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<document id="56498709382B5B9B8F8C74D5BF656C6C" ID-DOI="10.35463/j.apr.2020.01.04" ID-ISSN="1842-371x" ID-Zenodo-Dep="10520716" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1705450337321" checkinUser="felipe" docAuthor="Schlagintweit, Felix" docDate="2020" docId="A007EF1AFF91FFFAFF45E89DFD2FF8E8" docLanguage="en" docName="ActaPalaeontolRom.16.1.37-41.pdf" docOrigin="Acta Palaeontologica Romaniae 16 (1)" docSource="http://dx.doi.org/10.35463/j.apr.2020.01.04" docStyle="DocumentStyle:CA474AAD92E97C533D63E0C5C0F159F1.1:ActaPalaeontolRom.2019-.journal_article" docStyleId="CA474AAD92E97C533D63E0C5C0F159F1" docStyleName="ActaPalaeontolRom.2019-.journal_article" docStyleVersion="1" docTitle="Paracoskinolina prestati Peybernes &amp; Delmas 1981" docType="treatment" docVersion="3" lastPageNumber="39" masterDocId="5C3E9762FF93FFF8FFD2EB0EFFFAFFFD" masterDocTitle="Cribellopsis Moulladei (Saint-Marc, 1974) Nov. Comb. (Foraminiferida, Orbitolinidae): An Albian Marker Taxon Of The Southern Neotethysian Margin" masterLastPageNumber="41" masterPageNumber="37" pageNumber="39" updateTime="1708369347788" updateUser="juliana" zenodo-license-document="CC-BY-4.0">
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<mods:title id="23F419979343DF62D8CF8FB1D0B2D2D1">Cribellopsis Moulladei (Saint-Marc, 1974) Nov. Comb. (Foraminiferida, Orbitolinidae): An Albian Marker Taxon Of The Southern Neotethysian Margin</mods:title>
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<taxonomicName id="EFAE258FFF91FFFAFF45E89DFD20FC56" ID-CoL="8N6TZ" authority="Peybernes &amp; Delmas, 1981" authorityName="Peybernes &amp; Delmas" authorityYear="1981" box="[151,730,915,939]" class="Globothalamea" family="Orbitolinidae" genus="Paracoskinolina" kingdom="Chromista" order="Loftusiida" pageId="2" pageNumber="39" phylum="Foraminifera" rank="species" species="prestati">
<emphasis id="1ADA821EFF91FFFAFF45E89DFD20FC56" bold="true" box="[151,730,915,939]" pageId="2" pageNumber="39">
<emphasis id="1ADA821EFF91FFFAFF45E89DFE59FC56" bold="true" box="[151,419,915,939]" italics="true" pageId="2" pageNumber="39">Paracoskinolina prestati</emphasis>
<bibRefCitation id="4C3F23FDFF91FFFAFE79E89DFD20FC56" author="Peybernes, B. &amp; Delmas, J." box="[427,730,915,939]" pageId="2" pageNumber="39" pagination="80 - 86" refId="ref4131" refString="Peybernes, B., Delmas, J., 1981. Description de Paracoskinolina prestati nov. sp., Orbitolinide nouveau du Cretace Moyen de la Republique de Somalie. Bulletin de la Societe d'histoire naturelle de Toulouse, 154: 80 - 86." type="journal article" year="1981">Peybernés &amp; Delmas, 1981</bibRefCitation>
</emphasis>
</taxonomicName>
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<subSubSection id="60B40D87FF91FFFAFF45E8CEFD2FF8E8" pageId="2" pageNumber="39" type="description">
<paragraph id="28115E0CFF91FFFAFF45E8CEFD2FF8E8" blockId="2.[151,771,960,1813]" pageId="2" pageNumber="39">
The axial sections provided by
<bibRefCitation id="4C3F23FDFF91FFFAFDD6E8CEFF25FC0A" author="Peybernes, B. &amp; Delmas, J." pageId="2" pageNumber="39" pagination="80 - 86" refId="ref4131" refString="Peybernes, B., Delmas, J., 1981. Description de Paracoskinolina prestati nov. sp., Orbitolinide nouveau du Cretace Moyen de la Republique de Somalie. Bulletin de la Societe d'histoire naturelle de Toulouse, 154: 80 - 86." type="journal article" year="1981">Peybernès and Delmas (1981)</bibRefCitation>
as well as those from
<bibRefCitation id="4C3F23FDFF91FFFAFE0DE8EEFD8AFC0A" author="Prestat, B." box="[479,624,991,1015]" pageId="2" pageNumber="39" pagination="273 - 309" refId="ref4178" refString="Prestat, B., 1977. Marqueurs micropaleontologiques du Jurassique et du Cretace de Somalie centrale. Ann. Mines Geol. (Actes VI Coll. Africain Micropaleontol. Tunis 1974), 28: 273 - 309." type="journal article" year="1977">Prestat (1977</bibRefCitation>
; see synonymy) clearly evidence that the &quot;pillars&quot; are not aligned between subsequent chambers (
<figureCitation id="B0954289FF91FFFAFE2CEF12FD4DFBC9" box="[510,695,1052,1077]" captionStart="Fig" captionStartId="1.[225,260,1004,1025]" captionTargetBox="[225,1378,166,967]" captionTargetId="figure-730@1.[225,1378,166,967]" captionTargetPageId="1" captionText="Fig. 1 Cribellopsis moulladei (Saint-Marc) nov. comb., Albian of Somalia (ab, jl, r), Italy (cg, m, pq, s), and Lebanon (hi, no; type material). af Axial sections of megalospheric specimens. g, r Subaxial sections of possibly microspheric specimens. hi, k Subaxial sections of megalospheric specimens. j, m Tangential-oblique sections. l, q Tangential sections. np, s Transverse sections. ab, k extracted from Peybernès &amp; Delmas (1981, pl. 1, fig. 12, h) as Paracoskinolina prestati; j, l extracted from Peybernès (1982, pl. 2, fig. 12, fig. 11) illustrated as Paracoskinolina prestati; c, fg, m, p, q, s extracted from Chiocchini (1989, pl. 1, fig. 14, fig. 5, pl. 2, fig. 8, pl. 1, fig. 11, fig. 10, fig. 12, fig. 4) illustrated as Cribellopsis arnaudae n. sp.; d extracted from Schmitt et al. (2019, in press, fig. 5d) illustrated as Cribellopsis arnaudae Chiocchini. e extracted from Cruz-Abad (2018, fig. 93J) illustrated as Cribellopsis aff. arnaudae Chiocchini; hi, no extracted from Saint-Marc (1974, pl. 2, figs. 6, 34, 7) illustrated as Simplorbitolina moulladei n. sp. r extracted from Prestat (1977, pl. 12, fig. 10) illustrated as Iraqia sp. Abbreviations: b = beam (or main partition), deu = deuteroconch, ib = intercalary beam, pr = proloculus" figureDoi="http://doi.org/10.5281/zenodo.10520718" httpUri="https://zenodo.org/record/10520718/files/figure.png" pageId="2" pageNumber="39">Fig. 1ab, jl, r</figureCitation>
). This feature, however, is one of the diagnostic characteristics of the genus
<taxonomicName id="EFAE258FFF91FFFAFEE9EF54FD9AFB8F" authority="Moullade" authorityName="Moullade" authorityYear="1965" box="[315,608,1114,1138]" class="Globothalamea" family="Orbitolinidae" genus="Paracoskinolina" kingdom="Chromista" order="Loftusiida" pageId="2" pageNumber="39" phylum="Foraminifera" rank="genus">
<emphasis id="1ADA821EFF91FFFAFEE9EF54FE11FB8F" box="[315,491,1114,1138]" italics="true" pageId="2" pageNumber="39">Paracoskinolina</emphasis>
Moullade
</taxonomicName>
(see
<bibRefCitation id="4C3F23FDFF91FFFAFD78EF54FEC6FB6D" author="Arnaud-Vanneau, A." pageId="2" pageNumber="39" pagination="1 - 874" refId="ref3447" refString="Arnaud-Vanneau, A., 1980. L´Urgonien du Vercors septentrional et de la Chartreuse. Geologie Alpine Memoire, 11 (3 volumes): 1 - 874." type="journal article" year="1980">Arnaud-Vanneau, 1980</bibRefCitation>
, or Clavel in
<bibRefCitation id="4C3F23FDFF91FFFAFE02EF76FD5EFB6D" author="Granier, B. &amp; Clavel, B. &amp; Moullade, M. &amp; Busnardo, R. &amp; Charollais, J. &amp; Tronchetti, G. &amp; Desjacques, P." box="[464,676,1144,1168]" pageId="2" pageNumber="39" pagination="2013" refId="ref3836" refString="Granier, B., Clavel, B., Moullade, M., Busnardo, R., Charollais, J., Tronchetti, G., Desjacques, P., 2013. L'Estellon (Baronnies, France), a &quot; Rosetta Stone &quot; for the Urgonian biostratigraphy. Carnets de Geologie (Notebooks on Geology), Article 2013 / 04 (CG 2013 _ A 04)." type="journal article" year="2013">Granier et al., 2013</bibRefCitation>
, p. 153). Therefore, the upper? Albian Somali taxon can not be a species of
<taxonomicName id="EFAE258FFF91FFFAFEDDEFB8FE45FB33" authorityName="Moullade" authorityYear="1965" box="[271,447,1206,1230]" class="Globothalamea" family="Orbitolinidae" genus="Paracoskinolina" kingdom="Chromista" order="Loftusiida" pageId="2" pageNumber="39" phylum="Foraminifera" rank="genus">
<emphasis id="1ADA821EFF91FFFAFEDDEFB8FE45FB33" box="[271,447,1206,1230]" italics="true" pageId="2" pageNumber="39">Paracoskinolina</emphasis>
</taxonomicName>
. In fact, the partitions of the central zone in
<taxonomicName id="EFAE258FFF91FFFAFE91EFDAFE3FFB11" authorityName="Arnaud-Vanneau" authorityYear="1980" box="[323,453,1236,1260]" class="Globothalamea" family="Orbitolinidae" genus="Cribellopsis" kingdom="Chromista" order="Loftusiida" pageId="2" pageNumber="39" phylum="Foraminifera" rank="genus">
<emphasis id="1ADA821EFF91FFFAFE91EFDAFE3FFB11" box="[323,453,1236,1260]" italics="true" pageId="2" pageNumber="39">Cribellopsis</emphasis>
</taxonomicName>
are thickening &quot;upwards&quot; (= towards the apex;
<bibRefCitation id="4C3F23FDFF91FFFAFE8FEFFDFDA1FAF6" author="Arnaud-Vanneau, A." box="[349,603,1267,1291]" pageId="2" pageNumber="39" pagination="1 - 874" refId="ref3447" refString="Arnaud-Vanneau, A., 1980. L´Urgonien du Vercors septentrional et de la Chartreuse. Geologie Alpine Memoire, 11 (3 volumes): 1 - 874." type="journal article" year="1980">Arnaud-Vanneau, 1980</bibRefCitation>
, p. 668, &quot;épaississent vers le haut&quot;). This feature observable in axial sections, was also reported by
<bibRefCitation id="4C3F23FDFF91FFFAFE35EE3EFD5CFAB5" author="Saint-Marc, P." box="[487,678,1328,1352]" pageId="2" pageNumber="39" pagination="1 - 342" refId="ref4278" refString="Saint-Marc, P., 1974. Etude stratigraphique et micropaleontologique de l'Albien, du Cenomanien et du Turonien du Liban. Notes et Memoires sur le Moyen- Orient, 13: 1 - 342." type="journal article" year="1974">Saint-Marc (1974</bibRefCitation>
, p. 225) for &quot;
<taxonomicName id="EFAE258FFF91FFFAFF1DEE41FE03FA9A" authorityName="Schlagintweit" authorityYear="2020" box="[207,505,1359,1383]" class="Globothalamea" family="Orbitolinidae" genus="Simplorbitolina" kingdom="Chromista" order="Loftusiida" pageId="2" pageNumber="39" phylum="Foraminifera" rank="species" species="moulladei">
<emphasis id="1ADA821EFF91FFFAFF1DEE41FE8CFA9A" box="[207,374,1359,1383]" italics="true" pageId="2" pageNumber="39">Simplorbitolina</emphasis>
&quot;
<emphasis id="1ADA821EFF91FFFAFE5DEE41FE03FA9A" box="[399,505,1359,1383]" italics="true" pageId="2" pageNumber="39">moulladei</emphasis>
</taxonomicName>
(&quot;cloisons radiales ont une section triangulaire, en cȏne renversé&quot;). It is worth mentioning here that this aspect of the central zone is described from several doubtful paracoskinolinas such as
<taxonomicName id="EFAE258FFF91FFFAFF45EEC5FE66FA1F" authority="Peybernes" authorityName="Peybernes" box="[151,412,1482,1506]" class="Globothalamea" family="Orbitolinidae" genus="Paracoskinolina" kingdom="Chromista" order="Loftusiida" pageId="2" pageNumber="39" phylum="Foraminifera" rank="species" species="tunesiana">
<emphasis id="1ADA821EFF91FFFAFF45EEC5FEDBFA1F" box="[151,289,1483,1506]" italics="true" pageId="2" pageNumber="39">P. tunesiana</emphasis>
Peybernès
</taxonomicName>
,
<taxonomicName id="EFAE258FFF91FFFAFE79EEC5FF39F9FD" authority="Canerot &amp; Moullade" authorityName="Canerot &amp; Moullade" authorityYear="1971" class="Globothalamea" family="Orbitolinidae" genus="Paracoskinolina" kingdom="Chromista" order="Loftusiida" pageId="2" pageNumber="39" phylum="Foraminifera" rank="species" species="pfenderae">
<emphasis id="1ADA821EFF91FFFAFE79EEC5FDCCFA1F" box="[427,566,1482,1506]" italics="true" pageId="2" pageNumber="39">P. pfenderae</emphasis>
Canérot &amp; Moullade
</taxonomicName>
,
<taxonomicName id="EFAE258FFF91FFFAFF07EEE7FE23F9FD" authority="Peybernes" authorityName="Peybernes" authorityYear="1976" box="[213,473,1512,1536]" class="Globothalamea" family="Orbitolinidae" genus="Paracoskinolina" kingdom="Chromista" order="Loftusiida" pageId="2" pageNumber="39" phylum="Foraminifera" rank="species" species="hispanica">
<emphasis id="1ADA821EFF91FFFAFF07EEE7FEA7F9FD" box="[213,349,1512,1536]" italics="true" pageId="2" pageNumber="39">P. hispanica</emphasis>
Peybernès
</taxonomicName>
, or
<taxonomicName id="EFAE258FFF91FFFAFDDAEEE7FED0F9E2" authority="Canerot &amp; Peybernes" authorityName="Canerot &amp; Peybernes" authorityYear="1981" class="Globothalamea" family="Orbitolinidae" genus="Paracoskinolina" kingdom="Chromista" order="Loftusiida" pageId="2" pageNumber="39" phylum="Foraminifera" rank="species" species="querolensis">
<emphasis id="1ADA821EFF91FFFAFDDAEEE7FD5EF9FD" box="[520,676,1512,1536]" italics="true" pageId="2" pageNumber="39">P. querolensis</emphasis>
Canérot &amp; Peybernès
</taxonomicName>
as &quot;hemi-pillars…pointe en haut&quot; (e.g., Peybrnès and Delmas, 1981, p. 82). Transverse sections of
<taxonomicName id="EFAE258FFF91FFFAFF67ED4AFECDF9A1" authorityName="Arnaud-Vanneau" authorityYear="1980" box="[181,311,1604,1628]" class="Globothalamea" family="Orbitolinidae" genus="Cribellopsis" kingdom="Chromista" order="Loftusiida" pageId="2" pageNumber="39" phylum="Foraminifera" rank="genus">
<emphasis id="1ADA821EFF91FFFAFF67ED4AFECDF9A1" box="[181,311,1604,1628]" italics="true" pageId="2" pageNumber="39">Cribellopsis</emphasis>
</taxonomicName>
show that the vertical elements of the central zone are not isolated columnar pillars but sections of a reticulate (or anastomosing), often loose and poorly constrained, network formed by laterally fused main partitions (e.g.,
<figureCitation id="B0954289FF91FFFAFECCEDB1FE9CF92A" box="[286,358,1727,1751]" captionStart="Fig" captionStartId="1.[225,260,1004,1025]" captionTargetBox="[225,1378,166,967]" captionTargetId="figure-730@1.[225,1378,166,967]" captionTargetPageId="1" captionText="Fig. 1 Cribellopsis moulladei (Saint-Marc) nov. comb., Albian of Somalia (ab, jl, r), Italy (cg, m, pq, s), and Lebanon (hi, no; type material). af Axial sections of megalospheric specimens. g, r Subaxial sections of possibly microspheric specimens. hi, k Subaxial sections of megalospheric specimens. j, m Tangential-oblique sections. l, q Tangential sections. np, s Transverse sections. ab, k extracted from Peybernès &amp; Delmas (1981, pl. 1, fig. 12, h) as Paracoskinolina prestati; j, l extracted from Peybernès (1982, pl. 2, fig. 12, fig. 11) illustrated as Paracoskinolina prestati; c, fg, m, p, q, s extracted from Chiocchini (1989, pl. 1, fig. 14, fig. 5, pl. 2, fig. 8, pl. 1, fig. 11, fig. 10, fig. 12, fig. 4) illustrated as Cribellopsis arnaudae n. sp.; d extracted from Schmitt et al. (2019, in press, fig. 5d) illustrated as Cribellopsis arnaudae Chiocchini. e extracted from Cruz-Abad (2018, fig. 93J) illustrated as Cribellopsis aff. arnaudae Chiocchini; hi, no extracted from Saint-Marc (1974, pl. 2, figs. 6, 34, 7) illustrated as Simplorbitolina moulladei n. sp. r extracted from Prestat (1977, pl. 12, fig. 10) illustrated as Iraqia sp. Abbreviations: b = beam (or main partition), deu = deuteroconch, ib = intercalary beam, pr = proloculus" figureDoi="http://doi.org/10.5281/zenodo.10520718" httpUri="https://zenodo.org/record/10520718/files/figure.png" pageId="2" pageNumber="39">Fig. 1s</figureCitation>
). This statement does not mean
<emphasis id="1ADA821EFF91FFFAFD11EDCEFCF9F92A" box="[707,771,1728,1751]" italics="true" pageId="2" pageNumber="39">per se</emphasis>
that all of the above cited species belong to
<taxonomicName id="EFAE258FFF91FFFAFDA8EDD0FD06F90B" authorityName="Arnaud-Vanneau" authorityYear="1980" box="[634,764,1758,1782]" class="Globothalamea" family="Orbitolinidae" genus="Cribellopsis" kingdom="Chromista" order="Loftusiida" pageId="2" pageNumber="39" phylum="Foraminifera" rank="genus">
<emphasis id="1ADA821EFF91FFFAFDA8EDD0FD06F90B" box="[634,764,1758,1782]" italics="true" pageId="2" pageNumber="39">Cribellopsis</emphasis>
</taxonomicName>
. Their taxonomic status must be clarified individually.
</paragraph>
</subSubSection>
</treatment>
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