455 lines
57 KiB
XML
455 lines
57 KiB
XML
<document id="E6955AC1C069812B3BA585DABF0C7C41" ID-CLB-Dataset="54548" ID-DOI="10.11646/zootaxa.5222.5.1" ID-GBIF-Dataset="a2aef7c8-09e1-48f1-983e-00132055a5a2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7471736" ID-ZooBank="F4983B58-7301-4A3F-AC08-3E5120450EA1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1671694447742" checkinUser="plazi" docAuthor="Evenhuis, Neal L. & Gharali, Babak" docDate="2022" docId="A1541B072610FFB2FF60F9121ED2FCA9" docLanguage="en" docName="zootaxa.5222.5.1.pdf" docOrigin="Zootaxa 5222 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Empidideicus (Anomaloptilus) rhinoclypeatus Evenhuis" docType="treatment" docVersion="7" lastPageNumber="418" masterDocId="5D6D637F261FFFA3FFF7FFEB1E24FFF4" masterDocTitle="The subgenus Empidideicus (Anomaloptilus) (Diptera: Mythicomyiidae) in Madagascar" masterLastPageNumber="426" masterPageNumber="401" pageNumber="416" updateTime="1699413895303" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="A286C44E959CB5957DAB2B76B915F038">
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<mods:title id="C9AF4EA193BF01375F757585E691E3C3">The subgenus Empidideicus (Anomaloptilus) (Diptera: Mythicomyiidae) in Madagascar</mods:title>
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<mods:name id="8D5DE614D3FA3B69548D80D8C9BB060E" type="personal">
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<mods:namePart id="9C925AA1C2BC316DB8FE15E73CD7B901">Evenhuis, Neal L.</mods:namePart>
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<mods:affiliation id="9B46951DE9303732CBAB8D83E1C99448">J. Linsley Gressitt Center for Research in Entomology, Bishop Museum, 1525 Bernice Street, Honolulu, Hawaii 96817 - 2704, USA neale @ bishopmuseum. org; https: // orcid. org / 0000 - 0002 - 1314 - 755 X</mods:affiliation>
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<mods:nameIdentifier id="C44D1E057A567E704BE64C02FF6E48A5" type="email">neale@bishopmuseum.org</mods:nameIdentifier>
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<mods:namePart id="1F6CA67667780D7D0DB3641130587490">Gharali, Babak</mods:namePart>
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<mods:affiliation id="31F642856FB3726C9A817FD16B8DBA00">Plant Protection Research Department, Qazvin Agricultural and Natural Resources Research and Education Center, AREEO, Qazvin, Iran. bgaharaei @ yahoo. com; https: // orcid. org / 0000 - 0003 - 2490 - 156 X</mods:affiliation>
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<mods:nameIdentifier id="2FA25C2F82BD303E16FA5669E9DB2759" type="email">bgaharaei@yahoo.com</mods:nameIdentifier>
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<mods:title id="A1D96352AC302F43E987D12BC60F7EF9">Zootaxa</mods:title>
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<mods:date id="52DD275CF2E8C370658854A8B858DD4E">2022</mods:date>
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<treatment id="A1541B072610FFB2FF60F9121ED2FCA9" ID-DOI="http://doi.org/10.5281/zenodo.7473306" ID-GBIF-Taxon="207927032" ID-Zenodo-Dep="7473306" LSID="urn:lsid:plazi:treatment:A1541B072610FFB2FF60F9121ED2FCA9" httpUri="http://treatment.plazi.org/id/A1541B072610FFB2FF60F9121ED2FCA9" lastPageId="17" lastPageNumber="418" pageId="15" pageNumber="416">
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<heading id="720A1D7D2610FFACFF60F9121D1AF8E0" bold="true" box="[151,830,1785,1812]" fontSize="11" level="1" pageId="15" pageNumber="416" reason="1">
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<taxonomicName id="EEFDD1922610FFACFF60F9121D1AF8E0" authority="Evenhuis" authorityName="Evenhuis" box="[151,830,1785,1812]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="rhinoclypeatus" subGenus="Anomaloptilus">
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<emphasis id="1B8976032610FFACFF60F9121D1AF8E0" bold="true" box="[151,830,1785,1812]" pageId="15" pageNumber="416">
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<emphasis id="1B8976032610FFACFF60F9121CE7F8E0" bold="true" box="[151,707,1785,1812]" italics="true" pageId="15" pageNumber="416">Empidideicus (Anomaloptilus) rhinoclypeatus</emphasis>
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Evenhuis
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<paragraph id="2942AA112610FFACFF60F8F51CB7F8CC" blockId="15.[151,830,1785,1849]" box="[151,659,1822,1849]" pageId="15" pageNumber="416">
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(
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<figureCitation id="B1C6B6942610FFACFF68F8F51ECCF8CC" box="[159,232,1822,1848]" captionStart="FIGURES 3–4" captionStartId="2.[151,264,1920,1944]" captionTargetBox="[157,1423,240,1880]" captionTargetId="figure-140@2.[151,1436,227,1896]" captionTargetPageId="2" captionText="FIGURES 3–4. Schematic illustration showing differences in clypeus lengths of Empidideicus (Anomaloptilus) species. 3, E. (A.) celluliferus Hesse; 4, E. (A.) rhinoclypeatus Evenhuis. Arrows point to clypeus. (Redrawn from Evenhuis 2007; figs 2–3)." figureDoi="http://doi.org/10.5281/zenodo.7471742" httpUri="https://zenodo.org/record/7471742/files/figure.png" pageId="15" pageNumber="416">Figs 4</figureCitation>
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,
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<figureCitation id="B1C6B6942610FFACFF03F8F51F18F8CC" box="[244,316,1822,1848]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[158,1429,181,1920]" captionTargetId="figure-17@5.[158,1429,181,1920]" captionTargetPageId="5" captionText="FIGURES 11–16. Malagasy Empidideicus (Anomaloptilus), habitus, lateral view. 11, E. (A.) regius, n. sp., male; 12, E. (A.) regius, n. sp., female; 13, E. (A.) rhinoclypeatus Evenhuis, male; 14, E. (A.) rhinoclypeatus Evenhuis, female; 15, E. (A.) rinhai, n. sp., male; 16, E. (A.) rinhai, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471748" httpUri="https://zenodo.org/record/7471748/files/figure.png" pageId="15" pageNumber="416">13, 14</figureCitation>
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,
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<figureCitation id="B1C6B6942610FFACFEBFF8F51F43F8CC" box="[328,359,1822,1848]" captionStart="FIGURES 20–22" captionStartId="8.[151,264,1965,1989]" captionTargetBox="[151,1363,1475,1942]" captionTargetId="figure-302@8.[151,1435,1475,1942]" captionTargetPageId="8" captionText="FIGURES 20–22. Malagasy Empidideicus (Anomaloptilus), heads. 20, E. (A.) contrasticus, n. sp., male, frontal view; 21, E. (A.) irwini, n. sp., male, oblique view; 22, E. (A.) rhinoclypeatus Evenhuis, male, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7471752" httpUri="https://zenodo.org/record/7471752/files/figure.png" pageId="15" pageNumber="416">22</figureCitation>
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,
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<figureCitation id="B1C6B6942610FFACFE85F8F51F9EF8CC" box="[370,442,1822,1848]" captionStart="FIGURES 34–39" captionStartId="14.[151,264,1264,1288]" captionTargetBox="[151,1435,181,1240]" captionTargetId="figure-17@14.[151,1435,181,1240]" captionTargetPageId="14" captionText="FIGURES 34–39. Malagasy Empidideicus (Anomaloptilus), thoraces, dorsal view. 34, E. (A.) regius, n. sp., male; 35, E. (A.) regius, n. sp., female; 36, E. (A.) rhinoclypeatus Evenhuis, male; 37, E. (A.) rhinoclypeatus Evenhuis, female; 38, E. (A.) rinhai, n. sp., male; 39, E. (A.) rinhai, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471760" httpUri="https://zenodo.org/record/7471760/files/figure.png" pageId="15" pageNumber="416">36, 37</figureCitation>
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,
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<figureCitation id="B1C6B6942610FFACFE31F8F51FC1F8CC" box="[454,485,1822,1848]" captionStart="FIGURES 40–47" captionStartId="15.[151,264,1607,1631]" captionTargetBox="[170,1416,309,1582]" captionTargetId="figure-81@15.[170,1416,309,1584]" captionTargetPageId="15" captionText="FIGURES 40–47. Malagasy Empidideicus (Anomaloptilus), wings. 40, E. (A.) andriambahomananii, n. sp.; 41, E. (A.) contrasticus, n. sp.; 42, E. (A.) hauseri, n. sp.; 43, E. (A.) irwini, n. sp.; 44, E. (A.) nanus, n. sp.; 45, E. (A.) regius, n. sp.; 46, E. (A.) rhinoclypeatus Evenhuis; 47, E. (A.) rinhai, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471764" httpUri="https://zenodo.org/record/7471764/files/figure.png" pageId="15" pageNumber="416">46</figureCitation>
