116 lines
9.9 KiB
XML
116 lines
9.9 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.777.23212" ID-GBIF-Dataset="516d998c-117e-4cbb-9e21-e38843778702" ID-PMC="PMC6079114" ID-Pensoft-Pub="1313-2970-777-57" ID-PubMed="30100790" ID-ZBK="D9F353643DCD4BA6B70D62FB275DEB1B" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-777-57" ModsDocOrigin="ZooKeys 777" ModsDocTitle="A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)" checkinTime="1533035384019" checkinUser="pensoft" docAuthor="Schilthuizen, Menno, Perreau, Michel & Njunjic, Iva" docDate="2018" docId="9F9D2431A8175D6BA4B04808E81C1B4B" docLanguage="en" docName="ZooKeys 777: 57-108" docOrigin="ZooKeys 777" docSource="http://dx.doi.org/10.3897/zookeys.777.23212" docTitle="Ptomaphaginus louis Schilthuizen & Perreau, sp. n." docType="treatment" docUuid="67975FC1-077B-4348-86EA-159B147E148C" docUuidSource="ZooBank" docVersion="5" lastPageNumber="73" masterDocId="F179FFE1BF6AFFEEEF68FFF0FFB5FFBD" masterDocTitle="A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)" masterLastPageNumber="108" masterPageNumber="57" pageNumber="73" updateTime="1668166102793" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Schilthuizen, Menno</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Perreau, Michel</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Njunjic, Iva</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2018</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>777</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>57</mods:start>
|
||
<mods:end>108</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zookeys.777.23212</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.777.23212</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1313-2970-777-57</mods:identifier>
|
||
<mods:identifier type="ZBK">D9F353643DCD4BA6B70D62FB275DEB1B</mods:identifier>
|
||
<mods:identifier type="ZooBank">D9F353643DCD4BA6B70D62FB275DEB1B</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="146746164" LSID="urn:lsid:zoobank.org:act:67975FC1-077B-4348-86EA-159B147E148C" httpUri="http://treatment.plazi.org/id/9F9D2431A8175D6BA4B04808E81C1B4B" lastPageNumber="73" pageId="16" pageNumber="73">
|
||
<subSubSection pageId="16" pageNumber="73" type="nomenclature">
|
||
<paragraph pageId="16" pageNumber="73">
|
||
<taxonomicName LSID="http://zoobank.org/67975FC1-077B-4348-86EA-159B147E148C" authority="Schilthuizen & Perreau" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphaginus louis" order="Coleoptera" pageId="16" pageNumber="73" phylum="Arthropoda" rank="species" species="louis">Ptomaphaginus louis Schilthuizen & Perreau</taxonomicName>
|
||
<taxonomicNameLabel pageId="16" pageNumber="73">sp. n.</taxonomicNameLabel>
|
||
Figure 5a, 6m, n, 9k, 10d
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="73" type="materials_examined">
|
||
<paragraph pageId="16" pageNumber="73">Material.</paragraph>
|
||
<paragraph pageId="16" pageNumber="73">
|
||
Holotype: Malaysia, Sarawak, Mulu National Park, TPS 7-13, 4.v.1978 (leg. P.M. Hammond & J.E. Marshall, NHMUK, B.M. 1978-49), male. Paratypes: Sarawak. Mulu National Park, TPS 7-13, 4.v.1978 (leg. P.M. Hammond & J.E. Marshall, NHMUK, B.M. 1978-49), 12 males, 14 females; Mulu National Park, Kerangas?, 7.iv.1978 (leg. P.M. Hammond & J.E. Marshall, NHMUK, B.M. 1978-49), 4 males, 2 females; Mulu National Park, mixed dipterocarp forest litter, TPS 1-2,
|
||
<normalizedToken originalValue="iv–viii">iv-viii</normalizedToken>
|
||
.1978 (leg. P.M. Hammond & J.E. Marshall, NHMUK, B.M. 1978-49), 3 males, 4 females; Mulu National Park, slope, baited traps, 7.iv.1978 (leg. P.M. Hammond & J.E. Marshall, NHMUK, B.M. 1978-49), 1 male, 1 female; Mulu National Park, mixed dipterocarp forest litter, TPS 3-4,
|
||
<normalizedToken originalValue="v–viii">v-viii</normalizedToken>
|
||
.1978 (leg. P.M. Hammond & J.E. Marshall, NHMUK, B.M. 1978-49), 2 females; Mulu National Park, mixed dipterocarp forest litter, TPS 7-10,
|
||
<normalizedToken originalValue="v–viii">v-viii</normalizedToken>
|
||
.1978 (leg. P.M. Hammond & J.E. Marshall, NHMUK, B.M. 1978-49), 1 female; Mulu National Park, alluvial forest 100 m,
|
||
<normalizedToken originalValue="iii–v">iii-v</normalizedToken>
|
||
.1978 (leg. I. Hanski, NHMUK, B.M. 1978-524), 5 females; Mulu National Park, Limestone 6.4, Tp 113, 400 m, 6.iv.1978 (leg. P.M. Hammond & J.E. Marshall, NHMUK, B.M. 1978-49), 1 female; Mulu National Park, mixed dipterocarp forest litter Tp 7, 21.iv.1978 (leg. P.M. Hammond & J.E. Marshall, NHMUK, B.M. 1978-49), 1 male. Other examined material (not included in the type series): Sarawak. Mulu National Park, slope, baited traps, 7.iv.1978 (leg. P.M. Hammond & J.E. Marshall, B.M. 1978-49, NHMUK), 2 females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="73" type="description">
