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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.284.3983" ID-PMC="PMC3677377" ID-Pensoft-Pub="1313-2970-284-1" ID-Pensoft-UUID="192EFF970E2BFFDF3161FFE53C4BFFAA" ID-PubMed="23794843" ID-Zenodo-Dep="578323" ID-ZooBank="33B15A74746D4A82A865EA1E7E55A9BB" ModsDocID="1313-2970-284-1" checkinTime="1451247469560" checkinUser="pensoft" docAuthor="Shi, Hongliang, Zhou, Hongzhang &amp; Liang, Hongbin" docDate="2013" docId="A1144401BB525A8993EE5A4420834296" docLanguage="en" docName="ZooKeys 284: 1-129" docOrigin="ZooKeys 284" docPubDate="2013-04-04" docSource="http://dx.doi.org/10.3897/zookeys.284.3983" docTitle="Anchista fenestrata subsp. fenestrata fenestrata (Schmidt-Goebel 1846" docType="treatment" docVersion="3" id="192EFF970E2BFFDF3161FFE53C4BFFAA" lastPageNumber="26" masterDocId="192EFF970E2BFFDF3161FFE53C4BFFAA" masterDocTitle="Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera" masterLastPageNumber="129" masterPageNumber="1" pageNumber="24" updateTime="1668155586682" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shi, Hongliang</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; Graduate School of Chinese Academy of Sciences, Beijing 100039, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhou, Hongzhang</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liang, Hongbin</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
<mods:nameIdentifier type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-04-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>284</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>129</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.284.3983</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.284.3983</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-284-1</mods:identifier>
<mods:identifier type="ZooBank">33B15A74746D4A82A865EA1E7E55A9BB</mods:identifier>
<mods:identifier type="Pensoft-UUID">192EFF970E2BFFDF3161FFE53C4BFFAA</mods:identifier>
<mods:identifier type="Zenodo-Dep">578323</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182235611" LSID="urn:lsid:plazi:treatment:A1144401BB525A8993EE5A4420834296" httpUri="http://treatment.plazi.org/id/A1144401BB525A8993EE5A4420834296" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
<subSubSection pageId="23" pageNumber="24" type="nomenclature">
<paragraph pageId="23" pageNumber="24">
<taxonomicName LSID="A1144401-BB52-5A89-93EE-5A4420834296" authority="fenestrata (Schmidt-Goebel, 1846)" authorityName="fenestrata (Schmidt-Goebel" authorityYear="1846" baseAuthorityName="Schmidt-Göbel" baseAuthorityYear="1846" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Anchista fenestrata subsp. fenestrata" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="subSpecies" species="fenestrata" subSpecies="fenestrata">
Anchista fenestrata fenestrata (
<normalizedToken originalValue="Schmidt-Göbel">Schmidt-Goebel</normalizedToken>
, 1846)
</taxonomicName>
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Habitus, scale bars = 2.0 mm: 1 Paraphaea binotata (Dejean), a specimen from Yunnan 2 Paraphaea formosana (Jedlicka), a specimen from Taiwan 3 Paraphaea minor sp. n., a paratype from Hainan 4 Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir from India 5 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 6 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal." httpUri="https://binary.pensoft.net/fig/29670" pageId="23" pageNumber="24">Habitus: Figs 5, 6</figureCitation>
<figureCitation captionStart="Figures 37–42" captionStartId="F7" captionText="Figures 37 - 42. Type materials, scale bars = 2.0 mm: 37 Lectotype of Lebia brunnea Wiedemann [= Anchista brunnea (Wiedemann)] 38 Lectotype of Anchista picea Chaudoir [= Anchista brunnea (Wiedemann)] 39 Holotype of Plochionus fenestratus Schmidt-Goebel [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 40 Holotype of Anchista glabra Chaudoir [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 41 Lectotype of Anchista subpubescens Chaudoir [= Anchista fenestrata subpubescens Chaudoir] 42 Lectotype of Anchista nubila Andrewes." httpUri="https://binary.pensoft.net/fig/29680" pageId="23" pageNumber="24">39, 40</figureCitation>
<figureCitation captionStart="Figure 71" captionStartId="F16" captionText="Figure 71. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Chota Nagpore." httpUri="https://binary.pensoft.net/fig/29689" pageId="23" pageNumber="24">male genitalia: Figs 71</figureCitation>
<figureCitation captionStart="Figures 95–110" captionStartId="F40" captionText="Figures 95 - 110. Left and right parameres of male genitalia, scale bars = 0.2 mm: 95 Paraphaea binotata (Dejean), a specimen from Guangxi 96 Paraphaea minor sp. n., holotype 97 Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir 98 Anchista pilosa sp. n., holotype 99 Metallanchista laticollis sp. n., holotype 100 Physodera amplicollis van de Poll, a specimen from Taiwan 101 Diamella cupreomicans (Oberthuer), a specimen from Yunnan 102 Diamella arrowi (Jedlicka), holotype 103 Allocota aurata (Bates), a specimen from Tonkin 104 Allocota aurata (Bates), a specimen from Shaanxi 105 Allocota bicolor sp. n., holotype 106 Lachnoderma asperum Bates, a specimen from Taiwan 107 Dasiosoma ivorense Basilewsky, a paratype from Cote d'Ivoire 108 Dasiosoma quadraticolle sp. n., holotype 109 Orionella kathmanduensis (Kirschenhofer), a specimen from Nepal 110 Endynomena pradieri (Fairmaire), a specimen from Fujian." httpUri="https://binary.pensoft.net/fig/29716" pageId="23" pageNumber="24">96</figureCitation>
<figureCitation captionStart="Figures 111–125" captionStartId="F41" captionText="Figures 111 - 125. Left ovipositors of females, ventral view, scale bars = 0.2 mm: 111 Paraphaea binotata (Dejean), a specimen from Guangxi 112 Paraphaea minor sp. n., a paratype from Hainan 113 Paraphaea philippinensis (Jedlicka), holotype 114 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 115 Physodera dejeani Eschscholtz, a specimen from Yunnan 116 Physodera eschscholtzii Parry, a specimen from Yunnan 117 Diamella cupreomicans (Oberthuer), a specimen from Yunnan 118 Allocota aurata (Bates), a specimen from Shaanxi 119 Allocota bicolor sp. n., a paratype from Guangdong 120 Lachnoderma asperum Bates, a specimen from Zhejiang 121 Dasiosoma bellum (Habu), a specimen from Guangdong 122 Dasiosoma maindroni (Tian &amp; Deuve), a specimen from Tonkin 123 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 124 Orionella lewisii (Bates), a specimen from Zhejiang 125 Endynomena pradieri (Fairmaire), a specimen from Taiwan." httpUri="https://binary.pensoft.net/fig/29719" pageId="23" pageNumber="24">female genitalia: Figs 114</figureCitation>
<figureCitation captionStart="Figures 126–131" captionStartId="F42" captionText="Figures 126 - 131. Internal reproductive system of females, scale bars = 0.2 mm: 126 Paraphaea binotata (Dejean), a specimen from Guangxi 127 Paraphaea minor sp. n., a paratype from Hainan 128 Paraphaea philippinensis (Jedlicka), holotype 129 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 130 Physodera dejeani Eschscholtz, a specimen from Yunnan 131 Physodera eschscholtzii Parry, a specimen from Hainan." httpUri="https://binary.pensoft.net/fig/29720" pageId="23" pageNumber="24">129</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="24" type="reference_group">
<paragraph pageId="23" pageNumber="24">
<taxonomicName authority="fenestrata" authorityName="fenestrata (Schmidt-Goebel" authorityYear="1846" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="subSpecies" species="fenestrata" subSpecies="fenestrata">Anchista fenestrata fenestrata</taxonomicName>
<bibRefCitation author="Schmidt-Goebel, HM" journalOrPublisher="The Transactions of the Entomological Society of London" pageId="88" pageNumber="89" refId="B80" refString="Schmidt-Goebel, HM, 1846. Faunula Coleopterum Birmaniae, 94 pp, pl. 1-3." title="Faunula Coleopterum Birmaniae, 94 pp, pl. 1 - 3." year="1846">
<normalizedToken originalValue="Schmidt-Göbel">Schmidt-Goebel</normalizedToken>
1846
</bibRefCitation>
: 42 (original:
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Hexapoda" family="Carabidae" genus="Plochionus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="23" pageNumber="24">Plochionus</emphasis>
</taxonomicName>
; type locality: Birma; holotype deposited in MNPC);
<bibRefCitation author="Chaudoir, BM" journalOrPublisher="Annales de Societe Entomologique de Belgique" pageId="86" pageNumber="87" pagination="97 - 204" refId="B34" refString="Chaudoir, BM, 1872. Monographie des Callidides. Annales de Societe Entomologique de Belgique 16: 97 - 204" title="Monographie des Callidides." volume="16" year="1872">Chaudoir 1872</bibRefCitation>
: 168 (excluded from
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Hexapoda" family="Carabidae" genus="Plochionus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="23" pageNumber="24">Plochionus</emphasis>
</taxonomicName>
);
<bibRefCitation author="Bates, HW" journalOrPublisher="The Transactions of the Entomological Society of London" pageId="85" pageNumber="86" refId="B27" refString="Bates, HW, 1892. Viaggio di Leonardo Fea in Birmania e Regioni Vicini XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova (2) 12: 267-428." title="Viaggio di Leonardo Fea in Birmania e Regioni Vicini XLIV. List of the Carabidae. Annali del Museo Civico di Storia Naturale di Genova (2) 12: 267 - 428." year="1892">Bates 1892</bibRefCitation>
: 424 (
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Hexapoda" family="Carabidae" genus="Endynomena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="23" pageNumber="24">Endynomena</emphasis>
</taxonomicName>
);
<bibRefCitation author="Andrewes, HE" journalOrPublisher="Transactions of the Entomological Society of London" pageId="84" pageNumber="85" pagination="1 - 63" refId="B3" refString="Andrewes, HE, 1923. On the types of Carabidae described by Schmidt-Goebel in his Faunula Coleopterorum Birmaniae. Transactions of the Entomological Society of London 1923: 1 - 63" title="On the types of Carabidae described by Schmidt-Goebel in his Faunula Coleopterorum Birmaniae." volume="1923" year="1923">Andrewes 1923</bibRefCitation>
: 20 (
<taxonomicName authorityName="Nietner" authorityYear="1856" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="23" pageNumber="24">Anchista</emphasis>
</taxonomicName>
);
<bibRefCitation author="Andrewes, HE" journalOrPublisher="Zoologische Mededeelingen, Leiden" pageId="85" pageNumber="86" refId="B14" refString="Andrewes, HE, 1930d. Catalogue of Indian Insects, Part 18-Carabidae. Calcutta. Government of India Central Publication Branch, 389 pp." title="Catalogue of Indian Insects, Part 18 - Carabidae. Calcutta. Government of India Central Publication Branch, 389 pp." year="1930 d">Andrewes 1930d</bibRefCitation>
: 23 (
<taxonomicName authorityName="Nietner" authorityYear="1856" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="23" pageNumber="24">Anchista</emphasis>
</taxonomicName>
, catalogue);
<bibRefCitation author="Csiki, E" editor="Junk, W" journalOrPublisher="Volumen III. Carabidae III. Berlin" pageId="86" pageNumber="87" pagination="1279 - 1598" refId="B37" refString="Csiki, E, 1932. Carabidae: Harpalinae VII (Pars 124). In: Junk, W, Schenkling, S, Eds., Coleopterorum catalogus. Volumen III. Carabidae III. Berlin: 1279 - 1598" title="Carabidae: Harpalinae VII (Pars 124)." volumeTitle="Coleopterorum catalogus." year="1932">Csiki 1932</bibRefCitation>
: 1456 (
<taxonomicName authorityName="Nietner" authorityYear="1856" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="23" pageNumber="24">Anchista</emphasis>
</taxonomicName>
, catalogue);
<bibRefCitation author="Jedlicka, A" journalOrPublisher="Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden" pageId="87" pageNumber="88" pagination="269 - 579" refId="B58" refString="Jedlicka, A, 1963. Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden 28: 269 - 579" title="Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae)." volume="28" year="1963">
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
1963
</bibRefCitation>
: 449 (
<taxonomicName authorityName="Nietner" authorityYear="1856" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="23" pageNumber="24">Anchista</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph pageId="23" pageNumber="24">
<taxonomicName authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="glabra">Anchista glabra</taxonomicName>
<bibRefCitation author="Chaudoir, BM" journalOrPublisher="Bulletin de la Societe Imperiale des Naturalistes de Moscou" pageId="86" pageNumber="87" pagination="188 - 268" refId="B35" refString="Chaudoir, BM, 1877. Genres nouveaux et especes inedites de la famille des Carabiques. Bulletin de la Societe Imperiale des Naturalistes de Moscou 52 (1): 188 - 268" title="Genres nouveaux et especes inedites de la famille des Carabiques." volume="52" year="1877">Chaudoir 1877</bibRefCitation>
: 237 (type locality:
<normalizedToken originalValue="Pondichéry">Pondichery</normalizedToken>
; holotype deposited in MNHN);
<bibRefCitation author="Andrewes, HE" journalOrPublisher="Zoologische Mededeelingen, Leiden" pageId="85" pageNumber="86" refId="B14" refString="Andrewes, HE, 1930d. Catalogue of Indian Insects, Part 18-Carabidae. Calcutta. Government of India Central Publication Branch, 389 pp." title="Catalogue of Indian Insects, Part 18 - Carabidae. Calcutta. Government of India Central Publication Branch, 389 pp." year="1930 d">Andrewes 1930d</bibRefCitation>
: 23 (
<taxonomicName authorityName="Nietner" authorityYear="1856" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="23" pageNumber="24">Anchista</emphasis>
</taxonomicName>
; catalogue);
<bibRefCitation author="Csiki, E" editor="Junk, W" journalOrPublisher="Volumen III. Carabidae III. Berlin" pageId="86" pageNumber="87" pagination="1279 - 1598" refId="B37" refString="Csiki, E, 1932. Carabidae: Harpalinae VII (Pars 124). In: Junk, W, Schenkling, S, Eds., Coleopterorum catalogus. Volumen III. Carabidae III. Berlin: 1279 - 1598" title="Carabidae: Harpalinae VII (Pars 124)." volumeTitle="Coleopterorum catalogus." year="1932">Csiki 1932</bibRefCitation>
: 1456 (
<taxonomicName authorityName="Nietner" authorityYear="1856" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="23" pageNumber="24">Anchista</emphasis>
</taxonomicName>
, catalogue).
<emphasis bold="true" pageId="23" pageNumber="24">syn. n.</emphasis>
[Synonym]
</paragraph>
<paragraph pageId="23" pageNumber="24">
<taxonomicName authorityName="Kirschenhofer" authorityYear="1994" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="nepalensis">Anchista nepalensis</taxonomicName>
<bibRefCitation author="Kirschenhofer, E" journalOrPublisher="Linzer Biologische Beitraege" pageId="87" pageNumber="88" pagination="999 - 1067" refId="B61" refString="Kirschenhofer, E, 1994. Neue und wenig bekannte Carabidae aus der palaarktischen und orientalischen Region (Col., Carabidae, Lebiinae, Odacanthinae, Brachininae, Panagaeinae). Linzer Biologische Beitraege 26 (2): 999 - 1067" title="Neue und wenig bekannte Carabidae aus der palaarktischen und orientalischen Region (Col., Carabidae, Lebiinae, Odacanthinae, Brachininae, Panagaeinae)." volume="26" year="1994">Kirschenhofer 1994</bibRefCitation>
: 1005 (type locality: Ostnepal; holotype deposited in NHMW).
<emphasis bold="true" pageId="23" pageNumber="24">syn. n.</emphasis>
</paragraph>
</subSubSection>
<subSubSection lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="type examined">
<paragraph pageId="23" pageNumber="24">Type examined.</paragraph>
<paragraph lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<emphasis bold="true" pageId="23" pageNumber="24">Holotype</emphasis>
of
<taxonomicName authorityName="Schmidt-Goebel" authorityYear="1846" class="Hexapoda" family="Carabidae" genus="Plochionus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="fenestratus">
<emphasis italics="true" pageId="23" pageNumber="24">Plochionus fenestratus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Schmidt-Göbel">Schmidt-Goebel</normalizedToken>
, monotypy (NMPC): male, body length = 7.7 mm, board mounted, &quot;MUS. PRAGENSE / COLL. HELFER&quot;;
<pageBreakToken pageId="24" pageNumber="25" start="start">&quot;</pageBreakToken>
<emphasis italics="true" pageId="24" pageNumber="25">Birma / Helfer</emphasis>
.&quot;; &quot;Typus! teste / Dr. J. Obenberger&quot; [red label]; &quot;
<emphasis italics="true" pageId="24" pageNumber="25">fenestratus</emphasis>
/
<emphasis italics="true" pageId="24" pageNumber="25">Sch. g</emphasis>
. /COL. HELFER&quot;[Fig. 39].
<emphasis bold="true" pageId="24" pageNumber="25">Holotype</emphasis>
of
<taxonomicName authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="glabra">
<emphasis italics="true" pageId="24" pageNumber="25">Anchista glabra</emphasis>
</taxonomicName>
Chaudoir, monotypy (MNHN): female, body length = 8.2 mm, pin mounted, &quot;Ex Musaeo / Chaudoir&quot; [red letter];
<normalizedToken originalValue="“TYPE”">&quot;TYPE&quot;</normalizedToken>
[red label]; &quot;Museum Paris / 1952 / Coll. R.
<normalizedToken originalValue="Oberthür”">Oberthuer&quot;</normalizedToken>
; &quot;
<emphasis italics="true" pageId="24" pageNumber="25">glabra Chaudoir</emphasis>
/
<emphasis italics="true" pageId="24" pageNumber="25">Coromandel</emphasis>
/
<emphasis italics="true" pageId="24" pageNumber="25">
<normalizedToken originalValue="Pondichéry">Pondichery</normalizedToken>
</emphasis>
/
<emphasis italics="true" pageId="24" pageNumber="25">Guerin</emphasis>
&quot; [box label but pinned under specimen]; &quot;HOLOTYPE ♀/ Anchista glabra / Chaudoir, 1877 / Des. SHI H. L. 2011&quot; [red label][Fig. 40].
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="25" type="notes on types">
<paragraph pageId="24" pageNumber="25">Notes on types.</paragraph>
<paragraph pageId="24" pageNumber="25">
<taxonomicName authorityName="Schmidt-Goebel" authorityYear="1846" class="Hexapoda" family="Carabidae" genus="Plochionus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fenestratus">
<emphasis bold="true" italics="true" pageId="24" pageNumber="25">Plochionus fenestratus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Schmidt-Göbel">Schmidt-Goebel</normalizedToken>
: One female was mentioned in the original description. In NMPC boxes of
<normalizedToken originalValue="Schmidt-Göbel’s">Schmidt-Goebel's</normalizedToken>
collection, we only found a male determined by him (
<figureCitation captionStart="Figures 37–42" captionStartId="F7" captionText="Figures 37 - 42. Type materials, scale bars = 2.0 mm: 37 Lectotype of Lebia brunnea Wiedemann [= Anchista brunnea (Wiedemann)] 38 Lectotype of Anchista picea Chaudoir [= Anchista brunnea (Wiedemann)] 39 Holotype of Plochionus fenestratus Schmidt-Goebel [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 40 Holotype of Anchista glabra Chaudoir [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 41 Lectotype of Anchista subpubescens Chaudoir [= Anchista fenestrata subpubescens Chaudoir] 42 Lectotype of Anchista nubila Andrewes." httpUri="https://binary.pensoft.net/fig/29680" pageId="24" pageNumber="25">Fig. 39</figureCitation>
). This male is the holotype, and
<normalizedToken originalValue="Schmidt-Göbel">Schmidt-Goebel</normalizedToken>
determined the sex erroneously. In the label or original description, no precise locality for this specimen was mentioned. From the record of
<normalizedToken originalValue="Helfer’s">Helfer's</normalizedToken>
expedition (
<bibRefCitation author="Reichardt, HW" journalOrPublisher="The Transactions of the Entomological Society of London" pageId="88" pageNumber="89" refId="B79" refString="Reichardt, HW, 1880. Artikel &quot;Helfer, Johann Wilhelm&quot; in: Allgemeine Deutsche Biographie, herausgegeben von der Historischen Kommission bei der Bayerischen Akademie der Wissenschaften 11(1880), 687." title="Artikel &quot; Helfer, Johann Wilhelm &quot; in: Allgemeine Deutsche Biographie, herausgegeben von der Historischen Kommission bei der Bayerischen Akademie der Wissenschaften 11 (1880), 687." year="1880">Reichardt 1880</bibRefCitation>
), it can be inferred that this specimen was collected by Helfer from Tenasserim in Burma (Myanmar).
</paragraph>
<paragraph pageId="24" pageNumber="25">
<taxonomicName authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="glabra">
<emphasis bold="true" italics="true" pageId="24" pageNumber="25">Anchista glabra</emphasis>
</taxonomicName>
Chaudoir: Only one specimen was mentioned in the original description. Therefore the one in
<normalizedToken originalValue="Chaudoir’s">Chaudoir's</normalizedToken>
collection is the holotype (
<figureCitation captionStart="Figures 37–42" captionStartId="F7" captionText="Figures 37 - 42. Type materials, scale bars = 2.0 mm: 37 Lectotype of Lebia brunnea Wiedemann [= Anchista brunnea (Wiedemann)] 38 Lectotype of Anchista picea Chaudoir [= Anchista brunnea (Wiedemann)] 39 Holotype of Plochionus fenestratus Schmidt-Goebel [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 40 Holotype of Anchista glabra Chaudoir [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 41 Lectotype of Anchista subpubescens Chaudoir [= Anchista fenestrata subpubescens Chaudoir] 42 Lectotype of Anchista nubila Andrewes." httpUri="https://binary.pensoft.net/fig/29680" pageId="24" pageNumber="25">Fig. 40</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="25" type="non-type material examined">
<paragraph pageId="24" pageNumber="25">Non-type material examined</paragraph>
<paragraph pageId="24" pageNumber="25">
(Total 25 specimens).
<emphasis bold="true" pageId="24" pageNumber="25">India</emphasis>
: 1 male and 1 female (MNHN), &quot;S. India, Nedungadu, P. S. Nathan, 1936&quot;. 1 male (MNHN), &quot;Ramnad, Hindonstan&quot;. 17 specimens (MNHN), &quot;Chota Nagpore, Nowatoli, R. P. Cardon, XI-XII 1896&quot;[Fig. 71]. 2 specimens (NHMB), &quot;India: Orissa state, Similipal N. P. Lulung
<geoCoordinate degrees="21" direction="north" minutes="56" orientation="latitude" precision="925" value="21.933332">21°56'N</geoCoordinate>
,
<geoCoordinate degrees="86" direction="east" minutes="32" orientation="longitude" precision="925" value="86.53333">86°32'E</geoCoordinate>
, 25.v.-13.vi.1998, Karel &amp; Simon Majer leg&quot;.
<emphasis bold="true" pageId="24" pageNumber="25">Nepal</emphasis>
: 2 males and 1 female (NHMB), &quot;E-Nepal, Koshi, M. Brancucci&quot;; &quot;Simraghat, 500m, 13.VI.85&quot;[Figs 5, 6, 114, 129].
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="25" type="diagnosis">
<paragraph pageId="24" pageNumber="25">Diagnosis.</paragraph>
<paragraph pageId="24" pageNumber="25">Head and pronotum glabrous; pronotum widest at middle, lateral margins slightly angulate in middle; elytra distinctly bicolored; 3rd interval of elytra with three to five setigerous pores, 5th interval with two to four setigerous pores near base.</paragraph>
<paragraph pageId="24" pageNumber="25">
By the glabrous surface and number of setigerous pores on 3rd and 5th intervals, this species can be readily distinguished from most allied species, except
<taxonomicName authorityName="Andrewes" authorityYear="1931" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="nubila">
<emphasis italics="true" pageId="24" pageNumber="25">Anchista nubila</emphasis>
</taxonomicName>
. Diagnosis between these two species is presented in the key.
</paragraph>
<paragraph pageId="24" pageNumber="25">
Male genitalia characters show this species to be very similar to
<taxonomicName baseAuthorityName="Wiedemann" baseAuthorityYear="1823" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis italics="true" pageId="24" pageNumber="25">Anchista brunnea</emphasis>
</taxonomicName>
, but the latter has the apical lamella wider, internal sac without strongly scaled area near middle, and small sclerite near to the base. From the external characters, these two species can be readily distinguished by the different number of setigerous pores on 3rd and 5th intervals.
</paragraph>
</subSubSection>
<subSubSection lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
<paragraph pageId="24" pageNumber="25">Description.</paragraph>
<paragraph lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
Body length 7.7-8.5 mm; dorsal side yellowish to reddish yellow, pronotum lateral explanate area paler; antennae yellowish brown to reddish brown, 1st antennomere slightly paler. Elytra with variable bicolored pattern: darker color brownish to black while the paler parts yellowish to reddish yellow; sometimes elytra mostly dark, but left lateral margin and outer part of apex paler, 2nd to 6th intervals with an elongate pale patch occupying basal three-fifths as in the holotype of
<taxonomicName authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="glabra">
<emphasis italics="true" pageId="24" pageNumber="25">Anchista glabra</emphasis>
</taxonomicName>
Chaudoir (
<figureCitation captionStart="Figures 37–42" captionStartId="F7" captionText="Figures 37 - 42. Type materials, scale bars = 2.0 mm: 37 Lectotype of Lebia brunnea Wiedemann [= Anchista brunnea (Wiedemann)] 38 Lectotype of Anchista picea Chaudoir [= Anchista brunnea (Wiedemann)] 39 Holotype of Plochionus fenestratus Schmidt-Goebel [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 40 Holotype of Anchista glabra Chaudoir [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 41 Lectotype of Anchista subpubescens Chaudoir [= Anchista fenestrata subpubescens Chaudoir] 42 Lectotype of Anchista nubila Andrewes." httpUri="https://binary.pensoft.net/fig/29680" pageId="24" pageNumber="25">Fig. 40</figureCitation>
); or elytra mostly pale, with a small ovaloid or sub-diamondoid dark patch near apex as in some specimens from Nepal (
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Habitus, scale bars = 2.0 mm: 1 Paraphaea binotata (Dejean), a specimen from Yunnan 2 Paraphaea formosana (Jedlicka), a specimen from Taiwan 3 Paraphaea minor sp. n., a paratype from Hainan 4 Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir from India 5 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 6 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal." httpUri="https://binary.pensoft.net/fig/29670" pageId="24" pageNumber="25">Fig. 6</figureCitation>
); but in most specimens, elytra with pattern between these two extremities (
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Habitus, scale bars = 2.0 mm: 1 Paraphaea binotata (Dejean), a specimen from Yunnan 2 Paraphaea formosana (Jedlicka), a specimen from Taiwan 3 Paraphaea minor sp. n., a paratype from Hainan 4 Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir from India 5 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 6 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal." httpUri="https://binary.pensoft.net/fig/29670" pageId="24" pageNumber="25">Fig. 5</figureCitation>
). Ventral side yellowish to reddish yellow; femora yellow, tarsi and tibiae yellowish brown.
<emphasis bold="true" pageId="24" pageNumber="25">Head</emphasis>
<pageBreakToken pageId="25" pageNumber="26" start="start">glabrous</pageBreakToken>
, without microsculpture, vertex with a few fine punctures.
<emphasis bold="true" pageId="25" pageNumber="26">Pronotum</emphasis>
glabrous, cordiform, widest slightly before middle; ratio PW/PL 1.41 to 1.51; pronotal base briefly but distinctly lobed; disc moderately convex, microsculpture indistinct, without punctures; front angles with a few moderately long setae; lateral margins slightly angulate in middle, strongly sinuate before hind angles; lateral explanate area wide and even, with a few coarse punctures; hind angles sharp, rectangular or slightly acute, with a few very short accessory setae; basal foveae moderately deep, with a few punctures; median line distinct, strongly depressed and punctate anteriorly and subbasally.
<emphasis bold="true" pageId="25" pageNumber="26">Elytra</emphasis>
with striae shallow but distinct, finely punctate; intervals slightly convex, without accessory setae, finely and sparsely punctate; microsculpture distinct, isodiametric; 3rd interval with three to five setigerous pores, the basal one usually placed at basal one-third approximately, adjacent to 3rd stria or on middle of interval, the rest adjacent to 2nd stria; 5th interval with two to four setigerous pores, the basal one near interval base, the last one usually at middle, all pores usually in center of 5th interval or slightly adjacent to 5th stria; basal margination nearly complete. Male with adhesive hairs rudimentary (two rows, weakly present near apex) on first three mesotarsomeres.
<emphasis bold="true" pageId="25" pageNumber="26">Male genitalia</emphasis>
with median lobe of aedeagus straight and flat, slightly expanded to apex; left lateral margin slightly curved in middle; apical orifice opened dorsally, basal margin of apical orifice setose; apical lamella placed on right side, flat, slightly elongate and rounded in dorsal view, slightly curved up dorsally in lateral view; internal sac with a small strongly sclerotized sclerite near base, two weakly sclerotized elongate and sinuate sclerites near apical orifice, a small and strongly scaled area present near middle of median lobe, internal sac in basal half sparsely and finely setose ventrally (
<figureCitation captionStart="Figure 71" captionStartId="F16" captionText="Figure 71. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Chota Nagpore." httpUri="https://binary.pensoft.net/fig/29689" pageId="25" pageNumber="26">Fig. 71</figureCitation>
).
<emphasis bold="true" pageId="25" pageNumber="26">Female genitalia</emphasis>
. Spermathecal gland inserted near apex of spermatheca, strongly expanded but not bent at base; spermatheca gradually expanded to apex, apex globular with a small papilla; ring-sculpture very faint, only present on subapical part of spermatheca (
<figureCitation captionStart="Figures 126–131" captionStartId="F42" captionText="Figures 126 - 131. Internal reproductive system of females, scale bars = 0.2 mm: 126 Paraphaea binotata (Dejean), a specimen from Guangxi 127 Paraphaea minor sp. n., a paratype from Hainan 128 Paraphaea philippinensis (Jedlicka), holotype 129 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 130 Physodera dejeani Eschscholtz, a specimen from Yunnan 131 Physodera eschscholtzii Parry, a specimen from Hainan." httpUri="https://binary.pensoft.net/fig/29720" pageId="25" pageNumber="26">Fig. 129</figureCitation>
). Apical segment of ovipositor scimitar-form, inner margin gradually curved; length nearly three times basal width; inner margin setose in apical half; apex rounded and slightly projected to outer side, with membranous extension slightly widened.
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="26" type="distribution">
<paragraph pageId="25" pageNumber="26">Distribution</paragraph>
<paragraph pageId="25" pageNumber="26">
(
<figureCitation captionStart="Map 3" captionStartId="F47" captionText="Map 3. Known distribution of genus Anchista: ◆ Anchista brunnea (Wiedemann) ★ Anchista nubila Andrewes ● Anchista pilosa sp. n. △ Anchista fenestrata fenestrata (Schmidt-Goebel) ▼ Anchista fenestrata subpubescens Chaudoir." httpUri="https://binary.pensoft.net/fig/29725" pageId="25" pageNumber="26">Map 3</figureCitation>
). India, Myanmar, Nepal.
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="26" type="notes on synonym">
<paragraph pageId="25" pageNumber="26">Notes on synonym.</paragraph>
<paragraph pageId="25" pageNumber="26">
We dissected males from Nedungadu (close to type locality of
<taxonomicName authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="glabra">
<emphasis italics="true" pageId="25" pageNumber="26">Anchista glabra</emphasis>
</taxonomicName>
), Chota Nagpore, Nepal Koshi (type locality of
<taxonomicName authorityName="Kirschenhofer" authorityYear="1994" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="nepalensis">
<emphasis italics="true" pageId="25" pageNumber="26">Anchista nepalensis</emphasis>
</taxonomicName>
), and the holotype of
<taxonomicName baseAuthorityName="Schmidt-Goebel" baseAuthorityYear="1846" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fenestrata">
<emphasis italics="true" pageId="25" pageNumber="26">Anchista fenestrata</emphasis>
</taxonomicName>
. There is no significant difference among these specimens from different localities, except for elytral pattern variation. We therefore herein synonymize
<taxonomicName authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="glabra">
<emphasis italics="true" pageId="25" pageNumber="26">Anchista glabra</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Kirschenhofer" authorityYear="1994" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="nepalensis">
<emphasis italics="true" pageId="25" pageNumber="26">Anchista nepalensis</emphasis>
</taxonomicName>
with
<taxonomicName baseAuthorityName="Schmidt-Goebel" baseAuthorityYear="1846" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fenestrata">
<emphasis italics="true" pageId="25" pageNumber="26">Anchista fenestrata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="26" type="geographical variation">
<paragraph pageId="25" pageNumber="26">Geographical variation.</paragraph>
<paragraph pageId="25" pageNumber="26">
In this species, specimens from one locality may be obviously different in elytral pattern. Sometimes, intrapopulational variation ranges even wider than interpopulational. But statistically interpopulational pattern variation exists: individuals from south India usually with dark pattern occupying more area on elytra (e. g. in the holotype of
<taxonomicName authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="glabra">
<emphasis italics="true" pageId="25" pageNumber="26">Anchista glabra</emphasis>
</taxonomicName>
Chaudoir,
<figureCitation captionStart="Figures 37–42" captionStartId="F7" captionText="Figures 37 - 42. Type materials, scale bars = 2.0 mm: 37 Lectotype of Lebia brunnea Wiedemann [= Anchista brunnea (Wiedemann)] 38 Lectotype of Anchista picea Chaudoir [= Anchista brunnea (Wiedemann)] 39 Holotype of Plochionus fenestratus Schmidt-Goebel [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 40 Holotype of Anchista glabra Chaudoir [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 41 Lectotype of Anchista subpubescens Chaudoir [= Anchista fenestrata subpubescens Chaudoir] 42 Lectotype of Anchista nubila Andrewes." httpUri="https://binary.pensoft.net/fig/29680" pageId="25" pageNumber="26">Fig. 40</figureCitation>
) than those from north India; while some specimens from Nepal have the paler color occupying the largest area, with only a small dark patch near apex (
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Habitus, scale bars = 2.0 mm: 1 Paraphaea binotata (Dejean), a specimen from Yunnan 2 Paraphaea formosana (Jedlicka), a specimen from Taiwan 3 Paraphaea minor sp. n., a paratype from Hainan 4 Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir from India 5 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 6 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal." httpUri="https://binary.pensoft.net/fig/29670" pageId="25" pageNumber="26">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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