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,
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<figureCitation id="B1C6B6942610FFACFE07F8F51C1CF8CC" box="[496,568,1822,1849]" captionStart="FIGURES 59–62" captionStartId="19.[151,264,694,718]" captionTargetBox="[152,1435,181,670]" captionTargetId="figure-17@19.[152,1435,181,670]" captionTargetPageId="19" captionText="FIGURES 59–62. Malagasy Empidideicus (Anomaloptilus), abdomens, dorsal view. 59, E. (A.) rhinoclypeatus Evenhuis, male; 60, E. (A.) rhinoclypeatus Evenhuis, female; 61, E. (A.) rinhai, n. sp., male; 62, E. (A.) rinhai, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471774" httpUri="https://zenodo.org/record/7471774/files/figure.png" pageId="15" pageNumber="416">59, 60</figureCitation>
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,
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<figureCitation id="B1C6B6942610FFACFDB3F8F51CAFF8CC" box="[580,651,1822,1849]" captionStart="FIGURES 75–76" captionStartId="22.[152,270,1430,1454]" captionTargetBox="[155,1432,838,1400]" captionTargetId="figure-57@22.[151,1436,832,1406]" captionTargetPageId="22" captionText="FIGURES 75–76. Empidideicus (Anomaloptilus) rhinoclypeatus Evenhuis, genitalia. 75, male, A, lateral view, B, phallic complex and gonocoxae, dorsal view; 76, female. Abbreviation: gcx = gonocoxa." figureDoi="http://doi.org/10.5281/zenodo.7471796" httpUri="https://zenodo.org/record/7471796/files/figure.png" pageId="15" pageNumber="416">75, 76</figureCitation>
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)
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</paragraph>
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<taxonomicName id="EEFDD1922610FFACFF60F8891CAFF88E" authority="Evenhuis, 2007: 56" authorityName="Evenhuis" authorityPageNumber="56" authorityYear="2007" box="[151,651,1890,1914]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="rhinoclypeatus">
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<emphasis id="1B8976032610FFACFF60F8891F99F88E" box="[151,445,1890,1914]" italics="true" pageId="15" pageNumber="416">Empidideicus rhinoclypeatus</emphasis>
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<bibRefCitation id="4D6CD7E02610FFACFE33F8891CAFF88E" author="Evenhuis, N. L." box="[452,651,1890,1914]" pageId="15" pageNumber="416" pagination="55 - 62" refId="ref15043" refString="Evenhuis, N. L. (2007) A remarkable new species of Empidideicus (Diptera: Mythicomyiidae) from Madagascar. Zootaxa, 1474 (1), 55 - 62. https: // doi. org / 10.11646 / zootaxa. 1474.1.2" type="journal article" year="2007">Evenhuis, 2007: 56</bibRefCitation>
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.
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</paragraph>
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<subSubSection id="61E7F99A2610FFB3FF60F8481D0BFD95" lastPageId="16" lastPageNumber="417" pageId="15" pageNumber="416" type="materials_examined">
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<paragraph id="2942AA112610FFB3FF60F8481D0BFD95" blockId="15.[151,1437,1955,2020]" lastBlockId="16.[151,1437,151,609]" lastPageId="16" lastPageNumber="417" pageId="15" pageNumber="416">
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<emphasis id="1B8976032610FFACFF60F8481FF7F849" bold="true" box="[151,467,1955,1981]" pageId="15" pageNumber="416">Material Examined. Types</emphasis>
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.
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<materialsCitation id="9995A04C2610FFACFE14F8481BB8F815" ID-GBIF-Occurrence="4003029335" collectingDate="2003-02-05" collectingDateMax="2003-02-15" collectingDateMin="2003-02-05" collectorName="M. Irwin" country="Madagascar" county="Tsishombe" elevation="37" latitude="-25.587667" location="Tulear Province" longLatPrecision="12" longitude="45.163" municipality="Cape Ste. Marie Special Reserve" pageId="15" pageNumber="416" specimenCount="23" specimenCount-female="22" typeStatus="holotype">
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<emphasis id="1B8976032610FFACFE14F8481C6EF849" box="[483,586,1955,1981]" italics="true" pageId="15" pageNumber="416">
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<typeStatus id="F64614B32610FFACFE14F8481C6EF849" box="[483,586,1955,1981]" pageId="15" pageNumber="416" type="holotype">Holotype</typeStatus>
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</emphasis>
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♂ and 43♂,
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<specimenCount id="3FFB61982610FFACFD2AF8481D2AF849" box="[733,782,1955,1981]" pageId="15" pageNumber="416" type="female">22♀</specimenCount>
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<emphasis id="1B8976032610FFACFCE1F84F1DA0F849" box="[790,900,1956,1981]" italics="true" pageId="15" pageNumber="416">
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<typeStatus id="F64614B32610FFACFCE1F84F1DA0F849" box="[790,900,1956,1981]" pageId="15" pageNumber="416" type="paratype">paratypes</typeStatus>
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</emphasis>
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from
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<collectingCountry id="51EAEA812610FFACFC3DF8481AB2F849" box="[970,1174,1955,1981]" name="Madagascar" pageId="15" pageNumber="416">MADAGASCAR</collectingCountry>
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:
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<location id="2C22FCCA2610FFACFB57F8481B7CF849" LSID="urn:lsid:plazi:treatment:A1541B072610FFB2FF60F9121ED2FCA9:2C22FCCA2610FFACFB57F8481B7CF849" box="[1184,1368,1955,1981]" country="Madagascar" county="Tsishombe" latitude="-25.587667" longLatPrecision="12" longitude="45.163" municipality="Cape Ste. Marie Special Reserve" name="Tulear Province" pageId="15" pageNumber="416">Tulear Province</location>
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:
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<collectingMunicipality id="C926306B2610FFACFA94F8481FF0F816" pageId="15" pageNumber="416">Cape Ste. Marie Special Reserve</collectingMunicipality>
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,
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<elevation id="A2D04D222610FFACFE15F82C1C05F816" box="[482,545,1991,2018]" metricMagnitude="1" metricUnit="m" metricValue="3.7" pageId="15" pageNumber="416" unit="m" value="37.0">37 m</elevation>
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</quantity>
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,
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<quantity id="EE0507F42610FFACFDC7F82C1C5FF815" box="[560,635,1991,2017]" metricMagnitude="4" metricUnit="m" metricValue="7.4" pageId="15" pageNumber="416" unit="km" value="74.0">74 km</quantity>
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S. of
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<collectingCounty id="C023D29D2610FFACFD30F82C1D67F815" box="[711,835,1991,2017]" pageId="15" pageNumber="416">Tsishombe</collectingCounty>
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,
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,
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,
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|
||
<collectingDate id="4D0775392610FFACFB99F82C1B04F815" box="[1134,1312,1991,2018]" pageId="15" pageNumber="416" value="2003-02-05" valueMax="2003-02-15" valueMin="2003-02-05">5–15 Feb 2003</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="8408CFC72610FFACFAD9F8231BBCF815" box="[1326,1432,1991,2018]" pageId="15" pageNumber="416">M. Irwin</collectorName>
|
||
,
|
||
</materialsCitation>
|
||
<materialsCitation id="9995A04C260FFFB3FF60FF7C1DD1FF21" ID-GBIF-Occurrence="4003029375" collectingDate="2002-04-18" collectingDateMax="2002-04-29" collectingDateMin="2002-04-18" collectionCode="R" collectorName="M. E. Irwin" country="Madagascar" latitude="-22.913334" location="Tulear Province" longLatPrecision="92" longitude="43.482166" municipality="Mikea Forest" pageId="16" pageNumber="417" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
|
||
<collectorName id="8408CFC7260FFFB3FF60FF7C1F24FF45" box="[151,256,151,177]" pageId="16" pageNumber="417">F. Parker</collectorName>
|
||
,
|
||
<collectingMethod id="F0BCD206260FFFB3FEF9FF7C1FBAFF45" box="[270,414,151,177]" pageId="16" pageNumber="417">Malaise trap</collectingMethod>
|
||
, spiny bush.