|
||
<paragraph pageId="16" pageNumber="73">Description.</paragraph>
|
||
<paragraph pageId="16" pageNumber="73">
|
||
Length 2.0-2.8 mm. Habitus: Light to dark reddish brown; flattened and relatively short and broad, head broad, pronotum 1.6-1.7 times as wide as long, narrower than the elytra, caudal angles almost not extended. Elytra short, gently convex, jointly ca. 1.2 times as long as wide (length measured from the caudal tip of the scutellum to the apex of the elytra). Body entirely covered in dense golden-yellow setation. Wings present. Antennae slender, 4th, 9th, and 10th antennomeres almost as long as wide. Male protarsi slightly dilated, the first four tarsomeres jointly ca. 3.5 times as long as wide. Aedeagus gently bent ventrad, flattened, apically broad and convex but subapically tapering in a rounded fashion into a broad but sharp upturned tip. Stylet long and thin, hair-like; stored in a wide loop in the basal part of the aedeagus. Spiculum gastrale elongate-ovoid, with the caudal part button-shaped, truncated. Spermatheca semicircular, thick, otherwise featureless,
|
||
<normalizedToken originalValue="“sausage-shaped”">"sausage-shaped"</normalizedToken>
|
||
. Spermiduct long, thin, consisting of ca. 4-6 360° coils.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="73" type="differential diagnosis">
|
||
<paragraph pageId="16" pageNumber="73">Differential diagnosis.</paragraph>
|
||
<paragraph pageId="16" pageNumber="73">
|
||
Aedeagus in dorsal view very similar to that of
|
||
<taxonomicName lsidName="P. tarsalis" pageId="16" pageNumber="73" rank="species" species="tarsalis">P. tarsalis</taxonomicName>
|
||
Symczakowski, 1964 from Sumatra, but in lateral view apically clearly more convex and with a shorter stylet. Moreover, the habitus and appendages of
|
||
<taxonomicName lsidName="P. tarsalis" pageId="16" pageNumber="73" rank="species" species="tarsalis">P. tarsalis</taxonomicName>
|
||
are very stout and thick, whereas those in
|
||
<taxonomicName lsidName="P. louis" pageId="16" pageNumber="73" rank="species" species="louis">P. louis</taxonomicName>
|
||
are much more slender. Also very similar to
|
||
<taxonomicName lsidName="P. muluensis" pageId="16" pageNumber="73" rank="species" species="muluensis">P. muluensis</taxonomicName>
|
||
, but externally distinguished by the smaller size and more stocky habitus, with shorter elytra, narrower pronotum and the absence of drawn-out caudal pronotal angles. Aedeagus in dorsal view tapering abruptly towards the apex, not as gradually as in
|
||
<taxonomicName lsidName="P. muluensis" pageId="16" pageNumber="73" rank="species" species="muluensis">P. muluensis</taxonomicName>
|
||
; in lateral view, the apex is more convex. Spermatheca distinguished from
|
||
<taxonomicName lsidName="P. muluensis" pageId="16" pageNumber="73" rank="species" species="muluensis">P. muluensis</taxonomicName>
|
||
by the semicircular shape without any distinctive rings.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="73" type="habitat">
|
||
<paragraph pageId="16" pageNumber="73">Habitat and distribution.</paragraph>
|
||
<paragraph pageId="16" pageNumber="73">Only known from Mulu National Park, Sarawak.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="73" type="remarks">
|
||
<paragraph pageId="16" pageNumber="73">Remarks.</paragraph>
|
||
<paragraph pageId="16" pageNumber="73">
|
||
Two females from the Mulu locality
|
||
<normalizedToken originalValue="“Slope”">"Slope"</normalizedToken>
|
||
have several rings at the basis of the spermatheca. As they are externally identical to other females of this species, they have been provisionally included in this species, but excluded from the type series. Several specimens infected on the elytra and pygidium with black
|
||
<taxonomicName lsidName="Staphylinoidea" order="Laboulbeniales" pageId="16" pageNumber="73" rank="order">Laboulbeniales</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="73" type="etymology">
|
||
<paragraph pageId="16" pageNumber="73">Etymology.</paragraph>
|
||
<paragraph pageId="16" pageNumber="73">We name this species after our friend and colleague Dr. Louis Deharveng (MNHN), in recognition for his logistic and emotional support during the preparation of this paper. The specific epithet is given as a noun in apposition.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |