malagasy Empidideicus (Anomaloptilus), antennae. 23, E. (A.) andriambahomananii, n. sp.; 24, E. (A.) nanus, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471754" httpUri="https://zenodo.org/record/7471754/files/figure.png" pageId="3" pageNumber="404">Fig. 24</figureCitation>
)......................................................................................... E.
<taxonomicName id="EEFDD192261CFFA0FBDCFE691B78FE6C" authority="Evenhuis &amp; Gharali" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[1067,1372,386,408]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov." subGenus="Anomaloptilus">
<emphasis id="1B897603261CFFA0FBDCFE691AB0FE63" bold="true" box="[1067,1172,386,408]" pageId="3" pageNumber="404">
(
<emphasis id="1B897603261CFFA0FBC4FE691A67FE63" bold="true" box="[1075,1091,386,407]" italics="true" pageId="3" pageNumber="404">A</emphasis>
.)
<emphasis id="1B897603261CFFA0FBA0FE691AB0FE63" bold="true" box="[1111,1172,386,407]" italics="true" pageId="3" pageNumber="404">nanus</emphasis>
</emphasis>
Evenhuis &amp; Gharali
</taxonomicName>
,
<emphasis id="1B897603261CFFA0FA91FE691BB9FE63" bold="true" box="[1382,1437,386,407]" pageId="3" pageNumber="404">
<taxonomicNameLabel id="00BACB78261CFFA0FA91FE691BB9FE63" box="[1382,1437,386,407]" pageId="3" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9209B2B4261CFFA0FF60FE741BB8FE04" pageId="3" pageNumber="404">
<paragraph id="2942AA11261CFFA0FF60FE741BB8FE26" blockId="3.[151,1437,151,819]" pageId="3" pageNumber="404">
<keyLead id="920C0924261CFFA0FF60FE741BB8FE26" pageId="3" pageNumber="404">
5. Prescutellar area with yellow color; scutellum all yellow in color, no darker color present; R 4+5 either upcurved (
<emphasis id="1B897603261CFFA0FAECFE4B1BA0FE40" box="[1307,1412,415,437]" italics="true" pageId="3" pageNumber="404">
<taxonomicName id="EEFDD192261CFFA0FAECFE4B1BA4FE40" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[1307,1408,415,437]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="hauseri" status="sp. nov.">E. hauseri</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="00BACB78261CFFA0FA7DFE4B1EDAFE26" pageId="3" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
) or not upcurved (
<emphasis id="1B897603261CFFA0FE46FE561C29FE26" box="[433,525,445,466]" italics="true" pageId="3" pageNumber="404">
<taxonomicName id="EEFDD192261CFFA0FE46FE561C2DFE26" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[433,521,445,466]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="rinhai" status="sp. nov.">E. rinhai</taxonomicName>
,
</emphasis>
<emphasis id="1B897603261CFFA0FDE5FE561C6DFE26" bold="true" box="[530,585,445,466]" pageId="3" pageNumber="404">
<taxonomicNameLabel id="00BACB78261CFFA0FDE5FE561C6DFE26" box="[530,585,445,466]" pageId="3" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
) at wing margin........................................................ 7
</keyLead>
</paragraph>
<paragraph id="2942AA11261CFFA0FF60FE301BB8FE04" blockId="3.[151,1437,151,819]" box="[151,1436,474,501]" pageId="3" pageNumber="404">
<keyLead id="920C0924261CFFA0FF60FE301BB8FE04" box="[151,1436,474,501]" pageId="3" pageNumber="404">
–. Prescutellar area black; scutellum with some dark brown to black color, not all yellow; R
<subScript id="B579A854261CFFA0FBCFFE021A69FE01" attach="left" box="[1080,1101,489,501]" fontSize="5" pageId="3" pageNumber="404">4+5</subScript>
not upcurved at wing margin... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9209B2B4261CFFA0FF60FE1C1BB9FD76" pageId="3" pageNumber="404">
<paragraph id="2942AA11261CFFA0FF60FE1C1BB9FDDE" blockId="3.[151,1437,151,819]" pageId="3" pageNumber="404">
<keyLead id="920C0924261CFFA0FF60FE1C1BB9FDDE" pageId="3" pageNumber="404">
6. Scutellum all black; anal cell narrowly open at wing margin (
<figureCitation id="B1C6B694261CFFA0FCD3FE1C1D4FFDF9" box="[804,875,503,525]" captionStart="FIGURES 40–47" captionStartId="15.[151,264,1607,1631]" captionTargetBox="[170,1416,309,1582]" captionTargetId="figure-81@15.[170,1416,309,1584]" captionTargetPageId="15" captionText="FIGURES 40–47. Malagasy Empidideicus (Anomaloptilus), wings. 40, E. (A.) andriambahomananii, n. sp.; 41, E. (A.) contrasticus, n. sp.; 42, E. (A.) hauseri, n. sp.; 43, E. (A.) irwini, n. sp.; 44, E. (A.) nanus, n. sp.; 45, E. (A.) regius, n. sp.; 46, E. (A.) rhinoclypeatus Evenhuis; 47, E. (A.) rinhai, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471764" httpUri="https://zenodo.org/record/7471764/files/figure.png" pageId="3" pageNumber="404">Fig. 45</figureCitation>
); females with pleura predominantly yellow, darker yellow to brown only on lower pleurites........................................