|
||
<emphasis id="1B897603260FFFB3FDCFFF7C1CD4FF44" box="[568,752,151,177]" italics="true" pageId="16" pageNumber="417">
|
||
<collectorName id="8408CFC7260FFFB3FDCFFF7C1C5DFF45" box="[568,633,151,177]" pageId="16" pageNumber="417">Other</collectorName>
|
||
<typeStatus id="F64614B3260FFFB3FD75FF7C1CD4FF44" box="[642,752,151,176]" pageId="16" pageNumber="417" type="paratype">paratypes</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<collectingCountry id="51EAEA81260FFFB3FCF5FF7C1DEAFF45" box="[770,974,151,177]" name="Madagascar" pageId="16" pageNumber="417">MADAGASCAR</collectingCountry>
|
||
:
|
||
<location id="2C22FCCA260FFFB3FC2DFF7C1AB0FF45" LSID="urn:lsid:plazi:treatment:A1541B072610FFB2FF60F9121ED2FCA9:2C22FCCA260FFFB3FC2DFF7C1AB0FF45" box="[986,1172,151,177]" country="Madagascar" latitude="-22.913334" longLatPrecision="92" longitude="43.482166" municipality="Mikea Forest" name="Tulear Province" pageId="16" pageNumber="417">Tulear Province</location>
|
||
: 2♂,
|
||
<specimenCount id="3FFB6198260FFFB3FB3BFF731ACAFF47" box="[1228,1262,152,179]" pageId="16" pageNumber="417" type="female">3♀</specimenCount>
|
||
,
|
||
<collectingMunicipality id="C926306B260FFFB3FB09FF7C1BBCFF45" box="[1278,1432,151,177]" pageId="16" pageNumber="417">Mikea Forest</collectingMunicipality>
|
||
,
|
||
<locationDeviation id="38F0B333260FFFB3FF60FF501F40FF21" box="[151,356,187,213]" location="Manombo" pageId="16" pageNumber="417">NW of Manombo</locationDeviation>
|
||
,
|
||
<geoCoordinate id="4CC9CCD6260FFFB3FE87FF571FC4FF23" box="[368,480,188,215]" degrees="22" direction="south" minutes="54.8" orientation="latitude" pageId="16" pageNumber="417" precision="92" value="-22.913334">22°54.8′S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="4CC9CCD6260FFFB3FE19FF571C4AFF23" box="[494,622,188,215]" degrees="43" direction="east" minutes="28.93" orientation="longitude" pageId="16" pageNumber="417" precision="9" value="43.482166">43°28.93′E</geoCoordinate>
|
||
,
|
||
<date id="5D438CD1260FFFB3FD8DFF501D62FF21" box="[634,838,187,213]" pageId="16" pageNumber="417" value="2002-04-18" valueMax="2002-04-29" valueMin="2002-04-18">
|
||
<collectingDate id="4D077539260FFFB3FD8DFF501D62FF21" box="[634,838,187,213]" pageId="16" pageNumber="417" value="2002-04-18" valueMax="2002-04-29" valueMin="2002-04-18">18–29 April 2002</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="8408CFC7260FFFB3FCA5FF501DF6FF21" box="[850,978,187,213]" pageId="16" pageNumber="417">M.E. Irwin</collectorName>
|
||
,
|
||
<collectionCode id="4FEC32D4260FFFB3FC28FF501DD1FF21" box="[991,1013,187,213]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="417">R</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="9995A04C260FFFB3FBF6FF501B2DFF0D" ID-GBIF-Occurrence="4003029324" collectingDate="2002-12-16" collectingDateMax="2002-12-17" collectingDateMin="2002-12-16" collectionCode="R" collectorName="M. E. Irwin & F. D. Parker" country="Madagascar" county="Parcel" location="Harin'Hala" municipality="Andohahela National Park" pageId="16" pageNumber="417" specimenCount="1" typeStatus="paratype">
|
||
<location id="2C22FCCA260FFFB3FBF6FF501AA6FF21" LSID="urn:lsid:plazi:treatment:A1541B072610FFB2FF60F9121ED2FCA9:2C22FCCA260FFFB3FBF6FF501AA6FF21" box="[1025,1154,187,213]" country="Madagascar" county="Parcel" municipality="Andohahela National Park" name="Harin'Hala" pageId="16" pageNumber="417">Harin’Hala</location>
|
||
,
|
||
<collectingMethod id="F0BCD206260FFFB3FB78FF501B3AFF21" box="[1167,1310,187,213]" pageId="16" pageNumber="417">Malaise trap</collectingMethod>
|
||
, deciduous dry forest; 3♂,
|
||
<collectingMunicipality id="C926306B260FFFB3FEBCFF341C59FF0D" box="[331,637,223,249]" pageId="16" pageNumber="417">Andohahela National Park</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="C023D29D260FFFB3FD7BFF341CF6FF0D" box="[652,722,223,249]" pageId="16" pageNumber="417">Parcel</collectingCounty>
|
||
II,
|
||
<date id="5D438CD1260FFFB3FCF7FF341DE6FF0D" box="[768,962,223,249]" pageId="16" pageNumber="417" value="2002-12-16" valueMax="2002-12-17" valueMin="2002-12-16">
|
||
<collectingDate id="4D077539260FFFB3FCF7FF341DE6FF0D" box="[768,962,223,249]" pageId="16" pageNumber="417" value="2002-12-16" valueMax="2002-12-17" valueMin="2002-12-16">16–17 Dec 2002</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="8408CFC7260FFFB3FC27FF341A76FF0D" box="[976,1106,223,249]" pageId="16" pageNumber="417">M.E. Irwin</collectorName>
|
||
,
|
||
<collectorName id="8408CFC7260FFFB3FB97FF341AC1FF0D" box="[1120,1253,223,249]" pageId="16" pageNumber="417">F.D. Parker</collectorName>
|
||
,
|
||
<collectionCode id="4FEC32D4260FFFB3FB04FF341B2DFF0D" box="[1267,1289,223,249]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="417">R</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="9995A04C260FFFB3FAE0FF341C1DFEB5" ID-GBIF-Occurrence="4003029371" collectingDate="2003-10-18" collectingDateMax="2003-10-26" collectingDateMin="2003-10-18" collectionCode="R" collectorName="Harin'Hala & M. Irwin" country="Madagascar" location="Tulear Province" pageId="16" pageNumber="417" specimenCount="1">
|
||
<collectorName id="8408CFC7260FFFB3FAE0FF341BBCFF0D" box="[1303,1432,223,249]" pageId="16" pageNumber="417">Harin’Hala</collectorName>
|
||
,
|
||
<collectingMethod id="F0BCD206260FFFB3FF60FEE81F0DFEE9" box="[151,297,259,285]" pageId="16" pageNumber="417">Malaise trap</collectingMethod>
|
||
, transitional forest.
|
||
<emphasis id="1B897603260FFFB3FDE4FEE91D6CFEE8" bold="true" box="[531,840,258,284]" pageId="16" pageNumber="417">Other material examined</emphasis>
|
||
.
|
||
<collectingCountry id="51EAEA81260FFFB3FCACFEE81A03FEE9" box="[859,1063,259,285]" name="Madagascar" pageId="16" pageNumber="417">MADAGASCAR</collectingCountry>
|
||
:
|
||
<location id="2C22FCCA260FFFB3FBC3FEE81ACBFEE9" LSID="urn:lsid:plazi:treatment:A1541B072610FFB2FF60F9121ED2FCA9:2C22FCCA260FFFB3FBC3FEE81ACBFEE9" box="[1076,1263,259,285]" country="Madagascar" name="Tulear Province" pageId="16" pageNumber="417">Tulear Province</location>
|
||
: 4♂, topotypic except,
|
||
<date id="5D438CD1260FFFB3FF19FECC1F82FEB5" box="[238,422,295,321]" pageId="16" pageNumber="417" value="2003-10-18" valueMax="2003-10-26" valueMin="2003-10-18">
|
||
<collectingDate id="4D077539260FFFB3FF19FECC1F82FEB5" box="[238,422,295,321]" pageId="16" pageNumber="417" value="2003-10-18" valueMax="2003-10-26" valueMin="2003-10-18">18–26 Oct 2003</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="8408CFC7260FFFB3FE46FECC1C33FEB5" box="[433,535,295,321]" pageId="16" pageNumber="417">M. Irwin</collectorName>
|
||
,
|
||
<collectionCode id="4FEC32D4260FFFB3FDD4FECC1C1DFEB5" box="[547,569,295,321]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="417">R</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="9995A04C260FFFB3FDB3FECC1B58FE7D" ID-GBIF-Occurrence="4003029369" collectingDate="2002-10-27" collectingDateMax="2003-06-29" collectingDateMin="2002-10-27" collectionCode="CASLOT" collectorName="Harin'Hala & M. Irwin" country="Madagascar" elevation="2" location="Harin'Hala" pageId="16" pageNumber="417" specimenCode="CASLOT 024745, CASLOT 024750, CASLOT 024746, CASLOT 024728" specimenCount="8" specimenCount-female="8">
|
||
<location id="2C22FCCA260FFFB3FDB3FECC1CE1FEB5" LSID="urn:lsid:plazi:treatment:A1541B072610FFB2FF60F9121ED2FCA9:2C22FCCA260FFFB3FDB3FECC1CE1FEB5" box="[580,709,295,321]" name="Harin'Hala" pageId="16" pageNumber="417">Harin’Hala</location>
|
||
,
|
||
<collectingMethod id="F0BCD206260FFFB3FD27FECC1D7AFEB5" box="[720,862,295,321]" pageId="16" pageNumber="417">Malaise trap</collectingMethod>
|
||
, spiny forest; 3♂,
|
||
<specimenCount id="3FFB6198260FFFB3FBDAFECC1A74FEB5" box="[1069,1104,295,321]" pageId="16" pageNumber="417" type="female">1♀</specimenCount>
|
||
, same data except
|
||
<date id="5D438CD1260FFFB3FAD2FECC1EEBFE91" pageId="16" pageNumber="417" value="2003-04-20" valueMax="2003-04-30" valueMin="2003-04-20">
|
||
<collectingDate id="4D077539260FFFB3FAD2FECC1EEBFE91" pageId="16" pageNumber="417" value="2003-04-20" valueMax="2003-04-30" valueMin="2003-04-20">20–30 Apr 2003</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode id="795B026A260FFFB3FF17FEA01F94FE91" box="[224,432,330,357]" collectionCode="CASLOT" pageId="16" pageNumber="417">CASLOT 024745</specimenCode>
|
||
); 2♂,
|
||
<specimenCount id="3FFB6198260FFFB3FE0EFEA01C3FFE91" box="[505,539,331,357]" pageId="16" pageNumber="417" type="female">1♀</specimenCount>
|
||
, same data except
|
||
<date id="5D438CD1260FFFB3FD0FFEA01D8BFE91" box="[760,943,331,357]" pageId="16" pageNumber="417" value="2003-06-22" valueMax="2003-06-29" valueMin="2003-06-22">
|
||
<collectingDate id="4D077539260FFFB3FD0FFEA01D8BFE91" box="[760,943,331,357]" pageId="16" pageNumber="417" value="2003-06-22" valueMax="2003-06-29" valueMin="2003-06-22">22–29 Jun 2003</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode id="795B026A260FFFB3FC37FEA01AB4FE91" box="[960,1168,330,357]" collectionCode="CASLOT" pageId="16" pageNumber="417">CASLOT 024750</specimenCode>
|
||
); 5♂, same data except
|
||
<date id="5D438CD1260FFFB3FF60FE841FABFE7D" box="[151,399,367,393]" pageId="16" pageNumber="417" value="2003-04-30" valueMax="2003-05-11" valueMin="2003-04-30">
|
||
<collectingDate id="4D077539260FFFB3FF60FE841FABFE7D" box="[151,399,367,393]" pageId="16" pageNumber="417" value="2003-04-30" valueMax="2003-05-11" valueMin="2003-04-30">30 Apr–11 May 2003</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode id="795B026A260FFFB3FE57FE841C55FE7D" box="[416,625,367,393]" collectionCode="CASLOT" pageId="16" pageNumber="417">CASLOT 024746</specimenCode>
|
||
); 11♂,
|
||
<specimenCount id="3FFB6198260FFFB3FD3FFE841CC8FE7D" box="[712,748,367,393]" pageId="16" pageNumber="417" type="female">6♀</specimenCount>
|
||
, same data except
|
||
<date id="5D438CD1260FFFB3FC3CFE841A7FFE7D" box="[971,1115,367,393]" pageId="16" pageNumber="417" value="2002-10-27">
|
||
<collectingDate id="4D077539260FFFB3FC3CFE841A7FFE7D" box="[971,1115,367,393]" pageId="16" pageNumber="417" value="2002-10-27">27 Oct 2002</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode id="795B026A260FFFB3FB9BFE841B19FE7D" box="[1132,1341,367,393]" collectionCode="CASLOT" pageId="16" pageNumber="417">CASLOT 024728</specimenCode>
|
||
);
|
||
<quantity id="EE0507F4260FFFB3FAA4FE841B58FE7D" box="[1363,1404,367,393]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="417" unit="m" value="2.0">
|
||
<elevation id="A2D04D22260FFFB3FAA4FE841B58FE7D" box="[1363,1404,367,393]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="417" unit="m" value="2.0">2m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="9995A04C260FFFB3FA76FE841A51FE59" ID-GBIF-Occurrence="4003029373" collectionCode="R" collectorName="Dec & M. E. Irwin" country="Madagascar" latitude="-22.913334" location="Mikea Forest" longLatPrecision="92" longitude="43.482166" pageId="16" pageNumber="417" specimenCount="1">
|
||
1f,
|
||
<location id="2C22FCCA260FFFB3FF60FE781F0AFE59" LSID="urn:lsid:plazi:treatment:A1541B072610FFB2FF60F9121ED2FCA9:2C22FCCA260FFFB3FF60FE781F0AFE59" box="[151,302,403,429]" country="Madagascar" latitude="-22.913334" longLatPrecision="92" longitude="43.482166" name="Mikea Forest" pageId="16" pageNumber="417">Mikea Forest</location>
|
||
,
|
||
<locationDeviation id="38F0B333260FFFB3FECEFE781C25FE59" box="[313,513,403,429]" location="Manombo" pageId="16" pageNumber="417">NW of Manombo</locationDeviation>
|
||
,
|
||
<geoCoordinate id="4CC9CCD6260FFFB3FDFCFE7F1C58FE5B" box="[523,636,404,431]" degrees="22" direction="south" minutes="54.8" orientation="latitude" pageId="16" pageNumber="417" precision="92" value="-22.913334">22°54.8′S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="4CC9CCD6260FFFB3FD70FE7F1D23FE5B" box="[647,775,404,431]" degrees="43" direction="east" minutes="28.93" orientation="longitude" pageId="16" pageNumber="417" precision="9" value="43.482166">43°28.93′E</geoCoordinate>
|
||
,
|
||
<date id="5D438CD1260FFFB3FCE6FE781DEFFE59" box="[785,971,403,429]" pageId="16" pageNumber="417" value="1001-12-16" valueMax="1001-12-26" valueMin="1001-12-16">
|
||
16–26
|
||
<collectorName id="8408CFC7260FFFB3FCA9FE781DAFFE59" box="[862,907,403,429]" pageId="16" pageNumber="417">Dec</collectorName>
|
||
1001
|
||
</date>
|
||
,
|
||
<collectorName id="8408CFC7260FFFB3FC20FE781A70FE59" box="[983,1108,403,429]" pageId="16" pageNumber="417">M.E. Irwin</collectorName>
|
||
,
|
||
<collectionCode id="4FEC32D4260FFFB3FBA8FE781A51FE59" box="[1119,1141,403,429]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="417">R</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="9995A04C260FFFB3FB88FE781FF6FD95" ID-GBIF-Occurrence="4003029339" collectingDate="2001-10-12" collectingDateMax="2002-08-06" collectingDateMin="2001-10-12" collectionCode="CAS" collectorName="Dec & M. E. Irwin" country="Madagascar" latitude="-22.913334" location="Harin'Hala" longLatPrecision="92" longitude="43.482166" pageId="16" pageNumber="417" specimenCode="CASLOT 024686, CASLOT 024697, CASLOT 024700, CASLOT 024667, CASLOT 024657, CASLOT 024656, CASLOT 024646" specimenCount="3" specimenCount-female="3" typeStatus="holotype">
|
||
<location id="2C22FCCA260FFFB3FB88FE781B24FE59" LSID="urn:lsid:plazi:treatment:A1541B072610FFB2FF60F9121ED2FCA9:2C22FCCA260FFFB3FB88FE781B24FE59" box="[1151,1280,403,429]" name="Harin'Hala" pageId="16" pageNumber="417">Harin’Hala</location>
|
||
,
|
||
<collectingMethod id="F0BCD206260FFFB3FAFCFE781BBCFE59" box="[1291,1432,403,429]" pageId="16" pageNumber="417">Malaise trap</collectingMethod>
|
||
, spiny forest (
|
||
<specimenCode id="795B026A260FFFB3FEDFFE5C1FD2FE25" box="[296,502,439,465]" collectionCode="CASLOT" pageId="16" pageNumber="417">CASLOT 024686</specimenCode>
|
||
); 2♂, same data except
|
||
<date id="5D438CD1260FFFB3FD08FE5C1DF9FE25" box="[767,989,439,465]" pageId="16" pageNumber="417" value="2002-05-29" valueMax="2002-06-08" valueMin="2002-05-29">
|
||
<collectingDate id="4D077539260FFFB3FD08FE5C1DF9FE25" box="[767,989,439,465]" pageId="16" pageNumber="417" value="2002-05-29" valueMax="2002-06-08" valueMin="2002-05-29">29 May–8 Jun 2002</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode id="795B026A260FFFB3FC1CFE5C1A9DFE25" box="[1003,1209,439,465]" collectionCode="CASLOT" pageId="16" pageNumber="417">CASLOT 024697</specimenCode>
|
||
); 1♂,
|
||
<specimenCount id="3FFB6198260FFFB3FB0AFE5C1B04FE25" box="[1277,1312,439,465]" pageId="16" pageNumber="417" type="female">1♀</specimenCount>
|
||
, same data except
|
||
<date id="5D438CD1260FFFB3FF10FE301F98FE01" box="[231,444,475,501]" pageId="16" pageNumber="417" value="2002-07-23" valueMax="2002-08-06" valueMin="2002-07-23">
|
||
<collectingDate id="4D077539260FFFB3FF10FE301F98FE01" box="[231,444,475,501]" pageId="16" pageNumber="417" value="2002-07-23" valueMax="2002-08-06" valueMin="2002-07-23">23 Jul–6 Aug 2002</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode id="795B026A260FFFB3FE3CFE301CBDFE01" box="[459,665,475,501]" collectionCode="CASLOT" pageId="16" pageNumber="417">CASLOT 024700</specimenCode>
|
||
); 1♂,
|
||
<specimenCount id="3FFB6198260FFFB3FD2AFE301D24FE01" box="[733,768,475,501]" pageId="16" pageNumber="417" type="female">1♀</specimenCount>
|
||
, same data except
|
||
<date id="5D438CD1260FFFB3FC22FE301AA6FE01" box="[981,1154,475,501]" pageId="16" pageNumber="417" value="2002-07-13" valueMax="2002-07-23" valueMin="2002-07-13">
|
||
<collectingDate id="4D077539260FFFB3FC22FE301AA6FE01" box="[981,1154,475,501]" pageId="16" pageNumber="417" value="2002-07-13" valueMax="2002-07-23" valueMin="2002-07-13">13–23 Jul 2002</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode id="795B026A260FFFB3FB67FE301B7BFE01" box="[1168,1375,475,501]" collectionCode="CASLOT" pageId="16" pageNumber="417">CASLOT 024667</specimenCode>
|
||
); 2♂,
|
||
<specimenCount id="3FFB6198260FFFB3FF60FE141E9EFDED" box="[151,186,511,537]" pageId="16" pageNumber="417" type="female">1♀</specimenCount>
|
||
, same data except
|
||
<date id="5D438CD1260FFFB3FE6EFE141C19FDED" box="[409,573,511,537]" pageId="16" pageNumber="417" value="2002-03-01" valueMax="2002-03-08" valueMin="2002-03-01">
|
||
<collectingDate id="4D077539260FFFB3FE6EFE141C19FDED" box="[409,573,511,537]" pageId="16" pageNumber="417" value="2002-03-01" valueMax="2002-03-08" valueMin="2002-03-01">1–8 Mar 2002</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode id="795B026A260FFFB3FDB9FE141D3BFDED" box="[590,799,510,537]" collectionCode="CASLOT" pageId="16" pageNumber="417">CASLOT 024657</specimenCode>
|
||
); 3♂, same data except
|
||
<date id="5D438CD1260FFFB3FBC0FE141B39FDED" box="[1079,1309,511,537]" pageId="16" pageNumber="417" value="2002-02-19" valueMax="2002-03-01" valueMin="2002-02-19">
|
||
<collectingDate id="4D077539260FFFB3FBC0FE141B39FDED" box="[1079,1309,511,537]" pageId="16" pageNumber="417" value="2002-02-19" valueMax="2002-03-01" valueMin="2002-02-19">19 Feb–1 Mar 2002</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode id="795B026A260FFFB3FAD9FE141ED4FDC9" collectionCode="CASLOT" pageId="16" pageNumber="417">CASLOT 024656</specimenCode>
|
||
); 1♂, same data except
|
||
<date id="5D438CD1260FFFB3FE02FDC81CFAFDC9" box="[501,734,547,573]" pageId="16" pageNumber="417" value="2001-10-12" valueMax="2001-11-12" valueMin="2001-10-12">
|
||
<collectingDate id="4D077539260FFFB3FE02FDC81CFAFDC9" box="[501,734,547,573]" pageId="16" pageNumber="417" value="2001-10-12" valueMax="2001-11-12" valueMin="2001-10-12">12 Oct–12 Nov 2001</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode id="795B026A260FFFB3FD1CFDC81D9CFDC9" box="[747,952,547,573]" collectionCode="CASLOT" pageId="16" pageNumber="417">CASLOT 024646</specimenCode>
|
||
); 2♂, same data except
|
||
<date id="5D438CD1260FFFB3FB4AFDC81BB8FDC9" box="[1213,1436,547,573]" pageId="16" pageNumber="417" value="2002-04-29" valueMax="2002-05-09" valueMin="2002-04-29">
|
||
<collectingDate id="4D077539260FFFB3FB4AFDC81BB8FDC9" box="[1213,1436,547,573]" pageId="16" pageNumber="417" value="2002-04-29" valueMax="2002-05-09" valueMin="2002-04-29">29 Apr–9 May 2002</collectingDate>
|
||
</date>
|
||
(024662).
|
||
<typeStatus id="F64614B3260FFFB3FEFBFDAC1F50FD95" box="[268,372,583,609]" pageId="16" pageNumber="417" type="holotype">Holotype</typeStatus>
|
||
in
|
||
<collectionCode id="4FEC32D4260FFFB3FE6FFDAC1FF6FD95" box="[408,466,583,609]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="16" pageNumber="417" type="Museum">CAS</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="9995A04C260FFFB3FE2BFDAC1D0FFD95" ID-GBIF-Occurrence="4003029334" box="[476,811,583,609]" collectionCode="CAS, BPBM" country="Madagascar" pageId="16" pageNumber="417" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="F64614B3260FFFB3FE2BFDAC1C6DFD95" box="[476,585,583,609]" pageId="16" pageNumber="417" type="paratype">Paratypes</typeStatus>
|
||
in
|
||
<collectionCode id="4FEC32D4260FFFB3FD9AFDAC1C87FD95" box="[621,675,583,609]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="16" pageNumber="417" type="Museum">CAS</collectionCode>
|
||
and
|
||
<collectionCode id="4FEC32D4260FFFB3FD2DFDAC1D0FFD95" box="[730,811,583,609]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="16" pageNumber="417" type="Museum">BPBM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="7D82FA99260FFFB3FF60FBD61DC4FB8E" ID-DOI="http://doi.org/10.5281/zenodo.7471768" ID-Zenodo-Dep="7471768" httpUri="https://zenodo.org/record/7471768/files/figure.png" pageId="16" pageNumber="417" startId="16.[151,264,1085,1109]" targetBox="[150,1437,633,1062]" targetPageId="16">
|
||
<paragraph id="2942AA11260FFFB3FF60FBD61DC4FB8E" blockId="16.[151,1436,1085,1146]" pageId="16" pageNumber="417">
|
||
<emphasis id="1B897603260FFFB3FF60FBD61F6AFBA1" bold="true" box="[151,334,1085,1109]" pageId="16" pageNumber="417">FIGURES 48–49</emphasis>
|
||
.
|
||
<taxonomicName id="EEFDD192260FFFB3FEABFBD51D50FBA2" authority="Evenhuis & Gharali, 2022" authorityName="Evenhuis & Gharali" authorityYear="2022" box="[348,884,1086,1110]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="andriambahomananii" status="sp. nov." subGenus="Anomaloptilus">
|
||
<emphasis id="1B897603260FFFB3FEABFBD51FC2FBA2" box="[348,486,1086,1110]" italics="true" pageId="16" pageNumber="417">Empidideicus</emphasis>
|
||
(
|
||
<emphasis id="1B897603260FFFB3FE02FBD51CADFBA2" box="[501,649,1086,1110]" italics="true" pageId="16" pageNumber="417">Anomaloptilus</emphasis>
|
||
)
|
||
<emphasis id="1B897603260FFFB3FD6FFBD51D50FBA2" box="[664,884,1086,1110]" italics="true" pageId="16" pageNumber="417">andriambahomananii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="1B897603260FFFB3FC76FBD51D9FFBA2" bold="true" box="[897,955,1086,1110]" pageId="16" pageNumber="417">
|
||
<taxonomicNameLabel id="00BACB78260FFFB3FC76FBD51D9FFBA2" box="[897,955,1086,1110]" pageId="16" pageNumber="417" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, showing differences in closure of cell dm.
|
||
<emphasis id="1B897603260FFFB3FA8AFBD61BB2FBA1" bold="true" box="[1405,1430,1085,1109]" pageId="16" pageNumber="417">48</emphasis>
|
||
, cruciate appearance due to absence of crossvein;
|
||
<emphasis id="1B897603260FFFB3FD7DFB8A1C87FB8D" bold="true" box="[650,675,1121,1145]" pageId="16" pageNumber="417">49</emphasis>
|
||
, closed with a small crossvein.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="61E7F99A260FFFB3FF30FB431FC5FADA" pageId="16" pageNumber="417" type="diagnosis">
|
||
<paragraph id="2942AA11260FFFB3FF30FB431FC5FADA" blockId="16.[151,1437,1192,2016]" pageId="16" pageNumber="417">
|
||
<emphasis id="1B897603260FFFB3FF30FB431F1EFB36" bold="true" box="[199,314,1192,1218]" pageId="16" pageNumber="417">Diagnosis</emphasis>
|
||
: It is easily separated from all
|
||
<taxonomicName id="EEFDD192260FFFB3FD7DFB431D06FB36" authorityName="Becker. A" authorityYear="1907" box="[650,802,1192,1218]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1B897603260FFFB3FD7DFB431D06FB36" box="[650,802,1192,1218]" italics="true" pageId="16" pageNumber="417">Empidideicus</emphasis>
|
||
</taxonomicName>
|
||
by the elongate clypeus, extending below the oral margin and one-half the length of the labrum. This one of the largest known species in the genus (most other species are less than
|
||
<quantity id="EE0507F4260FFFB3FF39FB1B1F00FAFE" box="[206,292,1264,1291]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="417" unit="mm" value="1.5">1.5 mm</quantity>
|
||
in body length) and is thus easily spotted in collections due to its size combined with the downward or backward reflexed proboscis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61E7F99A260FFFB2FF30FAD31B66FD05" lastPageId="17" lastPageNumber="418" pageId="16" pageNumber="417" type="description">
|
||
<paragraph id="2942AA11260FFFB3FF30FAD31F7EFAA6" blockId="16.[151,1437,1192,2016]" box="[199,346,1336,1362]" pageId="16" pageNumber="417">
|
||
<emphasis id="1B897603260FFFB3FF30FAD31F75FAA6" bold="true" box="[199,337,1336,1362]" pageId="16" pageNumber="417">Description</emphasis>
|
||
:
|
||
</paragraph>
|
||
<paragraph id="2942AA11260FFFB3FF30FAB71AF6F986" blockId="16.[151,1437,1192,2016]" pageId="16" pageNumber="417">
|
||
<emphasis id="1B897603260FFFB3FF30FAB71F20FA82" bold="true" box="[199,260,1372,1398]" pageId="16" pageNumber="417">Male</emphasis>
|
||
(
|
||
<figureCitation id="B1C6B694260FFFB3FEEFFAB71F54FA82" box="[280,368,1372,1398]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[158,1429,181,1920]" captionTargetId="figure-17@5.[158,1429,181,1920]" captionTargetPageId="5" captionText="FIGURES 11–16. Malagasy Empidideicus (Anomaloptilus), habitus, lateral view. 11, E. (A.) regius, n. sp., male; 12, E. (A.) regius, n. sp., female; 13, E. (A.) rhinoclypeatus Evenhuis, male; 14, E. (A.) rhinoclypeatus Evenhuis, female; 15, E. (A.) rinhai, n. sp., male; 16, E. (A.) rinhai, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471748" httpUri="https://zenodo.org/record/7471748/files/figure.png" pageId="16" pageNumber="417">Fig. 13</figureCitation>
|
||
). Length:
|
||
<quantity id="EE0507F4260FFFB3FE1BFAB71CB2FA82" box="[492,662,1372,1398]" metricMagnitude="-3" metricUnit="m" metricValue="2.24" metricValueMax="2.88" metricValueMin="1.6" pageId="16" pageNumber="417" unit="mm" value="2.24" valueMax="2.88" valueMin="1.6">1.60–2.88 mm</quantity>
|
||
. Wing:
|
||
<quantity id="EE0507F4260FFFB3FD02FAB71D84FA82" box="[757,928,1372,1399]" metricMagnitude="-3" metricUnit="m" metricValue="2.325" metricValueMax="2.95" metricValueMin="1.7" pageId="16" pageNumber="417" unit="mm" value="2.325" valueMax="2.95" valueMin="1.7">1.70–2.95 mm</quantity>
|
||
.
|
||
<emphasis id="1B897603260FFFB3FC58FAB71DC8FA82" box="[943,1004,1372,1398]" italics="true" pageId="16" pageNumber="417">Head</emphasis>
|
||
(
|
||
<figureCitation id="B1C6B694260FFFB3FC08FAB71A69FA82" box="[1023,1101,1372,1398]" captionStart="FIGURES 3–4" captionStartId="2.[151,264,1920,1944]" captionTargetBox="[157,1423,240,1880]" captionTargetId="figure-140@2.[151,1436,227,1896]" captionTargetPageId="2" captionText="FIGURES 3–4. Schematic illustration showing differences in clypeus lengths of Empidideicus (Anomaloptilus) species. 3, E. (A.) celluliferus Hesse; 4, E. (A.) rhinoclypeatus Evenhuis. Arrows point to clypeus. (Redrawn from Evenhuis 2007; figs 2–3)." figureDoi="http://doi.org/10.5281/zenodo.7471742" httpUri="https://zenodo.org/record/7471742/files/figure.png" pageId="16" pageNumber="417">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="B1C6B694260FFFB3FBAAFAB71A59FA82" box="[1117,1149,1372,1398]" captionStart="FIGURES 20–22" captionStartId="8.[151,264,1965,1989]" captionTargetBox="[151,1363,1475,1942]" captionTargetId="figure-302@8.[151,1435,1475,1942]" captionTargetPageId="8" captionText="FIGURES 20–22. Malagasy Empidideicus (Anomaloptilus), heads. 20, E. (A.) contrasticus, n. sp., male, frontal view; 21, E. (A.) irwini, n. sp., male, oblique view; 22, E. (A.) rhinoclypeatus Evenhuis, male, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7471752" httpUri="https://zenodo.org/record/7471752/files/figure.png" pageId="16" pageNumber="417">22</figureCitation>
|
||
): longer than high; eyes dichoptic, separated at vertex by 1.5× distance between lateral ocelli; occiput and postgena black; front slightly depressed medially, white with subrectangular brown area immediately below ocellar tubercle; face white, tip of oral margin tan to brown; antennae (cf.
