257 lines
36 KiB
XML
257 lines
36 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1179.107587" ID-Pensoft-Pub="1313-2970-1179-35" ID-Pensoft-UUID="07CEAFC9849852A1AC00767783926142" ID-ZooBank="D9BD50CBB127487981E99421E4F059BA" ModsDocID="1313-2970-1179-35" checkinTime="1694247957851" checkinUser="pensoft" docAuthor="Gao, Biao, Han, Hui-Lin, Kononenko, Vladimir S. & Pan, Zhao-Hui" docDate="2023" docId="A12C7C7AB7CA5D31894815776E3319ED" docLanguage="en" docName="ZooKeys 1179: 35-61" docOrigin="ZooKeys 1179" docPubDate="2023-09-08" docSource="http://dx.doi.org/10.3897/zookeys.1179.107587" docTitle="Hermonassa conusa Gao, Han & Kononenko 2023, sp. nov." docType="treatment" docUuid="A737AEAE-94E9-43AA-8F51-18DC02AA63FD" docUuidSource="ZooBank" docVersion="1" id="07CEAFC9849852A1AC00767783926142" lastPageNumber="35" masterDocId="07CEAFC9849852A1AC00767783926142" masterDocTitle="Five new species of the genus Hermonassa Walker, 1865 from Xizang Autonomous Region, China (Lepidoptera, Noctuidae, Noctuinae)" masterLastPageNumber="61" masterPageNumber="35" pageNumber="35" updateTime="1694247957851" updateUser="pensoft">
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<mods:titleInfo id="E7359366752FFB5AEC5BCC1D43524DB1">
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<mods:title id="E3CB513F1DD2D5598B3F1432BA22FA2A">Five new species of the genus Hermonassa Walker, 1865 from Xizang Autonomous Region, China (Lepidoptera, Noctuidae, Noctuinae)</mods:title>
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<mods:name id="8E1A03F1BD86D9F736BC729B6793D4FD" type="personal">
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<mods:roleTerm id="CF67432F0D2F462E0273A8E8DC2BA1BC">Author</mods:roleTerm>
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<mods:namePart id="E85E10A22C306EA3ED7FCBE8AECAD1E5">Gao, Biao</mods:namePart>
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<mods:affiliation id="4C0DBF00CD462E770D991169AAC17531">Northeast Forestry University, School of Forestry, Harbin 150040, China</mods:affiliation>
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<mods:name id="36EBA3C900219E9007E3C1F7EFF92E78" type="personal">
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<mods:namePart id="CB3AEB10F27917BE31C9DE2015F4C48F">Han, Hui-Lin</mods:namePart>
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<mods:nameIdentifier id="B9B218C4C4EB665DAC7B1AB8D39273A3" type="ORCID">https://orcid.org/0000-0002-2045-6182</mods:nameIdentifier>
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<mods:affiliation id="C328203C3813CAE375D2A9923E9CA59C">Northeast Forestry University, School of Forestry, Harbin 150040, China</mods:affiliation>
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</mods:name>
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<mods:name id="B1DDDA7F03942AC291AA065A575F8AA4" type="personal">
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<mods:namePart id="C4440C2447D8614095680B9A7E732245">Kononenko, Vladimir S.</mods:namePart>
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<mods:nameIdentifier id="957A8B5550AEB9C54015EC50FA62D2F6" type="ORCID">https://orcid.org/0000-0001-6103-4800</mods:nameIdentifier>
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<mods:affiliation id="74E6C04380ACC1E42DE169E450B72FFC">Northeast Asia Biodiversity Research Center, Northeast Forestry University, Harbin 15004, China</mods:affiliation>
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<mods:nameIdentifier id="746AAD81B68E3C901D8D8B978B029E3B" type="email">vsk528217@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A37D9FBD17046BAA2D9AD6D6C9B7C38F">Pan, Zhao-Hui</mods:namePart>
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<mods:affiliation id="EA46EF78CB96FD715EC25573600D0D55">Northeast Forestry University, Ministry of Education, Key Laboratory of Sustainable Forest Ecosystem Management, Harbin 150040, China</mods:affiliation>
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<mods:title id="21F06260E6C9399D94B46BBBE949E96F">ZooKeys</mods:title>
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<mods:date id="F1FBFECC02A586934D2C050784E886EE">2023</mods:date>
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<mods:number id="DB64753D3A4768681F80C99720040D33">2023-09-08</mods:number>
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<mods:identifier id="59BD030AFF796F06CE7D597AAE8C558B" type="DOI">http://dx.doi.org/10.3897/zookeys.1179.107587</mods:identifier>
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<treatment id="A12C7C7AB7CA5D31894815776E3319ED" LSID="urn:lsid:zoobank.org:act:A737AEAE-94E9-43AA-8F51-18DC02AA63FD" httpUri="http://treatment.plazi.org/id/A12C7C7AB7CA5D31894815776E3319ED" lastPageNumber="35" pageId="0" pageNumber="35">
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<subSubSection id="98CC14517F779D716ACAFA462EFA4F27" pageId="0" pageNumber="35" type="nomenclature">
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<paragraph id="F7668B0D1E1A71D9938F492DB7819559" pageId="0" pageNumber="35">
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<taxonomicName id="9D0F99EBC71B72D7EB1F5662F7F248B1" LSID="https://zoobank.org/A737AEAE-94E9-43AA-8F51-18DC02AA63FD" authority="Gao, Han & Kononenko" authorityName="Gao, Han & Kononenko" authorityYear="2023" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa conusa" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="conusa" status="sp. nov.">Hermonassa conusa Gao, Han & Kononenko</taxonomicName>
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<taxonomicNameLabel id="2066EDDD15ACAD5BA30ED6F6483E62E5" pageId="0" pageNumber="35">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="D612A3E7A84C22DE8DAC03C24136773E" pageId="0" pageNumber="35" type="description">
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<paragraph id="1CA827606EFC28E1ACD030F92A2F2927" pageId="0" pageNumber="35">
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<figureCitation id="1D999F8F66BE518A4E73C13D691CF4E8" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Hermonassa spp., adults 1 H. nigricans sp. nov., ♂, holotype, China 2 Ditto, ♀, paratype 3 H. anthracina, ♂, holotype, China (ZFMK) 4 H. anthracina, ♀, China 5 H. kalamantra, ♂, holotype, Nepal (after Kovacs et al. 2018) 6 H. kalamantra, ♂, China 7 H. conusa sp. nov., ♂, holotype, China 8 Ditto, ♀, paratype, China 9 H. renifera, ♂, holotype, China (NACR) 10 H. shizukoae, ♂, holotype, India (NIAES)." figureDoi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35">Figs 7</figureCitation>
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<figureCitation id="08B2504A25A7B6D691EFB57DA1453F5E" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Hermonassa spp., adults 1 H. nigricans sp. nov., ♂, holotype, China 2 Ditto, ♀, paratype 3 H. anthracina, ♂, holotype, China (ZFMK) 4 H. anthracina, ♀, China 5 H. kalamantra, ♂, holotype, Nepal (after Kovacs et al. 2018) 6 H. kalamantra, ♂, China 7 H. conusa sp. nov., ♂, holotype, China 8 Ditto, ♀, paratype, China 9 H. renifera, ♂, holotype, China (NACR) 10 H. shizukoae, ♂, holotype, India (NIAES)." figureDoi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35">, 8</figureCitation>
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<figureCitation id="1AE2C4A7EC155756CE14D777B1805F0C" captionStart="Figures 19–27" captionStartId="F3" captionText="Figures 19 - 27. Hermonassa spp., male (19 - 22) and female (23 - 27) genitalia 19 H. conusa sp. nov., holotype, genit. prep. hhl- 3492 - 1 20 H. renifera, paratype (after Chen 1991) 21 H. shizukoae, holotype, India (after Sugi 1995) 22 H. shizukoae, Nepal (after Kovacs et al. 2018) 23 H. nigricans, paratype, genit. prep. hhl- 3493 - 2 24 H. anthracina, China, (after Kovacs et al. 2018) 25 H. kalamantra, Nepal (after Kovacs et al. 2018) 26 H. conusa sp. nov., paratype, China, genit. prep. hhl- 3491 - 2 27 H. shizukoae, paratype, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures19-27" httpUri="https://binary.pensoft.net/fig/903265" pageId="0" pageNumber="35">, 19</figureCitation>
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<figureCitation id="B12177B83071BC48722D3DC4FBAE5232" captionStart="Figures 19–27" captionStartId="F3" captionText="Figures 19 - 27. Hermonassa spp., male (19 - 22) and female (23 - 27) genitalia 19 H. conusa sp. nov., holotype, genit. prep. hhl- 3492 - 1 20 H. renifera, paratype (after Chen 1991) 21 H. shizukoae, holotype, India (after Sugi 1995) 22 H. shizukoae, Nepal (after Kovacs et al. 2018) 23 H. nigricans, paratype, genit. prep. hhl- 3493 - 2 24 H. anthracina, China, (after Kovacs et al. 2018) 25 H. kalamantra, Nepal (after Kovacs et al. 2018) 26 H. conusa sp. nov., paratype, China, genit. prep. hhl- 3491 - 2 27 H. shizukoae, paratype, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures19-27" httpUri="https://binary.pensoft.net/fig/903265" pageId="0" pageNumber="35">, 26</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="B20251F51E0AF20DDE8881A8D02E97D6" pageId="0" pageNumber="35" type="type material">
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<paragraph id="31FE4A2F496BC378AEF51A09C1DACCC6" pageId="0" pageNumber="35">Type material.</paragraph>
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<paragraph id="D0B5536914E5480350B21645A7AC7102" pageId="0" pageNumber="35">
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<emphasis id="954D4C9C09FABD58B172ED22F140B1F0" bold="true" italics="true" pageId="0" pageNumber="35">Holotype</emphasis>
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: ♂, China, Aut. Reg. Xizang, Linzhi City, Mt. Sejila, 3300 m, 28 July 2015, H.L. Han leg., genit. prep. hhl-3492-1.
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<emphasis id="A21395CEF7FB5FCDAFC690B514B32E43" bold="true" italics="true" pageId="0" pageNumber="35">Paratypes</emphasis>
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: China: 1 ♂, Aut. Reg. Xizang, Linzhi City, Mt. Sejila, 3650 m, 22 Aug. 2014, H.L. Han leg., genit. prep. GB-66-1; 1 ♂, 2 ♀, Aut. Reg. Xizang, Linzhi City, Mt. Sejila, 3300 m, 28 July 2015, H.L. Han leg., genit. prep. hhl-3491-2; 1 ♂, Aut. Reg. Xizang, Linzhi City, Mt. Sejila, Yaguo, 3150 m, 24 Aug. 2015, H.L. Han leg., genit. prep. GB-29-1 (coll. NEFU).
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</paragraph>
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</subSubSection>
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<subSubSection id="C9BD68AE278372E0926F94237EE8EF1A" pageId="0" pageNumber="35" type="diagnosis">
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<paragraph id="6115CED02A5BDEC11FD1929F3948B706" pageId="0" pageNumber="35">Diagnosis.</paragraph>
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<paragraph id="8A6704B10F9207D247C791FF73339DCE" pageId="0" pageNumber="35">
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<taxonomicName id="22EDC59A36BDDB15329404FB3175D00E" authorityName="Gao, Han & Kononenko" authorityYear="2023" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa conusa" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="conusa">
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<emphasis id="112F86F5241BA8DED82061C4694FCF1C" italics="true" pageId="0" pageNumber="35">Hermonassa conusa</emphasis>
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</taxonomicName>
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sp. nov. (Figs
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<figureCitation id="78868BD790A7FC052E61CF4A23EE1F07" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Hermonassa spp., adults 1 H. nigricans sp. nov., ♂, holotype, China 2 Ditto, ♀, paratype 3 H. anthracina, ♂, holotype, China (ZFMK) 4 H. anthracina, ♀, China 5 H. kalamantra, ♂, holotype, Nepal (after Kovacs et al. 2018) 6 H. kalamantra, ♂, China 7 H. conusa sp. nov., ♂, holotype, China 8 Ditto, ♀, paratype, China 9 H. renifera, ♂, holotype, China (NACR) 10 H. shizukoae, ♂, holotype, India (NIAES)." figureDoi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35">7</figureCitation>
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,
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<figureCitation id="3805A3EA064F463A76C29836601C51AD" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Hermonassa spp., adults 1 H. nigricans sp. nov., ♂, holotype, China 2 Ditto, ♀, paratype 3 H. anthracina, ♂, holotype, China (ZFMK) 4 H. anthracina, ♀, China 5 H. kalamantra, ♂, holotype, Nepal (after Kovacs et al. 2018) 6 H. kalamantra, ♂, China 7 H. conusa sp. nov., ♂, holotype, China 8 Ditto, ♀, paratype, China 9 H. renifera, ♂, holotype, China (NACR) 10 H. shizukoae, ♂, holotype, India (NIAES)." figureDoi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35">8</figureCitation>
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,
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<figureCitation id="A80A9852DA997CB99E5F2340A4BDF4F0" captionStart="Figures 19–27" captionStartId="F3" captionText="Figures 19 - 27. Hermonassa spp., male (19 - 22) and female (23 - 27) genitalia 19 H. conusa sp. nov., holotype, genit. prep. hhl- 3492 - 1 20 H. renifera, paratype (after Chen 1991) 21 H. shizukoae, holotype, India (after Sugi 1995) 22 H. shizukoae, Nepal (after Kovacs et al. 2018) 23 H. nigricans, paratype, genit. prep. hhl- 3493 - 2 24 H. anthracina, China, (after Kovacs et al. 2018) 25 H. kalamantra, Nepal (after Kovacs et al. 2018) 26 H. conusa sp. nov., paratype, China, genit. prep. hhl- 3491 - 2 27 H. shizukoae, paratype, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures19-27" httpUri="https://binary.pensoft.net/fig/903265" pageId="0" pageNumber="35">19</figureCitation>
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,
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<figureCitation id="C8662DAB8F194D0D7E0A786BFB375928" captionStart="Figures 19–27" captionStartId="F3" captionText="Figures 19 - 27. Hermonassa spp., male (19 - 22) and female (23 - 27) genitalia 19 H. conusa sp. nov., holotype, genit. prep. hhl- 3492 - 1 20 H. renifera, paratype (after Chen 1991) 21 H. shizukoae, holotype, India (after Sugi 1995) 22 H. shizukoae, Nepal (after Kovacs et al. 2018) 23 H. nigricans, paratype, genit. prep. hhl- 3493 - 2 24 H. anthracina, China, (after Kovacs et al. 2018) 25 H. kalamantra, Nepal (after Kovacs et al. 2018) 26 H. conusa sp. nov., paratype, China, genit. prep. hhl- 3491 - 2 27 H. shizukoae, paratype, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures19-27" httpUri="https://binary.pensoft.net/fig/903265" pageId="0" pageNumber="35">26</figureCitation>
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) together with
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<taxonomicName id="4D352F5C28C5F03B01478432D3FAF917" family="Noctuidae" kingdom="Animalia" lsidName="H. renifera" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="renifera">
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<emphasis id="81C60002EE362195CB94022D2D5D6A54" italics="true" pageId="0" pageNumber="35">H. renifera</emphasis>
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</taxonomicName>
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(Figs
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<figureCitation id="7101CE593ECC510948F7001093DD2598" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Hermonassa spp., adults 1 H. nigricans sp. nov., ♂, holotype, China 2 Ditto, ♀, paratype 3 H. anthracina, ♂, holotype, China (ZFMK) 4 H. anthracina, ♀, China 5 H. kalamantra, ♂, holotype, Nepal (after Kovacs et al. 2018) 6 H. kalamantra, ♂, China 7 H. conusa sp. nov., ♂, holotype, China 8 Ditto, ♀, paratype, China 9 H. renifera, ♂, holotype, China (NACR) 10 H. shizukoae, ♂, holotype, India (NIAES)." figureDoi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35">9</figureCitation>
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,
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<figureCitation id="7310B43054791352ACD9D58CFCA22E72" captionStart="Figures 19–27" captionStartId="F3" captionText="Figures 19 - 27. Hermonassa spp., male (19 - 22) and female (23 - 27) genitalia 19 H. conusa sp. nov., holotype, genit. prep. hhl- 3492 - 1 20 H. renifera, paratype (after Chen 1991) 21 H. shizukoae, holotype, India (after Sugi 1995) 22 H. shizukoae, Nepal (after Kovacs et al. 2018) 23 H. nigricans, paratype, genit. prep. hhl- 3493 - 2 24 H. anthracina, China, (after Kovacs et al. 2018) 25 H. kalamantra, Nepal (after Kovacs et al. 2018) 26 H. conusa sp. nov., paratype, China, genit. prep. hhl- 3491 - 2 27 H. shizukoae, paratype, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures19-27" httpUri="https://binary.pensoft.net/fig/903265" pageId="0" pageNumber="35">20</figureCitation>
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) and
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<taxonomicName id="238A1AFCF41EC2730D2041883870D026" family="Noctuidae" kingdom="Animalia" lsidName="H. shizukoae" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="shizukoae">
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<emphasis id="1D3198069BFD5E81C3D1D4E6B3A9CFE1" italics="true" pageId="0" pageNumber="35">H. shizukoae</emphasis>
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</taxonomicName>
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(Figs
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<figureCitation id="5BE940F0D967C275B0CAC4806EBFEFDE" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Hermonassa spp., adults 1 H. nigricans sp. nov., ♂, holotype, China 2 Ditto, ♀, paratype 3 H. anthracina, ♂, holotype, China (ZFMK) 4 H. anthracina, ♀, China 5 H. kalamantra, ♂, holotype, Nepal (after Kovacs et al. 2018) 6 H. kalamantra, ♂, China 7 H. conusa sp. nov., ♂, holotype, China 8 Ditto, ♀, paratype, China 9 H. renifera, ♂, holotype, China (NACR) 10 H. shizukoae, ♂, holotype, India (NIAES)." figureDoi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35">10</figureCitation>
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,
|
||
<figureCitation id="970D301AE174B46B4C462CDF0D75235F" captionStart="Figures 19–27" captionStartId="F3" captionText="Figures 19 - 27. Hermonassa spp., male (19 - 22) and female (23 - 27) genitalia 19 H. conusa sp. nov., holotype, genit. prep. hhl- 3492 - 1 20 H. renifera, paratype (after Chen 1991) 21 H. shizukoae, holotype, India (after Sugi 1995) 22 H. shizukoae, Nepal (after Kovacs et al. 2018) 23 H. nigricans, paratype, genit. prep. hhl- 3493 - 2 24 H. anthracina, China, (after Kovacs et al. 2018) 25 H. kalamantra, Nepal (after Kovacs et al. 2018) 26 H. conusa sp. nov., paratype, China, genit. prep. hhl- 3491 - 2 27 H. shizukoae, paratype, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures19-27" httpUri="https://binary.pensoft.net/fig/903265" pageId="0" pageNumber="35">21</figureCitation>
|
||
,
|
||
<figureCitation id="58343373FB99E14EF1930DCF415B96C5" captionStart="Figures 19–27" captionStartId="F3" captionText="Figures 19 - 27. Hermonassa spp., male (19 - 22) and female (23 - 27) genitalia 19 H. conusa sp. nov., holotype, genit. prep. hhl- 3492 - 1 20 H. renifera, paratype (after Chen 1991) 21 H. shizukoae, holotype, India (after Sugi 1995) 22 H. shizukoae, Nepal (after Kovacs et al. 2018) 23 H. nigricans, paratype, genit. prep. hhl- 3493 - 2 24 H. anthracina, China, (after Kovacs et al. 2018) 25 H. kalamantra, Nepal (after Kovacs et al. 2018) 26 H. conusa sp. nov., paratype, China, genit. prep. hhl- 3491 - 2 27 H. shizukoae, paratype, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures19-27" httpUri="https://binary.pensoft.net/fig/903265" pageId="0" pageNumber="35">22</figureCitation>
|
||
,
|
||
<figureCitation id="6012B481BE8363FE70D643B9811DB22F" captionStart="Figures 19–27" captionStartId="F3" captionText="Figures 19 - 27. Hermonassa spp., male (19 - 22) and female (23 - 27) genitalia 19 H. conusa sp. nov., holotype, genit. prep. hhl- 3492 - 1 20 H. renifera, paratype (after Chen 1991) 21 H. shizukoae, holotype, India (after Sugi 1995) 22 H. shizukoae, Nepal (after Kovacs et al. 2018) 23 H. nigricans, paratype, genit. prep. hhl- 3493 - 2 24 H. anthracina, China, (after Kovacs et al. 2018) 25 H. kalamantra, Nepal (after Kovacs et al. 2018) 26 H. conusa sp. nov., paratype, China, genit. prep. hhl- 3491 - 2 27 H. shizukoae, paratype, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures19-27" httpUri="https://binary.pensoft.net/fig/903265" pageId="0" pageNumber="35">27</figureCitation>
|
||
) are the most similar species in the
|
||
<emphasis id="76D9777F08BF2C869095193639232FE9" italics="true" pageId="0" pageNumber="35">dispila</emphasis>
|
||
species group. Externally
|
||
<taxonomicName id="D41BF4920F56CF07F3C476F06FFC92D2" family="Noctuidae" kingdom="Animalia" lsidName="H. conusa" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="conusa">
|
||
<emphasis id="FA4E434E7CEA2F3CD4A6D7264B09D866" italics="true" pageId="0" pageNumber="35">H. conusa</emphasis>
|
||
</taxonomicName>
|
||
differs from related species by more a robust habit, a broader forewing shape, dark brown with pale violet tint colouration of the forewing (in
|
||
<taxonomicName id="BD1145B9997AA0354288AD0070880320" family="Noctuidae" kingdom="Animalia" lsidName="H. renifera" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="renifera">
|
||
<emphasis id="EC4B46FD4D2C37DC6DFF06CF2FC80996" italics="true" pageId="0" pageNumber="35">H. renifera</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D08E09CCDC87E99356F5F1CB0D1D2148" family="Noctuidae" kingdom="Animalia" lsidName="H. shizukoae" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="shizukoae">
|
||
<emphasis id="FD552861B73286C777B9E6E8660DE416" italics="true" pageId="0" pageNumber="35">H. shizukoae</emphasis>
|
||
</taxonomicName>
|
||
the forewing colouration is brown with a reddish tint, especially in the costal part) and by a more coarse forewing pattern with more distinct basal, ante-, and postmedial lines (they are weakly expressed in
|
||
<taxonomicName id="463BE5B68F147672AFAE675751ACCB6E" family="Noctuidae" kingdom="Animalia" lsidName="H. renifera" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="renifera">
|
||
<emphasis id="21F1925A9DA6A4236F65E518F002241C" italics="true" pageId="0" pageNumber="35">H. renifera</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="1907BC75DB78E020519AFB129C4BEFBD" family="Noctuidae" kingdom="Animalia" lsidName="H. shizukoae" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="shizukoae">
|
||
<emphasis id="9AAD8141FD54BCE03C317086E0A4DA30" italics="true" pageId="0" pageNumber="35">H. shizukoae</emphasis>
|
||
</taxonomicName>
|
||
); more distinct claviform, and larger drop-shaped orbicular than in related species (in
|
||
<taxonomicName id="E5479C307C797E482DA3D2069A77E0ED" family="Noctuidae" kingdom="Animalia" lsidName="H. renifera" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="renifera">
|
||
<emphasis id="D0FA27BC068FB0D9BF6719D461D3F96C" italics="true" pageId="0" pageNumber="35">H. renifera</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="DBCC9C4676EF3055EB6CB60AB36313C1" family="Noctuidae" kingdom="Animalia" lsidName="H. shizukoae" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="shizukoae">
|
||
<emphasis id="A90AA48C339DD9FA29E4940B87572EBF" italics="true" pageId="0" pageNumber="35">H. shizukoae</emphasis>
|
||
</taxonomicName>
|
||
the claviform is weekly expressed and the orbicular is small drop-shaped). The male genitalia of
|
||
<taxonomicName id="49FD2C62F0747FD1990B03F85F3FC58F" family="Noctuidae" kingdom="Animalia" lsidName="H. conusa" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="conusa">
|
||
<emphasis id="C8A70962DC200C5E969316916EEC812E" italics="true" pageId="0" pageNumber="35">H. conusa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differ from those of the related species by shorter tegumen (it is ~ 1/3
|
||
<normalizedToken id="AF47503822C24DD9B05E0DA95EB6521C" originalValue="×">x</normalizedToken>
|
||
height of vinculum vs ~ 1/2
|
||
<normalizedToken id="51A44FCF19E1354F93BED3A95E6D35D9" originalValue="×">x</normalizedToken>
|
||
in
|
||
<taxonomicName id="36F3548EA3079BCBBFC69AD8EB850BBB" family="Noctuidae" kingdom="Animalia" lsidName="H. renifera" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="renifera">
|
||
<emphasis id="530C32B0C5814F7569E7A0182D4C3B19" italics="true" pageId="0" pageNumber="35">H. renifera</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="959266416DBFAD365DCC4C182C40CC3D" family="Noctuidae" kingdom="Animalia" lsidName="H. shizukoae" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="shizukoae">
|
||
<emphasis id="99C4F07B305CD020F6539A92083A9C7A" italics="true" pageId="0" pageNumber="35">H. shizukoae</emphasis>
|
||
</taxonomicName>
|
||
), the broad valva with prominent subbasal extensions of the costa (in the related species the valva are much narrower, without extension of the costa); the harpe with a large and broad base, which reaches but does not exceed the dorsal margin of valva (in the related species the base of harpe is narrower and exceed the dorsal margin of valva); the top of harpe is short, straight and broad (vs long, narrow and curved in
|
||
<taxonomicName id="8BA59E76826D044E44ED4CBB9A5A1F26" family="Noctuidae" kingdom="Animalia" lsidName="H. renifera" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="renifera">
|
||
<emphasis id="427C002CE2E48BDAE5C860217D8BD626" italics="true" pageId="0" pageNumber="35">H. renifera</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4DF138BBB04A9EF45E56278B5EE19918" family="Noctuidae" kingdom="Animalia" lsidName="H. shizukoae" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="shizukoae">
|
||
<emphasis id="61E65764B05234D5E82D210C972ECE14" italics="true" pageId="0" pageNumber="35">H. shizukoae</emphasis>
|
||
</taxonomicName>
|
||
). The juxta is wide, rectangular, with a medium process, inverted nail-shaped (in
|
||
<taxonomicName id="115A73414BA088195349B04ECB53FED5" family="Noctuidae" kingdom="Animalia" lsidName="H. renifera" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="renifera">
|
||
<emphasis id="0B8C3FCEFD50CF1D26D8AAAF0C392C99" italics="true" pageId="0" pageNumber="35">H. renifera</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="90F303F3327690191ADA77F4BEECE1B5" family="Noctuidae" kingdom="Animalia" lsidName="H. shizukoae" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="shizukoae">
|
||
<emphasis id="D6ACC3EDC15E86BDFD078F6CAEF432B0" italics="true" pageId="0" pageNumber="35">H. shizukoae</emphasis>
|
||
</taxonomicName>
|
||
the juxta is tongue-shaped, or nearly heart-shaped with a smaller and shorter process); the sacculus is much broader than in related species. The aedeagus is much longer and narrower than in
|
||
<taxonomicName id="A91E7E866E36DE1EECA641CA78E9A531" family="Noctuidae" kingdom="Animalia" lsidName="H. renifera" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="renifera">
|
||
<emphasis id="D815ECF4853F3EDC755F2F9EA7DDD936" italics="true" pageId="0" pageNumber="35">H. renifera</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="87A42B88F25ADA851890F9AF57D66D60" family="Noctuidae" kingdom="Animalia" lsidName="H. shizukoae" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="shizukoae">
|
||
<emphasis id="90FE5194DE6359E40C62238FFDB9B930" italics="true" pageId="0" pageNumber="35">H. shizukoae</emphasis>
|
||
</taxonomicName>
|
||