<emphasis id="1B897603261CFFA0FBF8FDFE1A3BFDDE" bold="true" box="[1039,1055,533,554]" italics="true" pageId="3" pageNumber="404">E</emphasis>
.
<taxonomicName id="EEFDD192261CFFA0FBDCFDFF1B78FDDF" authority="Evenhuis &amp; Gharali" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[1067,1372,532,555]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="regius" status="sp. nov." subGenus="Anomaloptilus">
<emphasis id="1B897603261CFFA0FBDCFDFF1AB0FDDE" bold="true" box="[1067,1172,532,554]" pageId="3" pageNumber="404">
(
<emphasis id="1B897603261CFFA0FBC4FDFE1A67FDDE" bold="true" box="[1075,1091,533,554]" italics="true" pageId="3" pageNumber="404">A</emphasis>
.)
<emphasis id="1B897603261CFFA0FBA0FDFE1AB0FDDE" bold="true" box="[1111,1172,533,554]" italics="true" pageId="3" pageNumber="404">regius</emphasis>
</emphasis>
Evenhuis &amp; Gharali
</taxonomicName>
,
<emphasis id="1B897603261CFFA0FA91FDFE1BB9FDDE" bold="true" box="[1382,1437,533,554]" pageId="3" pageNumber="404">
<taxonomicNameLabel id="00BACB78261CFFA0FA91FDFE1BB9FDDE" box="[1382,1437,533,554]" pageId="3" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="2942AA11261CFFA0FF60FDD81BB9FD76" blockId="3.[151,1437,151,819]" pageId="3" pageNumber="404">
<keyLead id="920C0924261CFFA0FF60FDD81BB9FD76" pageId="3" pageNumber="404">
–. Scutellum predominantly yellow with brown basomedially; anal cell closed at wing margin (
<figureCitation id="B1C6B694261CFFA0FB8CFDD91AE1FDB3" box="[1147,1221,562,584]" captionStart="FIGURES 40–47" captionStartId="15.[151,264,1607,1631]" captionTargetBox="[170,1416,309,1582]" captionTargetId="figure-81@15.[170,1416,309,1584]" captionTargetPageId="15" captionText="FIGURES 40–47. Malagasy Empidideicus (Anomaloptilus), wings. 40, E. (A.) andriambahomananii, n. sp.; 41, E. (A.) contrasticus, n. sp.; 42, E. (A.) hauseri, n. sp.; 43, E. (A.) irwini, n. sp.; 44, E. (A.) nanus, n. sp.; 45, E. (A.) regius, n. sp.; 46, E. (A.) rhinoclypeatus Evenhuis; 47, E. (A.) rinhai, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471764" httpUri="https://zenodo.org/record/7471764/files/figure.png" pageId="3" pageNumber="404">Fig. 40</figureCitation>
); females with pleura predominantly black, yellow only on upper portions of katepimeron, katepisternum and propleuron...........................................................................
<emphasis id="1B897603261CFFA0FC83FD861DA0FD76" bold="true" box="[884,900,621,642]" italics="true" pageId="3" pageNumber="404">E</emphasis>
.
<taxonomicName id="EEFDD192261CFFA0FC67FD871B78FD77" authority="Evenhuis &amp; Gharali" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[912,1372,620,643]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="andriambahomananii" status="sp. nov." subGenus="Anomaloptilus">
<emphasis id="1B897603261CFFA0FC67FD871AB0FD76" bold="true" box="[912,1172,620,642]" pageId="3" pageNumber="404">
(
<emphasis id="1B897603261CFFA0FC6FFD861D8CFD76" bold="true" box="[920,936,621,642]" italics="true" pageId="3" pageNumber="404">A</emphasis>
.)
<emphasis id="1B897603261CFFA0FC4BFD871AB0FD76" bold="true" box="[956,1172,620,642]" italics="true" pageId="3" pageNumber="404">andriambahomananii</emphasis>
</emphasis>
Evenhuis &amp; Gharali
</taxonomicName>
,
<emphasis id="1B897603261CFFA0FA91FD861BB9FD76" bold="true" box="[1382,1437,621,642]" pageId="3" pageNumber="404">
<taxonomicNameLabel id="00BACB78261CFFA0FA91FD861BB9FD76" box="[1382,1437,621,642]" pageId="3" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9209B2B4261CFFA0FF60FD601BB9FCC6" pageId="3" pageNumber="404">
<paragraph id="2942AA11261CFFA0FF60FD601BB9FD2E" blockId="3.[151,1437,151,819]" pageId="3" pageNumber="404">
<keyLead id="920C0924261CFFA0FF60FD601BB9FD2E" pageId="3" pageNumber="404">
7. Prescutellar area with some dark color medially, yellow marks not broadly coalesced (some orange color medially in females) (
<figureCitation id="B1C6B694261CFFA0FF1EFD4C1F7DFD49" box="[233,345,679,701]" captionStart="FIGURES 59–62" captionStartId="19.[151,264,694,718]" captionTargetBox="[152,1435,181,670]" captionTargetId="figure-17@19.[152,1435,181,670]" captionTargetPageId="19" captionText="FIGURES 59–62. Malagasy Empidideicus (Anomaloptilus), abdomens, dorsal view. 59, E. (A.) rhinoclypeatus Evenhuis, male; 60, E. (A.) rhinoclypeatus Evenhuis, female; 61, E. (A.) rinhai, n. sp., male; 62, E. (A.) rinhai, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471774" httpUri="https://zenodo.org/record/7471774/files/figure.png" pageId="3" pageNumber="404">Figs 61–62</figureCitation>
); anal cell open at wing margin; wing vein R 4+5 straight to wing margin; abdominal tergite I with white band along posterior margin..................................................... E.
<taxonomicName id="EEFDD192261CFFA0FBDCFD2F1B78FD2F" authority="Evenhuis &amp; Gharali" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[1067,1372,708,731]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="rinhai" status="sp. nov." subGenus="Anomaloptilus">
<emphasis id="1B897603261CFFA0FBDCFD2F1AB0FD2E" bold="true" box="[1067,1172,708,730]" pageId="3" pageNumber="404">
(
<emphasis id="1B897603261CFFA0FBC4FD2E1A67FD2E" bold="true" box="[1075,1091,709,730]" italics="true" pageId="3" pageNumber="404">A</emphasis>
.)
<emphasis id="1B897603261CFFA0FBA0FD2F1AB0FD2E" bold="true" box="[1111,1172,708,730]" italics="true" pageId="3" pageNumber="404">rinhai</emphasis>
</emphasis>
Evenhuis &amp; Gharali
</taxonomicName>
,
<emphasis id="1B897603261CFFA0FA91FD2E1BB9FD2E" bold="true" box="[1382,1437,709,730]" pageId="3" pageNumber="404">
<taxonomicNameLabel id="00BACB78261CFFA0FA91FD2E1BB9FD2E" box="[1382,1437,709,730]" pageId="3" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="2942AA11261CFFA0FF60FD081BB9FCC6" blockId="3.[151,1437,151,819]" pageId="3" pageNumber="404">
<keyLead id="920C0924261CFFA0FF60FD081BB9FCC6" pageId="3" pageNumber="404">
–. Prescutellar area yellow, without dark color medially anterior to scutellum, yellow marks broadly coalesced (
<figureCitation id="B1C6B694261CFFA0FAE9FD091BB6FD0C" box="[1310,1426,738,760]" captionStart="FIGURES 50–58" captionStartId="18.[151,264,1889,1913]" captionTargetBox="[239,1348,181,1865]" captionTargetId="figure-17@18.[239,1348,181,1865]" captionTargetPageId="18" captionText="FIGURES 50–58. Malagasy Empidideicus (Anomaloptilus), abdomens, dorsal view. 50, E. (A.) andriambahomananii, n. sp., female; 51, E. (A.) contrasticus, n. sp., male; 52, E. (A.) contrasticus, n. sp., female; 53, E. (A.) hauseri, n. sp., male; 54, E. (A.) hauseri, n. sp., female; 55, E. (A.) irwini, n. sp., male; 56, E. (A.) irwini, n. sp., female; 57, E. (A.) nanus, n. sp., male; 58, E. (A.) regius, n. sp., male." figureDoi="http://doi.org/10.5281/zenodo.7471772" httpUri="https://zenodo.org/record/7471772/files/figure.png" pageId="3" pageNumber="404">Figs 53–54</figureCitation>
); anal cell closed in wing margin; wing vein R 4+5 upcurved at wing margin; abdominal tergite I all brown, without white band posteriorly......................................................... E.
<taxonomicName id="EEFDD192261CFFA0FBE9FCF71B78FCC7" authority="Evenhuis &amp; Gharali" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[1054,1372,796,819]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="hauseri" status="sp. nov." subGenus="Anomaloptilus">
<emphasis id="1B897603261CFFA0FBE9FCF71AB1FCC6" bold="true" box="[1054,1173,796,818]" pageId="3" pageNumber="404">
(
<emphasis id="1B897603261CFFA0FBD1FCF61A12FCC6" bold="true" box="[1062,1078,797,818]" italics="true" pageId="3" pageNumber="404">A</emphasis>
.)
<emphasis id="1B897603261CFFA0FBBDFCF71AB1FCC6" bold="true" box="[1098,1173,796,818]" italics="true" pageId="3" pageNumber="404">hauseri</emphasis>
</emphasis>
Evenhuis &amp; Gharali
</taxonomicName>
,
<emphasis id="1B897603261CFFA0FA91FCF61BB9FCC6" bold="true" box="[1382,1437,797,818]" pageId="3" pageNumber="404">
<taxonomicNameLabel id="00BACB78261CFFA0FA91FCF61BB9FCC6" box="[1382,1437,797,818]" pageId="3" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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