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<document id="7BF186A474D8DA9CC3FED43E67FEBB0A" ID-CLB-Dataset="55793" ID-DOI="10.1080/00222933.2013.791936" ID-GBIF-Dataset="6591e69b-631a-4c4d-bf83-db8c70d1ad05" ID-ISSN="1464-5262" ID-Zenodo-Dep="5198102" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628884845165" checkinUser="felipe" docAuthor="Zhou, Yu-Lingzi &amp; Zhou, Hong-Zhang" docDate="2013" docId="A10D4838C35737572630FA4219BCFA34" docLanguage="en" docName="JNatHist.47.45-46.2869-2904.pdf" docOrigin="Journal of Natural History 47 (45 - 46)" docSource="http://dx.doi.org/10.1080/00222933.2013.791936" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Liotesba recticeps Zhou &amp; Zhou 2013, sp. nov." docType="treatment" docVersion="8" lastPageNumber="2882" masterDocId="5D343040C35B37582413FFC11B0DFF84" masterDocTitle="Taxonomy and phylogeny of the genus Liotesba Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae: Xantholinini) with descriptions of four new species" masterLastPageNumber="2904" masterPageNumber="2869" pageNumber="2879" updateTime="1699079816598" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="E90CEEA8612ED5A69160A2199A357691" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9634ABDA92302C250231FD427D1BB00D">
<mods:title id="2F71A87CD84E3C56FA20DA6AC46D9E60">Taxonomy and phylogeny of the genus Liotesba Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae: Xantholinini) with descriptions of four new species</mods:title>
</mods:titleInfo>
<mods:name id="118F216F136D1780C38FE45E69F7EEB5" type="personal">
<mods:role id="4C4D258EB78F9899931FEEC2C5C4E681">
<mods:roleTerm id="11B560DCEDD8A4476A7DD8DFB2921166">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5BB2C8C12AF25E476A2A480CCF2DB619">Zhou, Yu-Lingzi</mods:namePart>
</mods:name>
<mods:name id="E82264D95D47D0962D246B388ABEC1CD" type="personal">
<mods:role id="AAFB9E19068BF8E77C93CB85AF3895CA">
<mods:roleTerm id="7D423448AD95713684F008168A722724">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1EEA3812167CF09AB9483D66AB8E2CD0">Zhou, Hong-Zhang</mods:namePart>
</mods:name>
<mods:typeOfResource id="98D12E379871B18ABC82C2DD67BBAC74">text</mods:typeOfResource>
<mods:relatedItem id="AB84CE95B9D6634CFBE1DB70FE8812B3" type="host">
<mods:titleInfo id="3E6EA810F483AC412117F081433727E7">
<mods:title id="B29F509C25406B2B25A4F05625DB9E6F">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="935EAEE9B2DFD5CE9C6573C8542CAC14">
<mods:date id="5E1D5971EA7B4B5CB97861166F541C27">2013</mods:date>
<mods:detail id="028DFCF467D231A3BBE9DBF2012E8C54" type="pubDate">
<mods:number id="4AE73A94A1B67BF79FCC752F70AA125A">2013-08-08</mods:number>
</mods:detail>
<mods:detail id="93B3741EDC048B2F6AABB6BBB25C5D18" type="volume">
<mods:number id="198D8B44D1B6EE4DF77B5AB757647FA0">47</mods:number>
</mods:detail>
<mods:detail id="AABFAEC262E32B9FE276B6D2EA31F947" type="issue">
<mods:number id="B9ACEEB273D322681895275F3DD439E2">45 - 46</mods:number>
</mods:detail>
<mods:extent id="A4697794F372C56799818FA271E3D59C" unit="page">
<mods:start id="84F601CDB0CBE6D32E2D5758F221D820">2869</mods:start>
<mods:end id="04FC399A405E8E8C318FA789A6612B08">2904</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="7EFBA1BBB365DBAA0C0A0B12DC6DCFB5">
<mods:url id="2E56B95C4C70C2AFAD85017532D9C11D">http://dx.doi.org/10.1080/00222933.2013.791936</mods:url>
</mods:location>
<mods:classification id="4EA0FBF5ADE711E9255984EEF10AAE20">journal article</mods:classification>
<mods:identifier id="D219A82D384E05E77270FED9C5DE6177" type="CLB-Dataset">55793</mods:identifier>
<mods:identifier id="C64A18FF6F695FFFC950A7CDDBC19853" type="DOI">10.1080/00222933.2013.791936</mods:identifier>
<mods:identifier id="9514E29B4694614FD8FBDB198E153D2E" type="GBIF-Dataset">6591e69b-631a-4c4d-bf83-db8c70d1ad05</mods:identifier>
<mods:identifier id="0FC1D2D125F144AA0ABE777ED1347882" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="98D86B03533DD268546901448B83CD47" type="Zenodo-Dep">5198102</mods:identifier>
</mods:mods>
<treatment id="A10D4838C35737572630FA4219BCFA34" ID-DOI="http://doi.org/10.5281/zenodo.5198084" ID-GBIF-Taxon="194873652" ID-Zenodo-Dep="5198084" LSID="urn:lsid:plazi:treatment:A10D4838C35737572630FA4219BCFA34" httpUri="http://treatment.plazi.org/id/A10D4838C35737572630FA4219BCFA34" lastPageId="15" lastPageNumber="2882" pageId="12" pageNumber="2879">
<subSubSection id="61BEAAA5C35737542630FA421849FA1F" box="[547,836,1411,1435]" pageId="12" pageNumber="2879" type="nomenclature">
