treatments-xml/data/A1/54/1B/A1541B072614FFAFFF60FD3D1B28FCA9.xml
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<document id="AFCBC731A179802D94C2BB27DBEB66EE" ID-CLB-Dataset="54548" ID-DOI="10.11646/zootaxa.5222.5.1" ID-GBIF-Dataset="a2aef7c8-09e1-48f1-983e-00132055a5a2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7471736" ID-ZooBank="F4983B58-7301-4A3F-AC08-3E5120450EA1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1671694447742" checkinUser="plazi" docAuthor="Evenhuis, Neal L. &amp; Gharali, Babak" docDate="2022" docId="A1541B072614FFAFFF60FD3D1B28FCA9" docLanguage="en" docName="zootaxa.5222.5.1.pdf" docOrigin="Zootaxa 5222 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Empidideicus (Anomaloptilus) nanus Evenhuis &amp; Gharali 2022, n. sp." docType="treatment" docVersion="7" lastPageNumber="413" masterDocId="5D6D637F261FFFA3FFF7FFEB1E24FFF4" masterDocTitle="The subgenus Empidideicus (Anomaloptilus) (Diptera: Mythicomyiidae) in Madagascar" masterLastPageNumber="426" masterPageNumber="401" pageNumber="412" updateTime="1699413895303" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8E1856209DA5105408ED60048A8B75F1">The subgenus Empidideicus (Anomaloptilus) (Diptera: Mythicomyiidae) in Madagascar</mods:title>
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<mods:namePart id="884369F5543D79F9E143C15072499C8D">Evenhuis, Neal L.</mods:namePart>
<mods:nameIdentifier id="B8BA929B893C5292566C0E51F816923C" type="ORCID">0000-0002-1314-755X</mods:nameIdentifier>
<mods:affiliation id="2A746D74270079A3F1EF0158960F93F6">J. Linsley Gressitt Center for Research in Entomology, Bishop Museum, 1525 Bernice Street, Honolulu, Hawaii 96817 - 2704, USA neale @ bishopmuseum. org; https: // orcid. org / 0000 - 0002 - 1314 - 755 X</mods:affiliation>
<mods:nameIdentifier id="5EB6FE9F8362E0B73D844F260BB958AF" type="email">neale@bishopmuseum.org</mods:nameIdentifier>
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<mods:namePart id="253286D71D090F44754D2E1DCCA5BECC">Gharali, Babak</mods:namePart>
<mods:nameIdentifier id="9B7F2ABD021075FB85C6B1E1DCE1C212" type="ORCID">0000-0003-2490-156X</mods:nameIdentifier>
<mods:affiliation id="5F84CBFA08E098FEE035CA43885E1C96">Plant Protection Research Department, Qazvin Agricultural and Natural Resources Research and Education Center, AREEO, Qazvin, Iran. bgaharaei @ yahoo. com; https: // orcid. org / 0000 - 0003 - 2490 - 156 X</mods:affiliation>
<mods:nameIdentifier id="9BFECEF493FBEC004B6618202DBF4F6F" type="email">bgaharaei@yahoo.com</mods:nameIdentifier>
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<mods:title id="3379ABF767D034E112659BE9659903BB">Zootaxa</mods:title>
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<mods:date id="DFEDFB568E34AB68382DB182DFD33FE2">2022</mods:date>
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<treatment id="A1541B072614FFAFFF60FD3D1B28FCA9" ID-DOI="http://doi.org/10.5281/zenodo.7471788" ID-GBIF-Taxon="207927030" ID-Zenodo-Dep="7471788" LSID="urn:lsid:plazi:treatment:A1541B072614FFAFFF60FD3D1B28FCA9" httpUri="http://treatment.plazi.org/id/A1541B072614FFAFFF60FD3D1B28FCA9" lastPageId="12" lastPageNumber="413" pageId="11" pageNumber="412">
<subSubSection id="61E7F99A2614FFA8FF60FD3D1D94FD05" box="[151,944,726,753]" pageId="11" pageNumber="412" type="nomenclature">
<paragraph id="2942AA112614FFA8FF60FD3D1D94FD05" blockId="11.[151,944,726,789]" box="[151,944,726,753]" pageId="11" pageNumber="412">
<heading id="720A1D7D2614FFA8FF60FD3D1D94FD05" bold="true" box="[151,944,726,753]" fontSize="11" level="1" pageId="11" pageNumber="412" reason="1">
<emphasis id="1B8976032614FFA8FF60FD3D1D94FD05" bold="true" box="[151,944,726,753]" pageId="11" pageNumber="412">
<taxonomicName id="EEFDD1922614FFA8FF60FD3D1D45FD04" authority="Evenhuis &amp; Gharali" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[151,865,726,753]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov." subGenus="Anomaloptilus">
<emphasis id="1B8976032614FFA8FF60FD3D1C7FFD05" bold="true" box="[151,603,726,753]" italics="true" pageId="11" pageNumber="412">Empidideicus (Anomaloptilus) nanus</emphasis>
Evenhuis &amp; Gharali
</taxonomicName>
,
<taxonomicNameLabel id="00BACB782614FFA8FC9AFD3C1D94FD05" box="[877,944,727,753]" pageId="11" pageNumber="412" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="61E7F99A2614FFA8FF60FD101FFBFB31" pageId="11" pageNumber="412" type="description">
<paragraph id="2942AA112614FFA8FF60FD101C07FCE1" blockId="11.[151,944,726,789]" box="[151,547,762,789]" pageId="11" pageNumber="412">
(
<figureCitation id="B1C6B6942614FFA8FF68FD101ED2FCE1" box="[159,246,763,789]" captionStart="FIGURES 510" captionStartId="4.[151,264,1768,1792]" captionTargetBox="[151,1435,181,1745]" captionTargetId="figure-17@4.[151,1435,181,1745]" captionTargetPageId="4" captionText="FIGURES 510. Malagasy Empidideicus (Anomaloptilus), habitus, lateral view. 5, E. (A.) andriambahomananii, n. sp., male; 6, E. (A.) andriambahomananii, n. sp., female; 7, E. (A.) contrasticus, n. sp., male; 8, E. (A.) hauseri, n. sp., male; 9, E. (A.) irwini, n. sp., male; 10, E. (A.) nanus, n. sp., male." figureDoi="http://doi.org/10.5281/zenodo.7471746" httpUri="https://zenodo.org/record/7471746/files/figure.png" pageId="11" pageNumber="412">Figs 10</figureCitation>
,
<figureCitation id="B1C6B6942614FFA8FEF5FD101F04FCE1" box="[258,288,763,789]" captionStart="FIGURES 2324" captionStartId="10.[151,264,1309,1333]" captionTargetBox="[151,1435,181,1284]" captionTargetId="figure-17@10.[151,1435,181,1285]" captionTargetPageId="10" captionText="FIGURES 2324. Malagasy Empidideicus (Anomaloptilus), antennae. 23, E. (A.) andriambahomananii, n. sp.; 24, E. (A.) nanus, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471754" httpUri="https://zenodo.org/record/7471754/files/figure.png" pageId="11" pageNumber="412">24</figureCitation>
,
<figureCitation id="B1C6B6942614FFA8FEDBFD101F50FCE1" box="[300,372,763,789]" captionStart="FIGURES 2533" captionStartId="13.[151,264,1878,1902]" captionTargetBox="[151,1435,273,1855]" captionTargetId="figure-65@13.[151,1435,273,1855]" captionTargetPageId="13" captionText="FIGURES 2533. Malagasy Empidideicus (Anomaloptilus), thoraces, dorsal view. 25, E. (A.) andriambahomananii, n. sp., male; 26, E. (A.) contrasticus, n. sp., male; 27, E. (A.) contrasticus, n. sp., female; 28, E. (A.) hauseri, n. sp., male; 29, E. (A.) hauseri, n. sp., female; 30, E. (A.) irwini, n. sp., male; 31, E. (A.) irwini, n. sp., female; 32, E. (A.) nanus, n. sp., male; 33, E. (A.) nanus, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471758" httpUri="https://zenodo.org/record/7471758/files/figure.png" pageId="11" pageNumber="412">32, 33</figureCitation>
