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<document ID-CLB-Dataset="6723" ID-DOI="http://dx.doi.org/10.3897/dez.67.56008" ID-GBIF-Dataset="377b4525-1708-4a83-9ad4-4458abeb5dd8" ID-Pensoft-Pub="1860-1324-2-141" ID-Pensoft-UUID="26E05E69CD4D58459F7CB2EFDD2F1CBA" ID-ZooBank="28F525EFC0304E04970674C64A59225F" ModsDocID="1860-1324-67-2-141" checkinTime="1621372317078" checkinUser="pensoft" docAuthor="Aspoeck, Ulrike, Aspoeck, Horst &amp; Gruppe, Axel" docDate="2020" docId="A0FCDD99D9825CFCB4F3C80995CC9818" docLanguage="en" docName="DeutEntomolZeit 67(2): 141-149" docOrigin="Deutsche Entomologische Zeitschrift 67 (2)" docPubDate="2020-08-14" docSource="http://dx.doi.org/10.3897/dez.67.56008" docTitle="Nusalala peruana Aspöck &amp; Aspöck &amp; Gruppe 2020, sp. nov." docType="treatment" docUuid="B551F33F-D24B-4F27-AC28-0BECD2CC11C5" docUuidSource="ZooBank" docVersion="5" id="26E05E69CD4D58459F7CB2EFDD2F1CBA" lastPageNumber="141" masterDocId="26E05E69CD4D58459F7CB2EFDD2F1CBA" masterDocTitle="Anchored between heaven and earth - a new flightless brown lacewing from Peru (Neuroptera, Hemerobiidae)" masterLastPageNumber="149" masterPageNumber="141" pageNumber="141" updateTime="1732858572708" updateUser="ExternalLinkService">
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<mods:title id="14C7246E6A796BECF508C3D2BD8C7BE2">Anchored between heaven and earth - a new flightless brown lacewing from Peru (Neuroptera, Hemerobiidae)</mods:title>
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<mods:namePart id="2DFC3E1CB81132CB4450C6167AABA5A9">Aspoeck, Ulrike</mods:namePart>
<mods:affiliation id="6848CC8519FD9BD9184228D68D763F65">Natural History Museum, 2 nd Zoological Department, Burgring 7, A- 1010 Vienna, Austria &amp; Department of Evolutionary Biology, University of Vienna, Althanstrasse 14, A- 1090 Vienna, Austria</mods:affiliation>
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<mods:namePart id="17E776F8F534AA016BE1C86F59C1E790">Aspoeck, Horst</mods:namePart>
<mods:affiliation id="59A90E748711A3F2E29CD711446180CF">Institute of Specific Prophylaxis and Tropical Medicine, Medical University Vienna (MUW), Kinderspitalgasse 15, A- 1090 Vienna, Austria</mods:affiliation>
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<mods:namePart id="28B6A41B4FA6C4A7F6866356C7442698">Gruppe, Axel</mods:namePart>
<mods:nameIdentifier id="C189BE7E14930351CFA3E04804B10426" type="ORCID">https://orcid.org/0000-0001-9104-7362</mods:nameIdentifier>
<mods:affiliation id="B35F81682F70A76D38C6D64852C4E107">Chair of Zoology - Entomological Division, Technical University of Munich, Hans-Carl-von-Carlowitz-Platz 2, D- 85354 Freising, Germany</mods:affiliation>
<mods:nameIdentifier id="FF0B7BCEB3ED768134EBF22AB9446EFC" type="email">gruppe@wzw.tum.de</mods:nameIdentifier>
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<mods:date id="C735CF556F2D24BE54D1802E5D99C8E0">2020</mods:date>
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<mods:number id="E4E9B75765EBEB9E95BF01C93C23361A">2020-08-14</mods:number>
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<treatment id="A0FCDD99D9825CFCB4F3C80995CC9818" ID-GBIF-Taxon="182223421" LSID="urn:lsid:zoobank.org:act:B551F33F-D24B-4F27-AC28-0BECD2CC11C5" httpUri="http://treatment.plazi.org/id/A0FCDD99D9825CFCB4F3C80995CC9818" lastPageNumber="141" pageId="0" pageNumber="141" scope_family="Hemerobiidae" scope_order="Neuroptera">
<subSubSection id="AEC0FCD3076811896F557FE427F211F5" pageId="0" pageNumber="141" type="nomenclature">
<paragraph id="E64CF40B3986ED7F216C0C7E7B7D7600" pageId="0" pageNumber="141">
<taxonomicName id="BC19D4AE1C0581113793BAA510510C70" LSID="http://zoobank.org/B551F33F-D24B-4F27-AC28-0BECD2CC11C5" authority="Aspöck &amp; Aspöck &amp; Gruppe, 2020" authorityName="Aspöck &amp; Aspöck &amp; Gruppe" authorityYear="2020" class="Insecta" family="Hemerobiidae" genus="Nusalala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nusalala peruana" order="Neuroptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="peruana" status="sp. nov.">Nusalala peruana</taxonomicName>
<taxonomicNameLabel id="1708AF159F412DA938AA7166E124A721" pageId="0" pageNumber="141">sp. nov.</taxonomicNameLabel>
<figureCitation id="C7AA42D0D2F9F9231FB209524AF29F15" captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1 - 3. Nusalala peruana sp. nov., holotype, male. 1. habitus, dorsal; 2. habitus, lateral; 3. head frontal." figureDoi="10.3897/dez.67.56008.figures1-3" httpUri="https://binary.pensoft.net/fig/442147" pageId="0" pageNumber="141">Figs 1-3</figureCitation>
<figureCitation id="29017DA1DEC2F8928FDF1D464B488467" captionStart="Figures 4–7" captionStartId="F2" captionText="Figures 4 - 7. Nusalala peruana sp. nov., paratype, male (from Hualgayoc). 4. habitus, dorsal; 5. habitus, ventral; 6. right forewing (broken by spreading under cover glass); 7. metathorax with hindwings (spread under cover glass)." figureDoi="10.3897/dez.67.56008.figures4-7" httpUri="https://binary.pensoft.net/fig/442148" pageId="0" pageNumber="141">, 4-7</figureCitation>
<figureCitation id="52E0BF81D8EC51108855232E47BEF05B" captionStart="Figures 8–11" captionStartId="F3" captionText="Figures 8 - 11. Nusalala peruana sp. nov., paratype, male (from Hualgayoc). 8. genital sclerites, lateral; 9. genital sclerites, ventral; 10. genital sclerites, lateral, partly dissected; 11. genital sclerites, dorsal, partly dissected; e - ectoproct; gp 10 - gonapophysis 10; gst 9 - fused gonostyli 9; gx 9 - gonocoxite 9; gx 10 - partly fused gonapophyses 10; hi - hypandrium internum; S - sternite; T - tergite." figureDoi="10.3897/dez.67.56008.figures8-11" httpUri="https://binary.pensoft.net/fig/442149" pageId="0" pageNumber="141">, 8-11</figureCitation>
<figureCitation id="FCAE351D0B46388C5710532A5E0F7F23" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Nusalala peruana sp. nov., paratype, female (from Hualgayoc). 12. habitus, dorsal with right forewing (left forewing missing) and left scale-like hindwing; 13. scale-like left hindwing; 14. head, frontal; 15. genital sclerites, lateral; 16. genital sclerites, ventral; e - ectoproct; gp 8 - gonapophysis 8; gp 9 - gonapophysis 9; gx 8 - fused gonocoxites 8; gx 9 - gonocoxite 9; T - tergite; tr - trichobothrium." figureDoi="10.3897/dez.67.56008.figures12-16" httpUri="https://binary.pensoft.net/fig/442150" pageId="0" pageNumber="141">, 12-16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E2F6F8440F21B94195E6C38B1A02589A" pageId="0" pageNumber="141" type="diagnosis">
<paragraph id="CB4CE045BA5AB3C7E66A588101D30224" pageId="0" pageNumber="141">Diagnosis.</paragraph>
<paragraph id="3F9FA88026E6381BA6F8084D18B17DA5" pageId="0" pageNumber="141">
The new species can be differentiated from most other species of
<taxonomicName id="1A2511B91B67A816450111B0442AAD4C" authorityName="Navas" authorityYear="1913" class="Insecta" family="Hemerobiidae" genus="Nusalala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nusalala" order="Neuroptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="4E86D2AF0CA23F371386683B093D7ED1" italics="true" pageId="0" pageNumber="141">Nusalala</emphasis>
</taxonomicName>
<normalizedToken id="A8C7465703C4457F99E68D9FACBDCE40" originalValue="Navás">Navas</normalizedToken>
, 1913, by the following combination of characters: Forewings coriaceous and domed, covering legs and body (Figs
<figureCitation id="4D7E641C521F2933263D836CBC2035F9" captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1 - 3. Nusalala peruana sp. nov., holotype, male. 1. habitus, dorsal; 2. habitus, lateral; 3. head frontal." figureDoi="10.3897/dez.67.56008.figures1-3" httpUri="https://binary.pensoft.net/fig/442147" pageId="0" pageNumber="141">1</figureCitation>
,
<figureCitation id="62D3E77E89406D44D612FED47590E8F2" captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1 - 3. Nusalala peruana sp. nov., holotype, male. 1. habitus, dorsal; 2. habitus, lateral; 3. head frontal." figureDoi="10.3897/dez.67.56008.figures1-3" httpUri="https://binary.pensoft.net/fig/442147" pageId="0" pageNumber="141">2</figureCitation>
,
<figureCitation id="8EE6A744097B2FD6A44FD54DC1E75DDC" captionStart="Figures 4–7" captionStartId="F2" captionText="Figures 4 - 7. Nusalala peruana sp. nov., paratype, male (from Hualgayoc). 4. habitus, dorsal; 5. habitus, ventral; 6. right forewing (broken by spreading under cover glass); 7. metathorax with hindwings (spread under cover glass)." figureDoi="10.3897/dez.67.56008.figures4-7" httpUri="https://binary.pensoft.net/fig/442148" pageId="0" pageNumber="141">4-6</figureCitation>
); hindwings long and ribbon-like in the male (Fig.
<figureCitation id="5341D9C303971DFDF3FC9E7A68889B44" captionStart="Figures 4–7" captionStartId="F2" captionText="Figures 4 - 7. Nusalala peruana sp. nov., paratype, male (from Hualgayoc). 4. habitus, dorsal; 5. habitus, ventral; 6. right forewing (broken by spreading under cover glass); 7. metathorax with hindwings (spread under cover glass)." figureDoi="10.3897/dez.67.56008.figures4-7" httpUri="https://binary.pensoft.net/fig/442148" pageId="0" pageNumber="141">7</figureCitation>
), micropterous and scale-like in the female (Figs
<figureCitation id="B7FC982EE6942DE860A5383F2824C5B0" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Nusalala peruana sp. nov., paratype, female (from Hualgayoc). 12. habitus, dorsal with right forewing (left forewing missing) and left scale-like hindwing; 13. scale-like left hindwing; 14. head, frontal; 15. genital sclerites, lateral; 16. genital sclerites, ventral; e - ectoproct; gp 8 - gonapophysis 8; gp 9 - gonapophysis 9; gx 8 - fused gonocoxites 8; gx 9 - gonocoxite 9; T - tergite; tr - trichobothrium." figureDoi="10.3897/dez.67.56008.figures12-16" httpUri="https://binary.pensoft.net/fig/442150" pageId="0" pageNumber="141">12</figureCitation>
,
<figureCitation id="657D1C3220E8DCAE87AA856902E9D9C6" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Nusalala peruana sp. nov., paratype, female (from Hualgayoc). 12. habitus, dorsal with right forewing (left forewing missing) and left scale-like hindwing; 13. scale-like left hindwing; 14. head, frontal; 15. genital sclerites, lateral; 16. genital sclerites, ventral; e - ectoproct; gp 8 - gonapophysis 8; gp 9 - gonapophysis 9; gx 8 - fused gonocoxites 8; gx 9 - gonocoxite 9; T - tergite; tr - trichobothrium." figureDoi="10.3897/dez.67.56008.figures12-16" httpUri="https://binary.pensoft.net/fig/442150" pageId="0" pageNumber="141">13</figureCitation>
). Venation in both wings reduced and partly vestigial in both sexes. The coriaceous forewings differentiate
<taxonomicName id="1CAAD09D51D1012A8B25BD7AB9012C45" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="N. peruana" order="Neuroptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="peruana">
<emphasis id="13663166B31613540FA914CB50E64417" italics="true" pageId="0" pageNumber="141">N. peruana</emphasis>
</taxonomicName>
sp. nov. from all other
<taxonomicName id="EC08D209D1473D8100030E0F2DF9AEC2" authorityName="Navas" authorityYear="1913" class="Insecta" family="Hemerobiidae" genus="Nusalala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nusalala" order="Neuroptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="26443B1B843CD81F508B8415606D04A4" italics="true" pageId="0" pageNumber="141">Nusalala</emphasis>
</taxonomicName>
species, except
<taxonomicName id="CF7DE0DF8E7F459B181FAA41D6B525BD" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="N. andina" order="Neuroptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="A10B726932F6798CB67FDAE2F713EFB7" italics="true" pageId="0" pageNumber="141">N. andina</emphasis>
</taxonomicName>
Penny &amp; Sturm, 1984, and
<taxonomicName id="DE639CB8653D791CEA5324402F530AE7" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="N. brachyptera" order="Neuroptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="brachyptera">
<emphasis id="D850AC1547657B8E26C0C97716E825F5" italics="true" pageId="0" pageNumber="141">N. brachyptera</emphasis>
</taxonomicName>
Oswald, 1997. The males of the latter two species have scale-like (not ribbon-like) hindwings. The female of
<taxonomicName id="088DDF4742E90EFDF9ED8481729655AF" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="N. andina" order="Neuroptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="AF19DC918FF782BB82BDE3417B12971A" italics="true" pageId="0" pageNumber="141">N. andina</emphasis>
</taxonomicName>
is unknown.
<bibRefCitation id="150B671BB5F699C11870CCF8907DEDE3" DOI="https://doi.org/10.1111/j.1365-3113.1996.tb00603.x" author="Oswald, JD" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="141" pagination="343 - 352" refId="B25" refString="Oswald, JD, 1996. [1997] A new brachypterous Nusalala species from Costa Rica, with comments on the evolution of flightlessness in brown lacewings (Neuroptera: Hemerobiidae). Systematic Entomology 21 (4): 343 - 352, DOI: https://doi.org/10.1111/j.1365-3113.1996.tb00603.x" title="[1997] A new brachypterous Nusalala species from Costa Rica, with comments on the evolution of flightlessness in brown lacewings (Neuroptera: Hemerobiidae)." url="https://doi.org/10.1111/j.1365-3113.1996.tb00603.x" volume="21" year="1996">Oswald (1996)</bibRefCitation>
[1997] mentioned a female
<taxonomicName id="260ED76A10C0244F4A2F8F6E16007E61" authorityName="Navas" authorityYear="1913" class="Insecta" family="Hemerobiidae" genus="Nusalala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nusalala" order="Neuroptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="69C9F5C7F744E0BD75984FA085EB2B60" italics="true" pageId="0" pageNumber="141">Nusalala</emphasis>
</taxonomicName>
which he tentatively identified as
<taxonomicName id="E50EBD221EA9D819C5F9960D553DDFC0" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="N. brachyptera" order="Neuroptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="brachyptera">
<emphasis id="EC9EEBA2CB9A09CCFAB03F0CA4E66461" italics="true" pageId="0" pageNumber="141">N. brachyptera</emphasis>
</taxonomicName>
due to similar wing modifications.
</paragraph>
</subSubSection>
<subSubSection id="88D0B78ECE0881636803790646FB3E71" pageId="0" pageNumber="141" type="description">
<paragraph id="E66FA1F4224657253CC2AAB9555884CC" pageId="0" pageNumber="141">Description of male holotype and male paratypes</paragraph>
<paragraph id="43423D82A5514066C973D6B0DB51C2F9" pageId="0" pageNumber="141">
(Figs
<figureCitation id="4DDA1C96F9C4152686B0968FE699150A" captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1 - 3. Nusalala peruana sp. nov., holotype, male. 1. habitus, dorsal; 2. habitus, lateral; 3. head frontal." figureDoi="10.3897/dez.67.56008.figures1-3" httpUri="https://binary.pensoft.net/fig/442147" pageId="0" pageNumber="141">1</figureCitation>
-
<figureCitation id="EBDA638A5056B03B0271B1F75F7B093B" captionStart="Figures 8–11" captionStartId="F3" captionText="Figures 8 - 11. Nusalala peruana sp. nov., paratype, male (from Hualgayoc). 8. genital sclerites, lateral; 9. genital sclerites, ventral; 10. genital sclerites, lateral, partly dissected; 11. genital sclerites, dorsal, partly dissected; e - ectoproct; gp 10 - gonapophysis 10; gst 9 - fused gonostyli 9; gx 9 - gonocoxite 9; gx 10 - partly fused gonapophyses 10; hi - hypandrium internum; S - sternite; T - tergite." figureDoi="10.3897/dez.67.56008.figures8-11" httpUri="https://binary.pensoft.net/fig/442149" pageId="0" pageNumber="141">11</figureCitation>
).
<emphasis id="060D6D968164B565B4BF76BCDF4DDF6B" bold="true" italics="true" pageId="0" pageNumber="141">Head</emphasis>
blackish-brown; frons, labrum and genae polished, without pilosity; occiput covered with golden brownish setae. Antennae with broad cylindrical scapus, slim cylindrical pedicellus, 40-45 moniliform flagellomeres, apically darker. Compound eyes dark, relatively small.
</paragraph>
<caption id="46DF1E4FCF8CD81B585305E0835B1FA5" doi="10.3897/dez.67.56008.figures1-3" httpUri="https://binary.pensoft.net/fig/442147" pageId="0" pageNumber="141" start="Figures 1–3" startId="F1">
<paragraph id="A2331701A3241544AB23B878C77A175B" pageId="0" pageNumber="141">
<emphasis id="0C05602AE3A651E8A2F1EA03E4D14130" bold="true" pageId="0" pageNumber="141">Figures 1-3.</emphasis>
<taxonomicName id="6E3755ABD0C609E823D6D2CDC8362681" authorityName="Aspöck &amp; Aspöck &amp; Gruppe" authorityYear="2020" class="Insecta" family="Hemerobiidae" genus="Nusalala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nusalala peruana" order="Neuroptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="peruana">
<emphasis id="95B546DD446531E53DEC4CFC0D53F15D" italics="true" pageId="0" pageNumber="141">Nusalala peruana</emphasis>
</taxonomicName>
sp. nov., holotype, male.
<emphasis id="690A4CE731C4DDAC2737182F230D8F71" bold="true" pageId="0" pageNumber="141">1.</emphasis>
habitus, dorsal;
<emphasis id="309B6B241BC2B453373F4965BF629338" bold="true" pageId="0" pageNumber="141">2.</emphasis>
habitus, lateral;
<emphasis id="5925E518E19B0C5B796BE2D119DF9DA2" bold="true" pageId="0" pageNumber="141">3.</emphasis>
head frontal.
</paragraph>
</caption>
<caption id="80783E5E17E76EB0A40BFB71F2C67E19" doi="10.3897/dez.67.56008.figures4-7" httpUri="https://binary.pensoft.net/fig/442148" pageId="0" pageNumber="141" start="Figures 4–7" startId="F2">
<paragraph id="9A2CC3D306756749363E172ACDDE7D16" pageId="0" pageNumber="141">
<emphasis id="12503ABFCA8FFE7087136776CC8DAD13" bold="true" pageId="0" pageNumber="141">Figures 4-7.</emphasis>
<taxonomicName id="1417276D82608CE88FE55EFCC57CE72D" authorityName="Aspöck &amp; Aspöck &amp; Gruppe" authorityYear="2020" class="Insecta" family="Hemerobiidae" genus="Nusalala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nusalala peruana" order="Neuroptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="peruana">
<emphasis id="688353EB3252F42E3C554B4A14A62E18" italics="true" pageId="0" pageNumber="141">Nusalala peruana</emphasis>
</taxonomicName>
sp. nov., paratype, male (from Hualgayoc).
<emphasis id="E67BE7B942B3609EF2C4D3705E5447E8" bold="true" pageId="0" pageNumber="141">4.</emphasis>
habitus, dorsal;
<emphasis id="06BBE6191BB2818001FB164A38986ECB" bold="true" pageId="0" pageNumber="141">5.</emphasis>
habitus, ventral;
<emphasis id="9719B54B073955884C3FE8CA4901F2E6" bold="true" pageId="0" pageNumber="141">6.</emphasis>
right forewing (broken by spreading under cover glass);
<emphasis id="7842C39E63E27F72275FCD9AD4474921" bold="true" pageId="0" pageNumber="141">7.</emphasis>
metathorax with hindwings (spread under cover glass).
</paragraph>
</caption>
<paragraph id="4D31C655289F6975DFFE102F83E57CEC" pageId="0" pageNumber="141">
<emphasis id="A8619F75D0B0385C7777E018D0DF35B1" bold="true" italics="true" pageId="0" pageNumber="141">Thorax</emphasis>
: Pronotum shield-like, blackish-brown, with dense brownish pilosity. Legs brownish, only tarsalia yellowish. Each leg with two simple apical claws.
</paragraph>
<paragraph id="498F2C233B09A6F76990B95B32C822C5" pageId="0" pageNumber="141">
<emphasis id="98E49D59B8F836A60D8B4CDE5852F096" bold="true" italics="true" pageId="0" pageNumber="141">Forewings</emphasis>
with significant concavo-convexity, heavily domed, length of domed wings 3.6 mm, length of spread (flattened) wing 3.8 mm, width 1.9 mm, coriaceous, suboval, apically slightly tapered; densely covered with short hairs; colouration spotted brownish with white markings and a brownish stripe in the middle of the wing, which is proximally accompanied by a whitish stripe; colouration, however, extremely variable. Venation variable and irregularly reduced in some specimens. Costal space proximally broad, veins irregularly connected with small interjacent veins, no pterostigma discernible; subcostal space with a few irregular cross veins; radius with 3-5 radial branches; gradate series of radial space not clearly discernible; bases of radius and media fused; venation of median, cubital and anal spaces not clearly ascertainable. Hindwings ribbon-like, length 2.7 mm, width 0.4 mm, with a short, oblique subcosta (?) basally. Longitudinal veins branched, cross veins absent.
</paragraph>
<paragraph id="13E25C95B773C943753A469F083AFDED" pageId="0" pageNumber="141">
<emphasis id="88301D62FFC40FA359BE714623EF67E2" bold="true" italics="true" pageId="0" pageNumber="141">Male genitalia</emphasis>
(Figs
<figureCitation id="AC23A9DA769940F79CA8ADF0DAB5552A" captionStart="Figures 8–11" captionStartId="F3" captionText="Figures 8 - 11. Nusalala peruana sp. nov., paratype, male (from Hualgayoc). 8. genital sclerites, lateral; 9. genital sclerites, ventral; 10. genital sclerites, lateral, partly dissected; 11. genital sclerites, dorsal, partly dissected; e - ectoproct; gp 10 - gonapophysis 10; gst 9 - fused gonostyli 9; gx 9 - gonocoxite 9; gx 10 - partly fused gonapophyses 10; hi - hypandrium internum; S - sternite; T - tergite." figureDoi="10.3897/dez.67.56008.figures8-11" httpUri="https://binary.pensoft.net/fig/442149" pageId="0" pageNumber="141">8-11</figureCitation>
): Tergite 9 divided into a pair of hemitergites, ventrolaterally connected with ectoproct and strongly sclerotised with a broad cephalic apodeme; ectoprocts basally strongly sclerotised, with blunt terminal process. Rosette of trichobothria reduced to a single trichobothrium. Gonocoxites 9 forming broad discs, dorsally fused by a small sclerotised bridge, laterally with two processus, the upper one representing the gonapophyses 9; the fused gonostyli 9 impressively shaped into a long, ventrally directed hook; gonocoxites 10 basally fused to an unpaired sclerite, terminally bilobed, connected with paired and slightly domed sclerites which are addressed as modified gonapophyses 10; hypandrium internum large and ordinarily shaped.
</paragraph>
<caption id="73EFB38469C800C7F258E5D30A6C4EB3" doi="10.3897/dez.67.56008.figures8-11" httpUri="https://binary.pensoft.net/fig/442149" pageId="0" pageNumber="141" start="Figures 8–11" startId="F3">
<paragraph id="BFEE0476325D4E5D0B2530C1EC18AF18" pageId="0" pageNumber="141">
<emphasis id="43D361B742749AC1FED913083779ABA6" bold="true" pageId="0" pageNumber="141">Figures 8-11.</emphasis>
<taxonomicName id="8E1A1E12CBA20C13262DDEA067BDC100" authorityName="Aspöck &amp; Aspöck &amp; Gruppe" authorityYear="2020" class="Insecta" family="Hemerobiidae" genus="Nusalala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nusalala peruana" order="Neuroptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="peruana">
<emphasis id="B6F24F5ED09C047A8FA13EB325DD793C" italics="true" pageId="0" pageNumber="141">Nusalala peruana</emphasis>
</taxonomicName>
sp. nov., paratype, male (from Hualgayoc).
<emphasis id="BB0F699D1605F7012FAE49BA1B20DC0A" bold="true" pageId="0" pageNumber="141">8.</emphasis>
genital sclerites, lateral;
<emphasis id="7C3C462C3E5C1009C2DC0DA2511FF774" bold="true" pageId="0" pageNumber="141">9.</emphasis>
genital sclerites, ventral;
<emphasis id="3CF189A3BEDC71B6556FF78FB8634593" bold="true" pageId="0" pageNumber="141">10.</emphasis>
genital sclerites, lateral, partly dissected;
<emphasis id="CA66FE6A055A0F04F1FC15E607B40E31" bold="true" pageId="0" pageNumber="141">11.</emphasis>
genital sclerites, dorsal, partly dissected;
<emphasis id="158D20ADAA3032A03E86A3B75FB760AF" bold="true" pageId="0" pageNumber="141">e</emphasis>
- ectoproct;
<emphasis id="F04275BF650CDAC2490ABFDA223260E3" bold="true" pageId="0" pageNumber="141">gp10</emphasis>
- gonapophysis 10;
<emphasis id="4E1C4A11BBF789F0AC5F200DAA23EE07" bold="true" pageId="0" pageNumber="141">gst9</emphasis>
- fused gonostyli 9;
<emphasis id="AC68D961EA4E6446360B32443C7B1AEB" bold="true" pageId="0" pageNumber="141">gx9</emphasis>
- gonocoxite 9;
<emphasis id="80EA0BDDB3AD0A15F5D6797FE3B2DBEC" bold="true" pageId="0" pageNumber="141">gx10</emphasis>
- partly fused gonapophyses 10;
<emphasis id="1366C245951F21F7283176EA4A83A45C" bold="true" pageId="0" pageNumber="141">hi</emphasis>
- hypandrium internum;
<emphasis id="F21B22590CFAEDB3BCC41B1F14876A69" bold="true" pageId="0" pageNumber="141">S</emphasis>
- sternite;
<emphasis id="45A5741D8040969052AD8EBBA57AADAD" bold="true" pageId="0" pageNumber="141">T</emphasis>
- tergite.
</paragraph>
</caption>
<paragraph id="A4F6E7B89A7E16E064521CAFC260A9E2" pageId="0" pageNumber="141">
<emphasis id="AE5A0B02E36BC7F304C12BF5F9DDD483" bold="true" pageId="0" pageNumber="141">Description of the female paratype</emphasis>
(Figs
<figureCitation id="96B5CCFDABC6A4BC010D81984B712142" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Nusalala peruana sp. nov., paratype, female (from Hualgayoc). 12. habitus, dorsal with right forewing (left forewing missing) and left scale-like hindwing; 13. scale-like left hindwing; 14. head, frontal; 15. genital sclerites, lateral; 16. genital sclerites, ventral; e - ectoproct; gp 8 - gonapophysis 8; gp 9 - gonapophysis 9; gx 8 - fused gonocoxites 8; gx 9 - gonocoxite 9; T - tergite; tr - trichobothrium." figureDoi="10.3897/dez.67.56008.figures12-16" httpUri="https://binary.pensoft.net/fig/442150" pageId="0" pageNumber="141">12-16</figureCitation>
). Length of the domed forewing approximately 3.2 mm, width 1.9 mm. Habitus similar to male, but specimen in poor condition, only right forewing preserved, left forewing missing. Hindwing scale-like (Fig.
<figureCitation id="1471ACD4613D42C8701B773517674E93" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Nusalala peruana sp. nov., paratype, female (from Hualgayoc). 12. habitus, dorsal with right forewing (left forewing missing) and left scale-like hindwing; 13. scale-like left hindwing; 14. head, frontal; 15. genital sclerites, lateral; 16. genital sclerites, ventral; e - ectoproct; gp 8 - gonapophysis 8; gp 9 - gonapophysis 9; gx 8 - fused gonocoxites 8; gx 9 - gonocoxite 9; T - tergite; tr - trichobothrium." figureDoi="10.3897/dez.67.56008.figures12-16" httpUri="https://binary.pensoft.net/fig/442150" pageId="0" pageNumber="141">13</figureCitation>
).
</paragraph>
<caption id="06811350D1B587EFF66F8485E3EB478E" doi="10.3897/dez.67.56008.figures12-16" httpUri="https://binary.pensoft.net/fig/442150" pageId="0" pageNumber="141" start="Figures 12–16" startId="F4">
<paragraph id="87CF1917CF5A5C5FBBC76D5CFBAEA8CC" pageId="0" pageNumber="141">
<emphasis id="FCD5E32E81F76211E02C92773D73C670" bold="true" pageId="0" pageNumber="141">Figures 12-16.</emphasis>
<taxonomicName id="5FC97D255BEEFFE6102E60B9868D18BC" authorityName="Aspöck &amp; Aspöck &amp; Gruppe" authorityYear="2020" class="Insecta" family="Hemerobiidae" genus="Nusalala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nusalala peruana" order="Neuroptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="peruana">
<emphasis id="6AC703D6FA6300CC575A5D56F20ADF2B" italics="true" pageId="0" pageNumber="141">Nusalala peruana</emphasis>
</taxonomicName>
sp. nov., paratype, female (from Hualgayoc).
<emphasis id="BCFC0954C6DC4F4EBA1218ECC95D29E7" bold="true" pageId="0" pageNumber="141">12.</emphasis>
habitus, dorsal with right forewing (left forewing missing) and left scale-like hindwing;
<emphasis id="466B066ABC2115DB07BF24C23C4DE050" bold="true" pageId="0" pageNumber="141">13.</emphasis>
scale-like left hindwing;
<emphasis id="A9807D26A6037C72A53EFCDD2EF8D3D6" bold="true" pageId="0" pageNumber="141">14.</emphasis>
head, frontal;
<emphasis id="00AD84988F0918C3C7A7EA2EC4C36056" bold="true" pageId="0" pageNumber="141">15.</emphasis>
genital sclerites, lateral;
<emphasis id="316F783033E774ABCB5EA55147AFB69C" bold="true" pageId="0" pageNumber="141">16.</emphasis>
genital sclerites, ventral;
<emphasis id="4C57A6D8805403CFFBFF60E61D06B66E" bold="true" pageId="0" pageNumber="141">e</emphasis>
- ectoproct;
<emphasis id="C342BC6187C4592CF10AF7F9095DC91B" bold="true" pageId="0" pageNumber="141">gp8</emphasis>
- gonapophysis 8;
<emphasis id="AAE87778BF7C810895946D501F994C8A" bold="true" pageId="0" pageNumber="141">gp9</emphasis>
- gonapophysis 9;
<emphasis id="F13A7DC0ADD7D4C7A99DDBDFC8C563FC" bold="true" pageId="0" pageNumber="141">gx8</emphasis>
- fused gonocoxites 8;
<emphasis id="6E4106D238A6F7970BD613B36CF6B0FF" bold="true" pageId="0" pageNumber="141">gx9</emphasis>
- gonocoxite 9;
<emphasis id="F96FA6263B02EBFC62EB621567868C4A" bold="true" pageId="0" pageNumber="141">T</emphasis>
- tergite;
<emphasis id="3995B34010FF5B9DCD9B6EBE1B58D37F" bold="true" pageId="0" pageNumber="141">tr</emphasis>
- trichobothrium.
</paragraph>
</caption>
<paragraph id="B9EAAE9E56163BE71945AB502DFD460E" pageId="0" pageNumber="141">
<emphasis id="EAE389372934F3699E9F8D7C2AE5B8A9" bold="true" italics="true" pageId="0" pageNumber="141">Female genitalia</emphasis>
(Figs
<figureCitation id="15F730A277160F009AC9FDC19F42712E" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Nusalala peruana sp. nov., paratype, female (from Hualgayoc). 12. habitus, dorsal with right forewing (left forewing missing) and left scale-like hindwing; 13. scale-like left hindwing; 14. head, frontal; 15. genital sclerites, lateral; 16. genital sclerites, ventral; e - ectoproct; gp 8 - gonapophysis 8; gp 9 - gonapophysis 9; gx 8 - fused gonocoxites 8; gx 9 - gonocoxite 9; T - tergite; tr - trichobothrium." figureDoi="10.3897/dez.67.56008.figures12-16" httpUri="https://binary.pensoft.net/fig/442150" pageId="0" pageNumber="141">15</figureCitation>
,
<figureCitation id="3028286FE2259E8B10B74C316B6F49A8" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Nusalala peruana sp. nov., paratype, female (from Hualgayoc). 12. habitus, dorsal with right forewing (left forewing missing) and left scale-like hindwing; 13. scale-like left hindwing; 14. head, frontal; 15. genital sclerites, lateral; 16. genital sclerites, ventral; e - ectoproct; gp 8 - gonapophysis 8; gp 9 - gonapophysis 9; gx 8 - fused gonocoxites 8; gx 9 - gonocoxite 9; T - tergite; tr - trichobothrium." figureDoi="10.3897/dez.67.56008.figures12-16" httpUri="https://binary.pensoft.net/fig/442150" pageId="0" pageNumber="141">16</figureCitation>
): Sternite 7 inconspicuous; tergite 8 reaching beyond spiraculum; gonocoxites 8 fused to a small strongly sclerotised sclerite (subgenitale); paired small ovoid internal sclerites are interpreted as the highly-reduced gonapophyses 8; tergite 9 small, dorsally divided, ventrolaterally broadened; gonocoxites 9 ovoid, connected with tiny (reduced) sclerites which are interpreted as gonapophyses 9; ectoproct dorsally divided, with a single trichobothrium.
</paragraph>
<paragraph id="79E81C6B475CEEF3D83A08075F866188" pageId="0" pageNumber="141">
Glandular systems, as described and figured in
<bibRefCitation id="B2B99076D10330414E9E3F4E57EC3390" author="Monserrat, VJ" journalOrPublisher="Fragmenta Entomologica" pageId="0" pageNumber="141" pagination="83 - 162" refId="B21" refString="Monserrat, VJ, 2000. Revision del genero Nusalala (Neuroptera, Hemerobiidae). Fragmenta Entomologica 32 (1): 83 - 162" title="Revision del genero Nusalala (Neuroptera, Hemerobiidae)." volume="32" year="2000">Monserrat (2000)</bibRefCitation>
, could not be found in the single available female specimen.
</paragraph>
</subSubSection>
<subSubSection id="1A21F13AB88A5795E118E1C53D942745" pageId="0" pageNumber="141" type="materials_examined">
<paragraph id="737360AB23A0CE45950DF2454E121CEB" pageId="0" pageNumber="141">Material studied.</paragraph>
<paragraph id="E3E65864FFFF8D767E205B33C5D7A8E7" pageId="0" pageNumber="141">
<specimenCount id="B244E9DEE094C49FB90236851002AEDD" type="male">13 males</specimenCount>
:
<typeStatus id="F2BDB4A8731A435FC3333079D116260C">Holotype</typeStatus>
and
<typeStatus id="A3B179C6B5705560511311982B426F1F">paratypes</typeStatus>
.
</paragraph>
<paragraph id="F7FD50E7E13E55540C1B3B8CA1DC4547" pageId="0" pageNumber="141">
<materialsCitation id="F16BCEE42E876D3499EB4CE16B355CAD" collectingDate="2017-09-07" collectingDateMax="2017-09-14" collectingDateMin="2017-09-07" collectorName="L. Figueroa" country="PERU" elevation="3851" latitude="-6.770761" location="Hualgayoc" longLatPrecision="1" longitude="-78.62513" specimenCount="1" specimenCount-male="1" typeStatus="Holotype">
<typeStatus id="438BE4AE70E2A83C1623290841F85B3A">Holotype</typeStatus>
and
<specimenCount id="D3FEBF6F08163B4A432A947068D95BD6" type="male">1 male</specimenCount>
<typeStatus id="118288676C3FF46D0978DEF28A8881DC">paratype</typeStatus>
: &quot;
<collectingCountry id="EA49594AFA19EDC2387DD30C34051BD5" name="Peru">PERU</collectingCountry>
. CA.
<location id="61C1FAC401B7350FDD56F095592E3410" LSID="urn:lsid:plazi:treatment:A0FCDD99D9825CFCB4F3C80995CC9818:61C1FAC401B7350FDD56F095592E3410" country="PERU" latitude="-6.770761" longLatPrecision="1" longitude="-78.62513" name="Hualgayoc">Hualgayoc</location>
,
<geoCoordinate id="7F1F9B726B3CA276D87AEBCCEA270A48" degrees="6" direction="south" minutes="46" orientation="latitude" precision="1" seconds="14.74" value="-6.770761">6°46'14.74&quot;S</geoCoordinate>
,
<geoCoordinate id="41778A7DF1D3B1CCD678F7B7A64089E9" degrees="78" direction="west" minutes="37" orientation="longitude" precision="1" seconds="30.45" value="-78.62513">78°37'30.45&quot;W</geoCoordinate>
,
<quantity id="029EDDCB835A810BCDEB6C506328775A" metricMagnitude="3" metricUnit="m" metricValue="3.851" unit="m" value="3851.0">
<elevation id="6CE6B54C15CBA4E3BE3552CA470D5C38" metricMagnitude="3" metricUnit="m" metricValue="3.851" unit="m" value="3851.0">3851 m</elevation>
</quantity>
alt.,
<collectingDate id="C54FFC71ABE75C0D662D929B3A372A9D" value="2017-09-07" valueMax="2017-09-14" valueMin="2017-09-07">7-14.iX.2017</collectingDate>
,
<collectorName id="4D6B774073199AE799002C2BC43A0874">L. Figueroa</collectorName>
leg.&quot;;
</materialsCitation>
</paragraph>
<paragraph id="72CE206EB940A39A67EF7AF1FAD8DCBB" pageId="0" pageNumber="141">
<materialsCitation id="1CE2599B7D6C4616D423CD95949B017A" collectingDate="2017-09-07" collectingDateMax="2017-09-14" collectingDateMin="2017-09-07" collectorName="L. Figueroa" country="PERU" elevation="3997" latitude="-6.758625" location="Hualgayoc" longLatPrecision="1" longitude="-78.622185" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="EFB9D871E3B09C6935343A9BE7016896" type="male">1 male</specimenCount>
<typeStatus id="E24B6BA55B7A921FD0F3A542AE47B2F5">paratype</typeStatus>
: &quot;
<collectingCountry id="D158EE00CA3DB2F987FDBFB880FE9FF7" name="Peru">PERU</collectingCountry>
. CA.
<location id="BBFD6A9A6B49A34C5E3BB46F514C7562" LSID="urn:lsid:plazi:treatment:A0FCDD99D9825CFCB4F3C80995CC9818:BBFD6A9A6B49A34C5E3BB46F514C7562" country="PERU" latitude="-6.758625" longLatPrecision="1" longitude="-78.622185" name="Hualgayoc">Hualgayoc</location>
,
<geoCoordinate id="0650978D55CAD58EBBB52B02E6B65594" degrees="6" direction="south" minutes="45" orientation="latitude" precision="1" seconds="31.05" value="-6.758625">6°45'31.05&quot;S</geoCoordinate>
,
<geoCoordinate id="58680E49CBCD05FEA6635FB462976EA1" degrees="78" direction="west" minutes="37" orientation="longitude" precision="1" seconds="19.85" value="-78.622185">78°37'19.85&quot;W</geoCoordinate>
,
<quantity id="77D199620EE8803BE654C72073AF0EDE" metricMagnitude="3" metricUnit="m" metricValue="3.997" unit="m" value="3997.0">
<elevation id="CA61B5823EA86DCA29D33F4F955CE548" metricMagnitude="3" metricUnit="m" metricValue="3.997" unit="m" value="3997.0">3997 m</elevation>
</quantity>
alt.,
<collectingDate id="BC8BED55EE4D96E62F1AB34AA66E2987" value="2017-09-07" valueMax="2017-09-14" valueMin="2017-09-07">7-14.iX.2017</collectingDate>
,
<collectorName id="87C48B098616261BDDD6085665331C70">L. Figueroa</collectorName>
leg.&quot;
</materialsCitation>
;
<materialsCitation id="A1BB8E32C054B833834375B1FC40CAD1" collectingDate="2018-07-19" collectingDateMax="2018-07-26" collectingDateMin="2018-07-19" collectorName="L. Figueroa" country="PERU" elevation="3968" latitude="-6.766328" location="Hualgayoc" longLatPrecision="1" longitude="-78.62608" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="7846C25032B61D5565070BFC60EBDB4C" type="male">1 male</specimenCount>
<typeStatus id="CD3CB178D53C6EC345BAA83C9E9C1D06">paratype</typeStatus>
: &quot;
<collectingCountry id="4B6F90BEAD86870DB15620D95C443D26" name="Peru">PERU</collectingCountry>
. CA.
<location id="75B161CF8B28E0B920E375B67FB1C9A2" LSID="urn:lsid:plazi:treatment:A0FCDD99D9825CFCB4F3C80995CC9818:75B161CF8B28E0B920E375B67FB1C9A2" country="PERU" latitude="-6.766328" longLatPrecision="1" longitude="-78.62608" name="Hualgayoc">Hualgayoc</location>
,
<geoCoordinate id="E17C5F8D4B8A08D0666651F6A3E984BE" degrees="6" direction="south" minutes="45" orientation="latitude" precision="1" seconds="58.78" value="-6.766328">6°45'58.78&quot;S</geoCoordinate>
,
<geoCoordinate id="2F68004F094D8CC6DB5D6ABC99759837" degrees="78" direction="west" minutes="37" orientation="longitude" precision="1" seconds="33.88" value="-78.62608">78°37'33.88&quot;W</geoCoordinate>
,
<quantity id="28AFD6339BC04657A199157A49BE12AC" metricMagnitude="3" metricUnit="m" metricValue="3.968" unit="m" value="3968.0">
<elevation id="06848E6C7DBD4C2377A11FC9B3AC58C0" metricMagnitude="3" metricUnit="m" metricValue="3.968" unit="m" value="3968.0">3968 m</elevation>
</quantity>
alt.,
<collectingDate id="7068A4666736604CFD9B93D8FD55DEE2" value="2018-07-19" valueMax="2018-07-26" valueMin="2018-07-19">19.-26.Vii.2018</collectingDate>
,
<collectorName id="A6F8A51E37A6442DC1208940B7B1CBF2">L. Figueroa</collectorName>
leg.&quot;
</materialsCitation>
;
<materialsCitation id="906B2CE3706A40F767AAEF25C62F0EC3" collectingDate="2018-07-19" collectingDateMax="2018-07-26" collectingDateMin="2018-07-19" collectorName="L. Figueroa" country="PERU" elevation="3875" latitude="-6.7563696" location="Hualgayoc" longLatPrecision="1" longitude="-78.62457" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="EB4044C064B109B71127144422702BBF" type="male">1 male</specimenCount>
<typeStatus id="4873B9F40E8EC61E58190F832A6D2402">paratype</typeStatus>
: &quot;
<collectingCountry id="F2BE6276B15E4E591447A8CB91CAE738" name="Peru">PERU</collectingCountry>
. CA.
<location id="AAB5F01584E0ADE1CF4432C5495F5A21" LSID="urn:lsid:plazi:treatment:A0FCDD99D9825CFCB4F3C80995CC9818:AAB5F01584E0ADE1CF4432C5495F5A21" country="PERU" latitude="-6.7563696" longLatPrecision="1" longitude="-78.62457" name="Hualgayoc">Hualgayoc</location>
,
<geoCoordinate id="954D03633C7E1BBC61ECD7321E87E846" degrees="6" direction="south" minutes="45" orientation="latitude" precision="1" seconds="22.93" value="-6.7563696">6°45'22.93&quot;S</geoCoordinate>
,
<geoCoordinate id="714AB2BF5DB763B66DE3B01F4D7FDDC1" degrees="78" direction="west" minutes="37" orientation="longitude" precision="1" seconds="28.45" value="-78.62457">78°37'28.45&quot;W</geoCoordinate>
,
<quantity id="322B33EBBE39CA51037F8867D5A2BE4C" metricMagnitude="3" metricUnit="m" metricValue="3.875" unit="m" value="3875.0">
<elevation id="20F567ADDACDC10C96099CFB58F72CB9" metricMagnitude="3" metricUnit="m" metricValue="3.875" unit="m" value="3875.0">3875 m</elevation>
</quantity>
alt.,
<collectingDate id="A44791CE79AD311FC4AB6A80AADA8F32" value="2018-07-19" valueMax="2018-07-26" valueMin="2018-07-19">19.-26.Vii.2018</collectingDate>
,
<collectorName id="585E72B2199825CFB5926A44574875DA">L. Figueroa</collectorName>
leg.&quot;
</materialsCitation>
;
<materialsCitation id="DDC357D2B9EE8E3D950BAD677A817A5A" collectingDate="2019-07-22" collectingDateMax="2019-07-27" collectingDateMin="2019-07-22" collectorName="P. Sanchez" country="PERU" elevation="3790" latitude="-6.7639027" location="Hualgayoc" longLatPrecision="1" longitude="-78.651726" specimenCount="4" specimenCount-male="4" typeStatus="paratypes">
<specimenCount id="D11895A0A0BBA369C49195BC9701A3A0" type="male">4 males</specimenCount>
<typeStatus id="A8B3D690ADA7E8CAA07B9DF833F9B688">paratypes</typeStatus>
: &quot;
<collectingCountry id="E8E1A017218D89F1DA6E617035CEF7B0" name="Peru">PERU</collectingCountry>
. CA.
<location id="1743CBAD8BC2295BD168EFAD3D6A8B4D" LSID="urn:lsid:plazi:treatment:A0FCDD99D9825CFCB4F3C80995CC9818:1743CBAD8BC2295BD168EFAD3D6A8B4D" country="PERU" latitude="-6.7639027" longLatPrecision="1" longitude="-78.651726" name="Hualgayoc">Hualgayoc</location>
,
<geoCoordinate id="FB5FB3FBDD7F10DCCF0A6FA66C5CAF81" degrees="6" direction="south" minutes="45" orientation="latitude" precision="1" seconds="50.05" value="-6.7639027">6°45'50.05&quot;S</geoCoordinate>
,
<geoCoordinate id="E6873312B2DC136A46A9970444A0F355" degrees="78" direction="west" minutes="39" orientation="longitude" precision="1" seconds="6.20" value="-78.651726">78°39'6.20&quot;W</geoCoordinate>
,
<quantity id="1A6DE19BEEA60CD787302D30B5A9B7C9" metricMagnitude="3" metricUnit="m" metricValue="3.79" unit="m" value="3790.0">
<elevation id="0069F5B6FCB6072D95BCFE187F45A9F4" metricMagnitude="3" metricUnit="m" metricValue="3.79" unit="m" value="3790.0">3790 m</elevation>
</quantity>
alt.,
<collectingDate id="1DD3A9F1CE3AD4AC2C440DAA5E0C3899" value="2019-07-22" valueMax="2019-07-27" valueMin="2019-07-22">22.-27.Vii.2019</collectingDate>
,
<collectorName id="E6EDCEADD33138362223CBB1B701200D">
P.
<normalizedToken id="3F5E84FFB1B0EFDD64834C7E50F8BE25" originalValue="Sánchez">Sanchez</normalizedToken>
</collectorName>
leg.&quot;
</materialsCitation>
;
<materialsCitation id="90EF9EB6A28978254E4B6B19D4B63B28" collectingDate="2019-03-15" collectingDateMax="2019-03-20" collectingDateMin="2019-03-15" collectorName="P. Sanchez" country="PERU" elevation="3790" latitude="-6.7639027" location="Hualgayoc" longLatPrecision="1" longitude="-78.651726" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="FB128DCFE83E5EF425FC77419CA24069" type="male">1 male</specimenCount>
<typeStatus id="1DEB9D526314FB2A13AD2223A8936412">paratype</typeStatus>
: &quot;
<collectingCountry id="4A7EBB171E48821B57713E7B62C84680" name="Peru">PERU</collectingCountry>
. CA.
<location id="39F597082B477B9EFD77A577549E9071" LSID="urn:lsid:plazi:treatment:A0FCDD99D9825CFCB4F3C80995CC9818:39F597082B477B9EFD77A577549E9071" country="PERU" latitude="-6.7639027" longLatPrecision="1" longitude="-78.651726" name="Hualgayoc">Hualgayoc</location>
,
<geoCoordinate id="892725C50752B5FD041E3148C2322B6F" degrees="6" direction="south" minutes="45" orientation="latitude" precision="1" seconds="50.05" value="-6.7639027">6°45'50.05&quot;S</geoCoordinate>
,
<geoCoordinate id="F64CF90CB3FC59CD5711D31DC71AA583" degrees="78" direction="west" minutes="39" orientation="longitude" precision="1" seconds="6.20" value="-78.651726">78°39'6.20&quot;W</geoCoordinate>
,
<quantity id="CE4558758FBC166FA910C104980C375D" metricMagnitude="3" metricUnit="m" metricValue="3.79" unit="m" value="3790.0">
<elevation id="BF1311A4AB98621F87C1512384A2A109" metricMagnitude="3" metricUnit="m" metricValue="3.79" unit="m" value="3790.0">3790 m</elevation>
</quantity>
alt.,
<collectingDate id="B015D6B5DFFE74048A7FD50791E0CE81" value="2019-03-15" valueMax="2019-03-20" valueMin="2019-03-15">15.-20.iii.2019</collectingDate>
,
<collectorName id="8212FFD0A81FE65FBDCD4E81592B25BB">
P.
<normalizedToken id="6A9BA77CF939097BB164B96C3398A93C" originalValue="Sánchez">Sanchez</normalizedToken>
</collectorName>
leg.&quot;
</materialsCitation>
;
<materialsCitation id="A5F38649C1D7D55C0CA13730BF9CDD9A" collectingDate="2019-03-15" collectingDateMax="2019-03-20" collectingDateMin="2019-03-15" collectorName="P. Sanchez" country="PERU" elevation="3756" latitude="-6.7617083" location="Hualgayoc" longLatPrecision="1" longitude="-78.64844" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="2033521B4C4B0FDBB0996C0868EC8176" type="male">1 male</specimenCount>
<typeStatus id="B1A07D0CDEC8213B40A2F40B65D65428">paratype</typeStatus>
: &quot;
<collectingCountry id="8B6AA52565C86CBDA356B153967184ED" name="Peru">PERU</collectingCountry>
. CA.
<location id="B9AB64EBB6C29790B65D1DA3A5AB70F5" LSID="urn:lsid:plazi:treatment:A0FCDD99D9825CFCB4F3C80995CC9818:B9AB64EBB6C29790B65D1DA3A5AB70F5" country="PERU" latitude="-6.7617083" longLatPrecision="1" longitude="-78.64844" name="Hualgayoc">Hualgayoc</location>
,
<geoCoordinate id="6D6BE17239D807B7661BD31E3C3DAE20" degrees="6" direction="south" minutes="45" orientation="latitude" precision="1" seconds="42.15" value="-6.7617083">6°45'42.15&quot;S</geoCoordinate>
,
<geoCoordinate id="CA90966C5D6D462800EBB952897D2FE8" degrees="78" direction="west" minutes="38" orientation="longitude" precision="1" seconds="54.39" value="-78.64844">78°38' 54.39&quot;W</geoCoordinate>
,
<quantity id="C79FC082050A3B2129E16708081F81E8" metricMagnitude="3" metricUnit="m" metricValue="3.756" unit="m" value="3756.0">
<elevation id="DDB9ACAAD4DED46306C8F99C7C0FAAFF" metricMagnitude="3" metricUnit="m" metricValue="3.756" unit="m" value="3756.0">3756 m</elevation>
</quantity>
alt.,
<collectingDate id="80147DF348F25BA0AA7E2F14236D00E2" value="2019-03-15" valueMax="2019-03-20" valueMin="2019-03-15">15.-20.iii.2019</collectingDate>
,
<collectorName id="A809BC34FCEE8EB12049774B7C4AFA81">
P.
<normalizedToken id="0681FE333B25C163E7B8448BB3A8690E" originalValue="Sánchez">Sanchez</normalizedToken>
</collectorName>
leg.&quot;
</materialsCitation>
;
<materialsCitation id="2EA7AD842022F3AA7E5DFFC0567A434A" collectingDate="2019-07-22" collectingDateMax="2019-07-27" collectingDateMin="2019-07-22" collectorName="P. Sanchez" country="PERU" elevation="3875" latitude="-6.7563696" location="Hualgayoc" longLatPrecision="1" longitude="-78.62457" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="66477A6E132220EB44A0FA0DF3D86A62" type="male">1 male</specimenCount>
<typeStatus id="8098743D24CBFCD43660B0A64A3483C4">paratype</typeStatus>
: &quot;
<collectingCountry id="28BA549CA91D39620197D2AE7FCC43C5" name="Peru">PERU</collectingCountry>
. CA.
<location id="541EF7A6D3951E9DB057FC9716F058D6" LSID="urn:lsid:plazi:treatment:A0FCDD99D9825CFCB4F3C80995CC9818:541EF7A6D3951E9DB057FC9716F058D6" country="PERU" latitude="-6.7563696" longLatPrecision="1" longitude="-78.62457" name="Hualgayoc">Hualgayoc</location>
,
<geoCoordinate id="2D3477DB9469CD4BB5484FC5C9DD53B4" degrees="6" direction="south" minutes="45" orientation="latitude" precision="1" seconds="22.93" value="-6.7563696">6°45'22.93&quot;S</geoCoordinate>
,
<geoCoordinate id="E027FC5D01E3C40F2672515F42849BD4" degrees="78" direction="west" minutes="37" orientation="longitude" precision="1" seconds="28.45" value="-78.62457">78°37'28.45&quot;W</geoCoordinate>
,
<quantity id="91B65B116A972D8099184F520607C3B3" metricMagnitude="3" metricUnit="m" metricValue="3.875" unit="m" value="3875.0">
<elevation id="19724FED698910FDC5563ABE8296071D" metricMagnitude="3" metricUnit="m" metricValue="3.875" unit="m" value="3875.0">3875 m</elevation>
</quantity>
alt.,
<collectingDate id="EBF4D225DE4DB85D5A15F02316769C18" value="2019-07-22" valueMax="2019-07-27" valueMin="2019-07-22">22.-27.Vii.2019</collectingDate>
,
<collectorName id="5564036A05E52B9D9E7146307A98A35F">
P.
<normalizedToken id="79C4D6C830EDDDB6FF865AFA49F61F26" originalValue="Sánchez">Sanchez</normalizedToken>
</collectorName>
leg.&quot;
</materialsCitation>
;
<materialsCitation id="E693512D3220116946E999A34C78FD7E" collectingDate="2019-07-22" collectingDateMax="2019-07-27" collectingDateMin="2019-07-22" collectorName="P. Sanchez" country="PERU" elevation="3805" latitude="-6.7820773" location="Hualgayoc" longLatPrecision="1" longitude="-78.62925" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="187100EE1991664B3635775EEE2046D9" type="male">1 male</specimenCount>
<typeStatus id="6594B34B3B99BD2FB0D67E4FC90B5B6F">paratype</typeStatus>
: &quot;
<collectingCountry id="9A6FEE853A5085A3A7CA514EAA4B3B68" name="Peru">PERU</collectingCountry>
. CA.
<location id="91AAE3DB5C9D6CFEACF0E21336A9A892" LSID="urn:lsid:plazi:treatment:A0FCDD99D9825CFCB4F3C80995CC9818:91AAE3DB5C9D6CFEACF0E21336A9A892" country="PERU" latitude="-6.7820773" longLatPrecision="1" longitude="-78.62925" name="Hualgayoc">Hualgayoc</location>
,
<geoCoordinate id="A947CE29D44F29DB46C26F67DDD141E6" degrees="6" direction="south" minutes="46" orientation="latitude" precision="1" seconds="55.48" value="-6.7820773">6°46'55.48&quot;S</geoCoordinate>
,
<geoCoordinate id="0BE52B897662F21B7B924962ADD25403" degrees="78" direction="west" minutes="37" orientation="longitude" precision="1" seconds="45.29" value="-78.62925">78°37'45.29&quot;W</geoCoordinate>
,
<quantity id="03E53DF7DCDA0C31DC660056CA3DCE36" metricMagnitude="3" metricUnit="m" metricValue="3.8049999999999997" unit="m" value="3805.0">
<elevation id="531BA006294856999C66B951FE9203CA" metricMagnitude="3" metricUnit="m" metricValue="3.8049999999999997" unit="m" value="3805.0">3805 m</elevation>
</quantity>
alt.,
<collectingDate id="BDA1E242EE08C8C9F7369FC5E96BCE54" value="2019-07-22" valueMax="2019-07-27" valueMin="2019-07-22">22.-27.Vii.2019</collectingDate>
,
<collectorName id="7E596DF7845B9090DB541F562B96BEC1">
P.
<normalizedToken id="2D4FC7B0ECEC1AB661C508724A2EAF49" originalValue="Sánchez">Sanchez</normalizedToken>
</collectorName>
leg.&quot;
</materialsCitation>
</paragraph>
<paragraph id="29E8C2BD49AAE5204EA018CF6F81E11A" pageId="0" pageNumber="141">
<materialsCitation id="DF6F99CC0B378F499DB28BA5AEF01924" collectingDate="2017-09-07" collectingDateMax="2017-09-14" collectingDateMin="2017-09-07" collectorName="L. Figueroa" country="PERU" elevation="3790" latitude="-6.7639027" location="Hualgayoc" longLatPrecision="1" longitude="-78.651726" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="C90829B1B2427010018A31AC63636A56" type="female">1 female</specimenCount>
<typeStatus id="8EF18239AC5F3FDE73BB68E97C93C5B8">paratype</typeStatus>
: &quot;
<collectingCountry id="32BC121CE005FE1B3EE9188E14740159" name="Peru">PERU</collectingCountry>
. CA.
<location id="43B6D6F228260DDEC7C769A0351D5D38" LSID="urn:lsid:plazi:treatment:A0FCDD99D9825CFCB4F3C80995CC9818:43B6D6F228260DDEC7C769A0351D5D38" country="PERU" latitude="-6.7639027" longLatPrecision="1" longitude="-78.651726" name="Hualgayoc">Hualgayoc</location>
,
<geoCoordinate id="B525F72B2A671FEAE3C3FE27C3C62B34" degrees="6" direction="south" minutes="45" orientation="latitude" precision="1" seconds="50.05" value="-6.7639027">6°45'50.05&quot;S</geoCoordinate>
,
<geoCoordinate id="3DB463EE55CCE180F1723066FC945A6F" degrees="78" direction="west" minutes="39" orientation="longitude" precision="1" seconds="6.20" value="-78.651726">78°39'6.20&quot;W</geoCoordinate>
,
<quantity id="F86441DB269D1E4B5AD5BFD8209318D4" metricMagnitude="3" metricUnit="m" metricValue="3.79" unit="m" value="3790.0">
<elevation id="34889ABCC9E0FEC099BEF4E9849A9E2F" metricMagnitude="3" metricUnit="m" metricValue="3.79" unit="m" value="3790.0">3790 m</elevation>
</quantity>
alt.,
<collectingDate id="91F39DD12E903FC9762B49BA19C5DF96" value="2017-09-07" valueMax="2017-09-14" valueMin="2017-09-07">7.-14.iX.2017</collectingDate>
,
<collectorName id="CD7A900E3BA71DCCA41791B78CE7EB1D">L. Figueroa</collectorName>
leg.&quot;
</materialsCitation>
</paragraph>
<paragraph id="2335D6298FF5B1DC21B0E2DA59917556" pageId="0" pageNumber="141">
<materialsCitation id="1CADE8223B31347B61095BF796336AC8" collectingDate="2020-01-01" collectingDateMax="2020-12-31" collectingDateMin="2020-01-01" collectorName="Natural History Museum, Vienna." location="Bavarian State Collection of Zoology" municipality="Museo de Historia Natural" specimenCount="20" specimenCount-female="1" specimenCount-male="12" stateProvince="Lima" typeStatus="Holotype">
<typeStatus id="0571F91753A9EC79CABCCF985A596B26">Holotype</typeStatus>
male,
<specimenCount id="6096232CC32C8577133936C64A6A3B91" type="generic" typeStatus="paratypes">7 paratypes</specimenCount>
(
<specimenCount id="2847569DE7B39A2224389FEF702E588D" type="male">6 males</specimenCount>
,
<specimenCount id="B2283EF4EC339FE880F7D5335C8B8588" type="female">1 female</specimenCount>
) will be deposited in coll.
<collectingMunicipality id="B231A38A611EAE1324EDAC1AA89956FD">Museo de Historia Natural</collectingMunicipality>
<collectingRegion id="D90D30219B663AD10922274C6295E117" country="Peru" name="Lima">Lima</collectingRegion>
,
<specimenCount id="9DF324100DF8911E3370F92A09AE663D" type="male">3 male</specimenCount>
<typeStatus id="F15B604BEC10235E2EF76024D118CB67">paratypes</typeStatus>
in coll.
<location id="C30E6C3E23A758A65C84C712ED94BEFD" LSID="urn:lsid:plazi:treatment:A0FCDD99D9825CFCB4F3C80995CC9818:C30E6C3E23A758A65C84C712ED94BEFD" municipality="Museo de Historia Natural" name="Bavarian State Collection of Zoology" stateProvince="Lima">Bavarian State Collection of Zoology</location>
, Munich and
<specimenCount id="9D4E475008E8706B704790D189BAB643" type="male">3 male</specimenCount>
<typeStatus id="C7F6908D15E76B6868F7087B428804D3">paratypes</typeStatus>
in coll.
<collectorName id="3FDA14EED49772D8DA28F6B83DD92DCC">Natural History Museum</collectorName>
,
<collectorName id="5E6B1F88B6D0EB1379BDCF872FDAD392">Vienna.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="66BAE1BEAC061D2772E2F5C501410CE2" pageId="0" pageNumber="141" type="distribution">
<paragraph id="25BDC3F71643551D05B0A2A35602C7A9" pageId="0" pageNumber="141">Distribution.</paragraph>
<paragraph id="9F7BEDC30FD27F94D7096945589DE25B" pageId="0" pageNumber="141">Currently known only from the type locality in the province of Hualgayoc, Cajamarca region, Peru.</paragraph>
</subSubSection>
<subSubSection id="48BF6CC66AF10EDE57D2A2053BC0EDED" pageId="0" pageNumber="141" type="ecology">
<paragraph id="AF2467A8F6ECD96D4738200901AA16CD" pageId="0" pageNumber="141">Ecology.</paragraph>
<paragraph id="31147E8AAF6E4984B3B92151E31EF779" pageId="0" pageNumber="141">
The specimens were collected in a Puna grassland (Figs
<figureCitation id="BC7ABFB59F853D1041F0F32EF1F2B64A" captionStart="Figures 17–19" captionStartId="F5" captionText="Figures 17 - 19. Type locality of Nusalala peruana sp. nov. 17 - landscape; 18, 19 - microhabitat with yellow pans in which Nusalala peruana sp. nov. was trapped." figureDoi="10.3897/dez.67.56008.figures17-19" httpUri="https://binary.pensoft.net/fig/442151" pageId="0" pageNumber="141">17-19</figureCitation>
). Altitudes of findings range from 3756 m to 3997 m, time of collecting from March to September in the years 2017 to 2019. The specimens were collected around the mine Cerro Corona within an area of 3 km
<normalizedToken id="A14D7AD6E17E605FBAF0D5C6484C8B00" originalValue="×">x</normalizedToken>
3.5 km; the collecting protocol included pitfall and yellow pan traps; however, the specimens were found solely in the yellow pan traps.
</paragraph>
</subSubSection>
<subSubSection id="525D17B66ECD56FCE8731D1493E2AB5E" pageId="0" pageNumber="141" type="etymology">
<paragraph id="E1CA038C1D8194A2285976EBA48CB0E3" pageId="0" pageNumber="141">Etymology.</paragraph>
<paragraph id="28C6D29F8A86F32C6A3D4B4D47AE8D38" pageId="0" pageNumber="141">
<taxonomicName id="943DAE160E5588BDFA0A1519A1D3F593" authorityName="Aspöck &amp; Aspöck &amp; Gruppe" authorityYear="2020" class="Insecta" family="Hemerobiidae" genus="Nusalala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nusalala peruana" order="Neuroptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="peruana">
<emphasis id="AE45F58BFC9FD2098FC4895EB7169BA4" italics="true" pageId="0" pageNumber="141">Nusalala peruana</emphasis>
</taxonomicName>
= Peruvian, adjective, feminine. The epithet is an adjective apposition to the genus
<taxonomicName id="32EC204561A7C95D348DC89A3378A7FD" authorityName="Navas" authorityYear="1913" class="Insecta" family="Hemerobiidae" genus="Nusalala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nusalala" order="Neuroptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="7542041778327780E5AF63047F602015" italics="true" pageId="0" pageNumber="141">Nusalala</emphasis>
</taxonomicName>
(feminine).
</paragraph>
<caption id="FC6FE69532D930AA7AF1156ED487147F" doi="10.3897/dez.67.56008.figures17-19" httpUri="https://binary.pensoft.net/fig/442151" pageId="0" pageNumber="141" start="Figures 17–19" startId="F5">
<paragraph id="1735D51D46C9431AE92BBD7AF7D67D62" pageId="0" pageNumber="141">
<emphasis id="A3C6946048F77D6B97040EC0F1A6F9C7" bold="true" pageId="0" pageNumber="141">Figures 17-19.</emphasis>
Type locality of
<taxonomicName id="4F08B029F14893E1BF4828026C8E4773" authorityName="Aspöck &amp; Aspöck &amp; Gruppe" authorityYear="2020" class="Insecta" family="Hemerobiidae" genus="Nusalala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nusalala peruana" order="Neuroptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="peruana">
<emphasis id="24F9D9AF7AB2C30194A48D75F3E5A805" italics="true" pageId="0" pageNumber="141">Nusalala peruana</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="101D32F7013690F80BD6F85253AE830F" bold="true" pageId="0" pageNumber="141">17</emphasis>
- landscape;
<emphasis id="58477110E7E46A1DE66712E0F6C0D138" bold="true" pageId="0" pageNumber="141">18, 19</emphasis>
- microhabitat with yellow pans in which
<taxonomicName id="CF30ADF44C46752C5BA5D8B1D9DDD73F" authorityName="Aspöck &amp; Aspöck &amp; Gruppe" authorityYear="2020" class="Insecta" family="Hemerobiidae" genus="Nusalala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nusalala peruana" order="Neuroptera" pageId="0" pageNumber="141" phylum="Arthropoda" rank="species" species="peruana">
<emphasis id="9C951F284FA2FD0001C30AD236DE55B2" italics="true" pageId="0" pageNumber="141">Nusalala peruana</emphasis>
</taxonomicName>
sp. nov. was trapped.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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