treatments-xml/data/A0/23/87/A02387D80055FFD7FF56FB4DFCC5EC63.xml

<document id="4087495496E795FB4D3FB1613BF701AF" ID-CLB-Dataset="7342" ID-DOI="10.11646/zootaxa.4950.3.3" ID-GBIF-Dataset="45cb427a-19fd-4068-b314-2630db74b60e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4649999" ID-ZooBank="790FA467-0BCA-4F7E-9428-74D6315DA34B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617176317065" checkinUser="plazi" docAuthor="Rodcharoen, Eknarin &amp; Bruce, Niel L." docDate="2021" docId="A02387D80055FFD7FF56FB4DFCC5EC63" docLanguage="en" docName="zootaxa.4950.3.3.pdf" docOrigin="Zootaxa 4950 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cirolana khamensis Rodcharoen &amp; Bruce 2021, sp. nov." docType="treatment" docVersion="9" lastPageNumber="478" masterDocId="5C1AFFA00057FFDEFFC1FF82FFEAED06" masterDocTitle="Two new species of the marine isopod genus Cirolana Leach, 1818 (Crustacea Isopoda: Cirolanidae) from the coast of the western Gulf of Thailand" masterLastPageNumber="486" masterPageNumber="469" pageNumber="471" updateTime="1701999093242" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="EA45C71A516A0C9B2C0A6FC4759FCF28" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="052A579B776510DB4438B1B92A2B171B">
<mods:title id="934F41BF87B8C897ACAC2A117E0AC100">Two new species of the marine isopod genus Cirolana Leach, 1818 (Crustacea Isopoda: Cirolanidae) from the coast of the western Gulf of Thailand</mods:title>
</mods:titleInfo>
<mods:name id="8B131A0BD7707792AA48CA54D4D90204" type="personal">
<mods:role id="80B15F414582A2D9D8D17203C7253828">
<mods:roleTerm id="0C5E9528FDB2ACDFE53418AEC4B5CCA6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EF0150F9DE21B805C0CF6B776571C584">Rodcharoen, Eknarin</mods:namePart>
<mods:affiliation id="7CA4E7133A52D3A1CE9696DFC6680879">Aquatic Science and Innovative Management Division, Faculty of Natural Resources; and Discipline of Excellence for Sustainable Aquaculture, Prince of Songkla University, Hat Yai, Songkhla 90110, Thailand.</mods:affiliation>
</mods:name>
<mods:name id="C132EDFD85B335B90C2B7DDED38EB381" type="personal">
<mods:role id="292196D0E6EF89FCA24AEFA60A2359C8">
<mods:roleTerm id="3521EF8DD4942CB19916859512EC65C4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E4E8800FA2D87EABA632B33A8EA75BB1">Bruce, Niel L.</mods:namePart>
<mods:nameIdentifier id="12B5567A59895D7C2BB43A70926739BA" type="ORCID">0000-0003-4745-5048</mods:nameIdentifier>
<mods:affiliation id="EE2C847A406C3168884DA5E49CBDBD86">Biodiversity &amp; Geosciences Program, Queensland Museum, PO Box: 3300, South Brisbane BC, Queensland 4101, Australia; and Water Research Group, Unit for Environmental Sciences and Management, North-West University, Private Bag X 6001, Potchefstroom 2520, South Africa. niel. bruce @ qm. qld. gov. au; https: // orcid. org / 0000 - 0003 - 4745 - 5048</mods:affiliation>
<mods:nameIdentifier id="229242984F9FF0E2CA45A49D3FDFE905" type="email">niel.bruce@qm.qld.gov.au</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="F9A06B8718B9EF88B0222156582BBA91">text</mods:typeOfResource>
<mods:relatedItem id="F962C531CCC34CEEA313B8B869C2AEDB" type="host">
<mods:titleInfo id="10E899CFBA475DAE007CC1CCAB124D3A">
<mods:title id="64B0777CFE06778E993C74315588B13B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="311D88E4281D1FA8C682D7EE10CABE9B">
<mods:date id="0657DE1A469EB7ECD55412EEB0266F0E">2021</mods:date>
<mods:detail id="D66928B01DF66F71D31958770528736C" type="pubDate">
<mods:number id="EB87E29456936759F6CD37343D6F342D">2021-03-31</mods:number>
</mods:detail>
<mods:detail id="E1B693580EACB452FBA1429D1D421758" type="volume">
<mods:number id="2C5DEBDF8E2322E6C3431CD46B74D441">4950</mods:number>
</mods:detail>
<mods:detail id="856ABCA0D1D46B2E7F04B6440EA087B9" type="issue">
<mods:number id="E10E5EE642DCCE9C7094309CAECB2D97">3</mods:number>
</mods:detail>
<mods:extent id="264405D53595F0E90C1832ED29E51EB4" unit="page">
<mods:start id="F9A79010BB681B5DE6C8A509CCD334C7">469</mods:start>
<mods:end id="5D42DF3CE9A9FFFD6656B86C0DE73FB7">486</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="C5626ED6F70E806296D6DD4378EC8A4C">journal article</mods:classification>
<mods:identifier id="879EE001789AF22CE5E5C26798424662" type="CLB-Dataset">7342</mods:identifier>
<mods:identifier id="368B745095E5D967C33AC758E99BDAFA" type="DOI">10.11646/zootaxa.4950.3.3</mods:identifier>
<mods:identifier id="91E2FDAD443117574FF95790585938D0" type="GBIF-Dataset">45cb427a-19fd-4068-b314-2630db74b60e</mods:identifier>
<mods:identifier id="2FFF9DD476BA1A6F57597EBC4AAA8C57" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EB0DCC21BA17B27AC485A5AB09CFDE40" type="Zenodo-Dep">4649999</mods:identifier>
<mods:identifier id="693B7C8CDFB3CA34AAC2D00E313B75DE" type="ZooBank">790FA467-0BCA-4F7E-9428-74D6315DA34B</mods:identifier>
</mods:mods>
<treatment id="A02387D80055FFD7FF56FB4DFCC5EC63" ID-DOI="http://doi.org/10.5281/zenodo.4694596" ID-GBIF-Taxon="181010796" ID-Zenodo-Dep="4694596" LSID="urn:lsid:plazi:treatment:A02387D80055FFD7FF56FB4DFCC5EC63" httpUri="http://treatment.plazi.org/id/A02387D80055FFD7FF56FB4DFCC5EC63" lastPageId="9" lastPageNumber="478" pageId="2" pageNumber="471">
<subSubSection id="609065450055FFDCFF56FB4DFE1AE9EF" box="[151,496,1231,1258]" pageId="2" pageNumber="471" type="nomenclature">
<paragraph id="283536CE0055FFDCFF56FB4DFE1AE9EF" blockId="2.[151,496,1231,1293]" box="[151,496,1231,1258]" pageId="2" pageNumber="471">
<heading id="737D81A20055FFDCFF56FB4DFE1AE9EF" bold="true" box="[151,496,1231,1258]" fontSize="11" level="1" pageId="2" pageNumber="471" reason="1">
<emphasis id="1AFEEADC0055FFDCFF56FB4DFE1AE9EF" bold="true" box="[151,496,1231,1258]" pageId="2" pageNumber="471">
<taxonomicName id="EF8A4D4D0055FFDCFF56FB4DFE61E9EC" ID-CoL="86YDD" authority="Rodcharoen &amp; Bruce, 2021" authorityName="Rodcharoen &amp; Bruce" authorityYear="2021" box="[151,395,1231,1258]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="471" phylum="Arthropoda" rank="species" species="khamensis" status="sp. nov.">
<emphasis id="1AFEEADC0055FFDCFF56FB4DFE61E9EC" bold="true" box="[151,395,1231,1258]" italics="true" pageId="2" pageNumber="471">Cirolana khamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="01CD57A70055FFDCFE53FB4DFE1AE9EF" box="[402,496,1231,1257]" pageId="2" pageNumber="471" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="609065450055FFDCFF56FB71FEE1E80B" box="[151,267,1267,1293]" pageId="2" pageNumber="471" type="description">
<paragraph id="283536CE0055FFDCFF56FB71FEE1E80B" blockId="2.[151,496,1231,1293]" box="[151,267,1267,1293]" pageId="2" pageNumber="471">
(
<figureCitation id="B0B12A4B0055FFDCFF5EFB71FEEEE80B" box="[159,260,1267,1293]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="4.[151,250,1908,1932]" captionStartId-1="5.[151,250,1943,1967]" captionStartId-2="6.[151,250,1943,1967]" captionStartId-3="7.[151,250,1943,1967]" captionStartId-4="8.[151,250,1378,1402]" captionStartId-5="9.[151,250,1964,1988]" captionTargetBox-0="[203,1378,191,1877]" captionTargetBox-1="[184,1403,183,1911]" captionTargetBox-2="[207,1380,189,1917]" captionTargetBox-3="[249,1339,189,1918]" captionTargetBox-4="[152,1426,187,1344]" captionTargetBox-5="[165,1421,387,1937]" captionTargetId-0="figure-27@4.[197,1389,181,1884]" captionTargetId-1="figure-17@5.[182,1405,181,1920]" captionTargetId-2="figure-27@6.[205,1382,181,1919]" captionTargetId-3="figure-17@7.[244,1343,181,1920]" captionTargetId-4="figure-27@8.[151,1435,181,1353]" captionTargetId-5="figure-111@9.[156,1432,381,1940]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionText-0="FIGURE 2. Cirolana khamensis sp. nov., holotype male (6.5 mm, PSUZC–CR0288-01), A, D; paratype male (8.4 mm, PSU- ZC–CR0288-02),B, E); paratype female (7.8 mm, PSUZC–CR0288-02), C; paratype male (7.9 mm, PSUZC–CR0288-02), F, G, H. A, dorsal view; B, lateral view; C, head; D, frons; E, pleon; F, antennula; G, antenna peduncle; H, penial opening, sternite 7." captionText-1="FIGURE 3. Cirolana khamensis sp. nov., paratype male (7.9 mm, PSUZC-CR2088-02). A, left mandible; B, distal part of mandible; C, maxillule; D, maxilla E, maxilliped; G, maxilliped endite." captionText-2="FIGURE 4. Cirolana khamensis sp. nov., paratype male (7.9 mm, PSUZC-CR2088-02). A, pereopod 1; B, dactylus of pereopod 1; C, mesial view of pereopod 1; D, pereopod 2; E, pereopod 4." captionText-3="FIGURE 5. Cirolana khamensis sp. nov., paratype male (7.9 mm, PSUZC-CR2088-02), A, lateral of pereopod 6; B, mesial view of pereopod 6; C, lateral view of pereopod 7; D, mesial of pereopod 7." captionText-4="FIGURE 6. Cirolana khamensis sp. nov., paratype male (7.9 mm, PSUZC-CR2088-02). A–E, pleopods 1–5 respectively." captionText-5="FIGURE 7. Cirolana khamensis sp. nov., paratype male (7.9 mm, PSUZC-CR2088-02).A, uropod; B, ventral view of uropodal exopod; C, pleotelson; D, uropodal exopod apex." figureDoi-0="http://doi.org/10.5281/zenodo.4650005" figureDoi-1="http://doi.org/10.5281/zenodo.4650009" figureDoi-2="http://doi.org/10.5281/zenodo.4650011" figureDoi-3="http://doi.org/10.5281/zenodo.4650013" figureDoi-4="http://doi.org/10.5281/zenodo.4650015" figureDoi-5="http://doi.org/10.5281/zenodo.4650017" httpUri-0="https://zenodo.org/record/4650005/files/figure.png" httpUri-1="https://zenodo.org/record/4650009/files/figure.png" httpUri-2="https://zenodo.org/record/4650011/files/figure.png" httpUri-3="https://zenodo.org/record/4650013/files/figure.png" httpUri-4="https://zenodo.org/record/4650015/files/figure.png" httpUri-5="https://zenodo.org/record/4650017/files/figure.png" pageId="2" pageNumber="471">Figs 2–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="609065450055FFDCFF56FAB9FCB6E8C7" pageId="2" pageNumber="471" type="materials_examined">
<paragraph id="283536CE0055FFDCFF56FAB9FB8BE87F" blockId="2.[151,1437,1339,2014]" pageId="2" pageNumber="471">
<emphasis id="1AFEEADC0055FFDCFF56FAB9FE68E853" bold="true" box="[151,386,1339,1365]" pageId="2" pageNumber="471">Material examined.</emphasis>
<materialsCitation id="98E23C930055FFDCFE4BFAB9FB8BE87F" ID-GBIF-Occurrence="3072681305" collectingDate="2011-07-09" collectionCode="PSUZC-CR" collectorName="E. Rodcharoen" elevation="8" latitude="6.9719" location="Koh Kham" longLatPrecision="1" longitude="100.85709" pageId="2" pageNumber="471" specimenCode="PSUZC-CR2087-01" specimenCount="1" specimenCount-male="1" stateProvince="Songkhla" typeStatus="holotype">
<emphasis id="1AFEEADC0055FFDCFE4BFAB9FDEBE853" bold="true" box="[394,513,1339,1365]" pageId="2" pageNumber="471">
<typeStatus id="F731886C0055FFDCFE4BFAB9FE11E853" box="[394,507,1339,1365]" pageId="2" pageNumber="471" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="3E8CFD470055FFDCFDC8FABEFDF4E850" box="[521,542,1340,1366]" pageId="2" pageNumber="471" type="male">♂</specimenCount>
(
<quantity id="EF729B2B0055FFDCFDEEFAB9FD61E850" box="[559,651,1339,1366]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="471" unit="mm" value="6.5">6.5 mm</quantity>
), 
<location id="2D5560150055FFDCFD61FAB9FCF5E853" LSID="urn:lsid:plazi:treatment:A02387D80055FFD7FF56FB4DFCC5EC63:2D5560150055FFDCFD61FAB9FCF5E853" box="[672,799,1339,1365]" latitude="6.9719" longLatPrecision="1" longitude="100.85709" name="Koh Kham" pageId="2" pageNumber="471" stateProvince="Songkhla">Koh Kham</location>
, 
<collectingRegion id="EA4EF82C0055FFDCFCEDFAB9FBEFE853" box="[812,1029,1339,1365]" country="Thailand" name="Songkhla" pageId="2" pageNumber="471">Songkhla Province</collectingRegion>
, 
<geoCoordinate id="4DBE50090055FFDCFBD3FAB9FB51E850" box="[1042,1211,1339,1366]" degrees="6" direction="north" minutes="58" orientation="latitude" pageId="2" pageNumber="471" precision="1" seconds="18.84" value="6.9719">6°58’18.84” N</geoCoordinate>
, 
<geoCoordinate id="4DBE50090055FFDCFB09FAB9FA6BE850" box="[1224,1409,1339,1366]" degrees="100" direction="east" minutes="51" orientation="longitude" pageId="2" pageNumber="471" precision="1" seconds="25.54" value="100.85709">100°51’25.54”E</geoCoordinate>
, 
<date id="5C34100E0055FFDCFA4FFAB9FEEDE87F" pageId="2" pageNumber="471" value="2011-07-09">
<collectingDate id="4C70E9E60055FFDCFA4FFAB9FEEDE87F" pageId="2" pageNumber="471" value="2011-07-09">9 July 2011</collectingDate>
</date>
, trapped at 
<elevation id="A3A7D1FD0055FFDCFE49FADDFE11E87F" box="[392,507,1375,1402]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="471" unit="m" value="8.0">
<quantity id="EF729B2B0055FFDCFE49FADDFE59E87C" box="[392,435,1375,1402]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="471" unit="m" value="8.0">8 m</quantity>
depth
</elevation>
, coral reef, coll. 
<collectorName id="857F53180055FFDCFD77FAE2FCB7E87F" box="[694,861,1375,1402]" pageId="2" pageNumber="471">E. Rodcharoen</collectorName>
(
<specimenCode id="782C9EB50055FFDCFCADFADDFBBCE87F" box="[876,1110,1375,1401]" collectionCode="PSUZC-CR" pageId="2" pageNumber="471">PSUZC-CR2087-01</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="283536CE0055FFDCFF06FA01FCB6E8C7" blockId="2.[151,1437,1339,2014]" pageId="2" pageNumber="471">
<emphasis id="1AFEEADC0055FFDCFF06FA01FED5E89B" bold="true" box="[199,319,1411,1437]" pageId="2" pageNumber="471">
<typeStatus id="F731886C0055FFDCFF06FA01FED5E89B" box="[199,319,1411,1437]" pageId="2" pageNumber="471" type="paratype">Paratypes</typeStatus>
</emphasis>
: 
<specimenCount id="3E8CFD470055FFDCFE8EFA01FE93E898" box="[335,377,1411,1438]" pageId="2" pageNumber="471" type="male">9 ♂</specimenCount>
(7.9, 8.2, 7.3, 7.7, 8.3, 7.1, 7.6, 8.2, 
<quantity id="EF729B2B0055FFDCFCD8FA01FC9BE898" box="[793,881,1411,1438]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="2" pageNumber="471" unit="mm" value="8.4">8.4 mm</quantity>
) [dissected]), 
<materialsCitation id="98E23C930055FFDCFBD3FA06FCB3E8C7" ID-GBIF-Occurrence="3072681302" collectingDate="2011-07-09" collectionCode="PSUZC-CR" collectorName="E. Rodcharoen" elevation="8" latitude="6.9719" location="Koh Kham" longLatPrecision="1" longitude="100.85709" pageId="2" pageNumber="471" specimenCode="PSUZC-CR2087-02" specimenCount="7" specimenCount-female="7" stateProvince="Songkhla" typeStatus="paratype">
<specimenCount id="3E8CFD470055FFDCFBD3FA06FBD7E898" box="[1042,1085,1412,1438]" pageId="2" pageNumber="471" type="female">7 ♀</specimenCount>
(7.4, 7.8, 8.5, 7.9, 8.3, 8.5, 
<quantity id="EF729B2B0055FFDCFAB8FA01FF29E8C4" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="2" pageNumber="471" unit="mm" value="7.8">7.8 mm</quantity>
[dissected]), same data as holotype, (
<specimenCode id="782C9EB50055FFDCFDA7FA25FCBAE8C7" box="[614,848,1447,1473]" collectionCode="PSUZC-CR" pageId="2" pageNumber="471">PSUZC-CR2087-02</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="609065450055FFDDFF06FA49FAFFEABF" lastPageId="3" lastPageNumber="472" pageId="2" pageNumber="471" type="description">
<paragraph id="283536CE0055FFDCFF06FA49FE9BEA90" blockId="2.[151,1437,1339,2014]" pageId="2" pageNumber="471">
<emphasis id="1AFEEADC0055FFDCFF06FA49FE5DE8E3" bold="true" box="[199,439,1483,1509]" pageId="2" pageNumber="471">Description of male.</emphasis>
Body dorsal surfaces punctuate, widest at pereonite 5 or pereonite 6, lateral margins subparallel. 
<emphasis id="1AFEEADC0055FFDCFF39FA6DFE61EB0F" box="[248,395,1519,1545]" italics="true" pageId="2" pageNumber="471">Rostral point</emphasis>
absent. 
<emphasis id="1AFEEADC0055FFDCFE27FA72FDF1EB0F" box="[486,539,1520,1545]" italics="true" pageId="2" pageNumber="471">Eyes</emphasis>
colour brown in alcohol; eyes separated by about 80% width of head. Pereonites 1–3 each with a single transverse impressed line, pereonites 4–6 each with 2 transverse impressed lines; pereonite 7 with 3 tranverse lines with posterior line forming nodulose ridge. 
<emphasis id="1AFEEADC0055FFDCFC57F9BAFB78EB57" box="[918,1170,1591,1617]" italics="true" pageId="2" pageNumber="471">Pereonite 1 and coxae</emphasis>
2–3 each with posteroventral angle rounded; coxae 5–7 with entire oblique carina. 
<emphasis id="1AFEEADC0055FFDCFCF8F9D9FC90EB73" box="[825,890,1627,1653]" italics="true" pageId="2" pageNumber="471">Pleon</emphasis>
with pleonite 1 largely concealed by pereonite 7; pleonites 3–5 with tubercles; posterolateral angles of pleonite 2 rounded, extending posteriorly to anterior of pleonite 4; pleonite 3 with row of 11–13 tubercles, posterolateral margins extending to but not beyond posterior margin of pleonite 5, narrow rounded; pleonite 4 with prominent median tubercles and 3–4 sublateral tubercles on each side, posterolater margins rounded, clearly extending beyond posterior margin of pleonite 5; pleonite 5 with prominent median tubercles and 2–3 sublateral tubercles on each side, posterolateral angles overlapped by lateral margins of pleonite 4. 
<emphasis id="1AFEEADC0055FFDCFED4F8B1FE61EA4B" box="[277,395,1843,1869]" italics="true" pageId="2" pageNumber="471">Pleotelson</emphasis>
0.7 times as long as anterior width, dorsal surface with paired submedian logitudinal carina, each of which has 4 tubercles; lateral margins weakly concave; margins smooth, posterior margin evenly rounded, with 8 robust setae.
</paragraph>
<paragraph id="283536CE0055FFDDFF06F81DFB92EC1B" blockId="2.[151,1437,1339,2014]" lastBlockId="3.[151,1437,151,1977]" lastPageId="3" lastPageNumber="472" pageId="2" pageNumber="471">
<emphasis id="1AFEEADC0055FFDCFF06F81DFED0EABF" box="[199,314,1951,1977]" italics="true" pageId="2" pageNumber="471">Antennula</emphasis>
peduncle articles 1 and 2 distinct articulated; article 2 0.5 times as long as article1; article 3 and 4 0.9 times as long as combined lengths of articles 1 and 2, article 3 1.8 times as long as wide; flagellum with 9 articles, extending to posterior margin of eye. 
<emphasis id="1AFEEADC0054FFDDFD88FF15FD4CEDB7" box="[585,678,151,177]" italics="true" pageId="3" pageNumber="472">Antenna</emphasis>
peduncle article 4 1.8 times as long as wide, 2.6 times as long as article 3, posterodistal angle with 1 short simple seta; article 5 1 times as long as article 4, 2.5 times as long as wide, anterodistal angle with cluster of 4 setae (2 simple and 2 palmate setae), anterodistal angle with cluster of 2 simple setae and 2 palmate setae; flagellum with 20 articles, extending to middle of pereonite 3.
</paragraph>
<paragraph id="283536CE0054FFDDFF06FEA5FDBBEC63" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="472">
<emphasis id="1AFEEADC0054FFDDFF06FEA5FE9BEC47" box="[199,369,295,321]" italics="true" pageId="3" pageNumber="472">Frontal lamina</emphasis>
pentagonal, 2.2 times as longer than greatest width, lateral margins straight, diverging slightly towards anterior, anterior margin acute.
</paragraph>
<paragraph id="283536CE0054FFDDFF06FEEDFCC4EF80" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="472">
<emphasis id="1AFEEADC0054FFDDFF06FEEDFE88EC8F" box="[199,354,367,393]" italics="true" pageId="3" pageNumber="472">Left mandible</emphasis>
molar process anterior margin with 15 flat teeth; with proximal cluster of long simple setae; left mandible spine row composed of 8 spines; palp articles 2 with 14 distolateral setae, palp article 3 with 17 biserrate RS. 
<emphasis id="1AFEEADC0054FFDDFF08FE35FE8CECD7" box="[201,358,439,465]" italics="true" pageId="3" pageNumber="472">Left maxillula</emphasis>
mesial lobe with 3 large and circumplumose RS; lateral lobe with 12 RS. 
<emphasis id="1AFEEADC0054FFDDFB74FE35FAA4ECD7" box="[1205,1358,439,465]" italics="true" pageId="3" pageNumber="472">Right maxilla</emphasis>
lateral lobe with 5 long setae; middle lobe with 13 long setae; mesial lobe with 3 distal long setae and 12 proximal long setae. 
<emphasis id="1AFEEADC0054FFDDFF1CFE7DFE7DEF1F" box="[221,407,511,537]" italics="true" pageId="3" pageNumber="472">Right maxilliped</emphasis>
palp article 2 mesial margin with 7 slender setae, lateral margin distally with 1 slender seta; article 3 mesial margin with 14 slender setae, lateral margin with 9 slender setae; article 4 mesial margin with 13 slender setae, lateral margin with 5 slender setae; article 5 distal margin 17 setae, lateral margin with 4 setae; endite with 5 long CPS, and 1 coupling setae (both left and right).
</paragraph>
<paragraph id="283536CE0054FFDDFF06FD0DFECFE80B" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="472">
<emphasis id="1AFEEADC0054FFDDFF06FD0DFE67EFAF" box="[199,397,655,681]" italics="true" pageId="3" pageNumber="472">Pereopod 1 basis</emphasis>
2.3 times as long as greatest width, infererior distal angle with cluster of 2 acute setae; 
<emphasis id="1AFEEADC0054FFDDFA41FD0DFF31EFCB" italics="true" pageId="3" pageNumber="472">ischium</emphasis>
times 0.6 times as long as basis, inferior margin with 2 setae, superior distal margin with 1 RS; 
<emphasis id="1AFEEADC0054FFDDFB3FFD31FAA8EFCA" box="[1278,1346,691,716]" italics="true" pageId="3" pageNumber="472">merus</emphasis>
inferior margin with 6 molariform RS (set as in row of 4+2), superior distal angle with 4 setae (slender); 
<emphasis id="1AFEEADC0054FFDDFB32FD55FAD5EFF6" box="[1267,1343,727,752]" italics="true" pageId="3" pageNumber="472">carpus</emphasis>
inferior distal margin with 2 setae (1 RS and 1 slender seta). 
<emphasis id="1AFEEADC0054FFDDFD39FD79FC8EEE13" box="[760,868,763,789]" italics="true" pageId="3" pageNumber="472">Propodus</emphasis>
2.0 times as long as wide, inferior margin with 3 RS; dactylus 0.7 times as long as propodus. 
<emphasis id="1AFEEADC0054FFDDFD4BFC9DFD1CEE3F" box="[650,758,799,825]" italics="true" pageId="3" pageNumber="472">Pereopod</emphasis>
2 ischium inferior margin with 2 stout, bluntly rounded RS, superior distal margin with 3 RS (2 simple and 1 biserrate); 
<emphasis id="1AFEEADC0054FFDDFCFDFCC1FC6AEE5A" box="[828,896,835,860]" italics="true" pageId="3" pageNumber="472">merus</emphasis>
inferior margin with 8 stout RS (set in row of 5 molariform and 3 acute), superior distal margin with 4 acute RS; 
<emphasis id="1AFEEADC0054FFDDFCA8FCE5FC5FEE86" box="[873,949,871,896]" italics="true" pageId="3" pageNumber="472">carpus</emphasis>
inferodistal angle with 3 RS (plus 1 slender seta); 
<emphasis id="1AFEEADC0054FFDDFF1BFC09FEA9EEA3" box="[218,323,907,933]" italics="true" pageId="3" pageNumber="472">propodus</emphasis>
1.7 times as long as wide; dactylus 0.9 times as long as propodus. 
<emphasis id="1AFEEADC0054FFDDFBE2FC09FB49EEA3" box="[1059,1187,907,933]" italics="true" pageId="3" pageNumber="472">Pereopod 3</emphasis>
similar to pereopod 2. 
<emphasis id="1AFEEADC0054FFDDFF56FC2DFEF2EECF" box="[151,280,943,969]" italics="true" pageId="3" pageNumber="472">Pereopod 4</emphasis>
intermediate in form between pereopod 3 and pereopod 5. 
<emphasis id="1AFEEADC0054FFDDFC6CFC2DFBF3EECF" box="[941,1049,943,969]" italics="true" pageId="3" pageNumber="472">Pereopod</emphasis>
6 similar to pereopod 7. 
<emphasis id="1AFEEADC0054FFDDFAF1FC2DFF0DEEEB" italics="true" pageId="3" pageNumber="472">Pereopod 7 basis</emphasis>
2.8 times as long as greatest width, superior margin convex, inferior margin with 4 palmate setae; 
<emphasis id="1AFEEADC0054FFDDFA84FC51FA76EEEB" box="[1349,1436,979,1005]" italics="true" pageId="3" pageNumber="472">ischium</emphasis>
0.6 times as long as basis. inferior margin with 7 RS (set in groups of 3, 1 and 3), superior distal angle with 6 RS, inferior distal angle with 2 RS; 
<emphasis id="1AFEEADC0054FFDDFE3BFB99FDD4E932" box="[506,574,1051,1076]" italics="true" pageId="3" pageNumber="472">merus</emphasis>
0.7 times as long as ischium, 1.3 times as long as wide, inferior margin with 4 RS (set in groups of 1 and 3), superior distal angle with 13 RS (2 simple and 11 biserrate), inferior distal angle with 6 RS; 
<emphasis id="1AFEEADC0054FFDDFED5FBE1FE8AE97A" box="[276,352,1123,1148]" italics="true" pageId="3" pageNumber="472">carpus</emphasis>
times 0.9 times as long as ischium, 1.7 times as long as wide, inferior margin with 3 RS, superior distal angle with 13 RS (simple and biserrate), inferior distal angle with 10 RS (6 simple and 4 biserrate); 
<emphasis id="1AFEEADC0054FFDDFAF2FB05FA76E9A7" box="[1331,1436,1159,1185]" italics="true" pageId="3" pageNumber="472">propodus</emphasis>
1.0 time as long as ischium, 4.4 times as long as wide, inferior margin with 4 RS (set in group 2 and 2), superior distal angle with 3 RS, 1 slender seta and 1 palmate seta, inferior distal angle with 2 RS; 
<emphasis id="1AFEEADC0054FFDDFBBBFB4DFB3DE9EF" box="[1146,1239,1231,1257]" italics="true" pageId="3" pageNumber="472">dactylus</emphasis>
0.4 times as long as propodus.
</paragraph>
<paragraph id="283536CE0054FFDDFF06FA95FC34E837" blockId="3.[151,1437,151,1977]" box="[199,990,1302,1329]" pageId="3" pageNumber="472">
<emphasis id="1AFEEADC0054FFDDFF06FA95FE96E836" box="[199,380,1303,1329]" italics="true" pageId="3" pageNumber="472">Penial openings</emphasis>
two low tubercles, separated by 25% of sternal width.
</paragraph>
<paragraph id="283536CE0054FFDDFF06FAB9FD65EB2B" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="472">
<emphasis id="1AFEEADC0054FFDDFF06FAB9FECEE853" box="[199,292,1339,1365]" italics="true" pageId="3" pageNumber="472">Pleopod</emphasis>
1 exopod 1.4 times as long as wide, lateral margin straight, distally broadly rounded, mesial margin strongly convex, with 30 PMS from distal one-third; endopod 2.2 times as long as wide, distally broadly rounded, lateral margin sinuate, with 10 PMS on distal margin only; peduncle 1.6 times as wide as long, mesial margin with 4 coupling hook. 
<emphasis id="1AFEEADC0054FFDDFE87FA25FE52E8C7" box="[326,440,1447,1473]" italics="true" pageId="3" pageNumber="472">Pleopod 2</emphasis>
exopod with 38 PMS, endopod with 13 PMS; 
<emphasis id="1AFEEADC0054FFDDFC00FA25FB48E8C7" box="[961,1186,1447,1473]" italics="true" pageId="3" pageNumber="472">appendix masculina</emphasis>
with parallel margins, 1.1 times as long as endopod, distally narrowly rounded. 
<emphasis id="1AFEEADC0054FFDDFCDEFA49FC78E8E3" box="[799,914,1483,1509]" italics="true" pageId="3" pageNumber="472">Pleopod 3</emphasis>
exopod with 40 PMS, endopod with 11 PMS. 
<emphasis id="1AFEEADC0054FFDDFF56FA6DFEE1EB0F" box="[151,267,1519,1545]" italics="true" pageId="3" pageNumber="472">Pleopod 4</emphasis>
exopod with 38 PMS, endopod with 9 PMS. 
<emphasis id="1AFEEADC0054FFDDFCD6FA6DFC9EEB0F" box="[791,884,1519,1545]" italics="true" pageId="3" pageNumber="472">Pleopod</emphasis>
5 exopod with 34 PMS. Pleopods 2–5 peduncle distolateral margin with prominent acute RS.
</paragraph>
<paragraph id="283536CE0054FFDDFF06F9B5FF2BEA03" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="472">
<emphasis id="1AFEEADC0054FFDDFF06F9B5FEF7EB57" box="[199,285,1591,1617]" italics="true" pageId="3" pageNumber="472">Uropod</emphasis>
peduncle ventrolateral margin with 2 RS, lateral margin with 1 mesial short acute RS, posterior lobe about one-half as long as endopod; rami extending beyond pleotelson, marginal setae in single tier, apices broadly round. 
<emphasis id="1AFEEADC0054FFDDFF29F9FDFEA7EB9F" box="[232,333,1663,1689]" italics="true" pageId="3" pageNumber="472">Endopod</emphasis>
apically not bifid; lateral margin distally convex, without prominent excision, with 2 RS, mesial margin strongly convex, with 7 RS. 
<emphasis id="1AFEEADC0054FFDDFDF3F921FD62EBBB" box="[562,648,1699,1725]" italics="true" pageId="3" pageNumber="472">Exopod</emphasis>
not extending to end of endopod, 1.5 times as long as greatest width, apically shallowly and equally sub-bifid; lateral margin weakly convex, with 5 RS; mesial margin convex, with 4 RS.
</paragraph>
<paragraph id="283536CE0054FFDDFF06F88DFC42EA2C" blockId="3.[151,1437,151,1977]" box="[199,936,1807,1834]" pageId="3" pageNumber="472">
<emphasis id="1AFEEADC0054FFDDFF06F88DFECFEA2F" bold="true" box="[199,293,1807,1833]" pageId="3" pageNumber="472">Female.</emphasis>
Similar to male but on average a little larger (see ‘Size’).
</paragraph>
<paragraph id="283536CE0054FFDDFF06F8B0FAFEEA93" blockId="3.[151,1437,151,1977]" pageId="3" pageNumber="472">
<emphasis id="1AFEEADC0054FFDDFF06F8B0FED5EA4A" bold="true" box="[199,319,1842,1868]" pageId="3" pageNumber="472">Variation.</emphasis>
Pleotelson (n=17 [
<specimenCount id="3E8CFD470054FFDDFDD3F8B1FDAFEA4B" box="[530,581,1843,1869]" pageId="3" pageNumber="472" type="male">10♂</specimenCount>
and 
<specimenCount id="3E8CFD470054FFDDFDBAF8B0FD77EA4B" box="[635,669,1842,1869]" pageId="3" pageNumber="472" type="female">7♀</specimenCount>
]) with 7–8 RS, with 8 RS (4+4) most frequent (76%). Uropod endopod mesial margin with 6–8 RS, with 7 (76%) and 6 (18%) most frequent and 8 (6%) occurring only once, lateral margin with 2 RS (100%); exopod mesial margin with 4 RS (100%), lateral margin with 5 RS (100%).
</paragraph>
<paragraph id="283536CE0054FFDDFF06F81CFAFFEABF" blockId="3.[151,1437,151,1977]" box="[199,1301,1950,1977]" pageId="3" pageNumber="472">
<emphasis id="1AFEEADC0054FFDDFF06F81CFF14EABE" bold="true" box="[199,254,1950,1976]" pageId="3" pageNumber="472">Size.</emphasis>
Adult males (n=10) 
<quantity id="EF729B2B0054FFDDFE29F81CFD85EABF" box="[488,623,1950,1977]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" metricValueMax="8.3" metricValueMin="6.5" pageId="3" pageNumber="472" unit="mm" value="7.4" valueMax="8.3" valueMin="6.5">6.5–8.3 mm</quantity>
(mean 
<quantity id="EF729B2B0054FFDDFD03F81DFCF6EABF" box="[706,796,1951,1977]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="3" pageNumber="472" unit="mm" value="7.7">7.7 mm</quantity>
); females (n=7) 
<quantity id="EF729B2B0054FFDDFC15F81CFBB1EABF" box="[980,1115,1950,1977]" metricMagnitude="-3" metricUnit="m" metricValue="7.95" metricValueMax="8.5" metricValueMin="7.4" pageId="3" pageNumber="472" unit="mm" value="7.95" valueMax="8.5" valueMin="7.4">7.4–8.5 mm</quantity>
(mean 8.0 mm).
</paragraph>
</subSubSection>
<caption id="7CF566460053FFDAFF56F8F6FF40EAFE" ID-DOI="http://doi.org/10.5281/zenodo.4650005" ID-Zenodo-Dep="4650005" httpUri="https://zenodo.org/record/4650005/files/figure.png" pageId="4" pageNumber="473" startId="4.[151,250,1908,1932]" targetBox="[203,1378,191,1877]" targetPageId="4">
<paragraph id="283536CE0053FFDAFF56F8F6FF40EAFE" blockId="4.[151,1437,1908,2040]" pageId="4" pageNumber="473">
<emphasis id="1AFEEADC0053FFDAFF56F8F6FEF9EA8A" bold="true" box="[151,275,1908,1932]" pageId="4" pageNumber="473">FIGURE 2.</emphasis>
<taxonomicName id="EF8A4D4D0053FFDAFEDBF8F6FE0CEA8A" authority="Rodcharoen &amp; Bruce, 2021" authorityName="Rodcharoen &amp; Bruce" authorityYear="2021" box="[282,486,1908,1932]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="473" phylum="Arthropoda" rank="species" species="khamensis" status="sp. nov.">
<emphasis id="1AFEEADC0053FFDAFEDBF8F6FE0CEA8A" box="[282,486,1908,1932]" italics="true" pageId="4" pageNumber="473">Cirolana khamensis</emphasis>
</taxonomicName>
<emphasis id="1AFEEADC0053FFDAFE2DF8F6FDD4EA8A" bold="true" box="[492,574,1908,1932]" pageId="4" pageNumber="473">
<taxonomicNameLabel id="01CD57A70053FFDAFE2DF8F6FDD4EA8A" box="[492,574,1908,1932]" pageId="4" pageNumber="473" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (6.5 mm, PSUZC–CR0288-01), A, D; paratype male (8.4 mm, PSUZC–CR0288-02),B, E); paratype female (7.8 mm, PSUZC–CR0288-02), C; paratype male (7.9 mm, PSUZC–CR0288-02), F, G, H. A, dorsal view; B, lateral view; C, head; D, frons; E, pleon; F, antennula; G, antenna peduncle; H, penial opening, sternite 7.
</paragraph>
</caption>
<caption id="7CF566460052FFDBFF56F815FC86EAD2" ID-DOI="http://doi.org/10.5281/zenodo.4650009" ID-Zenodo-Dep="4650009" httpUri="https://zenodo.org/record/4650009/files/figure.png" pageId="5" pageNumber="474" startId="5.[151,250,1943,1967]" targetBox="[184,1403,183,1911]" targetPageId="5">
<paragraph id="283536CE0052FFDBFF56F815FC86EAD2" blockId="5.[151,1436,1943,2004]" pageId="5" pageNumber="474">
<emphasis id="1AFEEADC0052FFDBFF56F815FEFFEAA9" bold="true" box="[151,277,1943,1967]" pageId="5" pageNumber="474">FIGURE 3.</emphasis>
<taxonomicName id="EF8A4D4D0052FFDBFEDFF81AFE06EAB6" authority="Rodcharoen &amp; Bruce, 2021" authorityName="Rodcharoen &amp; Bruce" authorityYear="2021" box="[286,492,1944,1968]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="khamensis" status="sp. nov.">
<emphasis id="1AFEEADC0052FFDBFEDFF81AFE06EAB6" box="[286,492,1944,1968]" italics="true" pageId="5" pageNumber="474">Cirolana khamensis</emphasis>
</taxonomicName>
<emphasis id="1AFEEADC0052FFDBFE35F81AFDADEAB6" bold="true" box="[500,583,1944,1968]" pageId="5" pageNumber="474">
<taxonomicNameLabel id="01CD57A70052FFDBFE35F81AFDADEAB6" box="[500,583,1944,1968]" pageId="5" pageNumber="474" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype male (7.9 mm, PSUZC-CR2088-02). A, left mandible; B, distal part of mandible; C, maxillule; D, maxilla E, maxilliped; G, maxilliped endite.
</paragraph>
</caption>
<caption id="7CF566460051FFD8FF56F815FCBEEAD5" ID-DOI="http://doi.org/10.5281/zenodo.4650011" ID-Zenodo-Dep="4650011" httpUri="https://zenodo.org/record/4650011/files/figure.png" pageId="6" pageNumber="475" startId="6.[151,250,1943,1967]" targetBox="[207,1380,189,1917]" targetPageId="6">
<paragraph id="283536CE0051FFD8FF56F815FCBEEAD5" blockId="6.[151,1436,1943,2003]" pageId="6" pageNumber="475">
<emphasis id="1AFEEADC0051FFD8FF56F815FEFEEAA9" bold="true" box="[151,276,1943,1967]" pageId="6" pageNumber="475">FIGURE 4.</emphasis>
<taxonomicName id="EF8A4D4D0051FFD8FEDDF815FE02EAA9" authority="Rodcharoen &amp; Bruce, 2021" authorityName="Rodcharoen &amp; Bruce" authorityYear="2021" box="[284,488,1943,1967]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="475" phylum="Arthropoda" rank="species" species="khamensis" status="sp. nov.">
<emphasis id="1AFEEADC0051FFD8FEDDF815FE02EAA9" box="[284,488,1943,1967]" italics="true" pageId="6" pageNumber="475">Cirolana khamensis</emphasis>
</taxonomicName>
<emphasis id="1AFEEADC0051FFD8FE2EF815FDABEAA9" bold="true" box="[495,577,1943,1967]" pageId="6" pageNumber="475">
<taxonomicNameLabel id="01CD57A70051FFD8FE2EF815FDABEAA9" box="[495,577,1943,1967]" pageId="6" pageNumber="475" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype male (7.9 mm, PSUZC-CR2088-02). A, pereopod 1; B, dactylus of pereopod 1; C, mesial view of pereopod 1; D, pereopod 2; E, pereopod 4.
</paragraph>
</caption>
<caption id="7CF566460050FFD9FF56F815FC72EAD5" ID-DOI="http://doi.org/10.5281/zenodo.4650013" ID-Zenodo-Dep="4650013" httpUri="https://zenodo.org/record/4650013/files/figure.png" pageId="7" pageNumber="476" startId="7.[151,250,1943,1967]" targetBox="[249,1339,189,1918]" targetPageId="7">
<paragraph id="283536CE0050FFD9FF56F815FC72EAD5" blockId="7.[151,1436,1943,2004]" pageId="7" pageNumber="476">
<emphasis id="1AFEEADC0050FFD9FF56F815FEFFEAB6" bold="true" box="[151,277,1943,1968]" pageId="7" pageNumber="476">FIGURE 5.</emphasis>
<taxonomicName id="EF8A4D4D0050FFD9FEDCF81AFE03EAB6" authority="Rodcharoen &amp; Bruce, 2021" authorityName="Rodcharoen &amp; Bruce" authorityYear="2021" box="[285,489,1944,1968]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="476" phylum="Arthropoda" rank="species" species="khamensis" status="sp. nov.">
<emphasis id="1AFEEADC0050FFD9FEDCF81AFE03EAB6" box="[285,489,1944,1968]" italics="true" pageId="7" pageNumber="476">Cirolana khamensis</emphasis>
</taxonomicName>
<emphasis id="1AFEEADC0050FFD9FE30F81AFDA9EAB6" bold="true" box="[497,579,1944,1968]" pageId="7" pageNumber="476">
<taxonomicNameLabel id="01CD57A70050FFD9FE30F81AFDA9EAB6" box="[497,579,1944,1968]" pageId="7" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype male (7.9 mm, PSUZC-CR2088-02), A, lateral of pereopod 6; B, mesial view of pereopod 6; C, lateral view of pereopod 7; D, mesial of pereopod 7.
</paragraph>
</caption>
<caption id="7CF56646005FFFD6FF56FAE0FA80E87C" ID-DOI="http://doi.org/10.5281/zenodo.4650015" ID-Zenodo-Dep="4650015" box="[151,1386,1378,1403]" httpUri="https://zenodo.org/record/4650015/files/figure.png" pageId="8" pageNumber="477" startId="8.[151,250,1378,1402]" targetBox="[152,1426,187,1344]" targetPageId="8">
<paragraph id="283536CE005FFFD6FF56FAE0FA80E87C" blockId="8.[151,1386,1378,1403]" box="[151,1386,1378,1403]" pageId="8" pageNumber="477">
<emphasis id="1AFEEADC005FFFD6FF56FAE0FEF9E87C" bold="true" box="[151,275,1378,1402]" pageId="8" pageNumber="477">FIGURE 6.</emphasis>
<taxonomicName id="EF8A4D4D005FFFD6FED8FAE0FE0EE87C" authority="Rodcharoen &amp; Bruce, 2021" authorityName="Rodcharoen &amp; Bruce" authorityYear="2021" box="[281,484,1378,1402]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="477" phylum="Arthropoda" rank="species" species="khamensis" status="sp. nov.">
<emphasis id="1AFEEADC005FFFD6FED8FAE0FE0EE87C" box="[281,484,1378,1402]" italics="true" pageId="8" pageNumber="477">Cirolana khamensis</emphasis>
</taxonomicName>
<emphasis id="1AFEEADC005FFFD6FE2BFAE0FDD1E87C" bold="true" box="[490,571,1378,1402]" pageId="8" pageNumber="477">
<taxonomicNameLabel id="01CD57A7005FFFD6FE2BFAE0FDD1E87C" box="[490,571,1378,1402]" pageId="8" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype male (7.9 mm, PSUZC-CR2088-02). A–E, pleopods 1–5 respectively.
</paragraph>
</caption>
<subSubSection id="60906545005FFFD7FF06FA2AFD37EC47" lastPageId="9" lastPageNumber="478" pageId="8" pageNumber="477" type="discussion">
<paragraph id="283536CE005FFFD6FF06FA2AFDF7EB9C" blockId="8.[151,1437,1448,2015]" pageId="8" pageNumber="477">
<emphasis id="1AFEEADC005FFFD6FF06FA2AFED1E8C4" bold="true" box="[199,315,1448,1474]" pageId="8" pageNumber="477">Remarks.</emphasis>
<taxonomicName id="EF8A4D4D005FFFD6FE85FA2AFDCCE8C4" authority="Rodcharoen &amp; Bruce, 2021" authorityName="Rodcharoen &amp; Bruce" authorityYear="2021" box="[324,550,1448,1474]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="477" phylum="Arthropoda" rank="species" species="khamensis" status="sp. nov.">
<emphasis id="1AFEEADC005FFFD6FE85FA2AFDCCE8C4" box="[324,550,1448,1474]" italics="true" pageId="8" pageNumber="477">Cirolana khamensis</emphasis>
</taxonomicName>
<emphasis id="1AFEEADC005FFFD6FDEEFA2AFD61E8C4" bold="true" box="[559,651,1448,1474]" pageId="8" pageNumber="477">
<taxonomicNameLabel id="01CD57A7005FFFD6FDEEFA2AFD61E8C4" box="[559,651,1448,1474]" pageId="8" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is characterized by pereonites 1–3 each with a single transverse impressed line; pereonites 4–6 with each with two transverse impressed lines; pereonite 7 with three transverse lines with the posterior line forming a nodulose ridge; the antennula peduncular articles 1 and 2 are distinct and articulated; pleotelson dorsal surface with two sub-median longitudinal carinae, each of which has four prominent tubercles, posterior margin evenly rounded, with eight robust setae; pereopod 1 merus inferior margin with six molariform RS; lateral margin of uropodal exopod having continuous row of PMS; and penes in the form of two low tubercles separated by 25% of sternal width.
</paragraph>
<paragraph id="283536CE005FFFD6FF06F927FBC7EA75" blockId="8.[151,1437,1448,2015]" pageId="8" pageNumber="477">
We place 
<taxonomicName id="EF8A4D4D005FFFD6FEF4F926FDFCEBB8" authorityName="Rodcharoen &amp; Bruce" authorityYear="2021" box="[309,534,1700,1726]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="477" phylum="Arthropoda" rank="species" species="khamensis">
<emphasis id="1AFEEADC005FFFD6FEF4F926FDFCEBB8" box="[309,534,1700,1726]" italics="true" pageId="8" pageNumber="477">Cirolana khamensis</emphasis>
</taxonomicName>
into the 
<emphasis id="1AFEEADC005FFFD6FDB8F926FC85EBB8" box="[633,879,1700,1726]" italics="true" pageId="8" pageNumber="477">
<taxonomicName id="EF8A4D4D005FFFD6FDB8F926FD37EBB8" authorityName="Leach" authorityYear="1818" box="[633,733,1700,1726]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="477" phylum="Arthropoda" rank="genus">Cirolana</taxonomicName>
‘pleonastica
</emphasis>
-group’ as the species shows all the characters for that group, with the exception of the uropodal exopod lateral margin, which has setae along the distal two-thirds of its length, though not the entire length. Nonetheless, the morphology and setation of the uropodal exopod is closer to that of the 
<emphasis id="1AFEEADC005FFFD6FEEEF892FDCFEA2C" box="[303,549,1808,1834]" italics="true" pageId="8" pageNumber="477">
<taxonomicName id="EF8A4D4D005FFFD6FEEEF892FE79EA2C" authorityName="Leach" authorityYear="1818" box="[303,403,1808,1834]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="477" phylum="Arthropoda" rank="genus">Cirolana</taxonomicName>
‘pleonastica
</emphasis>
-group’ than to the other clades, and furthermore 
<taxonomicName id="EF8A4D4D005FFFD6FB90F893FB0DEA2C" authorityName="Rodcharoen &amp; Bruce" authorityYear="2021" box="[1105,1255,1808,1835]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="477" phylum="Arthropoda" rank="species" species="khamensis">
<emphasis id="1AFEEADC005FFFD6FB90F893FB0DEA2C" box="[1105,1255,1808,1835]" italics="true" pageId="8" pageNumber="477">C. khamensis</emphasis>
</taxonomicName>
lacks the defining characters of the other groups (see 
<bibRefCitation id="4C1B4B3F005FFFD6FD8AF8B6FD15EA48" author="Sidabalok, C. M." box="[587,767,1844,1870]" pageId="8" pageNumber="477" refId="ref8329" refString="Sidabalok, C. M. (2019) Systematics, phylogeny and distribution of Cirolanidae (Crustacea: Isopoda) of Indonesia and adjacent waters. Vol. PhD. Centre for Sustainable Tropical Fisheries &amp; Aquaculture. James Cook University, Townsville, xxii + 295 pp." type="book" year="2019">Sidabalok 2019</bibRefCitation>
). In their diagnosis of the 
<emphasis id="1AFEEADC005FFFD6FBE2F8B6FB47EA48" box="[1059,1197,1844,1870]" italics="true" pageId="8" pageNumber="477">‘pleonastica</emphasis>
-group’, 
<bibRefCitation id="4C1B4B3F005FFFD6FACEF8B6FED2EA74" author="Sidabalok, C. &amp; Bruce, N. L." pageId="8" pageNumber="477" pagination="177 - 207" refId="ref8553" refString="Sidabalok, C. M &amp; Bruce, N. L. (2018 a) Review of the Cirolana ' pleonastica - group' (Crustacea: Isopoda: Cirolanidae) with description of four new species from the Indo-Malaysian region. Raffles Bulletin of Zoology, 66, 177 - 207." type="journal article" year="2018">Sidabalok &amp; Bruce (2018a)</bibRefCitation>
state the uropod exopod lateral margin character is usually present.
</paragraph>
<paragraph id="283536CE005FFFD7FF06F8FEFD37EC47" blockId="8.[151,1437,1448,2015]" lastBlockId="9.[151,1437,151,357]" lastPageId="9" lastPageNumber="478" pageId="8" pageNumber="477">
Similar South-East Asian species in the 
<emphasis id="1AFEEADC005FFFD6FD4BF8FEFC6AEA90" box="[650,896,1916,1942]" italics="true" pageId="8" pageNumber="477">
<taxonomicName id="EF8A4D4D005FFFD6FD4BF8FEFD04EA90" authorityName="Leach" authorityYear="1818" box="[650,750,1916,1942]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="477" phylum="Arthropoda" rank="genus">Cirolana</taxonomicName>
‘pleonastica
</emphasis>
-group’ are 
<taxonomicName id="EF8A4D4D005FFFD6FBC2F8FEFE97EABD" authority="Rodcharoen et al., 2017" authorityName="Rodcharoen" authorityYear="2017" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="477" phylum="Arthropoda" rank="species" species="phuketensis">
<emphasis id="1AFEEADC005FFFD6FBC2F8FEFB18EA90" box="[1027,1266,1916,1942]" italics="true" pageId="8" pageNumber="477">Cirolana phuketensis</emphasis>
<bibRefCitation id="4C1B4B3F005FFFD6FB38F8FEFF16EABC" author="Rodcharoen, E. &amp; Bruce, N. L. &amp; Pholpunthin, P." pageId="8" pageNumber="477" pagination="1 - 17" refId="ref8219" refString="Rodcharoen, E., Bruce, N. L. &amp; Pholpunthin, P. (2017) Cirolana phuketensis, a new species of marine isopod (Crustacea, Isopoda, Cirolanidae) from the Andaman Sea coast of Thailand. ZooKeys, 694, 1 - 17. https: // doi. org / 10.3897 / zookeys. 695.13771" type="journal article" year="2017">
Rodcharoen 
<emphasis id="1AFEEADC005FFFD6FA49F8FFFF47EABC" italics="true" pageId="8" pageNumber="477">et al</emphasis>
., 2017
</bibRefCitation>
(
<collectingCountry id="509D765E005FFFD6FECDF822FE9FEABD" box="[268,373,1952,1979]" name="Thailand" pageId="8" pageNumber="477">Thailand</collectingCountry>
)
</taxonomicName>
, 
<taxonomicName id="EF8A4D4D005FFFD6FE4BF823FC56EABD" authority="Sidabalok &amp; Bruce, 2018" authorityName="Sidabalok &amp; Bruce" authorityYear="2018" box="[394,956,1952,1979]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="477" phylum="Arthropoda" rank="species" species="merlion">
<emphasis id="1AFEEADC005FFFD6FE4BF823FDE9EABC" box="[394,515,1952,1979]" italics="true" pageId="8" pageNumber="477">C. merlion</emphasis>
Sidabalok &amp; Bruce, 2018 (
<collectingCountry id="509D765E005FFFD6FCFCF822FC5EEABD" box="[829,948,1952,1979]" name="Singapore" pageId="8" pageNumber="477">Singapore</collectingCountry>
)
</taxonomicName>
, 
<taxonomicName id="EF8A4D4D005FFFD6FC08F823FF30EAD9" authority="Sidabalok &amp; Bruce 2018" authorityName="Sidabalok &amp; Bruce" authorityYear="2018" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="477" phylum="Arthropoda" rank="species" species="fasfes">
<emphasis id="1AFEEADC005FFFD6FC08F823FBC1EABC" box="[969,1067,1952,1979]" italics="true" pageId="8" pageNumber="477">C. fasfes</emphasis>
Sidabalok &amp; Bruce 2018 (
<collectingCountry id="509D765E005FFFD6FA9FF822FF38EAD9" name="Indonesia" pageId="8" pageNumber="477">Indonesia</collectingCountry>
)
</taxonomicName>
(
<bibRefCitation id="4C1B4B3F005FFFD6FF2AF846FDF0EAD8" author="Sidabalok, C. &amp; Bruce, N. L." box="[235,538,1988,2015]" pageId="8" pageNumber="477" pagination="177 - 207" refId="ref8553" refString="Sidabalok, C. M &amp; Bruce, N. L. (2018 a) Review of the Cirolana ' pleonastica - group' (Crustacea: Isopoda: Cirolanidae) with description of four new species from the Indo-Malaysian region. Raffles Bulletin of Zoology, 66, 177 - 207." type="journal article" year="2018">Sidabalok &amp; Bruce 2018a</bibRefCitation>
). However, the new species can be distinguished from those species by lateral margin of uropodal exopod having a more continuous row of PMS (vs uropodal exopod lateral margin with widelyspaced and acute robust setae). Moreover, 
<taxonomicName id="EF8A4D4D005EFFD7FD41FF39FCF2EDD3" authority="Rodcharoen &amp; Bruce, 2021" authorityName="Rodcharoen &amp; Bruce" authorityYear="2021" box="[640,792,187,213]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="478" phylum="Arthropoda" rank="species" species="khamensis" status="sp. nov.">
<emphasis id="1AFEEADC005EFFD7FD41FF39FCF2EDD3" box="[640,792,187,213]" italics="true" pageId="9" pageNumber="478">C. khamensis</emphasis>
</taxonomicName>
<emphasis id="1AFEEADC005EFFD7FCE1FF39FC96EDD3" bold="true" box="[800,892,187,213]" pageId="9" pageNumber="478">
<taxonomicNameLabel id="01CD57A7005EFFD7FCE1FF39FC96EDD3" box="[800,892,187,213]" pageId="9" pageNumber="478" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be separated from the allopatric species 
<taxonomicName id="EF8A4D4D005EFFD7FA42FF39FE01EDFF" authority="Bruce &amp; Olesen, 2002" authorityName="Bruce &amp; Olesen" authorityYear="2002" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="478" phylum="Arthropoda" rank="species" species="bruscai">
<emphasis id="1AFEEADC005EFFD7FA42FF39FF01EDFF" italics="true" pageId="9" pageNumber="478">C. bruscai</emphasis>
<bibRefCitation id="4C1B4B3F005EFFD7FF33FF5DFE01EDFF" author="Bruce, N. L. &amp; Olesen, J." box="[242,491,223,249]" pageId="9" pageNumber="478" pagination="109 - 131" refId="ref7585" refString="Bruce, N. L. &amp; Olesen, J. (2002) Cirolanid isopods from the Andaman Sea off Phuket, Thailand, with description of two new species. In: Bruce, N. L., Berggren, M. &amp; Bussawarit, S. (Eds.), Proceedings of the International Workshop on the Crustacea in the Andaman Sea, Phuket Marine Biological Center, 29 November - 20 December 1998, PMBC Special Publication 23. Phuket Marine Biological Center, Phuket, pp. 109 - 131." type="book chapter" year="2002">Bruce &amp; Olesen, 2002</bibRefCitation>
</taxonomicName>
by the head not being produced and overriding the antennular bases; and the inferior margins of pereopods 6 and 7 are smooth (vs head weakly produced and overriding the antennular bases; inferior margins of pereopods 6 and 7 serrate in 
<taxonomicName id="EF8A4D4D005EFFD7FD99FEA5FD27EC47" authorityName="Bruce &amp; Olesen" authorityYear="2002" box="[600,717,295,321]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="478" phylum="Arthropoda" rank="species" species="bruscai">
<emphasis id="1AFEEADC005EFFD7FD99FEA5FD27EC47" box="[600,717,295,321]" italics="true" pageId="9" pageNumber="478">C. bruscai</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="60906545005EFFD7FF06FEC9FCC5EC63" box="[199,815,331,357]" pageId="9" pageNumber="478" type="etymology">
<paragraph id="283536CE005EFFD7FF06FEC9FCC5EC63" blockId="9.[151,1437,151,357]" box="[199,815,331,357]" pageId="9" pageNumber="478">
<emphasis id="1AFEEADC005EFFD7FF06FEC9FEA4EC63" bold="true" box="[199,334,331,357]" pageId="9" pageNumber="478">Etymology.</emphasis>
The epithet is taken from the 
<typeStatus id="F731886C005EFFD7FD5FFEC9FD24EC63" box="[670,718,331,357]" pageId="9" pageNumber="478">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>