,
<figureCitation id="B1C6B6942614FFA8FE77FD101FBBFCE1" box="[384,415,763,789]" captionStart="FIGURES 4047" captionStartId="15.[151,264,1607,1631]" captionTargetBox="[170,1416,309,1582]" captionTargetId="figure-81@15.[170,1416,309,1584]" captionTargetPageId="15" captionText="FIGURES 4047. Malagasy Empidideicus (Anomaloptilus), wings. 40, E. (A.) andriambahomananii, n. sp.; 41, E. (A.) contrasticus, n. sp.; 42, E. (A.) hauseri, n. sp.; 43, E. (A.) irwini, n. sp.; 44, E. (A.) nanus, n. sp.; 45, E. (A.) regius, n. sp.; 46, E. (A.) rhinoclypeatus Evenhuis; 47, E. (A.) rinhai, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471764" httpUri="https://zenodo.org/record/7471764/files/figure.png" pageId="11" pageNumber="412">44</figureCitation>
,
<figureCitation id="B1C6B6942614FFA8FE5DFD111FECFCE1" box="[426,456,762,789]" captionStart="FIGURES 5058" captionStartId="18.[151,264,1889,1913]" captionTargetBox="[239,1348,181,1865]" captionTargetId="figure-17@18.[239,1348,181,1865]" captionTargetPageId="18" captionText="FIGURES 5058. Malagasy Empidideicus (Anomaloptilus), abdomens, dorsal view. 50, E. (A.) andriambahomananii, n. sp., female; 51, E. (A.) contrasticus, n. sp., male; 52, E. (A.) contrasticus, n. sp., female; 53, E. (A.) hauseri, n. sp., male; 54, E. (A.) hauseri, n. sp., female; 55, E. (A.) irwini, n. sp., male; 56, E. (A.) irwini, n. sp., female; 57, E. (A.) nanus, n. sp., male; 58, E. (A.) regius, n. sp., male." figureDoi="http://doi.org/10.5281/zenodo.7471772" httpUri="https://zenodo.org/record/7471772/files/figure.png" pageId="11" pageNumber="412">57</figureCitation>
,
<figureCitation id="B1C6B6942614FFA8FE23FD101C3FFCE1" box="[468,539,763,789]" captionStart="FIGURES 7172" captionStartId="21.[152,270,1956,1980]" captionTargetBox="[152,1436,1100,1933]" captionTargetId="figure-168@21.[151,1436,1100,1933]" captionTargetPageId="21" captionText="FIGURES 7172. Empidideicus (Anomaloptilus) nanus, n. sp., genitalia. 71, male, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view; 72, female." figureDoi="http://doi.org/10.5281/zenodo.7471790" httpUri="https://zenodo.org/record/7471790/files/figure.png" pageId="11" pageNumber="412">71, 72</figureCitation>
)
</paragraph>
<paragraph id="2942AA112614FFA8FF60FCA81FFBFB31" blockId="11.[151,1437,835,2019]" pageId="11" pageNumber="412">
<emphasis id="1B8976032614FFA8FF60FCA81EF8FCA9" bold="true" box="[151,220,835,861]" pageId="11" pageNumber="412">Types</emphasis>
.
<materialsCitation id="9995A04C2614FFA8FF1CFCA81F9FFC51" ID-GBIF-Occurrence="4003029301" collectingDate="1986-09" collectingDateMax="2003-09-28" collectingDateMin="1986-09" collectionCode="R, CAS, ANGAP, V, BMNH" collectorName="M. E. Irwin &amp; Harin'Hala" country="Madagascar" county="Ampijora (Andranofasika) National Park" elevation="46" latitude="-12.95" location="Mahafely Reserve" longLatPrecision="775" longitude="49.14" municipality="Zombitse National Park" pageId="11" pageNumber="412" specimenCode="MG 25-16, CASLOT 018549, CASLOT 019832" specimenCount="20" specimenCount-female="20" stateProvince="Toliara" typeStatus="holotype">
<emphasis id="1B8976032614FFA8FF1CFCA81F76FCA9" box="[235,338,835,861]" italics="true" pageId="11" pageNumber="412">
<typeStatus id="F64614B32614FFA8FF1CFCA81F76FCA9" box="[235,338,835,861]" pageId="11" pageNumber="412" type="holotype">Holotype</typeStatus>
</emphasis>
♂ and 5♂,
<specimenCount id="3FFB61982614FFA8FE2FFCA81FDFFCA9" box="[472,507,835,861]" pageId="11" pageNumber="412" type="female">3♀</specimenCount>
<emphasis id="1B8976032614FFA8FDF4FCA81C55FCA8" box="[515,625,835,860]" italics="true" pageId="11" pageNumber="412">
<typeStatus id="F64614B32614FFA8FDF4FCA81C55FCA8" box="[515,625,835,860]" pageId="11" pageNumber="412" type="paratype">paratypes</typeStatus>
</emphasis>
from:
<collectingCountry id="51EAEA812614FFA8FD48FCA81DAFFCA9" box="[703,907,835,861]" name="Madagascar" pageId="11" pageNumber="412">MADAGASCAR</collectingCountry>
: Majunga Province:
<collectingCounty id="C023D29D2614FFA8FB85FCA81F12FC75" pageId="11" pageNumber="412">Ampijora (Andranofasika) National Park</collectingCounty>
,
<geoCoordinate id="4CC9CCD62614FFA8FEB4FC831FE6FC77" box="[323,450,872,899]" degrees="16" direction="south" minutes="19.16" orientation="latitude" pageId="11" pageNumber="412" precision="9" value="-16.319334">16°19.16S</geoCoordinate>
,
<geoCoordinate id="4CC9CCD62614FFA8FE39FC831C6BFC77" box="[462,591,872,899]" degrees="46" direction="east" minutes="48.90" orientation="longitude" pageId="11" pageNumber="412" precision="9" value="46.815">46°48.90E</geoCoordinate>
,
<quantity id="EE0507F42614FFA8FDACFC8C1CBCFC75" box="[603,664,871,897]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="11" pageNumber="412" unit="m" value="46.0">
<elevation id="A2D04D222614FFA8FDACFC8C1CBCFC75" box="[603,664,871,897]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="11" pageNumber="412" unit="m" value="46.0">46 m</elevation>
</quantity>
, 22/
<date id="5D438CD12614FFA8FD3EFC8C1D7EFC75" box="[713,858,871,897]" pageId="11" pageNumber="412" value="2003-09-28">
<collectingDate id="4D0775392614FFA8FD3EFC8C1D7EFC75" box="[713,858,871,897]" pageId="11" pageNumber="412" value="2003-09-28">28 Sep 2003</collectingDate>
</date>
,
<collectorName id="8408CFC72614FFA8FC90FC8C1DC2FC75" box="[871,998,871,897]" pageId="11" pageNumber="412">M.E. Irwin</collectorName>
,
<collectionCode id="4FEC32D42614FFA8FC04FC8C1A2DFC75" box="[1011,1033,871,897]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="412">R</collectionCode>
. HarinHala,
<collectingMethod id="F0BCD2062614FFA8FB54FC8C1B20FC75" box="[1187,1284,871,897]" pageId="11" pageNumber="412">hand net</collectingMethod>
in deciduous forest,
<specimenCode id="795B026A2614FFA8FF11FC601F46FC51" box="[230,354,906,933]" collectionCode="MG" country="China" name="Museum of Zoology" pageId="11" pageNumber="412">MG 2516</specimenCode>
(
<collectionCode id="4FEC32D42614FFA8FE84FC601F8BFC51" box="[371,431,907,933]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="11" pageNumber="412" type="Museum">CAS</collectionCode>
).
</materialsCitation>
<materialsCitation id="9995A04C2614FFA8FE34FC601CE9FC3D" ID-GBIF-Occurrence="4003029308" collectingDate="1986-09" collectingDateMax="2003-09-28" collectingDateMin="1986-09" collectionCode="R, CAS, ANGAP, V, BMNH" collectorName="Harin'Hala" country="Madagascar" county="Ampijora (Andranofasika) National Park" elevation="46" latitude="-12.95" location="Mahafely Reserve" longLatPrecision="775" longitude="49.14" municipality="Zombitse National Park" pageId="11" pageNumber="412" specimenCode="MG 25-16, CASLOT 018549, CASLOT 019832" specimenCount="20" specimenCount-female="20" stateProvince="Toliara" typeStatus="paratype">
<emphasis id="1B8976032614FFA8FE34FC601C5FFC50" box="[451,635,907,933]" italics="true" pageId="11" pageNumber="412">
Other
<typeStatus id="F64614B32614FFA8FDFAFC601C5FFC50" box="[525,635,907,932]" pageId="11" pageNumber="412" type="paratype">paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="51EAEA812614FFA8FD7BFC601D7CFC51" box="[652,856,907,933]" name="Madagascar" pageId="11" pageNumber="412">MADAGASCAR</collectingCountry>
: 9♂,
<specimenCount id="3FFB61982614FFA8FC61FC601D9FFC51" box="[918,955,907,933]" pageId="11" pageNumber="412" type="female">4♀</specimenCount>
, topotypic,
<date id="5D438CD12614FFA8FBB5FC601AC9FC51" box="[1090,1261,907,933]" pageId="11" pageNumber="412" value="2003-09-07" valueMax="2003-09-14" valueMin="2003-09-07">
<collectingDate id="4D0775392614FFA8FBB5FC601AC9FC51" box="[1090,1261,907,933]" pageId="11" pageNumber="412" value="2003-09-07" valueMax="2003-09-14" valueMin="2003-09-07">714 Sep 2003</collectingDate>
</date>
<collectionCode id="4FEC32D42614FFA8FB02FC601B2FFC51" box="[1269,1291,907,933]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="412">R</collectionCode>
.
<collectorName id="8408CFC72614FFA8FAE0FC601BBCFC51" box="[1303,1432,907,933]" pageId="11" pageNumber="412">HarinHala</collectorName>
,
<collectorName id="8408CFC72614FFA8FF60FC441ED4FC3D" box="[151,240,943,969]" pageId="11" pageNumber="412">Malaise</collectorName>
in deciduous forest (
<specimenCode id="795B026A2614FFA8FE19FC441CE5FC3D" box="[494,705,942,969]" collectionCode="CASLOT" pageId="11" pageNumber="412">CASLOT 018549</specimenCode>
);
</materialsCitation>
<materialsCitation id="9995A04C2614FFA8FD2EFC441C84FBE5" ID-GBIF-Occurrence="4003029342" collectingDate="1986-09" collectingDateMax="2003-09-28" collectingDateMin="1986-09" collectionCode="R, CAS, ANGAP, V, BMNH" collectorName="M. E. Irwin &amp; F. Parker &amp; Harin'Hala" country="Madagascar" county="Ampijora (Andranofasika) National Park" elevation="46" latitude="-12.95" location="Mahafely Reserve" longLatPrecision="775" longitude="49.14" municipality="Zombitse National Park" pageId="11" pageNumber="412" specimenCode="MG 25-16, CASLOT 018549, CASLOT 019832" specimenCount="20" specimenCount-female="20" stateProvince="Toliara" typeStatus="holotype">
Tulear Province:
<specimenCount id="3FFB61982614FFA8FC53FC441DE3FC3D" box="[932,967,943,969]" pageId="11" pageNumber="412" type="female">1♀</specimenCount>
,
<collectingMunicipality id="C926306B2614FFA8FC2DFC441AD6FC3D" box="[986,1266,943,969]" pageId="11" pageNumber="412">Zombitse National Park</collectingMunicipality>
, near
<collectionCode id="4FEC32D42614FFA8FACAFC441BB8FC3D" box="[1341,1436,943,969]" pageId="11" pageNumber="412">ANGAP</collectionCode>
<collectorName id="8408CFC72614FFA8FF60FC381EC5FC19" box="[151,225,979,1005]" pageId="11" pageNumber="412">Office</collectorName>
,
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,
<geoCoordinate id="4CC9CCD62614FFA8FE81FC3F1FD3FC1B" box="[374,503,980,1007]" degrees="44" direction="east" minutes="41.53" orientation="longitude" pageId="11" pageNumber="412" precision="9" value="44.692165">44°41.53E</geoCoordinate>
,
<quantity id="EE0507F42614FFA8FDF5FC381C6FFC19" box="[514,587,979,1005]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="11" pageNumber="412" unit="m" value="840.0">
<elevation id="A2D04D222614FFA8FDF5FC381C6FFC19" box="[514,587,979,1005]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="11" pageNumber="412" unit="m" value="840.0">840 m</elevation>
</quantity>
,
<date id="5D438CD12614FFA8FDA0FC381CC2FC19" box="[599,742,979,1005]" pageId="11" pageNumber="412" value="2001-12-29">
<collectingDate id="4D0775392614FFA8FDA0FC381CC2FC19" box="[599,742,979,1005]" pageId="11" pageNumber="412" value="2001-12-29">29 Dec 2001</collectingDate>
</date>
<date id="5D438CD12614FFA8FD0BFC391D5DFC19" box="[764,889,978,1005]" pageId="11" pageNumber="412" value="2003-01-05">
<collectingDate id="4D0775392614FFA8FD0BFC391D5DFC19" box="[764,889,978,1005]" pageId="11" pageNumber="412" value="2003-01-05">5 Jan 2003</collectingDate>
</date>
,
<collectorName id="8408CFC72614FFA8FC73FC381A26FC19" box="[900,1026,979,1005]" pageId="11" pageNumber="412">M.E. Irwin</collectorName>
,
<collectorName id="8408CFC72614FFA8FBF9FC381A56FC19" box="[1038,1138,979,1005]" pageId="11" pageNumber="412">F. Parker</collectorName>
&amp;
<collectionCode id="4FEC32D42614FFA8FB60FC381A89FC19" box="[1175,1197,979,1005]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="412">R</collectionCode>
.
<collectorName id="8408CFC72614FFA8FB4FFC381B1DFC19" box="[1208,1337,979,1005]" pageId="11" pageNumber="412">HarinHala</collectorName>
,
<collectorName id="8408CFC72614FFA8FAB3FC381BB9FC19" box="[1348,1437,979,1005]" pageId="11" pageNumber="412">Malaise</collectorName>
in deciduous spiny forest (
<specimenCode id="795B026A2614FFA8FE33FC1C1CB0FBE5" box="[452,660,1015,1041]" collectionCode="CASLOT" pageId="11" pageNumber="412">CASLOT 019832</specimenCode>
);
</materialsCitation>
<materialsCitation id="9995A04C2614FFA8FD5EFC131A85FBC1" ID-GBIF-Occurrence="4003029336" collectingDate="1986-09" collectingDateMax="2003-09-28" collectingDateMin="1986-09" collectionCode="R, CAS, ANGAP, V, BMNH" collectorName="M. E. Irwin &amp; Harin'Hala" country="Madagascar" county="Ampijora (Andranofasika) National Park" elevation="46" latitude="-12.95" location="Mahafely Reserve" longLatPrecision="775" longitude="49.14" municipality="Zombitse National Park" pageId="11" pageNumber="412" specimenCode="MG 25-16, CASLOT 018549, CASLOT 019832" specimenCount="20" specimenCount-female="20" stateProvince="Toliara" typeStatus="holotype">
10♂,
<specimenCount id="3FFB61982614FFA8FD1EFC1C1D28FBE5" box="[745,780,1015,1041]" pageId="11" pageNumber="412" type="female">6♀</specimenCount>
<location id="2C22FCCA2614FFA8FCE2FC1C1DC2FBE5" LSID="urn:lsid:plazi:treatment:A1541B072614FFAFFF60FD3D1B28FCA9:2C22FCCA2614FFA8FCE2FC1C1DC2FBE5" box="[789,998,1015,1041]" country="Madagascar" county="Ampijora (Andranofasika) National Park" latitude="-12.95" longLatPrecision="775" longitude="49.14" municipality="Zombitse National Park" name="Mahafely Reserve" pageId="11" pageNumber="412" stateProvince="Toliara">Mahafely Reserve</location>
,
<collectorName id="8408CFC72614FFA8FC05FC1C1A68FBE5" box="[1010,1100,1015,1041]" pageId="11" pageNumber="412">Parcel I</collectorName>
,
<geoCoordinate id="4CC9CCD62614FFA8FBADFC131AFCFBE7" box="[1114,1240,1016,1043]" degrees="23" direction="south" minutes="41.19" orientation="latitude" pageId="11" pageNumber="412" precision="9" value="-23.6865">23°41.19S</geoCoordinate>
,
<geoCoordinate id="4CC9CCD62614FFA8FB12FC131B42FBE7" box="[1253,1382,1016,1043]" degrees="44" direction="east" minutes="35.46" orientation="longitude" pageId="11" pageNumber="412" precision="9" value="44.591">44°35.46E</geoCoordinate>
,
<quantity id="EE0507F42614FFA8FA85FC1C1E8BFBC1" metricMagnitude="2" metricUnit="m" metricValue="1.64" pageId="11" pageNumber="412" unit="m" value="164.0">
<elevation id="A2D04D222614FFA8FA85FC1C1E8BFBC1" metricMagnitude="2" metricUnit="m" metricValue="1.64" pageId="11" pageNumber="412" unit="m" value="164.0">164 m</elevation>
</quantity>
,
<date id="5D438CD12614FFA8FF4BFBF11F43FBC1" box="[188,359,1050,1077]" pageId="11" pageNumber="412" value="2002-10-05" valueMax="2002-10-20" valueMin="2002-10-05">
<collectingDate id="4D0775392614FFA8FF4BFBF11F43FBC1" box="[188,359,1050,1077]" pageId="11" pageNumber="412" value="2002-10-05" valueMax="2002-10-20" valueMin="2002-10-05">520 Oct 2002</collectingDate>
</date>
,
<collectorName id="8408CFC72614FFA8FE85FBF01FD5FBC1" box="[370,497,1051,1077]" pageId="11" pageNumber="412">M.E. Irwin</collectorName>
,
<collectionCode id="4FEC32D42614FFA8FE09FBF01C30FBC1" box="[510,532,1051,1077]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="412">R</collectionCode>
.
<collectorName id="8408CFC72614FFA8FDE8FBF01C84FBC1" box="[543,672,1051,1077]" pageId="11" pageNumber="412">HarinHala</collectorName>
,
<collectorName id="8408CFC72614FFA8FD5BFBF01D21FBC1" box="[684,773,1051,1077]" pageId="11" pageNumber="412">Malaise</collectorName>
across trail in gallery forest (
<collectionCode id="4FEC32D42614FFA8FBA5FBF01AB4FBC1" box="[1106,1168,1051,1077]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="11" pageNumber="412" type="Museum">CAS</collectionCode>
).;
</materialsCitation>
<materialsCitation id="9995A04C2614FFA8FB5FFBF01B0CFB89" ID-GBIF-Occurrence="4003029374" collectingDate="1986-09" collectingDateMax="2003-09-28" collectingDateMin="1986-09" collectionCode="R, CAS, ANGAP, V, BMNH" collectorName="Griswold et al." country="Madagascar" county="Ampijora (Andranofasika) National Park" elevation="46" latitude="-12.95" location="Mahafely Reserve" longLatPrecision="775" longitude="49.14" municipality="Zombitse National Park" pageId="11" pageNumber="412" specimenCode="MG 25-16, CASLOT 018549, CASLOT 019832" specimenCount="20" specimenCount-female="20" stateProvince="Toliara" typeStatus="holotype">
<collectingRegion id="EB3964F32614FFA8FB5FFBF01B43FBC1" box="[1192,1383,1051,1077]" country="Madagascar" name="Toliara" pageId="11" pageNumber="412">Toliara Province</collectingRegion>
: 2♂,
<specimenCount id="3FFB61982614FFA8FF60FBD41E98FBAD" box="[151,188,1087,1113]" pageId="11" pageNumber="412" type="female">2♀</specimenCount>
,
<location id="2C22FCCA2614FFA8FF3FFBD41C7DFBAD" LSID="urn:lsid:plazi:treatment:A1541B072614FFAFFF60FD3D1B28FCA9:2C22FCCA2614FFA8FF3FFBD41C7DFBAD" box="[200,601,1087,1113]" country="Madagascar" county="Ampijora (Andranofasika) National Park" latitude="-12.95" longLatPrecision="775" longitude="49.14" municipality="Zombitse National Park" name="Parc National de Tsimanampetsotsa" pageId="11" pageNumber="412" stateProvince="Toliara">Parc National de Tsimanampetsotsa</location>
,
<location id="2C22FCCA2614FFA8FD93FBD41D76FBAD" LSID="urn:lsid:plazi:treatment:A1541B072614FFAFFF60FD3D1B28FCA9:2C22FCCA2614FFA8FD93FBD41D76FBAD" box="[612,850,1087,1113]" country="Madagascar" county="Ampijora (Andranofasika) National Park" latitude="-12.95" longLatPrecision="775" longitude="49.14" municipality="Zombitse National Park" name="Foret de Bemanateza" pageId="11" pageNumber="412" stateProvince="Toliara">
Forêt
<collectorName id="8408CFC72614FFA8FD52FBD41D76FBAD" box="[677,850,1087,1113]" pageId="11" pageNumber="412">de Bemanateza</collectorName>
</location>
,
<quantity id="EE0507F42614FFA8FCAAFBD41D9CFBAD" box="[861,952,1087,1113]" metricMagnitude="4" metricUnit="m" metricValue="2.07" pageId="11" pageNumber="412" unit="km" value="20.7">20.7 km</quantity>
<collectorName id="8408CFC72614FFA8FC48FBD41A10FBAD" box="[959,1076,1087,1113]" pageId="11" pageNumber="412">E. Efoetse</collectorName>
,
<geoCoordinate id="4CC9CCD62614FFA8FBC8FBAB1AE6FBAF" box="[1087,1218,1088,1115]" degrees="23" direction="south" minutes="59" orientation="latitude" pageId="11" pageNumber="412" precision="15" seconds="32" value="-23.992222">23°5932″S</geoCoordinate>
,
<geoCoordinate id="4CC9CCD62614FFA8FB3AFBAB1B76FBAF" box="[1229,1362,1088,1115]" degrees="43" direction="east" minutes="52" orientation="longitude" pageId="11" pageNumber="412" precision="15" seconds="50" value="43.880554">43°5250″E</geoCoordinate>
,
<quantity id="EE0507F42614FFA8FAAAFBD41BB3FBAD" box="[1373,1431,1087,1113]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="412" unit="m" value="90.0">
<elevation id="A2D04D222614FFA8FAAAFBD41BB3FBAD" box="[1373,1431,1087,1113]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="412" unit="m" value="90.0">90 m</elevation>
</quantity>
,
<date id="5D438CD12614FFA8FF60FB881F77FB89" box="[151,339,1123,1149]" pageId="11" pageNumber="412" value="2002-03-22" valueMax="2002-03-26" valueMin="2002-03-22">
<collectingDate id="4D0775392614FFA8FF60FB881F77FB89" box="[151,339,1123,1149]" pageId="11" pageNumber="412" value="2002-03-22" valueMax="2002-03-26" valueMin="2002-03-22">2226 Mar 2002</collectingDate>
</date>
,
<collectorName id="8408CFC72614FFA8FEABFB881F80FB89" box="[348,420,1123,1149]" pageId="11" pageNumber="412">Fisher</collectorName>
,
<collectorName id="8408CFC72614FFA8FE59FB881C32FB89" box="[430,534,1123,1149]" pageId="11" pageNumber="412">Griswold</collectorName>
<emphasis id="1B8976032614FFA8FDEBFB881C68FB89" box="[540,588,1123,1149]" italics="true" pageId="11" pageNumber="412">et al</emphasis>
.,
<collectingMethod id="F0BCD2062614FFA8FD97FB881CCEFB89" box="[608,746,1123,1149]" pageId="11" pageNumber="412">Malaise trap</collectingMethod>
in spiny forest thicket, coll. code: BLF6257 (
<collectionCode id="4FEC32D42614FFA8FB28FB881B3FFB89" box="[1247,1307,1123,1149]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="11" pageNumber="412" type="Museum">CAS</collectionCode>
);
</materialsCitation>
<materialsCitation id="9995A04C2614FFA8FADBFB881A7FFB55" ID-GBIF-Occurrence="4003029358" collectingDate="1986-09" collectingDateMax="2003-09-28" collectingDateMin="1986-09" collectionCode="R, CAS, ANGAP, V, BMNH" collectorName="S. V. Fowler" country="Madagascar" county="Ampijora (Andranofasika) National Park" elevation="46" latitude="-12.95" location="Mahafely Reserve" longLatPrecision="775" longitude="49.14" municipality="Zombitse National Park" pageId="11" pageNumber="412" specimenCode="MG 25-16, CASLOT 018549, CASLOT 019832" specimenCount="20" specimenCount-female="20" stateProvince="Toliara" typeStatus="holotype">
Ambilobe Province: 1♂,
<specimenCount id="3FFB61982614FFA8FECFFB6C1F78FB55" box="[312,348,1159,1185]" pageId="11" pageNumber="412" type="female">4♀</specimenCount>
,
<collectorName id="8408CFC72614FFA8FE92FB6C1FF6FB55" box="[357,466,1159,1185]" pageId="11" pageNumber="412">Ankarana</collectorName>
[
<geoCoordinate id="4CC9CCD62614FFA8FE28FB6D1C19FB55" box="[479,573,1158,1185]" degrees="12.95" direction="south" orientation="latitude" pageId="11" pageNumber="412" precision="555" value="-12.95">12.95°S</geoCoordinate>
,
<geoCoordinate id="4CC9CCD62614FFA8FDB0FB6C1C8CFB55" box="[583,680,1159,1185]" degrees="49.14" direction="east" orientation="longitude" pageId="11" pageNumber="412" precision="555" value="49.14">49.14°E</geoCoordinate>
],
<collectorName id="8408CFC72614FFA8FD40FB6C1CCEFB55" box="[695,746,1159,1185]" pageId="11" pageNumber="412">Aug</collectorName>
,
<date id="5D438CD12614FFA8FD03FB6C1D7AFB55" box="[756,862,1159,1185]" pageId="11" pageNumber="412" value="1986-09">
<collectingDate id="4D0775392614FFA8FD03FB6C1D7AFB55" box="[756,862,1159,1185]" pageId="11" pageNumber="412" value="1986-09">Sep 1986</collectingDate>
</date>
, S.
<collectionCode id="4FEC32D42614FFA8FC8CFB6C1DB6FB55" box="[891,914,1159,1185]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="412" type="Museum">V</collectionCode>
.
<collectorName id="8408CFC72614FFA8FC6CFB6C1DCEFB55" box="[923,1002,1159,1185]" pageId="11" pageNumber="412">Fowler</collectorName>
(
<collectionCode id="4FEC32D42614FFA8FC0FFB6C1A76FB55" box="[1016,1106,1159,1185]" pageId="11" pageNumber="412">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="9995A04C2614FFA8FB94FB6C1B02FB55" ID-GBIF-Occurrence="4003029333" box="[1123,1318,1159,1185]" collectionCode="CAS" country="Madagascar" pageId="11" pageNumber="412" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
<typeStatus id="F64614B32614FFA8FB94FB6C1AEFFB55" box="[1123,1227,1159,1185]" pageId="11" pageNumber="412" type="holotype">Holotype</typeStatus>
in
<collectionCode id="4FEC32D42614FFA8FB1BFB6C1B02FB55" box="[1260,1318,1159,1185]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="11" pageNumber="412" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<materialsCitation id="9995A04C2614FFA8FAD8FB6C1F6BFB31" ID-GBIF-Occurrence="4003029372" collectionCode="BMNH, CAS" country="Madagascar" pageId="11" pageNumber="412" specimenCount="1" stateProvince="Toliara" typeStatus="paratype">
<typeStatus id="F64614B32614FFA8FAD8FB6C1BB8FB55" box="[1327,1436,1159,1185]" pageId="11" pageNumber="412" type="paratype">Paratypes</typeStatus>
in
<collectionCode id="4FEC32D42614FFA8FF43FB401F2FFB31" box="[180,267,1195,1221]" pageId="11" pageNumber="412">BMNH</collectionCode>
,
<collectionCode id="4FEC32D42614FFA8FEE1FB401F6BFB31" box="[278,335,1195,1221]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="11" pageNumber="412" type="Museum">CAS</collectionCode>
</materialsCitation>
, and BPBM.
</paragraph>
</subSubSection>
<subSubSection id="61E7F99A2614FFA8FF30FB251B52FAC5" pageId="11" pageNumber="412" type="diagnosis">
<paragraph id="2942AA112614FFA8FF30FB251B52FAC5" blockId="11.[151,1437,835,2019]" pageId="11" pageNumber="412">
<emphasis id="1B8976032614FFA8FF30FB251F66FB1C" bold="true" box="[199,322,1230,1256]" pageId="11" pageNumber="412">Diagnosis.</emphasis>
This species is easily distinguished from other species of the subgenus in
<collectingCountry id="51EAEA812614FFA8FB7FFB241B2BFB1D" box="[1160,1295,1231,1257]" name="Madagascar" pageId="11" pageNumber="412">Madagascar</collectingCountry>
by the short second flagellomere (one fifth length of the first flagellomere), a brown spot on halter knob, the brown scutellum with a yellow apex, and the deep apical invagination of the spermatheca (depth about half the reservoir height).
</paragraph>
</subSubSection>
<subSubSection id="61E7F99A2614FFAFFF30FAD11A9CFCCD" lastPageId="12" lastPageNumber="413" pageId="11" pageNumber="412" type="description">
<paragraph id="2942AA112614FFA8FF30FAD11F75FAA0" blockId="11.[151,1437,835,2019]" box="[199,337,1338,1364]" pageId="11" pageNumber="412">
<emphasis id="1B8976032614FFA8FF30FAD11F75FAA0" bold="true" box="[199,337,1338,1364]" pageId="11" pageNumber="412">Description</emphasis>
</paragraph>
<paragraph id="2942AA112614FFA8FF30FAB41A2DF981" blockId="11.[151,1437,835,2019]" pageId="11" pageNumber="412">
<emphasis id="1B8976032614FFA8FF30FAB41F20FA8D" bold="true" box="[199,260,1375,1401]" pageId="11" pageNumber="412">Male</emphasis>
(
<figureCitation id="B1C6B6942614FFA8FEE5FAB41F40FA8D" box="[274,356,1375,1401]" captionStart="FIGURES 510" captionStartId="4.[151,264,1768,1792]" captionTargetBox="[151,1435,181,1745]" captionTargetId="figure-17@4.[151,1435,181,1745]" captionTargetPageId="4" captionText="FIGURES 510. Malagasy Empidideicus (Anomaloptilus), habitus, lateral view. 5, E. (A.) andriambahomananii, n. sp., male; 6, E. (A.) andriambahomananii, n. sp., female; 7, E. (A.) contrasticus, n. sp., male; 8, E. (A.) hauseri, n. sp., male; 9, E. (A.) irwini, n. sp., male; 10, E. (A.) nanus, n. sp., male." figureDoi="http://doi.org/10.5281/zenodo.7471746" httpUri="https://zenodo.org/record/7471746/files/figure.png" pageId="11" pageNumber="412">Fig. 10</figureCitation>
). Length:
<quantity id="EE0507F42614FFA8FE23FAB51C5CFA8D" box="[468,632,1374,1401]" metricMagnitude="-3" metricUnit="m" metricValue="1.035" metricValueMax="1.12" metricValueMin="0.95" pageId="11" pageNumber="412" unit="mm" value="1.035" valueMax="1.12" valueMin="0.95">0.951.12 mm</quantity>
. Wing: 1.00
<quantity id="EE0507F42614FFA8FCFAFAB41D54FA8D" box="[781,880,1375,1401]" metricMagnitude="-3" metricUnit="m" metricValue="1.21" pageId="11" pageNumber="412" unit="mm" value="1.21">1.21 mm</quantity>
.
<emphasis id="1B8976032614FFA8FC8DFAB41D93FA8D" box="[890,951,1375,1401]" italics="true" pageId="11" pageNumber="412">Head</emphasis>
: longer than high; eyes dichoptic, separated at vertex by 1.7× distance between lateral ocelli; occiput and postgena black; front slightly depressed medially, white with subtriangular light brown area immediately below ocellar tubercle; face white; antennae (
<figureCitation id="B1C6B6942614FFA8FAFFFA4C1B7EFA35" box="[1288,1370,1447,1473]" captionStart="FIGURES 2324" captionStartId="10.[151,264,1309,1333]" captionTargetBox="[151,1435,181,1284]" captionTargetId="figure-17@10.[151,1435,181,1285]" captionTargetPageId="10" captionText="FIGURES 2324. Malagasy Empidideicus (Anomaloptilus), antennae. 23, E. (A.) andriambahomananii, n. sp.; 24, E. (A.) nanus, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471754" httpUri="https://zenodo.org/record/7471754/files/figure.png" pageId="11" pageNumber="412">Fig. 24</figureCitation>
) with scape and pedicel yellow, flagellomeres dark brown; scape short, subtrapezoidal; pedicel subconical, wider than long; first flagellomere lanceolate, length 2× width; second flagellomere one-fifth length of first flagellomere, with minute transparent apical style; mentum white; clypeus white, basally light brown, length subequal to labrum height; proboscis dorsally yellow ventrally brown, shorter than head length; labrum sclerotized, stiff, pointed apically, length subequal to clypeus, yellow basally, brown apically; palpus not evident.
</paragraph>
<paragraph id="2942AA112614FFA8FF30F9941CFDF8B9" blockId="11.[151,1437,835,2019]" pageId="11" pageNumber="412">
<emphasis id="1B8976032614FFA8FF30F9941F3CF96D" box="[199,280,1663,1689]" italics="true" pageId="11" pageNumber="412">Thorax</emphasis>
(
<figureCitation id="B1C6B6942614FFA8FED1F9941F5DF96D" box="[294,377,1663,1689]" captionStart="FIGURES 2533" captionStartId="13.[151,264,1878,1902]" captionTargetBox="[151,1435,273,1855]" captionTargetId="figure-65@13.[151,1435,273,1855]" captionTargetPageId="13" captionText="FIGURES 2533. Malagasy Empidideicus (Anomaloptilus), thoraces, dorsal view. 25, E. (A.) andriambahomananii, n. sp., male; 26, E. (A.) contrasticus, n. sp., male; 27, E. (A.) contrasticus, n. sp., female; 28, E. (A.) hauseri, n. sp., male; 29, E. (A.) hauseri, n. sp., female; 30, E. (A.) irwini, n. sp., male; 31, E. (A.) irwini, n. sp., female; 32, E. (A.) nanus, n. sp., male; 33, E. (A.) nanus, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471758" httpUri="https://zenodo.org/record/7471758/files/figure.png" pageId="11" pageNumber="412">Fig. 32</figureCitation>
). Mesonotum matte brown, with admedian yellow longitudinal vittae not reaching to scutellum, leaving two small yellow marks in prescutellar area, with two brown spots just behind transverse suture; yellow dorsolaterally from humeral callus to postalar callus and two interhumeral marks yellow; scutellum brown except posterior margin narrowly yellow; pleura yellow with dark brown on following: anteroventral corner of anepisternum and anepimeron, lower half of meron, and lower three-fourths of katepisternum; propleuron white; halter stem yellow, knob yellow with light brown spot dorsally.
</paragraph>
<paragraph id="2942AA112614FFA8FF30F8BC1F0BF861" blockId="11.[151,1437,835,2019]" pageId="11" pageNumber="412">
<emphasis id="1B8976032614FFA8FF30F8BC1ED5F884" box="[199,241,1879,1904]" italics="true" pageId="11" pageNumber="412">Leg</emphasis>
. Fore coxa yellow, mid and hind coxae brown; femora and tibia yellow; basitarsus yellow, remainder of tarsus brown.
</paragraph>
<paragraph id="2942AA112614FFAFFF30F8741C82FF0D" blockId="11.[151,1437,835,2019]" lastBlockId="12.[151,1437,151,861]" lastPageId="12" lastPageNumber="413" pageId="11" pageNumber="412">
<emphasis id="1B8976032614FFA8FF30F8741F25F84C" box="[199,257,1951,1976]" italics="true" pageId="11" pageNumber="412">Wing</emphasis>
(
<figureCitation id="B1C6B6942614FFA8FEF8F8741F47F84D" box="[271,355,1951,1977]" captionStart="FIGURES 4047" captionStartId="15.[151,264,1607,1631]" captionTargetBox="[170,1416,309,1582]" captionTargetId="figure-81@15.[170,1416,309,1584]" captionTargetPageId="15" captionText="FIGURES 4047. Malagasy Empidideicus (Anomaloptilus), wings. 40, E. (A.) andriambahomananii, n. sp.; 41, E. (A.) contrasticus, n. sp.; 42, E. (A.) hauseri, n. sp.; 43, E. (A.) irwini, n. sp.; 44, E. (A.) nanus, n. sp.; 45, E. (A.) regius, n. sp.; 46, E. (A.) rhinoclypeatus Evenhuis; 47, E. (A.) rinhai, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471764" httpUri="https://zenodo.org/record/7471764/files/figure.png" pageId="11" pageNumber="412">Fig. 44</figureCitation>
). Elongate, length 2.7× width; hyaline; veins brown, well sclerotized except vein separating basal cells and A
<subScript id="B579A8542614FFA8FEE3F83E1F38F817" attach="left" box="[276,284,2005,2019]" fontSize="6" pageId="11" pageNumber="412">1</subScript>
; costa ends slightly beyond end of R
<subScript id="B579A8542614FFA8FD4CF83E1CF1F817" attach="left" box="[699,725,2005,2019]" fontSize="6" pageId="11" pageNumber="412">4+5</subScript>
; vein Sc incomplete; Rs evanescent at connection with R
<subScript id="B579A8542614FFA8FAAEF83E1B45F817" attach="left" box="[1369,1377,2005,2019]" fontSize="6" pageId="11" pageNumber="412">1</subScript>
; R
<subScript id="B579A8542614FFA8FA74F83E1BB9F817" attach="left" box="[1411,1437,2005,2019]" fontSize="6" pageId="11" pageNumber="412">4+5</subScript>
slightly bowed; vein M
<subScript id="B579A8542613FFAFFE6EFF421F85FF43" attach="left" box="[409,417,169,183]" fontSize="6" pageId="12" pageNumber="413">1</subScript>
curved; M
<subScript id="B579A8542613FFAFFDEDFF421C06FF43" attach="left" box="[538,546,169,183]" fontSize="6" pageId="12" pageNumber="413">2</subScript>
straight; cell dm closed by apical crossvein; A
<subScript id="B579A8542613FFAFFBD2FF421A09FF43" attach="left" box="[1061,1069,169,183]" fontSize="6" pageId="12" pageNumber="413">1</subScript>
effaced, straight to wing margin; anal cell open very slightly at wing margin; fringe of hair on posterior margin of wing well shortest at wing tip becoming longer and more sparse toward base.
</paragraph>
<paragraph id="2942AA112613FFAFFF30FEE81FFCFE91" blockId="12.[151,1437,151,861]" pageId="12" pageNumber="413">
<emphasis id="1B8976032613FFAFFF30FEE81F15FEE9" box="[199,305,259,285]" italics="true" pageId="12" pageNumber="413">Abdomen</emphasis>
(
<figureCitation id="B1C6B6942613FFAFFEB6FEE81FB2FEE9" box="[321,406,258,285]" captionStart="FIGURES 5058" captionStartId="18.[151,264,1889,1913]" captionTargetBox="[239,1348,181,1865]" captionTargetId="figure-17@18.[239,1348,181,1865]" captionTargetPageId="18" captionText="FIGURES 5058. Malagasy Empidideicus (Anomaloptilus), abdomens, dorsal view. 50, E. (A.) andriambahomananii, n. sp., female; 51, E. (A.) contrasticus, n. sp., male; 52, E. (A.) contrasticus, n. sp., female; 53, E. (A.) hauseri, n. sp., male; 54, E. (A.) hauseri, n. sp., female; 55, E. (A.) irwini, n. sp., male; 56, E. (A.) irwini, n. sp., female; 57, E. (A.) nanus, n. sp., male; 58, E. (A.) regius, n. sp., male." figureDoi="http://doi.org/10.5281/zenodo.7471772" httpUri="https://zenodo.org/record/7471772/files/figure.png" pageId="12" pageNumber="413">Fig. 57</figureCitation>
). Tergites dark brown, posterior margins narrowly yellow, brown mark laterally interrupted with wide yellow band; last tergite with two long dark brown bands; tergites with sparsely scattered minute yellowish white hairs; sternites yellow.
</paragraph>
<paragraph id="2942AA112613FFAFFF30FE841C44FD95" blockId="12.[151,1437,151,861]" pageId="12" pageNumber="413">
<emphasis id="1B8976032613FFAFFF30FE841F15FE7D" box="[199,305,367,393]" italics="true" pageId="12" pageNumber="413">Genitalia</emphasis>
(
<figureCitation id="B1C6B6942613FFAFFEB4FE841FBCFE7D" box="[323,408,367,393]" captionStart="FIGURES 7172" captionStartId="21.[152,270,1956,1980]" captionTargetBox="[152,1436,1100,1933]" captionTargetId="figure-168@21.[151,1436,1100,1933]" captionTargetPageId="21" captionText="FIGURES 7172. Empidideicus (Anomaloptilus) nanus, n. sp., genitalia. 71, male, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view; 72, female." figureDoi="http://doi.org/10.5281/zenodo.7471790" httpUri="https://zenodo.org/record/7471790/files/figure.png" pageId="12" pageNumber="413">Fig. 71</figureCitation>
). Epandrium in lateral view subquadrate, with long, narrow apicoventral process, slightly clubbed and darkly sclerotized apically (
<figureCitation id="B1C6B6942613FFAFFDA4FE781C9EFE59" box="[595,698,403,429]" captionStart="FIGURES 7172" captionStartId="21.[152,270,1956,1980]" captionTargetBox="[152,1436,1100,1933]" captionTargetId="figure-168@21.[151,1436,1100,1933]" captionTargetPageId="21" captionText="FIGURES 7172. Empidideicus (Anomaloptilus) nanus, n. sp., genitalia. 71, male, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view; 72, female." figureDoi="http://doi.org/10.5281/zenodo.7471790" httpUri="https://zenodo.org/record/7471790/files/figure.png" pageId="12" pageNumber="413">Fig. 71A</figureCitation>
); cerci small, not exerted; gonocoxa in lateral view linear-ellipsoid, with small circular gonostylus, in ventral view with relatively short medial cleft, basally with sclerotized band (
<figureCitation id="B1C6B6942613FFAFFA98FE5C1EEDFE01" captionStart="FIGURES 7172" captionStartId="21.[152,270,1956,1980]" captionTargetBox="[152,1436,1100,1933]" captionTargetId="figure-168@21.[151,1436,1100,1933]" captionTargetPageId="21" captionText="FIGURES 7172. Empidideicus (Anomaloptilus) nanus, n. sp., genitalia. 71, male, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view; 72, female." figureDoi="http://doi.org/10.5281/zenodo.7471790" httpUri="https://zenodo.org/record/7471790/files/figure.png" pageId="12" pageNumber="413">Fig. 71B</figureCitation>
); lateral arms of parameral sheath extending well beyond gonocoxa, longer than aedeagal bulb, flared apically; apical portion of parameral sheath cylindrical with sharp bulge subapically; aedeagal bulge spheroid, half length of lateral parameral arms (
<figureCitation id="B1C6B6942613FFAFFE39FDC81C1DFDC9" box="[462,569,547,573]" captionStart="FIGURES 7172" captionStartId="21.[152,270,1956,1980]" captionTargetBox="[152,1436,1100,1933]" captionTargetId="figure-168@21.[151,1436,1100,1933]" captionTargetPageId="21" captionText="FIGURES 7172. Empidideicus (Anomaloptilus) nanus, n. sp., genitalia. 71, male, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view; 72, female." figureDoi="http://doi.org/10.5281/zenodo.7471790" httpUri="https://zenodo.org/record/7471790/files/figure.png" pageId="12" pageNumber="413">Fig. 71C</figureCitation>
); aedeagal apodeme in lateral view peanut-shaped, truncate ventrally, with relatively small downturned lateral rami.
</paragraph>
<paragraph id="2942AA112613FFAFFF30FD801A9CFCCD" blockId="12.[151,1437,151,861]" pageId="12" pageNumber="413">
<emphasis id="1B8976032613FFAFFF30FD801F3AFD71" bold="true" box="[199,286,619,645]" pageId="12" pageNumber="413">Female</emphasis>
. Similar to male; the mesonotal vittae (
<figureCitation id="B1C6B6942613FFAFFD2FFD801D0DFD71" box="[728,809,619,645]" captionStart="FIGURES 2533" captionStartId="13.[151,264,1878,1902]" captionTargetBox="[151,1435,273,1855]" captionTargetId="figure-65@13.[151,1435,273,1855]" captionTargetPageId="13" captionText="FIGURES 2533. Malagasy Empidideicus (Anomaloptilus), thoraces, dorsal view. 25, E. (A.) andriambahomananii, n. sp., male; 26, E. (A.) contrasticus, n. sp., male; 27, E. (A.) contrasticus, n. sp., female; 28, E. (A.) hauseri, n. sp., male; 29, E. (A.) hauseri, n. sp., female; 30, E. (A.) irwini, n. sp., male; 31, E. (A.) irwini, n. sp., female; 32, E. (A.) nanus, n. sp., male; 33, E. (A.) nanus, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471758" httpUri="https://zenodo.org/record/7471758/files/figure.png" pageId="12" pageNumber="413">Fig. 33</figureCitation>
) are more evident than in male. Genitalia (
<figureCitation id="B1C6B6942613FFAFFAFDFD801B7FFD71" box="[1290,1371,619,645]" captionStart="FIGURES 7172" captionStartId="21.[152,270,1956,1980]" captionTargetBox="[152,1436,1100,1933]" captionTargetId="figure-168@21.[151,1436,1100,1933]" captionTargetPageId="21" captionText="FIGURES 7172. Empidideicus (Anomaloptilus) nanus, n. sp., genitalia. 71, male, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view; 72, female." figureDoi="http://doi.org/10.5281/zenodo.7471790" httpUri="https://zenodo.org/record/7471790/files/figure.png" pageId="12" pageNumber="413">Fig. 72</figureCitation>
) with genital fork U-shaped with two inwardly directed processes on lateral arms, lateral arms much more sclerotized than medial portion; spermatheca spheroid, sclerotized brown, with striations externally, apical invagination deep, about half length of reservoir height; apical spermathecal duct ca. 6.5× length of spermatheca, thin, transparent, basally (0.1× its length) slightly sclerotized, brown; sperm pump 0.8× length of apical duct, transparent, not sclerotized apically or basally; basal duct confluent with sperm pump, common spermathecal duct absent.
</paragraph>
</subSubSection>
<subSubSection id="61E7F99A2613FFAFFF30FCA81B28FCA9" box="[199,1292,835,861]" pageId="12" pageNumber="413" type="etymology">
<paragraph id="2942AA112613FFAFFF30FCA81B28FCA9" blockId="12.[151,1437,151,861]" box="[199,1292,835,861]" pageId="12" pageNumber="413">
<emphasis id="1B8976032613FFAFFF30FCA81F6CFCA9" bold="true" box="[199,328,835,861]" pageId="12" pageNumber="413">Etymology</emphasis>
. The specific epithet derives from the Latin
<taxonomicName id="EEFDD1922613FFAFFCCDFCA81D59FCA8" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[826,893,835,860]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="1B8976032613FFAFFCCDFCA81D59FCA8" box="[826,893,835,860]" italics="true" pageId="12" pageNumber="413">nanus</emphasis>
</taxonomicName>
= small; referring to the small size.
</paragraph>
</subSubSection>
</treatment>
</document>