|
||
<figureCitation id="B1C6B694260FFFB3FDB8FA231CBBFA16" box="[591,671,1480,1506]" captionStart="FIGURES 23–24" captionStartId="10.[151,264,1309,1333]" captionTargetBox="[151,1435,181,1284]" captionTargetId="figure-17@10.[151,1435,181,1285]" captionTargetPageId="10" captionText="FIGURES 23–24. Malagasy Empidideicus (Anomaloptilus), antennae. 23, E. (A.) andriambahomananii, n. sp.; 24, E. (A.) nanus, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471754" httpUri="https://zenodo.org/record/7471754/files/figure.png" pageId="16" pageNumber="417">Fig. 23</figureCitation>
|
||
) dark brown, set in deep pocket; scape short, subtrapezoidal; pedicel subconical, wider than long; first flagellomere lanceolate, length 3.5× width; second flagellomere one-half length of first flagellomere, with minute transparent apical style; mentum black; clypeus dark brown, elongate, extending well beyond tip of oral margin, length subequal to head height; proboscis dark brown to black, length 2.5× head length; labrum sclerotized, stiff, pointed apically, length subequal to clypeus; palpus not evident.
|
||
</paragraph>
|
||
<paragraph id="2942AA11260FFFB3FF30F9971F38F866" blockId="16.[151,1437,1192,2016]" pageId="16" pageNumber="417">
|
||
<emphasis id="1B897603260FFFB3FF30F9971F3CF962" box="[199,280,1660,1686]" italics="true" pageId="16" pageNumber="417">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="B1C6B694260FFFB3FED0F9971F5FF962" box="[295,379,1660,1686]" captionStart="FIGURES 34–39" captionStartId="14.[151,264,1264,1288]" captionTargetBox="[151,1435,181,1240]" captionTargetId="figure-17@14.[151,1435,181,1240]" captionTargetPageId="14" captionText="FIGURES 34–39. Malagasy Empidideicus (Anomaloptilus), thoraces, dorsal view. 34, E. (A.) regius, n. sp., male; 35, E. (A.) regius, n. sp., female; 36, E. (A.) rhinoclypeatus Evenhuis, male; 37, E. (A.) rhinoclypeatus Evenhuis, female; 38, E. (A.) rinhai, n. sp., male; 39, E. (A.) rinhai, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471760" httpUri="https://zenodo.org/record/7471760/files/figure.png" pageId="16" pageNumber="417">Fig. 36</figureCitation>
|
||
). Mesonotum shiny black with yellow markings in following areas: prescutellar area extending anteriorly for a short distance as traces of admedian vittae; dorsolaterally from humeral callus to transverse suture; postalar callus as anterolateral extension of prescutellar mark; scutellum yellow; mesonotal disc and dorsum of scutellum with short fine brown hairs, small patch of fine minute brown hairs on dorsolateral corner of anepisternum and upper half of propleuron, otherwise, pleura bare; pleura dark brown with pale yellow to white spots on following: posterodorsal corner of anepisternum and anepimeron, laterally on propleuron, upper half of meron; coxae and basal three-fourths of femora and tarsal segments 4 and 5 brown, remainder of legs yellow; halter stem grayish brown, knob white.
|
||
</paragraph>
|
||
<paragraph id="2942AA11260FFFB2FF30F8761AFCFF0D" blockId="16.[151,1437,1192,2016]" lastBlockId="17.[151,1437,151,861]" lastPageId="17" lastPageNumber="418" pageId="16" pageNumber="417">
|
||
<emphasis id="1B897603260FFFB3FF30F8761F25F842" box="[199,257,1949,1974]" italics="true" pageId="16" pageNumber="417">Wing</emphasis>
|
||
(
|
||
<figureCitation id="B1C6B694260FFFB3FEE4F8771F4EF842" box="[275,362,1948,1974]" captionStart="FIGURES 40–47" captionStartId="15.[151,264,1607,1631]" captionTargetBox="[170,1416,309,1582]" captionTargetId="figure-81@15.[170,1416,309,1584]" captionTargetPageId="15" captionText="FIGURES 40–47. Malagasy Empidideicus (Anomaloptilus), wings. 40, E. (A.) andriambahomananii, n. sp.; 41, E. (A.) contrasticus, n. sp.; 42, E. (A.) hauseri, n. sp.; 43, E. (A.) irwini, n. sp.; 44, E. (A.) nanus, n. sp.; 45, E. (A.) regius, n. sp.; 46, E. (A.) rhinoclypeatus Evenhuis; 47, E. (A.) rinhai, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471764" httpUri="https://zenodo.org/record/7471764/files/figure.png" pageId="16" pageNumber="417">Fig. 46</figureCitation>
|
||
). Elongate, length 3.5× width; subhyaline with pale brownish tinge; veins brown; costa ends slightly beyond end of R
|
||
<subScript id="B579A854260FFFB3FE43F8391FEAF814" attach="left" box="[436,462,2002,2016]" fontSize="6" pageId="16" pageNumber="417">4+5</subScript>
|
||
; vein Sc incomplete; Rs evanescent at connection with R
|
||
<subScript id="B579A854260FFFB3FB93F8391A48F814" attach="left" box="[1124,1132,2002,2016]" fontSize="6" pageId="16" pageNumber="417">1</subScript>
|
||
; R
|
||
<subScript id="B579A854260FFFB3FB78F8391A8DF814" attach="left" box="[1167,1193,2002,2016]" fontSize="6" pageId="16" pageNumber="417">4+5</subScript>
|
||
very slightly bowed; vein M
|
||
<subScript id="B579A854260EFFB2FF10FF421ECBFF43" attach="left" box="[231,239,169,183]" fontSize="6" pageId="17" pageNumber="418">1</subScript>
|
||
curved to wing margin; M
|
||
<subScript id="B579A854260EFFB2FDEDFF421C06FF43" attach="left" box="[538,546,169,183]" fontSize="6" pageId="17" pageNumber="418">2</subScript>
|
||
straight to wing margin; cell dm closed by apical crossvein, crossvein subequal in length to crossvein r-m; anal cell closed in wing margin; A
|
||
<subScript id="B579A854260EFFB2FCA2FF261D79FF2F" attach="left" box="[853,861,205,219]" fontSize="6" pageId="17" pageNumber="418">1</subScript>
|
||
effaced, straight to wing margin; fringe of hair on posterior margin of wing well shortest at wing tip becoming longer and more sparse toward base.
|
||
</paragraph>
|
||
<paragraph id="2942AA11260EFFB2FF30FEE81FD3FE7D" blockId="17.[151,1437,151,861]" pageId="17" pageNumber="418">
|
||
<emphasis id="1B897603260EFFB2FF30FEE81F15FEE9" box="[199,305,259,285]" italics="true" pageId="17" pageNumber="418">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="B1C6B694260EFFB2FEB7FEE81FB0FEE9" box="[320,404,258,285]" captionStart="FIGURES 59–62" captionStartId="19.[151,264,694,718]" captionTargetBox="[152,1435,181,670]" captionTargetId="figure-17@19.[152,1435,181,670]" captionTargetPageId="19" captionText="FIGURES 59–62. Malagasy Empidideicus (Anomaloptilus), abdomens, dorsal view. 59, E. (A.) rhinoclypeatus Evenhuis, male; 60, E. (A.) rhinoclypeatus Evenhuis, female; 61, E. (A.) rinhai, n. sp., male; 62, E. (A.) rinhai, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471774" httpUri="https://zenodo.org/record/7471774/files/figure.png" pageId="17" pageNumber="418">Fig. 59</figureCitation>
|
||
). Dorsum black, concolorous with central mesonotal color; tergite I white laterally; posterior margins of tergites II–VII and lateral portions white; when viewed laterally abdomen with lateral black band consisting of black color on extreme lateral tips of tergites and sternites; remainder of sternites white; tergites with sparsely scattered minute hairs.
|
||
</paragraph>
|
||
<paragraph id="2942AA11260EFFB2FF30FE781DC3FDC9" blockId="17.[151,1437,151,861]" pageId="17" pageNumber="418">
|
||
<emphasis id="1B897603260EFFB2FF30FE781F15FE59" box="[199,305,403,429]" italics="true" pageId="17" pageNumber="418">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="B1C6B694260EFFB2FEBDFE781F83FE59" box="[330,423,402,429]" captionStart="FIGURES 75–76" captionStartId="22.[152,270,1430,1454]" captionTargetBox="[155,1432,838,1400]" captionTargetId="figure-57@22.[151,1436,832,1406]" captionTargetPageId="22" captionText="FIGURES 75–76. Empidideicus (Anomaloptilus) rhinoclypeatus Evenhuis, genitalia. 75, male, A, lateral view, B, phallic complex and gonocoxae, dorsal view; 76, female. Abbreviation: gcx = gonocoxa." figureDoi="http://doi.org/10.5281/zenodo.7471796" httpUri="https://zenodo.org/record/7471796/files/figure.png" pageId="17" pageNumber="418">Fig. 75</figureCitation>
|
||
). Hypopygium shiny brown. Gonocoxa subtriangular in lateral view with rounded subrectangular gonostylus (
|
||
<figureCitation id="B1C6B694260EFFB2FE3BFE5C1C11FE25" box="[460,565,438,465]" captionStart="FIGURES 75–76" captionStartId="22.[152,270,1430,1454]" captionTargetBox="[155,1432,838,1400]" captionTargetId="figure-57@22.[151,1436,832,1406]" captionTargetPageId="22" captionText="FIGURES 75–76. Empidideicus (Anomaloptilus) rhinoclypeatus Evenhuis, genitalia. 75, male, A, lateral view, B, phallic complex and gonocoxae, dorsal view; 76, female. Abbreviation: gcx = gonocoxa." figureDoi="http://doi.org/10.5281/zenodo.7471796" httpUri="https://zenodo.org/record/7471796/files/figure.png" pageId="17" pageNumber="418">Fig. 75A</figureCitation>
|
||
); lateral arms of parameral sheath extending below gonocoxa, flared apically; aedeagal bulb large, rounded; aedeagal apodeme subequal in length to gonocoxal apodeme, peanut-shaped with short lanceolate lateral rami (
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||
<figureCitation id="B1C6B694260EFFB2FE10FE141C74FDED" box="[487,592,510,537]" captionStart="FIGURES 75–76" captionStartId="22.[152,270,1430,1454]" captionTargetBox="[155,1432,838,1400]" captionTargetId="figure-57@22.[151,1436,832,1406]" captionTargetPageId="22" captionText="FIGURES 75–76. Empidideicus (Anomaloptilus) rhinoclypeatus Evenhuis, genitalia. 75, male, A, lateral view, B, phallic complex and gonocoxae, dorsal view; 76, female. Abbreviation: gcx = gonocoxa." figureDoi="http://doi.org/10.5281/zenodo.7471796" httpUri="https://zenodo.org/record/7471796/files/figure.png" pageId="17" pageNumber="418">Fig. 75B</figureCitation>
|
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); epiphallus slightly longer than gonocoxa, with hooked apex; epandrium subrectangular with long, thin, clubbed apicoventral process in lateral view.
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||
</paragraph>
|
||
<paragraph id="2942AA11260EFFB2FF30FDAC1B66FD05" blockId="17.[151,1437,151,861]" pageId="17" pageNumber="418">
|
||
<emphasis id="1B897603260EFFB2FF30FDAC1F3AFD95" bold="true" box="[199,286,583,609]" pageId="17" pageNumber="418">Female</emphasis>
|
||
: As in male except dark coloration of thorax (
|
||
<figureCitation id="B1C6B694260EFFB2FCE9FDAC1D4BFD95" box="[798,879,583,609]" captionStart="FIGURES 34–39" captionStartId="14.[151,264,1264,1288]" captionTargetBox="[151,1435,181,1240]" captionTargetId="figure-17@14.[151,1435,181,1240]" captionTargetPageId="14" captionText="FIGURES 34–39. Malagasy Empidideicus (Anomaloptilus), thoraces, dorsal view. 34, E. (A.) regius, n. sp., male; 35, E. (A.) regius, n. sp., female; 36, E. (A.) rhinoclypeatus Evenhuis, male; 37, E. (A.) rhinoclypeatus Evenhuis, female; 38, E. (A.) rinhai, n. sp., male; 39, E. (A.) rinhai, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471760" httpUri="https://zenodo.org/record/7471760/files/figure.png" pageId="17" pageNumber="418">Fig. 37</figureCitation>
|
||
) and abdomen brown. Abdominal dark coloration interrupted medially by longitudinal white stripe (
|
||
<figureCitation id="B1C6B694260EFFB2FD3CFD801D04FD71" box="[715,800,619,645]" captionStart="FIGURES 59–62" captionStartId="19.[151,264,694,718]" captionTargetBox="[152,1435,181,670]" captionTargetId="figure-17@19.[152,1435,181,670]" captionTargetPageId="19" captionText="FIGURES 59–62. Malagasy Empidideicus (Anomaloptilus), abdomens, dorsal view. 59, E. (A.) rhinoclypeatus Evenhuis, male; 60, E. (A.) rhinoclypeatus Evenhuis, female; 61, E. (A.) rinhai, n. sp., male; 62, E. (A.) rinhai, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471774" httpUri="https://zenodo.org/record/7471774/files/figure.png" pageId="17" pageNumber="418">Fig. 60</figureCitation>
|
||
). Genitalia (
|
||
<figureCitation id="B1C6B694260EFFB2FC5DFD801A25FD71" box="[938,1025,619,645]" captionStart="FIGURES 75–76" captionStartId="22.[152,270,1430,1454]" captionTargetBox="[155,1432,838,1400]" captionTargetId="figure-57@22.[151,1436,832,1406]" captionTargetPageId="22" captionText="FIGURES 75–76. Empidideicus (Anomaloptilus) rhinoclypeatus Evenhuis, genitalia. 75, male, A, lateral view, B, phallic complex and gonocoxae, dorsal view; 76, female. Abbreviation: gcx = gonocoxa." figureDoi="http://doi.org/10.5281/zenodo.7471796" httpUri="https://zenodo.org/record/7471796/files/figure.png" pageId="17" pageNumber="418">Fig. 76</figureCitation>
|
||
): genital fork spermatheca spheroidellipsoid, sclerotized brown, longer than wide, with striations on basal half, large hole apically; apical spermathecal duct ca. 1.5× length of spermatheca, thin, transparent; sperm pump 3–4× length of apical duct, transparent, not sclerotized apically or basally, slightly flared and rounded apically; basal duct confluent with sperm pump.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61E7F99A260EFFB2FF30FD101ED2FCA9" pageId="17" pageNumber="418" type="etymology">
|
||
<paragraph id="2942AA11260EFFB2FF30FD101ED2FCA9" blockId="17.[151,1437,151,861]" pageId="17" pageNumber="418">
|
||
<emphasis id="1B897603260EFFB2FF30FD101F6CFCE1" bold="true" box="[199,328,763,789]" pageId="17" pageNumber="418">Etymology</emphasis>
|
||
. The specific epithet derives from Greek, ρηινοσ = “nose” + χλψπεατυσ = “provide with a shield”; referring to the elongate clypeus of this new species, which is reminiscent of the nose guards of ancient Greek helmets.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |