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<document id="1C5736D909997A22EFBA79E6FC86E6DA" ID-CLB-Dataset="34707" ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. &amp; Tishechkin, Alexey K." docDate="2013" docId="A0D50F9207A4F3F5D4AB6BFC82F63A91" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus angustisternus Wenzel 1944" docType="treatment" docVersion="5" lastPageNumber="372" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="368" updateTime="1732934305285" updateUser="ExternalLinkService">
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<mods:title id="7E0E53308D0343E6C415A32F7A341FF1">A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
</mods:titleInfo>
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<mods:namePart id="8B96D58D7FC5C29D80E32AEFD9238EA8">Caterino, Michael S.</mods:namePart>
</mods:name>
<mods:name id="C3874FDEF521F77DF1757FA9947DA062" type="personal">
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</mods:role>
<mods:namePart id="7E562317F7EF870EE9076112A2982508">Tishechkin, Alexey K.</mods:namePart>
</mods:name>
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<mods:date id="D69A50C9E40C2E476833E7E9B9E57E91">2013</mods:date>
<mods:detail id="F17A4A1A483771589EF96DEC6F3B1374" type="volume">
<mods:number id="BE4E4A1997AE83EB1834EE6519E59985">271</mods:number>
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<mods:url id="8B0732ABFFBC790E89BA46BD22C5C656">http://dx.doi.org/10.3897/zookeys.271.4062</mods:url>
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<mods:classification id="104022269402B1CD1324518D208FDEA3">journal article</mods:classification>
<mods:identifier id="2253B549BA6858825E6CE7DDBD54343A" type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
<mods:identifier id="449F54AAB1520CB11588D46B03B59178" type="Pensoft-Pub">1313-2970-271-1</mods:identifier>
</mods:mods>
<treatment id="A0D50F9207A4F3F5D4AB6BFC82F63A91" ID-GBIF-Taxon="152040877" LSID="urn:lsid:plazi:treatment:A0D50F9207A4F3F5D4AB6BFC82F63A91" httpUri="http://treatment.plazi.org/id/A0D50F9207A4F3F5D4AB6BFC82F63A91" lastPageId="371" lastPageNumber="372" pageId="367" pageNumber="368" scope_family="Histeridae" scope_order="Coleoptera" scope_tribe="Exosternini">
<subSubSection id="833D5810FF934C6F816088C92FBC9321" pageId="367" pageNumber="368" type="nomenclature">
<paragraph id="F946E7793FF9E903BF3D4768CEFF5335" pageId="367" pageNumber="368">
<taxonomicName id="A7CCCDBAD53EA6BCA80C62FA42BE0F4C" ID-CoL="6SNK5" LSID="http://species-id.net/wiki/Operclipygus_angustisternus" authority="Wenzel, 1944" authorityName="Wenzel" authorityYear="1944" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus angustisternus" order="Coleoptera" pageId="367" pageNumber="368" phylum="Arthropoda" rank="species" species="angustisternus">Operclipygus angustisternus (Wenzel, 1944)</taxonomicName>
Figs 105
<normalizedToken id="B2B2FE4D7EF2F9862C770CCFF016827B" originalValue="A–C106A–EMap">A-C106A-EMap</normalizedToken>
35
</paragraph>
</subSubSection>
<subSubSection id="29969FDF27A6AEF97817EE87FE80BDFD" pageId="367" pageNumber="368" type="reference_group">
<paragraph id="BCEEBC52E8C54B32C337DBC33CB123D5" pageId="367" pageNumber="368">
<taxonomicName id="07A35E13D5D2AE6D7320868CE497A22C" genus="Phelisteroides" lsidName="Phelisteroides angustisternus" pageId="367" pageNumber="368" rank="species" species="angustisternus">Phelisteroides angustisternus</taxonomicName>
Wenzel, 1944: 142;
<taxonomicName id="B1E1A9BDCD835025E40C7FD41FDEE15A" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus angustisternus" order="Coleoptera" pageId="367" pageNumber="368" phylum="Arthropoda" rank="species" species="angustisternus">Operclipygus angustisternus</taxonomicName>
:
<bibRefCitation id="04A142CDFF2091D9AA5FAA3AD576FE5E" author="Wenzel, RL" journalOrPublisher="Resultats Scientifiques" pageId="377" pageNumber="378" pagination="235 - 266" title="XXVII - Histeridae from Ecuador. Mission zoologique Belge aux Iles Galapagos et en Ecuador (N. et J. Leleup 1964 - 1965)." volume="3" year="1976">Wenzel 1976</bibRefCitation>
: 257.
</paragraph>
</subSubSection>
<subSubSection id="B5AE77CEC9050F3B15E8D10038ECE877" pageId="367" pageNumber="368" type="type locality">
<paragraph id="0F6A135A7A118C6A53612A59273B8B46" pageId="367" pageNumber="368">Type locality.</paragraph>
<paragraph id="C93DAEEE9B162F47211D42049610128E" pageId="367" pageNumber="368">
Not specified beyond
<normalizedToken id="6D89754888D2BBE99C075A9B0CEA1197" originalValue="Pará">Para</normalizedToken>
State, Brazil.
</paragraph>
</subSubSection>
<subSubSection id="25C5A495A535F0D64BE409419A7DD482" pageId="367" pageNumber="368" type="type material">
<paragraph id="2BABFC634A2C73BE69110919CC2CAFAA" pageId="367" pageNumber="368">Type material.</paragraph>
<paragraph id="0E38F2C6B39EACD082D7F3F409F4E958" pageId="367" pageNumber="368">
Holotype male, slightly teneral:
<normalizedToken id="88ABA73AC21D413AE662E91FC106F412" originalValue="“Para”">&quot;Para&quot;</normalizedToken>
/
<normalizedToken id="DA10C4FB898E6803D441107CDF7B1BAA" originalValue="“Brazil”">&quot;Brazil&quot;</normalizedToken>
/
<normalizedToken id="55CC04BC56ADA62297C7433981F38075" originalValue="“June”">&quot;June&quot;</normalizedToken>
/ &quot;Type
<taxonomicName id="1A97F4AA5E4E0AB1ABC4138FE4F8D9B8" genus="Phelisteroides" lsidName="Phelisteroides angustisternus" pageId="367" pageNumber="368" rank="species" species="angustisternus">Phelisteroides angustisternus</taxonomicName>
Wenzel&quot; (FMNH), examined 2006.
</paragraph>
</subSubSection>
<subSubSection id="BECE2E7676470987AE6D50373723F26D" lastPageId="370" lastPageNumber="371" pageId="367" pageNumber="368" type="other material">
<paragraph id="7F6A0AD4D9025FC94E5BCE66AA2CECC5" pageId="367" pageNumber="368">Other material.</paragraph>
<paragraph id="F0E03E51227B311CCF205FAA34002C81" lastPageId="370" lastPageNumber="371" pageId="367" pageNumber="368">
BRAZIL:
<normalizedToken id="65086D8E4353E68D1C807312E61EA54C" originalValue="Amapá">Amapa</normalizedToken>
: 2: Serra do Navio,
<geoCoordinate id="F3C13AE319EB735BE87FB6E1FF69CF04" direction="north" orientation="latitude" precision="925" value="0.98333335">0°59'N</geoCoordinate>
,
<geoCoordinate id="C963FC5AAFE63625163637BE4676C9A1" direction="west" orientation="longitude" precision="925" value="-52.0">52°00'W</geoCoordinate>
, 1-14.v.1991, FIT (CHND); Mato Grosso: 1: Sinop,
<geoCoordinate id="C898A729AC630A923D2350E1AA57AB5F" direction="south" orientation="latitude" precision="925" value="-11.816667">11°49'S</geoCoordinate>
,
<geoCoordinate id="2B8E662D7A96AFF294B6DF6F25AC1D99" direction="west" orientation="longitude" precision="925" value="-55.483334">55°29'W</geoCoordinate>
, 12-24.vi.1985, FIT (CHND);
<normalizedToken id="DC73ADFB72295188A8DDB51888F0F723" originalValue="Pará">Para</normalizedToken>
: 1:
<normalizedToken id="755B421579523CEB6607B0A1FB2625EA" originalValue="Tucuruí">Tucurui</normalizedToken>
,
<geoCoordinate id="DC7CDF94F63450343CD356287BAF47C3" direction="south" orientation="latitude" precision="925" value="-3.75">3°45'S</geoCoordinate>
,
<geoCoordinate id="5E8228C0E783E67B2B65E5820A9C24E7" direction="west" orientation="longitude" precision="925" value="-49.666668">49°40'W</geoCoordinate>
, iv.1986, FIT (CHND); 1: v.1986, FIT (CHND), 1: 20.v-15.vi.1987, FIT (CHND); 2: IPEAN,
<normalizedToken id="02E92EDA55A20E9BB75D0691408799F4" originalValue="Belém">Belem</normalizedToken>
, Ut
<pageBreakToken id="412741E393D37206127A4F983BEF2E83" pageId="368" pageNumber="369" start="start">inga</pageBreakToken>
,
<geoCoordinate id="DD34A1ECDD55B6C9E323322C468978BA" direction="south" orientation="latitude" precision="925" value="-1.45">1°27'S</geoCoordinate>
,
<geoCoordinate id="77D756B2CA0543BCA889D6CBE1B87144" direction="west" orientation="longitude" precision="925" value="-48.433334">48°26'W</geoCoordinate>
, ix.1984, FIT (CHND), 1: ix.1985, FIT (CHND); 1:
<normalizedToken id="BEE908452AB158EFD24DF0BC5B81791A" originalValue="Melgaço">Melgaco</normalizedToken>
, Rio
<normalizedToken id="B6C82357E7C31BDF48FB7C6B1DF7B57F" originalValue="Marinaú">Marinau</normalizedToken>
,
<geoCoordinate id="1D57C05E08596F622A0B2E26E75B421C" direction="south" orientation="latitude" precision="92" value="-1.8583333">1°51.5'S</geoCoordinate>
,
<geoCoordinate id="8DF20714AD67B41BD073228F4DC84638" direction="west" orientation="longitude" precision="925" value="-51.333332">51°20'W</geoCoordinate>
, 29.x-13.xi.1993, FIT (CHND), 1: 31.x-13.xi.1993, FIT (CHND). COLOMBIA:
<normalizedToken id="BBCB504EE134F7FE8B33E4DF65035B74" originalValue="Vaupés">Vaupes</normalizedToken>
: 1: Est. Biol.
<normalizedToken id="11F8D4B1DFDB4A84C88FD42A153656EB" originalValue="Caparú">Caparu</normalizedToken>
, Rio Apoporis,
<geoCoordinate id="578F1ED34C60A2962C78FFADC528754E" direction="south" orientation="latitude" precision="5555" value="-1.1">1.1°S</geoCoordinate>
,
<geoCoordinate id="E1D65D3F915611DC3689839AC79439CF" direction="west" orientation="longitude" precision="5555" value="-69.5">69.5°W</geoCoordinate>
, 27.ix-1.xii.1995, FIT, Black-water terrace forest on sandy soils, B.D. Gill (BDGC); 1: Parque Nac. Mosiro-Itajura (
<normalizedToken id="F2BF42330B5B9F6AE8D0C757DC47BD7E" originalValue="Caparú">Caparu</normalizedToken>
), Centro Ambiental,
<geoCoordinate id="DEEA654D57FAAE7455CEAC4C2FAAF473" direction="south" orientation="latitude" precision="925" value="-1.0666667">1°04'S</geoCoordinate>
,
<geoCoordinate id="B5EB85B42344D59B6354FE5CDA92BA57" direction="west" orientation="longitude" precision="925" value="-69.51667">69°31'W</geoCoordinate>
, 60m, 20-30.i.2003, FIT, D. Arias &amp; M. Sharkey (IAVH). ECUADOR: Orellana: 1:
<normalizedToken id="C671D73D442D19465F5E44B7ABD9F6E1" originalValue="Yasuní">Yasuni</normalizedToken>
Res. Stn., mid.Rio Tiputini,
<geoCoordinate id="BC1DE73F1B18E9861E78421D1A7B1AB0" direction="south" orientation="latitude" precision="92" value="-0.675">0°40.5'S</geoCoordinate>
,
<geoCoordinate id="51C927EEF4D98BCAC783F404FE8AF916" direction="west" orientation="longitude" precision="925" value="-76.4">76°24'W</geoCoordinate>
, 28.vi-5.vii.1999, FIT, C.E. Carlton &amp; A.K. Tishechkin (LSAM), 1: 12-20.vii.1999, FIT, A.K. Tishechkin (LSAM), 1: 5-11.vii.1999 (LSAM), 1: 20-23.vi.1999 (LSAM), 1: 23-30.vi.1999LSAM); 1:
<normalizedToken id="C4400433DDD5831A1301E00C4B0E8946" originalValue="Yasuní">Yasuni</normalizedToken>
Res. Stn.,
<geoCoordinate id="37D43FE86BF26AC5D571E567B53FC95F" direction="south" orientation="latitude" precision="15" value="-0.67444444">00°40'28&quot;S</geoCoordinate>
,
<geoCoordinate id="B302CD462F568F29FBFBCA958E47BA0A" direction="west" orientation="longitude" precision="15" value="-76.64722">76°38'50&quot;W</geoCoordinate>
, 215m, 5-10.ix.1999, FIT, primary forest, E.G. Riley (TAMU); 1: Parque Nac.
<normalizedToken id="2AEBEA0321F77E86A43F6214E4A14D51" originalValue="Yasuní">Yasuni</normalizedToken>
, Via Maxus at Puente
<normalizedToken id="AAFAC54471691709A6994C4F99000DD3" originalValue="Piraña">Pirana</normalizedToken>
,
<geoCoordinate id="51F723AD68F05695F6CAB33460CE9A4B" direction="south" orientation="latitude" precision="92" value="-1.6583333">1°39.5'S</geoCoordinate>
,
<geoCoordinate id="6AE8BCC244078840CEE1AA045B39889C" direction="west" orientation="longitude" precision="925" value="-76.433334">76°26'W</geoCoordinate>
, 14-20.vii.2008, FIT, A.K. Tishechkin (AKTC); 1: Payamino Research Station,
<geoCoordinate id="A5FAE670179830BE8F3AF8EB815EB6EC" direction="south" orientation="latitude" precision="1" value="-0.49333608">0°29'36.01&quot;S</geoCoordinate>
,
<geoCoordinate id="295FCBFA19A531D69D9DCE545E26D15E" direction="west" orientation="longitude" precision="1" value="-77.29143">77°17'29.15&quot;W</geoCoordinate>
, 300m, 30.vii-12.
<pageBreakToken id="D068EF89295F23BC96EB892AEC52E259" pageId="369" pageNumber="370" start="start">viii</pageBreakToken>
.2007, FIT, tropical rainforest, CPDT Gillett (BMNH); 1: Tiputini Biodiversity Station,
<geoCoordinate id="3C2036EFC074B3682CA240EBADAED49E" direction="south" orientation="latitude" precision="92" value="-0.63666666">0°38.2'S</geoCoordinate>
,
<geoCoordinate id="749FA4E836C6E2717BEEEEA804D160C6" direction="west" orientation="longitude" precision="92" value="-76.14833">76°8.9'W</geoCoordinate>
, 27-31.vii.2008, FIT, A.K. Tishechkin (AKTC); 1: Tiputini Biodiversity Station,
<geoCoordinate id="C7CA8F1C87FFAB15DB9DB428BB2FD5DD" direction="south" orientation="latitude" precision="15" value="-0.6333333">0°38'0&quot;S</geoCoordinate>
,
<geoCoordinate id="0E0400C969445ACBA7EB39669CA3C6E5" direction="west" orientation="longitude" precision="15" value="-76.15">76°9'0&quot;W</geoCoordinate>
, 220m, 5-25.ix.2000, D.J.
<pageBreakToken id="3A5174976642248FD4E59951BCCAF839" pageId="370" pageNumber="371" start="start">Inward</pageBreakToken>
&amp; K.A. Jackson (BMNH). PERU:
<normalizedToken id="C705C616F70B76D5989C69180757CBA0" originalValue="Junín">Junin</normalizedToken>
: 1: 11km NE Puerto Ocopa, Los Olivos,
<geoCoordinate id="D2C0D974C48C5796B20550BF272E907E" direction="south" orientation="latitude" precision="9" value="-11.116667">11°7.00'S</geoCoordinate>
,
<geoCoordinate id="83D357F46AA7594FCB14654C5A572022" direction="west" orientation="longitude" precision="9" value="-74.25867">74°15.52'W</geoCoordinate>
, 1200m, 30-31.iii.2009, Window trap, A.V. Petrov (AKTC); 1: 7.5km NE Puerto Ocopa, Cananeden,
<geoCoordinate id="5578CB5E9134D20C667E75935CFE38EA" direction="south" orientation="latitude" precision="92" value="-11.081667">11°4.9'S</geoCoordinate>
,
<geoCoordinate id="700986B6019172928857578A7EA4C2F4" direction="west" orientation="longitude" precision="92" value="-74.26833">74°16.1'W</geoCoordinate>
, 1180m, 6.ii.2008, A.V. Petrov (AKTC); 2: ~16km NW Satipo, Rio Venado,
<geoCoordinate id="3C6674811DBBF7DF750542EA5923F255" direction="south" orientation="latitude" precision="1" value="-11.194616">11°11.677'S</geoCoordinate>
,
<geoCoordinate id="E420E7272025AECAD71DDEDE17E4529F" direction="west" orientation="longitude" precision="1" value="-74.76895">74°46.137'W</geoCoordinate>
, 1150m, 3-8.iii.2010, A.V. Petrov (AKTC, MUSM); Loreto: 1: 68km SW Iquitos to Nauta, Rio Itaya,
<geoCoordinate id="5A4083223B805AB550715001297A03FE" direction="south" orientation="latitude" precision="925" value="-4.1833334">4°11'S</geoCoordinate>
,
<geoCoordinate id="81DD600B7F756C4FAB5203AEE0C1B6E9" direction="west" orientation="longitude" precision="925" value="-73.433334">73°26'W</geoCoordinate>
, 110m, 28.ii.2008, A.V. Petrov (AKTC); 1: km 63, rd. Iquitos - Nauta, Rio Itaya,
<geoCoordinate id="474CBD78A760B572210535ABCA653363" direction="south" orientation="latitude" precision="1" value="-4.2534165">4°15.205'S</geoCoordinate>
,
<geoCoordinate id="C118E8A837E8070E6A706D1FBBE178D6" direction="west" orientation="longitude" precision="1" value="-73.43457">73°26.074'W</geoCoordinate>
, 140m, 8-17.i.2010, A.V. Petrov (MUSM). SURINAME: Para: 2: nr. Overbridge River Resort,
<geoCoordinate id="ADA5300C90A6662D8FF76DE7E0F4C37C" direction="north" orientation="latitude" precision="92" value="5.5299997">5°31.8'N</geoCoordinate>
,
<geoCoordinate id="E287019FD6A7C507AD8B5B334C890463" direction="west" orientation="longitude" precision="92" value="-55.058334">55°3.5'W</geoCoordinate>
, 13-15.ii.2010, FIT, C. Gillet, P. Skelley, W. Warner (FSCA), 5: 15-18.ii.2010, FIT, C. Gillet, P. Skelley, W. Warner (FSCA); 2: near Overbridge,
<geoCoordinate id="828D9111DCF36D2D1FF80880D21098FB" direction="north" orientation="latitude" precision="15" value="5.519444">05°31'10&quot;N</geoCoordinate>
,
<geoCoordinate id="0F86A2C9700387C5D057B5E02C4E4F41" direction="west" orientation="longitude" precision="15" value="-55.069443">55°04'10&quot;W</geoCoordinate>
, 10-14.ii.2010, FIT, W.B. Warner (WBWC), 1: 14-18.ii.2010 (WBWC).
</paragraph>
</subSubSection>
<subSubSection id="CE707BAB7C24617A24A57926BFD121C2" lastPageId="371" lastPageNumber="372" pageId="370" pageNumber="371" type="description">
<paragraph id="A066E4E7F96DC523813D3EA64D90855D" pageId="370" pageNumber="371">Diagnostic description.</paragraph>
<paragraph id="0508463E5B496D0CC42CE54CDC9B0A15" lastPageId="371" lastPageNumber="372" pageId="370" pageNumber="371">
Length: 1.68-2.09 mm, width: 1.50-1.87 mm; body rufescent, elongate oval, sides weakly rounded; frons strongly but narrowly depressed at middle; frontal stria with sides divergent, weakly rounded, strongly sinuate over antennal bases, arcuate anterad; supraorbital stria finely impressed, but strongly arcuate dorsad onto vertex, weakly connected with sides of frontal stria; epistoma flat, weakly emarginate at apex; labrum about 3
<normalizedToken id="46FC3329917424FA0BF0B67843EA7AB0" originalValue="×">x</normalizedToken>
as wide as long, weakly emarginate, asymmetrical, left side more strongly produced than right, weak projection beneath; left mandible untoothed, right with small, acute basal tooth; pronotum with sides rounded, rather strongly convergent to front, with anterior margin markedly produced over head, lacking prescutellar impression; pronotal disk with fine, sparse ground punctation, with at least a few (up to ~12) coarser punctures at sides; lateral submarginal pronotal stria complete along side, curved inward anteriorly, nearly
<pageBreakToken id="22A71DE0DE9326922A4574979FDBCA56" pageId="371" pageNumber="372" start="start">meeting</pageBreakToken>
anterior submarginal stria, which is weakly arcuate, barely turned posterad at sides; median pronotal gland openings annulate, situated behind ends of recurved anterior stria, about two-thirds pronotal length from anterior margin; elytra with two complete epipleural striae, outer subhumeral stria present in posterior one-third to one-half, inner subhumeral stria absent, elytron swollen in subhumeral region, dorsal striae 1-3 complete, 4th stria present in apical two-thirds, 5th stria present in apical third, sutural present in apical three-fourths; prosternal keel rounded, projecting posterad, carinal striae variably shortened, meeting about one-third from presternal suture, anterior one-third of keel narrow; prosternal lobe rather short, marginal stria abbreviated at sides; anterior mesoventral margin emarginate, marginal stria complete; mesometaventral stria weakly arched forward at middle, sinuate at sides, continued by lateral metaventral stria posterad toward inner third of metacoxa; 1st abdominal ventrite with three lateral striae, all three usually abbreviated posteriorly, innermost extending mediad for some distance along anterior margin of ventrite, nearly continuously in some individuals, outermost shortest of the three, ending at an annulate postmetacoxal fovea; sides of propygidium and entire pygidium with dense, fine ground punctation; propygidium with irregular, moderately large punctures at middle, separated by about one-fourth their diameters; pygidium with few barely larger punctures standing out from ground punctation; marginal pygidial stria narrow but deeply impressed apically, becoming shallower, as an interconnected series of punctures toward base. Male genitalia (Figs 106
<normalizedToken id="1222BB09C5513F1007EC40BFD74857CF" originalValue="A–E">A-E</normalizedToken>
): accessory sclerites absent; T8 with sides subparallel in basal three-fourths, angulate to short, narrowly but shallowly emarginate apex, basal emargination deep, broad, reaching basal membrane attachment line, ventrolateral apodemes most strongly developed at base, narrowing toward apex; S8 with sides subparallel, apical guides narrowing from base to narrowly rounded apex; T9 with sides rounded, with narrow, convergent apices; T10 with halves separate; S9 with stem desclerotized along midline, widest basally, base truncate, with equilateral apical emargination, apical flanges separate; tegmen parallel-sided in basal three-fourths, narrowed to blunt apex, lacking medioventral process; basal piece about one-fourth tegmen length; median lobe narrow, about one-half tegmen length.
</paragraph>
</subSubSection>
<subSubSection id="A17CFDB157F64C3759C3CE0584952791" pageId="371" pageNumber="372" type="remarks">
<paragraph id="7F6A11EEDDA39FCA2C3F757EBBFFF8C8" pageId="371" pageNumber="372">Remarks.</paragraph>
<paragraph id="9D7DDAA79202309634E993191045B3B0" pageId="371" pageNumber="372">
The presence of dense ground punctation on the sides of the propygidium (Fig. 105C) is restricted to only a few species of
<taxonomicName id="DC0E750AB44F7AB177ED2B2ECA085D62" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="genus">Operclipygus</taxonomicName>
.
<taxonomicName id="DAAC11845C342C58080AC9C92B474FA6" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus angustisternus" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="species" species="angustisternus">Operclipygus angustisternus</taxonomicName>
is further distinguished by the strongly sinuate frontal and strongly arched supraorbital striae, the outwardly projecting anterior pronotal margin (Fig. 105A), and by the presence of three lateral striae on the 1st abdominal ventrite (Fig. 105B).
</paragraph>
<caption id="B0A873B7854F3E5F7F641E61C4CA1FCD" pageId="371" pageNumber="372">
<paragraph id="336499434FB04CB9888444EF2AF5341E" pageId="371" pageNumber="372">
Figure 105. Various
<taxonomicName id="9F4AE4609A4D5C13EC90B956CF11D4E1" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="genus">Operclipygus</taxonomicName>
species. A Dorsal habitus of
<taxonomicName id="B88267535C63797FDBB432180211C5EF" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus angustisternus" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="species" species="angustisternus">Operclipygus angustisternus</taxonomicName>
B Ventral habitus of
<taxonomicName id="B8512DAE62D1E13664C0DF6A65C3B5E4" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus angustisternus" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="species" species="angustisternus">Operclipygus angustisternus</taxonomicName>
C Pygidia of
<taxonomicName id="9C33A06F5668997050FDCB9851740FD9" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus angustisternus" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="species" species="angustisternus">Operclipygus angustisternus</taxonomicName>
D Dorsal habitus of
<taxonomicName id="ABA961E43027AE4CD92F59B4EF11FCBF" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus shorti" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="species" species="shorti">Operclipygus shorti</taxonomicName>
E Pygidia of
<taxonomicName id="49B93E7780757BCEE8D51FAA15B1A13F" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus shorti" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="species" species="shorti">Operclipygus shorti</taxonomicName>
.
</paragraph>
</caption>
<caption id="671E0F392E4AB2F2E492989A269DE53B" pageId="371" pageNumber="372">
<paragraph id="2A37AB4F934706D228D06C8A71C351DE" pageId="371" pageNumber="372">
Figure 106. Male genitalia of
<taxonomicName id="A1A0452F19D0AEDBB59B4A70E6A5D33C" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="genus">Operclipygus</taxonomicName>
species. A T8 of
<taxonomicName id="967FD0B2C2082B23BCA5918A16B66A8D" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus angustisternus" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="species" species="angustisternus">Operclipygus angustisternus</taxonomicName>
B S8 of
<taxonomicName id="D38F4EF332D2F7C30689470FCE2F1DF7" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus angustisternus" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="species" species="angustisternus">Operclipygus angustisternus</taxonomicName>
C T9 &amp; T10 of
<taxonomicName id="75D51D113BB91C53643C1531C853B2F0" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus angustisternus" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="species" species="angustisternus">Operclipygus angustisternus</taxonomicName>
D S9 of
<taxonomicName id="1C09A2B3ECD7C46FA1B15072442F002F" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus angustisternus" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="species" species="angustisternus">Operclipygus angustisternus</taxonomicName>
E Aedeagus, dorsal and lateral views, of
<taxonomicName id="09BDFD6DA9A24E58AEB840517EC5E436" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus angustisternus" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="species" species="angustisternus">Operclipygus angustisternus</taxonomicName>
F Aedeagus, dorsal and lateral views, of
<taxonomicName id="932D85F6DBBD7CA4F306284B57B42D6F" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus shorti" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="species" species="shorti">Operclipygus shorti</taxonomicName>
G T8 of
<taxonomicName id="66C74C2E2AB3B805027ECE27A54A00D5" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus shorti" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="species" species="shorti">Operclipygus shorti</taxonomicName>
H S8 of
<taxonomicName id="896AE6FC5DA68E2226A37AA3FD128C42" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus shorti" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="species" species="shorti">Operclipygus shorti</taxonomicName>
I T9 &amp; T10 of
<taxonomicName id="1A5334FF4BF6DA867243B16CF5D5113F" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus shorti" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="species" species="shorti">Operclipygus shorti</taxonomicName>
J S9 of
<taxonomicName id="33C6EB1281DD469F54561D6F280A9571" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus shorti" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="species" species="shorti">Operclipygus shorti</taxonomicName>
.
</paragraph>
</caption>
<caption id="7D610DA77DF50DD7DF126E7871C0C59D" pageId="371" pageNumber="372">
<paragraph id="452BB1F0ACC332BE382C252E42F6D837" pageId="371" pageNumber="372">
Map 35. Records of the
<taxonomicName id="3456A3E487259243E7598A57F7707C8D" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus" order="Coleoptera" pageId="371" pageNumber="372" phylum="Arthropoda" rank="genus">Operclipygus</taxonomicName>
incertae sedis spp.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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