<paragraph id="291BF92EC35737542630FA421849FA1F" blockId="12.[547,836,1411,1467]" box="[547,836,1411,1435]" pageId="12" pageNumber="2879">
<heading id="72534E42C35737542630FA421849FA1F" box="[547,836,1411,1435]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="2879" reason="2">
<taxonomicName id="EEA482ADC35737542630FA4219E4FA1F" ID-CoL="3V87R" authority="Zhou &amp; Zhou, 2013" authorityName="Zhou &amp; Zhou" authorityYear="2013" box="[547,745,1411,1435]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2879" phylum="Arthropoda" rank="species" species="recticeps" status="sp. nov.">
<emphasis id="1BD0253CC35737542630FA4219E4FA1F" bold="true" box="[547,745,1411,1435]" italics="true" pageId="12" pageNumber="2879">Liotesba recticeps</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00E39847C357375426E3FA451849FA1F" box="[752,836,1412,1435]" pageId="12" pageNumber="2879" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C3573754264CFA62180AFA3F" box="[607,775,1443,1467]" pageId="12" pageNumber="2879" type="description">
<paragraph id="291BF92EC3573754264CFA62180AFA3F" blockId="12.[547,836,1411,1467]" box="[607,775,1443,1467]" pageId="12" pageNumber="2879">
(
<figureCitation id="B19FE5ABC35737542674FA62180DFA3F" box="[615,768,1443,1467]" captionStart="Figure 2" captionStartId="13.[143,210,1458,1480]" captionTargetBox="[144,1104,161,1425]" captionTargetId="figure-6@13.[144,1104,161,1425]" captionTargetPageId="13" captionText="Figure 2. Liotesba recticeps sp. nov., (A) head; (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male pleurites of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198108" httpUri="https://zenodo.org/record/5198108/files/figure.png" pageId="12" pageNumber="2879">Figure 2A–K</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C357375424C0FA231873F9DE" pageId="12" pageNumber="2879" type="materials_examined">
<paragraph id="291BF92EC357375424C0FA231873F9DE" blockId="12.[211,1171,1506,1626]" pageId="12" pageNumber="2879">
<emphasis id="1BD0253CC357375424C0FA231A71FA7E" bold="true" box="[211,380,1506,1530]" pageId="12" pageNumber="2879">Type material.</emphasis>
<materialsCitation id="99CCF373C3573754259FFA231879F99E" ID-GBIF-Occurrence="3757895310" collectingDate="2004-05-23" collectingDateMax="2004-05-26" collectingDateMin="2004-05-23" collectionCode="IZ-CAS" collectorName="Yu Xiaodong" country="China" elevation="2145" location="Wulidun village" municipality="Wolong co." pageId="12" pageNumber="2879" specimenCount="1" stateProvince="Sichuan" typeStatus="holotype">
<typeStatus id="F61F478CC3573754259FFA231AF6FA7E" box="[396,507,1506,1530]" pageId="12" pageNumber="2879" type="holotype">Holotype</typeStatus>
:
<specimenCount id="3FA232A7C3573754261EFA231948FA7E" box="[525,581,1506,1530]" pageId="12" pageNumber="2879" type="male">male</specimenCount>
,
<emphasis id="1BD0253CC3573754264BFA2319BFFA7E" bold="true" box="[600,690,1506,1530]" pageId="12" pageNumber="2879">
<collectingCountry id="51B3B9BEC3573754264BFA2319BFFA7E" box="[600,690,1506,1530]" name="China" pageId="12" pageNumber="2879">CHINA</collectingCountry>
</emphasis>
:
<emphasis id="1BD0253CC357375426DBFA23182DFA7E" bold="true" box="[712,800,1506,1530]" pageId="12" pageNumber="2879">
<collectingRegion id="EB6037CCC357375426DBFA23182DFA7E" box="[712,800,1506,1530]" country="China" name="Sichuan" pageId="12" pageNumber="2879">Sichuan</collectingRegion>
</emphasis>
:
<collectingMunicipality id="C97F6354C35737542725FA2318CDFA7E" box="[822,960,1506,1530]" pageId="12" pageNumber="2879">Wolong co.</collectingMunicipality>
,
<location id="2C7BAFF5C357375427C7FA231F82FA7E" LSID="urn:lsid:plazi:treatment:A10D4838C35737572630FA4219BCFA34:2C7BAFF5C357375427C7FA231F82FA7E" box="[980,1167,1506,1530]" country="China" municipality="Wolong co." name="Wulidun village" pageId="12" pageNumber="2879" stateProvince="Sichuan">Wulidun village</location>
,
<quantity id="EE5C54CBC357375424C0F9C31A25F99E" box="[211,296,1538,1562]" metricMagnitude="3" metricUnit="m" metricValue="2.145" pageId="12" pageNumber="2879" unit="m" value="2145.0">
<elevation id="A2891E1DC357375424C0F9C31A25F99E" box="[211,296,1538,1562]" metricMagnitude="3" metricUnit="m" metricValue="2.145" pageId="12" pageNumber="2879" unit="m" value="2145.0">2145 m</elevation>
</quantity>
,
<date id="5D1ADFEEC35737542520F9C31AFDF99E" box="[307,496,1538,1563]" pageId="12" pageNumber="2879" value="2004-05-23" valueMax="2004-05-26" valueMin="2004-05-23">
<collectingDate id="4D5E2606C35737542520F9C31AFDF99E" box="[307,496,1538,1563]" pageId="12" pageNumber="2879" value="2004-05-23" valueMax="2004-05-26" valueMin="2004-05-23">23–26 May 2004</collectingDate>
</date>
,
<collectorName id="84519CF8C357375425E8F9C2199EF99E" box="[507,659,1538,1562]" pageId="12" pageNumber="2879">Yu Xiaodong</collectorName>
collected (
<collectionCode id="4FB561EBC3573754271FF9C31861F99E" box="[780,876,1538,1562]" pageId="12" pageNumber="2879">IZ-CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="99CCF373C3573754276CF9C218C6F9BE" ID-GBIF-Occurrence="3757895303" collectingDate="1957-04-23" collectingDateMax="2004-07-08" collectingDateMin="1957-04-23" collectionCode="IZ-CAS" country="China" county="Genda co." elevation="1820" location="Emei Shan" municipality="Yu Xiaodong" pageId="12" pageNumber="2879" specimenCount="1" stateProvince="Sichuan" typeStatus="paratype">
<typeStatus id="F61F478CC3573754276CF9C218EAF99F" box="[895,999,1539,1563]" pageId="12" pageNumber="2879" type="paratype">Paratype</typeStatus>
: male,
<emphasis id="1BD0253CC35737542027F9C31F81F99E" bold="true" box="[1076,1164,1538,1562]" pageId="12" pageNumber="2879">
<collectingRegion id="EB6037CCC35737542027F9C31F81F99E" box="[1076,1164,1538,1562]" country="China" name="Sichuan" pageId="12" pageNumber="2879">Sichuan</collectingRegion>
</emphasis>
:
<collectingCounty id="C07A81A2C357375424C0F9E31A45F9BE" box="[211,328,1570,1594]" pageId="12" pageNumber="2879">Genda co.</collectingCounty>
,
<quantity id="EE5C54CBC35737542541F9E31AABF9BE" box="[338,422,1570,1594]" metricMagnitude="3" metricUnit="m" metricValue="1.8199999999999998" pageId="12" pageNumber="2879" unit="m" value="1820.0">
<elevation id="A2891E1DC35737542541F9E31AABF9BE" box="[338,422,1570,1594]" metricMagnitude="3" metricUnit="m" metricValue="1.8199999999999998" pageId="12" pageNumber="2879" unit="m" value="1820.0">1820 m</elevation>
</quantity>
,
<date id="5D1ADFEEC357375425A3F9E31945F9BE" box="[432,584,1570,1594]" pageId="12" pageNumber="2879" value="2004-07-05" valueMax="2004-07-08" valueMin="2004-07-05">
<collectingDate id="4D5E2606C357375425A3F9E31945F9BE" box="[432,584,1570,1594]" pageId="12" pageNumber="2879" value="2004-07-05" valueMax="2004-07-08" valueMin="2004-07-05">5–8 July 2004</collectingDate>
</date>
,
<collectingMunicipality id="C97F6354C35737542641F9E219E4F9BE" box="[594,745,1570,1594]" pageId="12" pageNumber="2879">Yu Xiaodong</collectingMunicipality>
collected (
<collectionCode id="4FB561EBC3573754274CF9E318B2F9BE" box="[863,959,1570,1594]" pageId="12" pageNumber="2879">IZ-CAS</collectionCode>
);
</materialsCitation>
<materialsCitation id="99CCF373C357375427C2F9E31877F9DE" ID-GBIF-Occurrence="3757895301" collectingDate="1957-04-23" collectingDateMax="2004-07-08" collectingDateMin="1957-04-23" collectionCode="IZ-CAS" country="China" county="Genda co." elevation="1820" location="Emei Shan" municipality="Yu Xiaodong" pageId="12" pageNumber="2879" specimenCount="1" stateProvince="Sichuan" typeStatus="paratype">
<location id="2C7BAFF5C357375427C2F9E31F40F9BE" LSID="urn:lsid:plazi:treatment:A10D4838C35737572630FA4219BCFA34:2C7BAFF5C357375427C2F9E31F40F9BE" box="[977,1101,1570,1594]" country="China" county="Genda co." municipality="Yu Xiaodong" name="Emei Shan" pageId="12" pageNumber="2879" stateProvince="Sichuan">Emei Shan</location>
, male,
<quantity id="EE5C54CBC357375424C0F9831A50F9DE" box="[211,349,1602,1626]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="12" pageNumber="2879" unit="m" value="900.0" valueMax="1000.0" valueMin="800.0">
<elevation id="A2891E1DC357375424C0F9831A50F9DE" box="[211,349,1602,1626]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="12" pageNumber="2879" unit="m" value="900.0" valueMax="1000.0" valueMin="800.0">800–1000 m</elevation>
</quantity>
,
<date id="5D1ADFEEC3573754257BF983190EF9DE" box="[360,515,1602,1626]" pageId="12" pageNumber="2879" value="1957-04-23">
<collectingDate id="4D5E2606C3573754257BF983190EF9DE" box="[360,515,1602,1626]" pageId="12" pageNumber="2879" value="1957-04-23">23 April 1957</collectingDate>
</date>
, Huangkeren collected (
<collectionCode id="4FB561EBC35737542701F983187EF9DE" box="[786,883,1602,1626]" pageId="12" pageNumber="2879">IZ-CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="7DDBA9A6C3563755249CFA7319E2F9A4" ID-DOI="http://doi.org/10.5281/zenodo.5198108" ID-Zenodo-Dep="5198108" httpUri="https://zenodo.org/record/5198108/files/figure.png" pageId="13" pageNumber="2880" startId="13.[143,210,1458,1480]" targetBox="[144,1104,161,1425]" targetPageId="13">
<paragraph id="291BF92EC3563755249CFA7319E2F9A4" blockId="13.[143,1104,1458,1568]" pageId="13" pageNumber="2880">
Figure 2.
<taxonomicName id="EEA482ADC356375524EBFA721AA1FA4C" authority="Zhou &amp; Zhou, 2013" authorityName="Zhou &amp; Zhou" authorityYear="2013" box="[248,428,1459,1481]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="2880" phylum="Arthropoda" rank="species" species="recticeps" status="sp. nov.">
<emphasis id="1BD0253CC356375524EBFA721AA1FA4C" box="[248,428,1459,1481]" italics="true" pageId="13" pageNumber="2880">Liotesba recticeps</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00E39847C356375525A7FA721909FA4C" box="[436,516,1459,1480]" pageId="13" pageNumber="2880" rank="species">sp. nov.</taxonomicNameLabel>
, (A) head; (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male pleurites of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm.
</paragraph>
</caption>
<subSubSection id="61BEAAA5C355375724C0FF691A68FBE5" lastPageId="15" lastPageNumber="2882" pageId="14" pageNumber="2881" type="description">
<paragraph id="291BF92EC355375624C0FF691A5FFF3B" blockId="14.[211,1172,168,423]" box="[211,338,168,191]" pageId="14" pageNumber="2881">
<heading id="72534E42C355375624C0FF691A5FFF3B" box="[211,338,168,191]" fontSize="10" level="3" pageId="14" pageNumber="2881" reason="5">
<emphasis id="1BD0253CC355375624C0FF691A5FFF3B" box="[211,338,168,191]" italics="true" pageId="14" pageNumber="2881">Description</emphasis>
</heading>
</paragraph>
<paragraph id="291BF92EC355375624C0FF0E18A3FE83" blockId="14.[211,1172,168,423]" pageId="14" pageNumber="2881">
<emphasis id="1BD0253CC355375624C0FF0E1A7DFF63" bold="true" box="[211,368,207,231]" pageId="14" pageNumber="2881">Measurement.</emphasis>
BL =
<quantity id="EE5C54CBC355375625DAFF0E193CFF63" box="[457,561,207,231]" metricMagnitude="-3" metricUnit="m" metricValue="8.66" pageId="14" pageNumber="2881" unit="mm" value="8.66">8.66 mm</quantity>
, FL =
<quantity id="EE5C54CBC3553756269EFF0E19FBFF63" box="[653,758,207,231]" metricMagnitude="-3" metricUnit="m" metricValue="4.46" pageId="14" pageNumber="2881" unit="mm" value="4.46">4.46 mm</quantity>
, HL =
<quantity id="EE5C54CBC35537562745FF0E18B3FF63" box="[854,958,207,231]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="2881" unit="mm" value="1.3">1.30 mm</quantity>
, HW =
<quantity id="EE5C54CBC35537562034FF0E1F82FF63" box="[1063,1167,207,231]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="14" pageNumber="2881" unit="mm" value="1.1">1.10 mm</quantity>
, PL =
<quantity id="EE5C54CBC35537562504FF2E1A71FE83" box="[279,380,239,263]" metricMagnitude="-3" metricUnit="m" metricValue="1.38" pageId="14" pageNumber="2881" unit="mm" value="1.38">1.38 mm</quantity>
, PW = 1.00 mm, EL =
<quantity id="EE5C54CBC35537562694FF2E19E1FE83" box="[647,748,239,263]" metricMagnitude="-3" metricUnit="m" metricValue="1.63" pageId="14" pageNumber="2881" unit="mm" value="1.63">1.63 mm</quantity>
, EW =
<quantity id="EE5C54CBC35537562756FF2E18A4FE83" box="[837,937,239,263]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="14" pageNumber="2881" unit="mm" value="1.25">1.25 mm</quantity>
.
</paragraph>
<paragraph id="291BF92EC355375624EDFECE18CAFE23" blockId="14.[211,1172,168,423]" pageId="14" pageNumber="2881">Body nearly cylindrical, medium-sized. Head, pronotum, mesoscutellum and abdomen black, except posterior margin of abdominal segment VII much lighter in colour. Elytra reddish brown. Legs entirely brown. Antennae brown, basal three antennomeres darker and smoother than the remaining antennomeres. Maxillary and labial palpi dark brown, apical portion of both last segments light brown.</paragraph>
<paragraph id="291BF92EC355375624C0FE0F1A90FC21" blockId="14.[211,1172,462,933]" pageId="14" pageNumber="2881">
<emphasis id="1BD0253CC355375624C0FE0F1A02FE62" bold="true" box="[211,271,462,486]" pageId="14" pageNumber="2881">Head</emphasis>
(
<figureCitation id="B19FE5ABC35537562532FE0F1A91FE62" box="[289,412,462,486]" captionStart="Figure 2" captionStartId="13.[143,210,1458,1480]" captionTargetBox="[144,1104,161,1425]" captionTargetId="figure-6@13.[144,1104,161,1425]" captionTargetPageId="13" captionText="Figure 2. Liotesba recticeps sp. nov., (A) head; (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male pleurites of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198108" httpUri="https://zenodo.org/record/5198108/files/figure.png" pageId="14" pageNumber="2881">Figure 2A</figureCitation>
). Subrectangular (HL to HW ratio 1.2), widest at middle of eyes, tempora slightly narrowed backwards, posterior angles rounded. Dorsal integument with numerous micropunctures; frontal region and disc smooth, without punctures; posterior half of head with large punctures, and becoming denser posterolaterally; deflexed portion of tempora with a row of small punctures. Each side of cranium with anterolateral puncture near antennal insertion, midlateral puncture about a distance of two to three punctures’ diameters from dorsal margin of eye. Frontal furrows relatively long and deep, anteocular furrows absent. Eye large, slightly shorter than the length of tempora (eye: tempora = 0.45:
<quantity id="EE5C54CBC35537562676FD0C19C6FD61" box="[613,715,717,741]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="2881" unit="mm" value="0.5">0.50 mm</quantity>
), distinctly protruding laterad; one large and one small puncture next to dorsal margin of eye. Epistoma narrow, subrectangular. Distance between antennal insertions equal to distance from antenna to eye (
<quantity id="EE5C54CBC35537562031FCCC1F8AFCA1" box="[1058,1159,781,805]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="14" pageNumber="2881" unit="mm" value="0.37">0.37 mm</quantity>
). Ventral side shiny, with scattered, larger punctures (than punctures on dorsal side) on each central surface, distance between punctures shorter than a puncture’s diameter, and also with smaller, denser punctures on sub-basal surface. Gular plate narrow; gular sutures separated.
</paragraph>
<paragraph id="291BF92EC355375624C0FC0D1A47FBE0" blockId="14.[211,1172,972,1124]" pageId="14" pageNumber="2881">
<emphasis id="1BD0253CC355375624C0FC0D1A4FFC60" bold="true" box="[211,322,972,996]" pageId="14" pageNumber="2881">Antennae.</emphasis>
Scape quite stout, distinctly widened apically, much longer than three subsequent antennomeres combined,
<quantity id="EE5C54CBC3553756264EFC2D19CBFB80" box="[605,710,1004,1028]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="14" pageNumber="2881" unit="mm" value="0.48">0.48 mm</quantity>
; antennomere II subglobular,
<quantity id="EE5C54CBC35537562034FC2D1F82FB80" box="[1063,1167,1004,1028]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="2881" unit="mm" value="0.13">0.13 mm</quantity>
; III distinctly elongate, observably longer than II,
<quantity id="EE5C54CBC35537562709FBCD188EFBA0" box="[794,899,1036,1060]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="14" pageNumber="2881" unit="mm" value="0.18">0.18 mm</quantity>
; IV and V subequal in length,
<quantity id="EE5C54CBC35537562535FBED1A87FBC0" box="[294,394,1068,1092]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="2881" unit="mm" value="0.08">0.08 mm</quantity>
; last antennomere
<quantity id="EE5C54CBC3553756264AFBED19B1FBC0" box="[601,700,1068,1092]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="14" pageNumber="2881" unit="mm" value="0.23">0.23 mm</quantity>
, shorter than two preceding antennomeres combined.
</paragraph>
<paragraph id="291BF92EC355375624C0FB4D1A51FAC7" blockId="14.[211,1172,1164,1347]" pageId="14" pageNumber="2881">
<emphasis id="1BD0253CC355375624C0FB4D1A5BFB20" bold="true" box="[211,342,1164,1188]" pageId="14" pageNumber="2881">Mouthparts</emphasis>
(
<figureCitation id="B19FE5ABC3553756257BFB4D190DFB20" box="[360,512,1164,1188]" captionStart="Figure 2" captionStartId="13.[143,210,1458,1480]" captionTargetBox="[144,1104,161,1425]" captionTargetId="figure-6@13.[144,1104,161,1425]" captionTargetPageId="13" captionText="Figure 2. Liotesba recticeps sp. nov., (A) head; (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male pleurites of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198108" httpUri="https://zenodo.org/record/5198108/files/figure.png" pageId="14" pageNumber="2881">Figure 2B–E</figureCitation>
). Labrum transverse, with a pair of median teeth, and a lateral tooth at each side (
<figureCitation id="B19FE5ABC355375625EBFB6D1965FB40" box="[504,616,1196,1220]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="14" pageNumber="2881">Figure 1B</figureCitation>
), which distinctly protrude. Mandibles falciform, right one with a broad tooth on inner edge (
<figureCitation id="B19FE5ABC355375626A6FB0D182BFB60" box="[693,806,1228,1252]" captionStart="Figure 2" captionStartId="13.[143,210,1458,1480]" captionTargetBox="[144,1104,161,1425]" captionTargetId="figure-6@13.[144,1104,161,1425]" captionTargetPageId="13" captionText="Figure 2. Liotesba recticeps sp. nov., (A) head; (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male pleurites of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198108" httpUri="https://zenodo.org/record/5198108/files/figure.png" pageId="14" pageNumber="2881">Figure 2C</figureCitation>
). Maxillary palpus elongate, second segment longest, last segment slender, fusiform, and longer than the penultimate (
<figureCitation id="B19FE5ABC355375624CFFACA1A5CFAA7" box="[220,337,1291,1315]" captionStart="Figure 2" captionStartId="13.[143,210,1458,1480]" captionTargetBox="[144,1104,161,1425]" captionTargetId="figure-6@13.[144,1104,161,1425]" captionTargetPageId="13" captionText="Figure 2. Liotesba recticeps sp. nov., (A) head; (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male pleurites of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198108" httpUri="https://zenodo.org/record/5198108/files/figure.png" pageId="14" pageNumber="2881">Figure 2D</figureCitation>
). Labial palpus elongate, last segment longest, slightly dilated subapically (
<figureCitation id="B19FE5ABC355375624CFFAEA1A40FAC7" box="[220,333,1323,1347]" captionStart="Figure 2" captionStartId="13.[143,210,1458,1480]" captionTargetBox="[144,1104,161,1425]" captionTargetId="figure-6@13.[144,1104,161,1425]" captionTargetPageId="13" captionText="Figure 2. Liotesba recticeps sp. nov., (A) head; (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male pleurites of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198108" httpUri="https://zenodo.org/record/5198108/files/figure.png" pageId="14" pageNumber="2881">Figure 2E</figureCitation>
).
</paragraph>
<paragraph id="291BF92EC355375624C0FAAA18D0FA07" blockId="14.[211,989,1387,1411]" box="[211,989,1387,1411]" pageId="14" pageNumber="2881">
<emphasis id="1BD0253CC355375624C0FAAA1A18FA07" bold="true" box="[211,277,1387,1411]" pageId="14" pageNumber="2881">Neck.</emphasis>
Wide (
<quantity id="EE5C54CBC35537562576FAAA1AC1FA07" box="[357,460,1387,1411]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="14" pageNumber="2881" unit="mm" value="0.53">0.53 mm</quantity>
), slightly narrower than half the width of head.
</paragraph>
<paragraph id="291BF92EC355375624C0FA6A1814F9E6" blockId="14.[211,1172,1451,1634]" pageId="14" pageNumber="2881">
<emphasis id="1BD0253CC355375624C0FA6A1A45FA47" bold="true" box="[211,328,1451,1475]" pageId="14" pageNumber="2881">Pronotum.</emphasis>
Subrectangular, distinctly elongate (PL to PW ratio 1.4), subequal to head length, but narrower. Widened anteriad, anterior angles well defined, lateral margins sinuate, posterior angles broadly rounded. Dorsal integument of pronotum extensively covered with micropunctures, with two large punctures on each anterolateral corner, and one large puncture on each posterolateral corner, also with irregular and small punctures located near anterior and lateral margins.
</paragraph>
<paragraph id="291BF92EC3543757249CFF661A09FF5B" blockId="15.[143,1103,167,223]" pageId="15" pageNumber="2882">
<emphasis id="1BD0253CC3543757249CFF661A36FF3B" bold="true" box="[143,315,167,191]" pageId="15" pageNumber="2882">Mesoscutellum.</emphasis>
Slightly shiny, rugose on basal surface; with two rows of scattered punctures.
</paragraph>
<paragraph id="291BF92EC3543757249CFEC31A2DFE3D" blockId="15.[143,1104,258,441]" pageId="15" pageNumber="2882">
<emphasis id="1BD0253CC3543757249CFEC31BD0FE9E" bold="true" box="[143,221,258,282]" pageId="15" pageNumber="2882">Elytra.</emphasis>
Subrectangular, distinctly elongate (EL to EW ratio 1.3), longer, wider than pronotum. Humeri well-developed, lateral margins divergent posteriorly, hind margin rounded. Integument shiny, flattened, without microsculpture; with two or three rows of small, dense punctures along median suture, a row of dense punctures along midwidth, and also with three or four rows of denser punctures on deflexed portion of each elytron.
</paragraph>
<paragraph id="291BF92EC3543757249CFE1D1F47FD97" blockId="15.[143,1104,476,532]" pageId="15" pageNumber="2882">
<emphasis id="1BD0253CC3543757249CFE1D1BC1FE70" bold="true" box="[143,204,476,500]" pageId="15" pageNumber="2882">Legs.</emphasis>
First four segments of protarsi heart-shaped, dilated, but slender in those of mesotarsi and metatarsi. Last segment shorter than the II–IV combined, respectively.
</paragraph>
<paragraph id="291BF92EC3543757249CFDF718F4FD49" blockId="15.[143,1103,566,717]" pageId="15" pageNumber="2882">
<emphasis id="1BD0253CC3543757249CFDF71BF3FDCA" bold="true" box="[143,254,566,590]" pageId="15" pageNumber="2882">Abdomen.</emphasis>
Cylindrical, broadest at segment VI. Tergites III–VII shiny, each segment covered with dense, transverse microstriae, and scattered with large punctures, distance between punctures being a puncture’s diameter. Each tergite with a punctate basal impression near anterior margin, which also bears transverse microstriae. All abdominal sternites shiny, with microstriae and punctures as those on tergites.
</paragraph>
<paragraph id="291BF92EC3543757249CFD311F0CFBA3" blockId="15.[143,1104,752,1063]" pageId="15" pageNumber="2882">
<emphasis id="1BD0253CC3543757249CFD311BC4FC8C" bold="true" box="[143,201,752,776]" pageId="15" pageNumber="2882">Male</emphasis>
(
<figureCitation id="B19FE5ABC354375724CAFD311A7EFC8C" box="[217,371,752,776]" captionStart="Figure 2" captionStartId="13.[143,210,1458,1480]" captionTargetBox="[144,1104,161,1425]" captionTargetId="figure-6@13.[144,1104,161,1425]" captionTargetPageId="13" captionText="Figure 2. Liotesba recticeps sp. nov., (A) head; (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male pleurites of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198108" httpUri="https://zenodo.org/record/5198108/files/figure.png" pageId="15" pageNumber="2882">Figure 2F–K</figureCitation>
). Abdominal segment VIII entirely covered with punctures, posterior margin of tergite very slightly rounded medially, posterior margin of sternite distinctly rounded (
<figureCitation id="B19FE5ABC35437572563FCF11901FCCC" box="[368,524,816,840]" captionStart="Figure 2" captionStartId="13.[143,210,1458,1480]" captionTargetBox="[144,1104,161,1425]" captionTargetId="figure-6@13.[144,1104,161,1425]" captionTargetPageId="13" captionText="Figure 2. Liotesba recticeps sp. nov., (A) head; (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male pleurites of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198108" httpUri="https://zenodo.org/record/5198108/files/figure.png" pageId="15" pageNumber="2882">Figure 2F, G</figureCitation>
). Tergite IX symmetrical, connected mediobasally. Sternite IX asymmetrical, widest near middle, with bluntly pointed base and straight lateral margins (
<figureCitation id="B19FE5ABC35437572554FCAE1AA2FC03" box="[327,431,879,903]" captionStart="Figure 2" captionStartId="13.[143,210,1458,1480]" captionTargetBox="[144,1104,161,1425]" captionTargetId="figure-6@13.[144,1104,161,1425]" captionTargetPageId="13" captionText="Figure 2. Liotesba recticeps sp. nov., (A) head; (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male pleurites of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198108" httpUri="https://zenodo.org/record/5198108/files/figure.png" pageId="15" pageNumber="2882">Figure 2I</figureCitation>
). Tergite X symmetrical, with relatively sharp pointed base, distinctly angulate at lateral margins (
<figureCitation id="B19FE5ABC35437572626FC4E19A5FC23" box="[565,680,911,935]" captionStart="Figure 2" captionStartId="13.[143,210,1458,1480]" captionTargetBox="[144,1104,161,1425]" captionTargetId="figure-6@13.[144,1104,161,1425]" captionTargetPageId="13" captionText="Figure 2. Liotesba recticeps sp. nov., (A) head; (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male pleurites of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198108" httpUri="https://zenodo.org/record/5198108/files/figure.png" pageId="15" pageNumber="2882">Figure 2H</figureCitation>
). Aedeagus pear-shaped, small in size (
<figureCitation id="B19FE5ABC3543757248BFC6E1A25FC43" box="[152,296,943,967]" captionStart="Figure 2" captionStartId="13.[143,210,1458,1480]" captionTargetBox="[144,1104,161,1425]" captionTargetId="figure-6@13.[144,1104,161,1425]" captionTargetPageId="13" captionText="Figure 2. Liotesba recticeps sp. nov., (A) head; (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male pleurites of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198108" httpUri="https://zenodo.org/record/5198108/files/figure.png" pageId="15" pageNumber="2882">Figure 2J, K</figureCitation>
), basal bulb
<quantity id="EE5C54CBC354375725ABFC6E1917FC43" box="[440,538,943,967]" metricMagnitude="-4" metricUnit="m" metricValue="7.7" pageId="15" pageNumber="2882" unit="mm" value="0.77">0.77 mm</quantity>
long; anterior portion short,
<quantity id="EE5C54CBC35437572775FC6E18C6FC43" box="[870,971,943,967]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="15" pageNumber="2882" unit="mm" value="0.16">0.16 mm</quantity>
, with a pair of black sclerites near apex in dorsal view. Parameres symmetrical, relatively strong, slightly shorter than half of basal bulb length. Internal sac thin, brown in colour, without spines, coiled several times within the base of median lobe (
<figureCitation id="B19FE5ABC35437572794FBCE18FDFBA3" box="[903,1008,1039,1063]" captionStart="Figure 2" captionStartId="13.[143,210,1458,1480]" captionTargetBox="[144,1104,161,1425]" captionTargetId="figure-6@13.[144,1104,161,1425]" captionTargetPageId="13" captionText="Figure 2. Liotesba recticeps sp. nov., (A) head; (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male pleurites of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198108" httpUri="https://zenodo.org/record/5198108/files/figure.png" pageId="15" pageNumber="2882">Figure 2J</figureCitation>
).
</paragraph>
<paragraph id="291BF92EC3543757249CFB881A68FBE5" blockId="15.[143,357,1097,1121]" box="[143,357,1097,1121]" pageId="15" pageNumber="2882">
<emphasis id="1BD0253CC3543757249CFB881BE8FBE5" bold="true" box="[143,229,1097,1121]" pageId="15" pageNumber="2882">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C3543757249CFB451AEEFB18" box="[143,483,1156,1180]" pageId="15" pageNumber="2882" type="distribution">
<paragraph id="291BF92EC3543757249CFB451AEEFB18" blockId="15.[143,483,1156,1180]" box="[143,483,1156,1180]" pageId="15" pageNumber="2882">
<emphasis id="1BD0253CC3543757249CFB451A11FB18" bold="true" box="[143,284,1156,1180]" pageId="15" pageNumber="2882">Distribution.</emphasis>
<collectingCountry id="51B3B9BEC35437572530FB451A65FB18" box="[291,360,1156,1180]" name="China" pageId="15" pageNumber="2882">China</collectingCountry>
(
<collectingRegion id="EB6037CCC35437572565FB451ADBFB18" box="[374,470,1156,1180]" country="China" name="Sichuan" pageId="15" pageNumber="2882">Sichuan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C3543757249CFB7F1BC0FA92" pageId="15" pageNumber="2882" type="etymology">
<paragraph id="291BF92EC3543757249CFB7F1BC0FA92" blockId="15.[143,1104,1214,1302]" pageId="15" pageNumber="2882">
<emphasis id="1BD0253CC3543757249CFB7F1A01FB52" bold="true" box="[143,268,1214,1238]" pageId="15" pageNumber="2882">Etymology.</emphasis>
The specific epithet is derived from the Latin words
<emphasis id="1BD0253CC35437572778FB7E18A1FB52" box="[875,940,1215,1238]" italics="true" pageId="15" pageNumber="2882">rectus</emphasis>
(straight) and
<emphasis id="1BD0253CC3543757249CFB1E1BB3FB72" box="[143,190,1247,1270]" italics="true" pageId="15" pageNumber="2882">ceps</emphasis>
(head, the short form of
<emphasis id="1BD0253CC354375725F5FB1E192FFB72" box="[486,546,1247,1270]" italics="true" pageId="15" pageNumber="2882">caput</emphasis>
) and refers to the parallel lateral margins of the head.
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C3543757249CFAF819BCFA34" pageId="15" pageNumber="2882" type="discussion">
<paragraph id="291BF92EC3543757249CFAF819BCFA34" blockId="15.[143,1103,1337,1456]" pageId="15" pageNumber="2882">
<emphasis id="1BD0253CC3543757249CFAF81BF5FAD5" bold="true" box="[143,248,1337,1361]" pageId="15" pageNumber="2882">Remarks.</emphasis>
This new species is similar in body size, elytral coloration and long frontal furrows to
<taxonomicName id="EEA482ADC35437572519FA9B1958FAF4" authority="(Sharp, 1889)" authorityName="Distance" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[266,597,1368,1393]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="2882" phylum="Arthropoda" rank="species" species="punctiventris">
<emphasis id="1BD0253CC35437572519FA9B1AB8FAF5" box="[266,437,1370,1393]" italics="true" pageId="15" pageNumber="2882">L. punctiventris</emphasis>
(
<bibRefCitation id="4D3584DFC354375725D0FA991940FAF4" author="Sharp DS" box="[451,589,1368,1392]" pageId="15" pageNumber="2882" pagination="249 - 267" refId="ref18004" refString="Sharp DS. 1889. The Staphylinidae of Japan. Ann Mag Nat History. 6: 249 - 267." type="journal article" year="1889">Sharp, 1889</bibRefCitation>
)
</taxonomicName>
, but it can be distinguished from the latter by slightly substraight tempora, larger eyes, subrectangular epistoma, arcuate posterior margin of sternite VIII and the unique inner sac.
</paragraph>
</subSubSection>
</treatment>
</document>
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