; the cornutus much smaller and thinner; the vesica is tube-like, with smaller subbasal diverticulum compared with
|
||
<taxonomicName id="F0CDD077F815F23E4378AB4205D8767A" family="Noctuidae" kingdom="Animalia" lsidName="H. renifera" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="renifera">
|
||
<emphasis id="52DE75CEE7533A9F37E88661EA993D0C" italics="true" pageId="0" pageNumber="35">H. renifera</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="197FE8D076FEB32C88F6FA1791F8555B" family="Noctuidae" kingdom="Animalia" lsidName="H. shizukoae." order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="shizukoae.">
|
||
<emphasis id="984B7BC83514FE5E339DA2097DE99AE3" italics="true" pageId="0" pageNumber="35">H. shizukoae.</emphasis>
|
||
</taxonomicName>
|
||
In the female genitalia, antevaginal plate is narrow and nearly heart-shaped vs broad and calyculate in
|
||
<taxonomicName id="9653D01F3AB8C18C02159AE7F668A104" family="Noctuidae" kingdom="Animalia" lsidName="H. shizukoae" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="shizukoae">
|
||
<emphasis id="D2D2A59CC99AF01792787C3B0B7317BB" italics="true" pageId="0" pageNumber="35">H. shizukoae</emphasis>
|
||
</taxonomicName>
|
||
; ductus bursae is long and slightly sclerotised vs short in
|
||
<taxonomicName id="DFB25D18159D2653310E4DC38BD4851E" family="Noctuidae" kingdom="Animalia" lsidName="H. shizukoae" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="shizukoae">
|
||
<emphasis id="2DCF2829832C348B1535C461E77013C8" italics="true" pageId="0" pageNumber="35">H. shizukoae</emphasis>
|
||
</taxonomicName>
|
||
; corpus bursae is shorter, with four band-like signa (in
|
||
<taxonomicName id="EAEF6DA0B06B7690D1FCD946E9A7BD7B" family="Noctuidae" kingdom="Animalia" lsidName="H. shizukoae" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="shizukoae">
|
||
<emphasis id="C649CEEB88CF2F6245D1A288EF8D5FB1" italics="true" pageId="0" pageNumber="35">H. shizukoae</emphasis>
|
||
</taxonomicName>
|
||
corpus bursae is longer and the signa are slighter and wrinkle-like.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D7CCF0C733B221555802E75460F85ACC" pageId="0" pageNumber="35" type="description">
|
||
<paragraph id="585EE3E676E853D1909F86C6B3547A99" pageId="0" pageNumber="35">Description.</paragraph>
|
||
<paragraph id="26C4407A1EE5BFF88BE4040466419DB8" pageId="0" pageNumber="35">
|
||
<emphasis id="6BCA5A60F8245CF562B719AA18C99ADA" bold="true" italics="true" pageId="0" pageNumber="35">Adult</emphasis>
|
||
(Figs
|
||
<figureCitation id="B67094732191C582230F9F88D2A8AF22" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Hermonassa spp., adults 1 H. nigricans sp. nov., ♂, holotype, China 2 Ditto, ♀, paratype 3 H. anthracina, ♂, holotype, China (ZFMK) 4 H. anthracina, ♀, China 5 H. kalamantra, ♂, holotype, Nepal (after Kovacs et al. 2018) 6 H. kalamantra, ♂, China 7 H. conusa sp. nov., ♂, holotype, China 8 Ditto, ♀, paratype, China 9 H. renifera, ♂, holotype, China (NACR) 10 H. shizukoae, ♂, holotype, India (NIAES)." figureDoi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35">7</figureCitation>
|
||
,
|
||
<figureCitation id="2FB61B051E444D4AB0C743BB624D0F80" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Hermonassa spp., adults 1 H. nigricans sp. nov., ♂, holotype, China 2 Ditto, ♀, paratype 3 H. anthracina, ♂, holotype, China (ZFMK) 4 H. anthracina, ♀, China 5 H. kalamantra, ♂, holotype, Nepal (after Kovacs et al. 2018) 6 H. kalamantra, ♂, China 7 H. conusa sp. nov., ♂, holotype, China 8 Ditto, ♀, paratype, China 9 H. renifera, ♂, holotype, China (NACR) 10 H. shizukoae, ♂, holotype, India (NIAES)." figureDoi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35">8</figureCitation>
|
||
). Wingspan 29-31 mm. Head, labial palps, thorax dark brown, patagia and tegulae black; abdomen black, mixed with greyish white at segments 1-3, abdominal tuft distinct. Forewing ground colour dark brown; basal line double, black; antemedial line double, black, slender, curved, slightly wavy; median line indistinct; postmedial line double, brown, mixed with black, excurved; subterminal line indistinct; terminal line greyish white, slender; orbicular spot drop-shaped and reniform spot crescent-shaped, both with pale brown edges. Hindwing pale, light greyish brown; discal spot arc-shaped; veins brown; fringe pale brown.
|
||
<emphasis id="7DE794C64BB1C134B71C69DB0BB8E8AF" bold="true" italics="true" pageId="0" pageNumber="35">Male genitalia</emphasis>
|
||
(Fig.
|
||
<figureCitation id="59D1A5395C40EB5049776AEF9F084670" captionStart="Figures 19–27" captionStartId="F3" captionText="Figures 19 - 27. Hermonassa spp., male (19 - 22) and female (23 - 27) genitalia 19 H. conusa sp. nov., holotype, genit. prep. hhl- 3492 - 1 20 H. renifera, paratype (after Chen 1991) 21 H. shizukoae, holotype, India (after Sugi 1995) 22 H. shizukoae, Nepal (after Kovacs et al. 2018) 23 H. nigricans, paratype, genit. prep. hhl- 3493 - 2 24 H. anthracina, China, (after Kovacs et al. 2018) 25 H. kalamantra, Nepal (after Kovacs et al. 2018) 26 H. conusa sp. nov., paratype, China, genit. prep. hhl- 3491 - 2 27 H. shizukoae, paratype, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures19-27" httpUri="https://binary.pensoft.net/fig/903265" pageId="0" pageNumber="35">19</figureCitation>
|
||
). Uncus slightly hooked apically. Tegumen broad, slightly longer than uncus, ~ 1/3
|
||
<normalizedToken id="BC12FE34D3D937F93CFE91110757F84C" originalValue="×">x</normalizedToken>
|
||
vinculum length. Juxta rectangular with medial inverted nail-shaped process. Valva broad medially, gradually narrower apically; cucullus flat, blunt round; harpe slightly drumstick-shaped, with straight top, extending to ventral margin of valva. Saccus broad, V-shaped, strongly sclerotised. Aedeagus straight; caecum narrow ~ 1/7
|
||
<normalizedToken id="32595E0E037C08FF30654F460019D377" originalValue="×">x</normalizedToken>
|
||
aedeagus length; carina long and thin serrated; vesica membranous, with short thin cornutus.
|
||
<emphasis id="98A84EE2147F6A2A9C546CB493584EEF" bold="true" italics="true" pageId="0" pageNumber="35">Female genitalia</emphasis>
|
||
(Fig.
|
||
<figureCitation id="79A9BA6251C07176C51EAFC32429F4AB" captionStart="Figures 19–27" captionStartId="F3" captionText="Figures 19 - 27. Hermonassa spp., male (19 - 22) and female (23 - 27) genitalia 19 H. conusa sp. nov., holotype, genit. prep. hhl- 3492 - 1 20 H. renifera, paratype (after Chen 1991) 21 H. shizukoae, holotype, India (after Sugi 1995) 22 H. shizukoae, Nepal (after Kovacs et al. 2018) 23 H. nigricans, paratype, genit. prep. hhl- 3493 - 2 24 H. anthracina, China, (after Kovacs et al. 2018) 25 H. kalamantra, Nepal (after Kovacs et al. 2018) 26 H. conusa sp. nov., paratype, China, genit. prep. hhl- 3491 - 2 27 H. shizukoae, paratype, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures19-27" httpUri="https://binary.pensoft.net/fig/903265" pageId="0" pageNumber="35">26</figureCitation>
|
||
). Papillae anales broad, slightly sclerotised. Apophyses anteriores slender, ~ 1/3
|
||
<normalizedToken id="D38CC4C1CA8AEB8A6A6F781CC6D9FC42" originalValue="×">x</normalizedToken>
|
||
length of apophyses posteriores. Ostium flat, antevaginal plate almost rounded-triangular with central narrow cut, strongly sclerotised. Ductus bursae broad, flat, with anterior half partially sclerotised, with two irregular plates. Corpus bursae long and swollen, ~ 6
|
||
<normalizedToken id="92D34ABA2E99936762A6DFE30A2C8E85" originalValue="×">x</normalizedToken>
|
||
length of ductus bursae, densely wrinkled, with four slender signa bands of different in lengths.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4A78DDAB09DC888C32D42DB271C7C40F" pageId="0" pageNumber="35" type="etymology">
|
||
<paragraph id="495977524C253C3A8F3C82A79910AE06" pageId="0" pageNumber="35">Etymology.</paragraph>
|
||
<paragraph id="03748FA17AD803CC3C3CE42989DE5977" pageId="0" pageNumber="35">The species name refers to the large, sclerotised, conical extension on the base of the juxta in the male genitalia of the new species.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEEF42A73A2022986DF8A77E2FF3A708" pageId="0" pageNumber="35" type="distribution">
|
||
<paragraph id="C5CD67C8A4FCE88CC6A8AA092768CC3E" pageId="0" pageNumber="35">Distribution and biology.</paragraph>
|
||
<paragraph id="71C41B136F2CD71C63EEE20416AD9504" pageId="0" pageNumber="35">
|
||
<taxonomicName id="D08DC2798D8836CA7B8A0E0AC0CD5AD2" authorityName="Gao, Han & Kononenko" authorityYear="2023" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa conusa" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="conusa">
|
||
<emphasis id="5D74DE9B410CA62A0DABA1079FAD9F49" italics="true" pageId="0" pageNumber="35">Hermonassa conusa</emphasis>
|
||
</taxonomicName>
|
||
is known only from Linzhi mountain range in Aut. Reg. Xizang, in southwest China. The species occurs in grassland meadows in the coniferous forest at an altitudinal range of 3150-3650 m. Adults have been collected between the end of July and the end of August.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |