173 lines
18 KiB
XML
173 lines
18 KiB
XML
<document id="6FC63150D9BE13DE9FF3A29C7176B0AD" ID-ISSN="2201-4349" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629583061888" checkinUser="felipe" docAuthor="Bickel, Daniel J." docDate="2006" docId="A03E2659FFD6C0778385C844FEBFF9C7" docLanguage="en" docName="RecAustMus.58.2.129-132.pdf" docOrigin="Records of the Australian Museum 58" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Munburra Bickel 2006, n.gen." docType="treatment" docUuid="7BE7515E-765E-425D-A299-7E4C817C2B1E" docUuidSource="ZooBank" docVersion="2" lastPageNumber="130" masterDocId="5C075E21FFD6C0768071CE62FFDCFC50" masterDocTitle="Munburra, A New Fly Genus from Tropical Australia (Empididae: Empidinae)" masterLastPageNumber="132" masterPageNumber="129" pageNumber="129" updateTime="1699081037041" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:titleInfo id="11E20551681D97E292BDC43A7240F886">
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<mods:title id="611E586993CF0666C212B93E073C5BF3">Munburra, A New Fly Genus from Tropical Australia (Empididae: Empidinae)</mods:title>
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<mods:name id="520FADDABCC1B33F97692F736D20D0F9" type="personal">
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<mods:roleTerm id="4DEFC7925406CDF13639834C7CE28F2C">Author</mods:roleTerm>
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<mods:namePart id="67D6B8FB4FBB2ADAFAF0F574E912B78E">Bickel, Daniel J.</mods:namePart>
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<mods:typeOfResource id="10251C20B23F071969BBE71AD5BF54AE">text</mods:typeOfResource>
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<mods:title id="6CE989617066D25F5F212D1D585B755E">Records of the Australian Museum</mods:title>
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<mods:part id="29E0BC9174C4AD5D9C3DB4D96F239DEC">
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<mods:date id="99A3FE5600B6B780341881A3523C6E4A">2006</mods:date>
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<mods:number id="48CE5D32D18C7C3A8B671550C1310615">58</mods:number>
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<mods:start id="B6325BC9F0A9444B6D324B95FF754208">129</mods:start>
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<mods:classification id="6A6C0E2E019991C9B2B7C78876A970DE">journal article</mods:classification>
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<mods:identifier id="EC4E39A59E035618BCB98B84A371B98C" type="ISSN">2201-4349</mods:identifier>
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<treatment id="A03E2659FFD6C0778385C844FEBFF9C7" LSID="urn:lsid:zoobank.org:act:7BE7515E-765E-425D-A299-7E4C817C2B1E" httpUri="http://treatment.plazi.org/id/A03E2659FFD6C0778385C844FEBFF9C7" lastPageId="1" lastPageNumber="130" pageId="0" pageNumber="129">
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<subSubSection id="608DC4C4FFD6C0768385C844FAD2FA6E" box="[1012,1294,1574,1598]" pageId="0" pageNumber="129" type="nomenclature">
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<paragraph id="2828974FFFD6C0768385C844FAD2FA6E" blockId="0.[1012,1294,1574,1598]" box="[1012,1294,1574,1598]" pageId="0" pageNumber="129">
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<heading id="73602023FFD6C0768385C844FAD2FA6E" box="[1012,1294,1574,1598]" centered="true" fontSize="11" level="2" pageId="0" pageNumber="129" reason="2">
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<emphasis id="1AE34B5DFFD6C0768385C844FAD2FA6E" bold="true" box="[1012,1294,1574,1598]" pageId="0" pageNumber="129">
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Genus
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<taxonomicName id="EF97ECCCFFD6C0768436C844FB63FA6E" ID-CoL="5VPZ" authority="Bickel, 2006" authorityName="Bickel" authorityYear="2006" box="[1095,1215,1574,1598]" class="Insecta" family="Empididae" genus="Munburra" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="1AE34B5DFFD6C0768436C844FB63FA6E" bold="true" box="[1095,1215,1574,1598]" italics="true" pageId="0" pageNumber="129">Munburra</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="01D0F626FFD6C07684B7C844FAD2FA6E" box="[1222,1294,1574,1598]" pageId="0" pageNumber="129" rank="genus">n.gen.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="608DC4C4FFD6C0768336C838FAB2FAFC" pageId="0" pageNumber="129" type="etymology">
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<paragraph id="2828974FFFD6C0768336C838FAB2FAFC" blockId="0.[839,1467,1626,1708]" pageId="0" pageNumber="129">
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<emphasis id="1AE34B5DFFD6C0768336C838FC62FA22" bold="true" box="[839,958,1626,1650]" pageId="0" pageNumber="129">Etymology</emphasis>
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.
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<taxonomicName id="EF97ECCCFFD6C07683BCC839FBEBFA22" authorityName="Bickel" authorityYear="2006" box="[973,1079,1627,1650]" class="Insecta" family="Empididae" genus="Munburra" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
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<emphasis id="1AE34B5DFFD6C07683BCC839FBEBFA22" box="[973,1079,1627,1650]" italics="true" pageId="0" pageNumber="129">Munburra</emphasis>
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</taxonomicName>
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is a geographic place name near the Mt Webb,
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<collectingRegion id="EA5359ADFFD6C07683C0C81AFBF0FADF" box="[945,1068,1656,1679]" country="Australia" name="Queensland" pageId="0" pageNumber="129">Queensland</collectingRegion>
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<typeStatus id="F72C29EDFFD6C0768441C81AFB81FADF" box="[1072,1117,1656,1679]" pageId="0" pageNumber="129">type</typeStatus>
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locality. The name is of aboriginal origin, and the gender is here designated as feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="608DC4C4FFD6C0768336C8ABFAD1FAB0" box="[839,1293,1737,1761]" pageId="0" pageNumber="129" type="type_taxon">
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<paragraph id="2828974FFFD6C0768336C8ABFAD1FAB0" blockId="0.[839,1293,1737,1761]" box="[839,1293,1737,1761]" pageId="0" pageNumber="129">
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<emphasis id="1AE34B5DFFD6C0768336C8ABFC0AFAB1" bold="true" box="[839,982,1737,1761]" pageId="0" pageNumber="129">
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<typeStatus id="F72C29EDFFD6C0768336C8ABFC5CFAB1" box="[839,896,1737,1761]" pageId="0" pageNumber="129">Type</typeStatus>
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species
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</emphasis>
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.
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<taxonomicName id="EF97ECCCFFD6C0768395C8A8FB0FFAB1" ID-CoL="44KD8" authority="Bickel, 2006" authorityName="Bickel" authorityYear="2006" box="[996,1235,1738,1761]" class="Insecta" family="Empididae" genus="Munburra" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="bulbicornis" status="sp. nov.">
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<emphasis id="1AE34B5DFFD6C0768395C8A8FB0FFAB1" box="[996,1235,1738,1761]" italics="true" pageId="0" pageNumber="129">Munburra bulbicornis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="01D0F626FFD6C07684ABC8ABFAD1FAB0" box="[1242,1293,1737,1760]" pageId="0" pageNumber="129" rank="species">n.sp.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="608DC4C4FFD6C0778336C89FFDCCFD86" lastPageId="1" lastPageNumber="130" pageId="0" pageNumber="129" type="diagnosis">
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<paragraph id="2828974FFFD6C0768336C89FFA67FB94" blockId="0.[839,1467,1789,1992]" pageId="0" pageNumber="129">
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<emphasis id="1AE34B5DFFD6C0768336C89FFC68FB45" bold="true" box="[839,948,1789,1813]" pageId="0" pageNumber="129">Diagnosis</emphasis>
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. Genus
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<taxonomicName id="EF97ECCCFFD6C0768462C89FFBA3FB44" authorityName="Bickel" authorityYear="2006" box="[1043,1151,1789,1812]" class="Insecta" family="Empididae" genus="Munburra" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
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<emphasis id="1AE34B5DFFD6C0768462C89FFBA3FB44" box="[1043,1151,1789,1812]" italics="true" pageId="0" pageNumber="129">Munburra</emphasis>
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</taxonomicName>
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: subfamily
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<taxonomicName id="EF97ECCCFFD6C0768574C89FFAA7FB44" box="[1285,1403,1789,1812]" class="Insecta" family="Empididae" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="subFamily" subFamily="Empidinae">Empidinae</taxonomicName>
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, tribe
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<taxonomicName id="EF97ECCCFFD6C0768336C979FC46FB62" authorityName="Collin" authorityYear="1961" box="[839,922,1819,1842]" class="Insecta" family="Empididae" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="tribe" tribe="Hilarini">Hilarini</taxonomicName>
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, with the following characters: laterotergite usually bare; tibia I with anteroapical comb of 8–10 short even setae; cercus usually small and desclerotized, and fused laterally with the surstylus and epandrium; hypandrium usually forming a curved convex hood over the aedeagus along the distal margin; costa circumambient; vein R
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<subScript id="B413950AFFD6C076841BC9DBFBAEFB98" attach="left" box="[1130,1138,1977,1992]" fontSize="6" pageId="0" pageNumber="129">1</subScript>
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swollen before it joins the costa.
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</paragraph>
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<paragraph id="2828974FFFD7C07780E6CEFBFDCCFD86" blockId="1.[121,749,153,470]" pageId="1" pageNumber="130">
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<materialsCitation id="98FF9D12FFD7C07780E6CEFBFDD0FD86" collectionCode="R" pageId="1" pageNumber="130" specimenCode="R4" specimenCount="1">
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Generic characters: ocellar triangle large and as rounded mound raised above surface of frons in both sexes; ocelli relatively large, well developed; male postpedicel slightly excavated at base, but basally inflated and tapering subtriangular; frons broad; thorax mostly orange yellow; male It
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<subScript id="B413950AFFD7C07780F8CF5AFF4DFD17" attach="left" box="[137,145,312,327]" fontSize="6" pageId="1" pageNumber="130">1</subScript>
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not swollen, but rather thin and slightly curved, with dorsal row of setae, and long curved dorsoapical seta, almost as long as tarsomere; Sc incomplete;
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<specimenCode id="78313F34FFD7C07781D0CF05FE67FDD2" box="[417,443,359,386]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="130">
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R
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<subScript id="B413950AFFD7C07781C2CF11FE67FDD2" attach="left" box="[435,443,371,386]" fontSize="6" pageId="1" pageNumber="130">4</subScript>
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</specimenCode>
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+5 forked and
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<specimenCode id="78313F34FFD7C0778233CF05FD87FDD2" box="[578,603,359,386]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="130">
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R
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<subScript id="B413950AFFD7C0778222CF11FD87FDD2" attach="left" box="[595,603,371,386]" fontSize="6" pageId="1" pageNumber="130">4</subScript>
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</specimenCode>
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with gentle Sshaped curve; hypopygium of the
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<emphasis id="1AE34B5DFFD7C07781B9CFE6FD59FDCB" box="[456,645,388,411]" italics="true" pageId="1" pageNumber="130">Hilara-Hilarempis</emphasis>
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<typeStatus id="F72C29EDFFD7C07782F8CFE6FD6BFDCB" box="[649,695,388,411]" pageId="1" pageNumber="130">type</typeStatus>
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, with hypandrial keel narrow and covering aedeagus, cercus small and setose, and epandrium deeply split
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="608DC4C4FFD7C0778008CF90FEBFF9C7" pageId="1" pageNumber="130" type="discussion">
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<paragraph id="2828974FFFD7C0778008CF90FD1DFF41" blockId="1.[121,749,498,1431]" pageId="1" pageNumber="130">
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<emphasis id="1AE34B5DFFD7C0778008CF90FF3EFE5A" bold="true" box="[121,226,498,522]" pageId="1" pageNumber="130">Remarks</emphasis>
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.
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<taxonomicName id="EF97ECCCFFD7C0778080CF91FE83FE5A" authorityName="Bickel" authorityYear="2006" box="[241,351,499,522]" class="Insecta" family="Empididae" genus="Munburra" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="genus">
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<emphasis id="1AE34B5DFFD7C0778080CF91FE83FE5A" box="[241,351,499,522]" italics="true" pageId="1" pageNumber="130">Munburra</emphasis>
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</taxonomicName>
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is a monotypic genus from tropical monsoonal
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<collectingCountry id="5080D7DFFFD7C0778088CC72FE81FE77" box="[249,349,528,551]" name="Australia" pageId="1" pageNumber="130">Australia</collectingCountry>
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with a single included species,
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<taxonomicName id="EF97ECCCFFD7C07782BECC72FF28FE14" authorityName="Bickel" authorityYear="2006" class="Insecta" family="Empididae" genus="Munburra" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="species" species="bulbicornis">
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<emphasis id="1AE34B5DFFD7C07782BECC72FF28FE14" italics="true" pageId="1" pageNumber="130">M. bulbicornis</emphasis>
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</taxonomicName>
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. It is included in the tribe
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<taxonomicName id="EF97ECCCFFD7C0778263CC4FFDBAFE14" authorityName="Collin" authorityYear="1961" box="[530,614,557,580]" class="Insecta" family="Empididae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="tribe" tribe="Hilarini">Hilarini</taxonomicName>
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(see Bickel, 1996) based on the characters listed in the Diagnosis, above. The genus is defined by the three apomorphies: a large, raised ocellar triangle with enlarged ocelli, and two distinctive male secondary sexual characters, a greatly swollen postpedicel, and a long dorsoapical seta on a narrow unswollen leg I first tarsomere. I have not seen these character states developed in other genera in the tribe.
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</paragraph>
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<paragraph id="2828974FFFD7C07780E6CD7AFD43F849" blockId="1.[121,749,498,1431]" pageId="1" pageNumber="130">
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Further, the
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<taxonomicName id="EF97ECCCFFD7C0778166CD7AFEB7FF7F" authorityName="Collin" authorityYear="1961" box="[279,363,792,815]" class="Insecta" family="Empididae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="tribe" tribe="Hilarini">Hilarini</taxonomicName>
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are more characteristic of temperate (both Northern and Southern Hemisphere) or montane habitats than the lowland tropics. Certainly many more taxa have been described from temperate regions, and although this may reflect regional collecting bias and/or the absence of netting along streams in the tropics, the Australian tropical fauna is indeed much poorer than that of southern
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<collectingCountry id="5080D7DFFFD7C07782F2CDAAFD38FF8F" box="[643,740,968,991]" name="Australia" pageId="1" pageNumber="130">Australia</collectingCountry>
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/
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<collectingRegion id="EA5359ADFFD7C0778008CD87FF3FFFAC" box="[121,227,997,1020]" country="Australia" name="Tasmania" pageId="1" pageNumber="130">Tasmania</collectingRegion>
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. Therefore the presence of such an unusual genus in the tropics of Cape York Peninsula is of interest.
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</paragraph>
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<paragraph id="2828974FFFD7C07780E6CA42FEBFF9C7" blockId="1.[121,749,498,1431]" pageId="1" pageNumber="130">
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In some respects,
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<taxonomicName id="EF97ECCCFFD7C0778125CA42FE62F867" authorityName="Bickel" authorityYear="2006" box="[340,446,1056,1079]" class="Insecta" family="Empididae" genus="Munburra" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="genus">
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<emphasis id="1AE34B5DFFD7C0778125CA42FE62F867" box="[340,446,1056,1079]" italics="true" pageId="1" pageNumber="130">Munburra</emphasis>
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</taxonomicName>
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appears similar to the genus
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<taxonomicName id="EF97ECCCFFD7C0778008CA5FFEF3F804" authority="Philippi" authorityName="Philippi" authorityYear="1865" box="[121,303,1085,1108]" class="Insecta" family="Empididae" genus="Sphicosa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="genus">
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<emphasis id="1AE34B5DFFD7C0778008CA5FFF0AF804" box="[121,214,1085,1108]" italics="true" pageId="1" pageNumber="130">Sphicosa</emphasis>
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Philippi
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</taxonomicName>
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, which comprises southern SouthAmerican species and undescribed Australian species.Although
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<taxonomicName id="EF97ECCCFFD7C07782E1CA39FD30F822" authorityName="Philippi" authorityYear="1865" box="[656,748,1115,1138]" class="Insecta" family="Empididae" genus="Sphicosa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="genus">
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<emphasis id="1AE34B5DFFD7C07782E1CA39FD30F822" box="[656,748,1115,1138]" italics="true" pageId="1" pageNumber="130">Sphicosa</emphasis>
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</taxonomicName>
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is definitely in the tribe
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<taxonomicName id="EF97ECCCFFD7C0778109CA1AFE07F8DF" authorityName="Latreille" authorityYear="1804" box="[376,475,1144,1167]" class="Insecta" family="Empididae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="tribe" tribe="Empidini">Empidini</taxonomicName>
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, the two genera appear to have a similar obliquely projecting proboscis (although in
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<taxonomicName id="EF97ECCCFFD7C0778008CAD1FF3FF89A" authorityName="Bickel" authorityYear="2006" box="[121,227,1203,1226]" class="Insecta" family="Empididae" genus="Munburra" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="genus">
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<emphasis id="1AE34B5DFFD7C0778008CAD1FF3FF89A" box="[121,227,1203,1226]" italics="true" pageId="1" pageNumber="130">Munburra</emphasis>
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</taxonomicName>
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it can be elevated and depressed through the oral cavity, and is normally held in a vertical position), a costa reduced in thickness beyond R
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<subScript id="B413950AFFD7C07781D5CA9BFE60F958" attach="left" box="[420,444,1273,1288]" fontSize="6" pageId="1" pageNumber="130">4+5</subScript>
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, and R
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<subScript id="B413950AFFD7C0778271CA9BFDD4F958" attach="left" box="[512,520,1273,1288]" fontSize="6" pageId="1" pageNumber="130">1</subScript>
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is not apically swollen. As well, the hypopygium of some male
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<taxonomicName id="EF97ECCCFFD7C0778267CB69FDAFF972" authorityName="Philippi" authorityYear="1865" box="[534,627,1291,1314]" class="Insecta" family="Empididae" genus="Sphicosa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="genus">
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<emphasis id="1AE34B5DFFD7C0778267CB69FDAFF972" box="[534,627,1291,1314]" italics="true" pageId="1" pageNumber="130">Sphicosa</emphasis>
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</taxonomicName>
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also have a keel-like hypandrium and appears almost hilarine-like. I regard these similarities as homoplasious, part of the suite of characters available for expression across the
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<taxonomicName id="EF97ECCCFFD7C0778184CB00FDB5F929" box="[501,617,1378,1401]" class="Insecta" family="Empididae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="subFamily" subFamily="Empidinae">Empidinae</taxonomicName>
|
||
, in both the
|
||
<taxonomicName id="EF97ECCCFFD7C0778008CBE2FF06F9C7" authorityName="Latreille" authorityYear="1804" box="[121,218,1408,1431]" class="Insecta" family="Empididae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="tribe" tribe="Empidini">Empidini</taxonomicName>
|
||
and
|
||
<taxonomicName id="EF97ECCCFFD7C077817ACBE2FE83F9C7" authorityName="Collin" authorityYear="1961" box="[267,351,1408,1431]" class="Insecta" family="Empididae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="tribe" tribe="Hilarini">Hilarini</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |