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<document id="7607D2091C5A790E413696615BA17BA4" ID-CLB-Dataset="55793" ID-DOI="10.1080/00222933.2013.791936" ID-GBIF-Dataset="6591e69b-631a-4c4d-bf83-db8c70d1ad05" ID-ISSN="1464-5262" ID-Zenodo-Dep="5198102" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628884845165" checkinUser="felipe" docAuthor="Zhou, Yu-Lingzi &amp; Zhou, Hong-Zhang" docDate="2013" docId="A10D4838C35137542611FBAC1A8FFADF" docLanguage="en" docName="JNatHist.47.45-46.2869-2904.pdf" docOrigin="Journal of Natural History 47 (45 - 46)" docSource="http://dx.doi.org/10.1080/00222933.2013.791936" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Liotesba fengyangshana Zhou &amp; Zhou 2013, sp. nov." docType="treatment" docVersion="9" lastPageNumber="2879" masterDocId="5D343040C35B37582413FFC11B0DFF84" masterDocTitle="Taxonomy and phylogeny of the genus Liotesba Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae: Xantholinini) with descriptions of four new species" masterLastPageNumber="2904" masterPageNumber="2869" pageNumber="2877" updateTime="1699079816598" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="771908994EE6B2DFD2D3AF51D6BD7EA5">Taxonomy and phylogeny of the genus Liotesba Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae: Xantholinini) with descriptions of four new species</mods:title>
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<mods:namePart id="7C80995CF6EB05A8890D1A8F483FAB0E">Zhou, Yu-Lingzi</mods:namePart>
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<mods:namePart id="8D25C747C75555644F94DF65CA01E3D1">Zhou, Hong-Zhang</mods:namePart>
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<mods:title id="E04E2E957B7B967D7933832681B7A8B5">Journal of Natural History</mods:title>
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<mods:date id="1A1B1BFFF0C34441031A56687D5346B9">2013</mods:date>
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<mods:number id="A5694365B642777B258098021411080D">2013-08-08</mods:number>
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<mods:number id="14E0E7E264A4CF8B0BD9C62B285A2BCC">47</mods:number>
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<mods:number id="B9D834B275ED90D749498E2E3C3AB9A8">45 - 46</mods:number>
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<treatment id="A10D4838C35137542611FBAC1A8FFADF" ID-DOI="http://doi.org/10.5281/zenodo.6505098" ID-GBIF-Taxon="194873654" ID-Zenodo-Dep="6505098" LSID="urn:lsid:plazi:treatment:A10D4838C35137542611FBAC1A8FFADF" httpUri="http://treatment.plazi.org/id/A10D4838C35137542611FBAC1A8FFADF" lastPageId="12" lastPageNumber="2879" pageId="10" pageNumber="2877">
<subSubSection id="61BEAAA5C35137522611FBAC1868FB01" box="[514,869,1133,1157]" pageId="10" pageNumber="2877" type="nomenclature">
<paragraph id="291BF92EC35137522611FBAC1868FB01" blockId="10.[514,869,1133,1189]" box="[514,869,1133,1157]" pageId="10" pageNumber="2877">
<heading id="72534E42C35137522611FBAC1868FB01" box="[514,869,1133,1157]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="2877" reason="2">
<taxonomicName id="EEA482ADC35137522611FBAC1806FB01" ID-CoL="3V87H" authority="Zhou &amp; Zhou, 2013" authorityName="Zhou &amp; Zhou" authorityYear="2013" box="[514,779,1133,1157]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2877" phylum="Arthropoda" rank="species" species="fengyangshana" status="sp. nov.">
<emphasis id="1BD0253CC35137522611FBAC1806FB01" bold="true" box="[514,779,1133,1157]" italics="true" pageId="10" pageNumber="2877">Liotesba fengyangshana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00E39847C35137522702FBAF1868FB01" box="[785,869,1134,1157]" pageId="10" pageNumber="2877" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C35137522673FB4C180AFB21" box="[608,775,1165,1189]" pageId="10" pageNumber="2877" type="description">
<paragraph id="291BF92EC35137522673FB4C180AFB21" blockId="10.[514,869,1133,1189]" box="[608,775,1165,1189]" pageId="10" pageNumber="2877">
(
<figureCitation id="B19FE5ABC35137522674FB4C19F2FB21" box="[615,767,1165,1189]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="10" pageNumber="2877">Figure 1AK</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C351375224C0FB0C19DDFA81" pageId="10" pageNumber="2877" type="materials_examined">
<paragraph id="291BF92EC351375224C0FB0C19DDFA81" blockId="10.[211,1171,1229,1286]" pageId="10" pageNumber="2877">
<emphasis id="1BD0253CC351375224C0FB0C1A7EFB61" bold="true" box="[211,371,1229,1253]" pageId="10" pageNumber="2877">Type material.</emphasis>
<materialsCitation id="99CCF373C3513752256AFB0C19DDFA81" ID-GBIF-Occurrence="3757895307" collectingDate="2007-07-31" collectionCode="IZ-CAS" collectorName="Yang Zhuo" country="China" elevation="1400" location="Fengyang Shan" municipality="Lanao co." pageId="10" pageNumber="2877" specimenCount="1" stateProvince="Zhejiang" typeStatus="holotype">
<typeStatus id="F61F478CC3513752256AFB0C1AE5FB61" box="[377,488,1229,1253]" pageId="10" pageNumber="2877" type="holotype">Holotype</typeStatus>
:
<specimenCount id="3FA232A7C351375225E3FB0C1925FB61" box="[496,552,1229,1253]" pageId="10" pageNumber="2877" type="male">male</specimenCount>
,
<emphasis id="1BD0253CC35137522622FB0C180CFB61" bold="true" box="[561,769,1229,1253]" pageId="10" pageNumber="2877">
<collectingCountry id="51B3B9BEC35137522622FB0C1980FB61" box="[561,653,1229,1253]" name="China" pageId="10" pageNumber="2877">CHINA</collectingCountry>
:
<collectingRegion id="EB6037CCC3513752268BFB0C19F0FB61" box="[664,765,1229,1253]" country="China" name="Zhejiang" pageId="10" pageNumber="2877">Zhejiang</collectingRegion>
:
</emphasis>
<collectingMunicipality id="C97F6354C35137522714FB0F1874FB61" box="[775,889,1230,1253]" pageId="10" pageNumber="2877">Lanao co.</collectingMunicipality>
:
<location id="2C7BAFF5C35137522791FB0F1F3FFB61" LSID="urn:lsid:plazi:treatment:A10D4838C35137542611FBAC1A8FFADF:2C7BAFF5C35137522791FB0F1F3FFB61" box="[898,1074,1229,1254]" country="China" municipality="Lanao co." name="Fengyang Shan" pageId="10" pageNumber="2877" stateProvince="Zhejiang">Fengyang Shan</location>
,
<quantity id="EE5C54CBC3513752202FFB0C1F82FB61" box="[1084,1167,1229,1253]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="10" pageNumber="2877" unit="m" value="1400.0">
<elevation id="A2891E1DC3513752202FFB0C1F82FB61" box="[1084,1167,1229,1253]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="10" pageNumber="2877" unit="m" value="1400.0">1400 m</elevation>
</quantity>
,
<date id="5D1ADFEEC351375224C0FB2C1A6CFA81" box="[211,353,1261,1285]" pageId="10" pageNumber="2877" value="2007-07-31">
<collectingDate id="4D5E2606C351375224C0FB2C1A6CFA81" box="[211,353,1261,1285]" pageId="10" pageNumber="2877" value="2007-07-31">31 July 2007</collectingDate>
</date>
,
<collectorName id="84519CF8C3513752257FFB2F1AE6FA81" box="[364,491,1261,1286]" pageId="10" pageNumber="2877">Yang Zhuo</collectorName>
collected (
<collectionCode id="4FB561EBC35137522670FB2C19C9FA81" box="[611,708,1261,1285]" pageId="10" pageNumber="2877">IZ-CAS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C351375424C0FAFD1A8FFADF" lastPageId="12" lastPageNumber="2879" pageId="10" pageNumber="2877" type="description">
<paragraph id="291BF92EC351375224C0FAFD1A5FFAD7" blockId="10.[211,1172,1340,1594]" box="[211,338,1340,1363]" pageId="10" pageNumber="2877">
<heading id="72534E42C351375224C0FAFD1A5FFAD7" box="[211,338,1340,1363]" fontSize="10" level="3" pageId="10" pageNumber="2877" reason="5">
<emphasis id="1BD0253CC351375224C0FAFD1A5FFAD7" box="[211,338,1340,1363]" italics="true" pageId="10" pageNumber="2877">Description</emphasis>
</heading>
</paragraph>
<paragraph id="291BF92EC351375224C0FAA218A3FA1F" blockId="10.[211,1172,1340,1594]" pageId="10" pageNumber="2877">
<emphasis id="1BD0253CC351375224C0FAA21A7DFAFF" bold="true" box="[211,368,1379,1403]" pageId="10" pageNumber="2877">Measurement.</emphasis>
BL =
<quantity id="EE5C54CBC351375225DAFAA2193CFAFF" box="[457,561,1379,1403]" metricMagnitude="-3" metricUnit="m" metricValue="8.74" pageId="10" pageNumber="2877" unit="mm" value="8.74">8.74 mm</quantity>
, FL =
<quantity id="EE5C54CBC3513752269EFAA219FBFAFF" box="[653,758,1379,1403]" metricMagnitude="-3" metricUnit="m" metricValue="4.49" pageId="10" pageNumber="2877" unit="mm" value="4.49">4.49 mm</quantity>
, HL =
<quantity id="EE5C54CBC35137522745FAA218B3FAFF" box="[854,958,1379,1403]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="10" pageNumber="2877" unit="mm" value="1.25">1.25 mm</quantity>
, HW =
<quantity id="EE5C54CBC35137522034FAA21F82FAFF" box="[1063,1167,1379,1403]" metricMagnitude="-3" metricUnit="m" metricValue="1.13" pageId="10" pageNumber="2877" unit="mm" value="1.13">1.13 mm</quantity>
, PL =
<quantity id="EE5C54CBC35137522504FA421A71FA1F" box="[279,380,1411,1435]" metricMagnitude="-3" metricUnit="m" metricValue="1.43" pageId="10" pageNumber="2877" unit="mm" value="1.43">1.43 mm</quantity>
, PW =
<quantity id="EE5C54CBC351375225C0FA42193AFA1F" box="[467,567,1411,1435]" metricMagnitude="-4" metricUnit="m" metricValue="9.3" pageId="10" pageNumber="2877" unit="mm" value="0.93">0.93 mm</quantity>
, EL =
<quantity id="EE5C54CBC35137522694FA4219E1FA1F" box="[647,748,1411,1435]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" pageId="10" pageNumber="2877" unit="mm" value="1.58">1.58 mm</quantity>
, EW =
<quantity id="EE5C54CBC35137522756FA4218A4FA1F" box="[837,937,1411,1435]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="10" pageNumber="2877" unit="mm" value="1.25">1.25 mm</quantity>
.
</paragraph>
<paragraph id="291BF92EC351375224EDFA621AD5F9BE" blockId="10.[211,1172,1340,1594]" pageId="10" pageNumber="2877">Body nearly cylindrical, medium-sized. Head, pronotum, mesoscutellum and abdomen black. Elytra reddish brown. Legs dark brown except tarsi lighter in colour. Antennae entirely dark brown, basal three antennomeres darker and smoother than the remaining antennomeres, apical half of last antennomere light brown. Maxillary and labial palpi brown.</paragraph>
<paragraph id="291BF92EC351375324C0F9A31BDEFDFA" blockId="10.[211,1172,1634,1690]" lastBlockId="11.[143,1104,167,638]" lastPageId="11" lastPageNumber="2878" pageId="10" pageNumber="2877">
<emphasis id="1BD0253CC351375224C0F9A31A02F9FE" bold="true" box="[211,271,1634,1658]" pageId="10" pageNumber="2877">Head</emphasis>
(
<figureCitation id="B19FE5ABC35137522530F9A31A92F9FE" box="[291,415,1634,1658]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="10" pageNumber="2877">Figure 1A</figureCitation>
). Subquadrate (HL to HW ratio 1.1), tempora slightly widened posteriorly, posterior angles rounded. Dorsal integument extensively bearing micropunctures; frontal region and disc smooth, without punctures; posterior half of head and temporal region with large punctures, distance between punctures nearly a punctures diameter; deflexed portion of tempora with three rows of small punctures. Each side of cranium with anterolateral puncture near antennal insertion, midlateral puncture approximately a distance of two punctures diameter from dorsal margin of eye, temporal puncture at the posterior one-fifth of head, and occipital puncture at the lateral one-fifth of head. Frontal furrows absent, anteocular furrows relatively long and observable. Eyes large, nearly three-quarters the length of tempora (eye: tempora = 0.45:
<quantity id="EE5C54CBC350375325B9FE67191EFE3A" box="[426,531,422,446]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="2878" unit="mm" value="0.6">0.60 mm</quantity>
), distinctly protruding laterad; one large and one small puncture next to posterior half of dorsal margin of eye (
<figureCitation id="B19FE5ABC35037532745FE0718C7FE5A" box="[854,970,454,478]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="11" pageNumber="2878">Figure 1A</figureCitation>
). Epistoma narrow, narrowed anteriorly. Distance between antennal insertions
<quantity id="EE5C54CBC3503753278FFE271F08FE7A" box="[924,1029,486,510]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="2878" unit="mm" value="0.45">0.45 mm</quantity>
, subequal to distance from antenna to eye (
<quantity id="EE5C54CBC35037532644FDC719CEFD9A" box="[599,707,518,542]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="11" pageNumber="2878" unit="mm" value="0.48">0.48 mm</quantity>
). Ventral side of head shiny, with scattered, larger punctures (than those on dorsal side), distance between punctures shorter than a puncture diameter. Gular plate narrow, subtriangular; gular sutures not fused.
</paragraph>
<paragraph id="291BF92EC3503753249CFD671AA7FCB9" blockId="11.[143,1103,678,829]" pageId="11" pageNumber="2878">
<emphasis id="1BD0253CC3503753249CFD671BF3FD3A" bold="true" box="[143,254,678,702]" pageId="11" pageNumber="2878">Antennae.</emphasis>
Scape stout, thickened apically, much longer than three subsequent antennomeres combined,
<quantity id="EE5C54CBC35037532563FD041AD8FD59" box="[368,469,709,733]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="11" pageNumber="2878" unit="mm" value="0.53">0.53 mm</quantity>
; antennomere II slightly elongate,
<quantity id="EE5C54CBC35037532746FD0418B4FD59" box="[853,953,709,733]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="2878" unit="mm" value="0.13">0.13 mm</quantity>
; III distinctly elongate, observably longer than II,
<quantity id="EE5C54CBC35037532634FD241981FD79" box="[551,652,741,765]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="11" pageNumber="2878" unit="mm" value="0.18">0.18 mm</quantity>
; IV and V subequal in length,
<quantity id="EE5C54CBC350375327F5FD241F46FD79" box="[998,1099,741,765]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="2878" unit="mm" value="0.09">0.09 mm</quantity>
; last antennomere medium in length,
<quantity id="EE5C54CBC3503753263BFCC41981FC99" box="[552,652,773,797]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="11" pageNumber="2878" unit="mm" value="0.23">0.23 mm</quantity>
, subequal to the length of two preceding antennomeres combined.
</paragraph>
<paragraph id="291BF92EC3503753249CFCA41A15FB98" blockId="11.[143,1103,869,1052]" pageId="11" pageNumber="2878">
<emphasis id="1BD0253CC3503753249CFCA41A1FFCF9" bold="true" box="[143,274,869,893]" pageId="11" pageNumber="2878">Mouthparts</emphasis>
(
<figureCitation id="B19FE5ABC35037532533FCA41AB9FCF9" box="[288,436,869,893]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="11" pageNumber="2878">Figure 1BE</figureCitation>
). Labrum transverse, with a pair of median teeth, and a lateral tooth at each side (
<figureCitation id="B19FE5ABC3503753258AFC441907FC19" box="[409,522,901,925]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="11" pageNumber="2878">Figure 1B</figureCitation>
), which distinctly protruding. Mandibles falciform, right one with a broad tooth on inner edge (
<figureCitation id="B19FE5ABC35037532681FC64180BFC39" box="[658,774,933,957]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="11" pageNumber="2878">Figure 1C</figureCitation>
). Maxillary palpus elongate, second segment longest, last segment slender, fusiform, longer than the penultimate (
<figureCitation id="B19FE5ABC3503753248BFC251A02FC78" box="[152,271,996,1020]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="11" pageNumber="2878">Figure 1D</figureCitation>
). Labial palpus slender, last segment longest, slightly dilated subapically (
<figureCitation id="B19FE5ABC3503753248BFBC51A05FB98" box="[152,264,1028,1052]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="11" pageNumber="2878">Figure 1E</figureCitation>
).
</paragraph>
<paragraph id="291BF92EC3503753249CFB851F17FBD8" blockId="11.[143,1050,1092,1116]" box="[143,1050,1092,1116]" pageId="11" pageNumber="2878">
<emphasis id="1BD0253CC3503753249CFB851BC4FBD8" bold="true" box="[143,201,1092,1116]" pageId="11" pageNumber="2878">Neck</emphasis>
(
<figureCitation id="B19FE5ABC350375324CBFB851A42FBD8" box="[216,335,1092,1116]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="11" pageNumber="2878">Figure 1A</figureCitation>
). Rather wide (
<quantity id="EE5C54CBC350375325EAFB85196DFBD8" box="[505,608,1092,1116]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="11" pageNumber="2878" unit="mm" value="0.53">0.53 mm</quantity>
), approximately half the width of head.
</paragraph>
<paragraph id="291BF92EC3503753249CFB451A05FADF" blockId="11.[143,1104,1156,1371]" pageId="11" pageNumber="2878">
<emphasis id="1BD0253CC3503753249CFB451BF0FB18" bold="true" box="[143,253,1156,1180]" pageId="11" pageNumber="2878">Pronotum</emphasis>
(
<figureCitation id="B19FE5ABC35037532519FB451A72FB18" box="[266,383,1156,1180]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="11" pageNumber="2878">Figure 1A</figureCitation>
). Subrectangular, distinctly elongate (PL to PW ratio 1.4), longer than head, but slightly narrower. Widened anteriad, lateral margins straight, anterior angles well defined, slightly deflexed downwards, posterior angles broadly rounded. Dorsal surface extensively covered with micropunctures, each side with two large punctures on each anterolateral corner, one large puncture on each posterolateral corner. Additional irregular punctures distributing near anterior and lateral margins of pronotum.
</paragraph>
<paragraph id="291BF92EC3503753249CFA421F3DFA1F" blockId="11.[143,1072,1411,1436]" box="[143,1072,1411,1436]" pageId="11" pageNumber="2878">
<emphasis id="1BD0253CC3503753249CFA421A36FA1F" bold="true" box="[143,315,1411,1435]" pageId="11" pageNumber="2878">Mesoscutellum.</emphasis>
Opaque, rugose, with several small, irregularly scattered punctures.
</paragraph>
<paragraph id="291BF92EC3503753249CFA021AF1F9FE" blockId="11.[143,1103,1475,1658]" pageId="11" pageNumber="2878">
<emphasis id="1BD0253CC3503753249CFA021BD8FA5F" bold="true" box="[143,213,1475,1499]" pageId="11" pageNumber="2878">Elytra</emphasis>
(
<figureCitation id="B19FE5ABC350375324F6FA021A50FA5F" box="[229,349,1475,1499]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="11" pageNumber="2878">Figure 1A</figureCitation>
). Subrectangular, distinctly elongate (EL to EW ratio 1.3), slightly longer, wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, hind margin rounded. Integument shiny, flattened, without microsculpture, each side with two or three rows of small, sparsely-scattered punctures along median suture, a row along midwidth, and also with three or four rows of denser punctures on deflexed portion of each elytron.
</paragraph>
<paragraph id="291BF92EC357375424C0FF6618BFFF7B" blockId="12.[211,1172,167,255]" pageId="12" pageNumber="2879">
<emphasis id="1BD0253CC357375424C0FF661A1DFF3B" bold="true" box="[211,272,167,191]" pageId="12" pageNumber="2879">Legs.</emphasis>
First four segments of protarsi slightly dilated, heart-shaped, last segment as long as the IIIV combined. Last segment of mesotarsi and metatarsi slender, shorter than the IIIV of mesotarsi and metatarsi combined, respectively.
</paragraph>
<paragraph id="291BF92EC357375424C0FEE61928FE7A" blockId="12.[211,1172,295,510]" pageId="12" pageNumber="2879">
<emphasis id="1BD0253CC357375424C0FEE61A4FFEBB" bold="true" box="[211,322,295,319]" pageId="12" pageNumber="2879">Abdomen.</emphasis>
Cylindrical, broadest at segment VI. Tergites IIIVII shiny, each segment with surface entirely covered with observable transverse microstriae, with sparse, medium-sized punctures, interspaces between punctures approximately the diameter of two to three punctures, and punctures being denser, smaller on its anterolateral portion. Each tergite with a punctate basal impression near anterior margin, which also bears transverse microstriae. All abdominal sternites shiny, with microstriae and punctures as those on tergites.
</paragraph>
<paragraph id="291BF92EC357375424C0FDE719F5FC19" blockId="12.[211,1172,550,925]" pageId="12" pageNumber="2879">
<emphasis id="1BD0253CC357375424C0FDE71A00FDBA" bold="true" box="[211,269,550,574]" pageId="12" pageNumber="2879">Male</emphasis>
(
<figureCitation id="B19FE5ABC3573754250DFDE71AB4FDBA" box="[286,441,550,574]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="12" pageNumber="2879">Figure 1FK</figureCitation>
). Tergite VIII entirely covered with punctures, posterior margin very slightly rounded medially; sternite VIII with punctures on posterior half, posterior margins very slightly emarginate medially (
<figureCitation id="B19FE5ABC357375426F1FDA7187AFDFA" box="[738,887,614,638]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="12" pageNumber="2879">Figure 1F, G</figureCitation>
). Tergite IX symmetrical, connected mediobasally. Sternite IX slightly asymmetrical, widest near the apical twothirds, with sharply pointed base, and rounded apical margin (
<figureCitation id="B19FE5ABC357375427B7FD671F02FD3A" box="[932,1039,678,702]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="12" pageNumber="2879">Figure 1I</figureCitation>
). Tergite X slightly asymmetrical, with obtusely pointed base, bluntly angulate at lateral margins (
<figureCitation id="B19FE5ABC35737542500FD241A84FD79" box="[275,393,741,765]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="12" pageNumber="2879">Figure 1H</figureCitation>
). Aedeagus pear-shaped, medium in size (
<figureCitation id="B19FE5ABC35737542762FD241F0BFD79" box="[881,1030,741,765]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="12" pageNumber="2879">Figure 1J, K</figureCitation>
), basal bulb
<quantity id="EE5C54CBC357375424C0FCC41A37FC99" box="[211,314,773,797]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="12" pageNumber="2879" unit="mm" value="0.95">0.95 mm</quantity>
long; anterior portion (from the apex of bulb to the base of parameres,
<figureCitation id="B19FE5ABC357375424C0FCE41A4DFCB9" box="[211,320,805,829]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="12" pageNumber="2879">Figure 1J</figureCitation>
: h) short,
<quantity id="EE5C54CBC357375425A4FCE41913FCB9" box="[439,542,805,829]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="2879" unit="mm" value="0.15">0.15 mm</quantity>
, with a pair of black sclerites near apex in dorsal view. Parameres symmetrical, thin, moderately curved inwards, and one-third the length of basal bulb. Internal sac thin, brown in colour, and soft, without spines, coiled several times within the base of median lobe (
<figureCitation id="B19FE5ABC3573754266DFC4419E5FC19" box="[638,744,901,925]" captionStart="Figure 1" captionStartId="9.[143,210,1472,1494]" captionTargetBox="[144,1104,162,1439]" captionTargetId="figure-6@9.[144,1104,162,1439]" captionTargetPageId="9" captionText="Figure 1. Liotesba fengyangshana sp. nov., (A) forebody (a. anterolateral puncture = antennal puncture; b. midlateral puncture = ocular puncture; c. temporal puncture; d. occipital puncture; e: anterolateral puncture; f. posterolateral puncture; g. anteocular furrow = ocular furrow; h. anterior portion of basal bulb); (B) labrum; (C) right mandible; (D) maxillary palpus; (E) labial palpus; (F) male tergite VIII; (G) male sternite VIII; (H) male tergite X = male tergite of genital segment; (I) male sternite IX = male sternite of genital segment; (J) aedeagus, dorsal view; (K) aedeagus, ventral view. Scale bars 0.30 mm." figureDoi="http://doi.org/10.5281/zenodo.5198104" httpUri="https://zenodo.org/record/5198104/files/figure.png" pageId="12" pageNumber="2879">Figure 1J</figureCitation>
).
</paragraph>
<paragraph id="291BF92EC357375424C0FC051AA4FC58" blockId="12.[211,425,964,988]" box="[211,425,964,988]" pageId="12" pageNumber="2879">
<emphasis id="1BD0253CC357375424C0FC051A24FC58" bold="true" box="[211,297,964,988]" pageId="12" pageNumber="2879">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="291BF92EC357375424C0FBC519A0FB98" blockId="12.[211,685,1028,1052]" box="[211,685,1028,1052]" pageId="12" pageNumber="2879">
<emphasis id="1BD0253CC357375424C0FBC51A6DFB98" bold="true" box="[211,352,1028,1052]" pageId="12" pageNumber="2879">Distribution.</emphasis>
<collectingCountry id="51B3B9BEC35737542574FBC51AA1FB98" box="[359,428,1028,1052]" name="China" pageId="12" pageNumber="2879">China</collectingCountry>
(
<collectingRegion id="EB6037CCC357375425A9FBC5192EFB98" box="[442,547,1028,1052]" country="China" name="Zhejiang" pageId="12" pageNumber="2879">Zhejiang</collectingRegion>
: Lanao co.).
</paragraph>
<paragraph id="291BF92EC357375424C0FB851AD0FBF8" blockId="12.[211,1171,1092,1148]" pageId="12" pageNumber="2879">
<emphasis id="1BD0253CC357375424C0FB851A5DFBD8" bold="true" box="[211,336,1092,1116]" pageId="12" pageNumber="2879">Etymology.</emphasis>
The specific epithet is derived from the Chinese name (Pin-yin) of the
<typeStatus id="F61F478CC35737542070FB841F9EFBD9" box="[1123,1171,1093,1117]" pageId="12" pageNumber="2879">type</typeStatus>
locality, Fengyangshan.
</paragraph>
<paragraph id="291BF92EC357375424C0FB651A8FFADF" blockId="12.[211,1172,1188,1371]" pageId="12" pageNumber="2879">
<emphasis id="1BD0253CC357375424C0FB651A31FB38" bold="true" box="[211,316,1188,1212]" pageId="12" pageNumber="2879">Remarks.</emphasis>
<taxonomicName id="EEA482ADC3573754255FFB651956FB38" authorityName="Zhou &amp; Zhou" authorityYear="2013" box="[332,603,1188,1212]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2879" phylum="Arthropoda" rank="species" species="fengyangshana">
<emphasis id="1BD0253CC3573754255FFB651956FB38" box="[332,603,1188,1212]" italics="true" pageId="12" pageNumber="2879">Liotesba fengyangshana</emphasis>
</taxonomicName>
is similar in structure to
<taxonomicName id="EEA482ADC357375427B7FB641F55FB38" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[932,1112,1189,1212]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="2879" phylum="Arthropoda" rank="species" species="punctiventris">
<emphasis id="1BD0253CC357375427B7FB641F55FB38" box="[932,1112,1189,1212]" italics="true" pageId="12" pageNumber="2879">L. punctiventris</emphasis>
</taxonomicName>
, but their labra and aedeagi are distinctly different from each other. This new species is distinguished from its congeners by the combination of the following characters: distributional pattern of punctures on head, long anteocular furrows, large eyes, a broad tooth of right mandible, and the distributional pattern of punctures on mesoscutellum.
</paragraph>
</subSubSection>
</treatment>
</document>