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<document id="C4CA6321ECE088148AF62A728E1A018D" ID-DOI="10.11646/zootaxa.3835.4.9" ID-GBIF-Dataset="040edf15-f51a-4b2d-a301-06f40e51909c" ID-ISSN="1175-5326" ID-Zenodo-Dep="225660" ID-ZooBank="A8B64E90-4FA4-41E9-9169-415BB4758AB7" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460457997776" checkinUser="plazi" docAuthor="Thomas, Donald B., Smith, Aaron D., Triplehorn, Charles A. &amp; Aalbu, Rolf A." docDate="2014" docId="9F4353615F7EFF83FF38FAB1FD36FE38" docLanguage="en" docName="zt03835p592.pdf" docOrigin="Zootaxa 3835 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eleodes binotatus Walker" docType="treatment" docVersion="8" lastPageNumber="587" masterDocId="637A2B195F7AFF86FFAFFFEDFFABFFB0" masterDocTitle="Walkers Eleodes (Coleoptera: Tenebrionidae)" masterLastPageNumber="592" masterPageNumber="583" pageNumber="587" updateTime="1698302759344" updateUser="plazi">
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<mods:title id="1E7054EEB6B69C5175A04C79EB644079">Walkers Eleodes (Coleoptera: Tenebrionidae)</mods:title>
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<mods:namePart id="7B75C40D828F24BACECE565D798E487B">Smith, Aaron D.</mods:namePart>
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<mods:namePart id="5FB126EF3ADE38E26733345739545CF3">Triplehorn, Charles A.</mods:namePart>
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<mods:namePart id="D3E20FFA7C835D53E87D7706C7B2BDAC">Aalbu, Rolf A.</mods:namePart>
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<subSubSection id="5FF0B1FC5F7EFF82FF38FAB1FD99FAC7" box="[151,562,1372,1399]" pageId="4" pageNumber="587" type="nomenclature">
<paragraph id="1755E2775F7EFF82FF38FAB1FD99FAC7" blockId="4.[151,562,1372,1399]" box="[151,562,1372,1399]" pageId="4" pageNumber="587">
<heading id="4C1D551B5F7EFF82FF38FAB1FD99FAC7" bold="true" box="[151,562,1372,1399]" fontSize="11" level="1" pageId="4" pageNumber="587" reason="1">
<emphasis id="259E3E655F7EFF82FF38FAB1FD99FAC7" bold="true" box="[151,562,1372,1399]" pageId="4" pageNumber="587">
<taxonomicName id="D0EA99F45F7EFF82FF38FAB1FE7AFAC7" authority="Walker" authorityName="Walker" box="[151,465,1372,1399]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="587" phylum="Arthropoda" rank="species" species="binotatus">
<emphasis id="259E3E655F7EFF82FF38FAB1FEC6FAC6" bold="true" box="[151,365,1372,1398]" italics="true" pageId="4" pageNumber="587">Eleodes binotatus</emphasis>
Walker
</taxonomicName>
(
<figureCitation id="8FD1FEF25F7EFF82FE4FFAB0FD82FAC7" box="[480,553,1373,1399]" captionStart="FIGURE 4" captionStartId="5.[151,244,1533,1555]" captionText="FIGURE 4. Eleodes binotatus Walker, holotype (a) habitus (b) labels." pageId="4" pageNumber="587">Fig. 4</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5FF0B1FC5F7EFF82FF38FA4EFDC3FA09" box="[151,616,1443,1465]" pageId="4" pageNumber="587" type="reference_group">
<paragraph id="1755E2775F7EFF82FF38FA4EFDC3FA09" blockId="4.[151,616,1443,1465]" box="[151,616,1443,1465]" pageId="4" pageNumber="587">
<taxonomicName id="D0EA99F45F7EFF82FF38FA4EFE7AFA09" authority="Walker, 1866" authorityName="Walker" authorityYear="1866" box="[151,465,1443,1465]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="587" phylum="Arthropoda" rank="species" species="binotata">
<emphasis id="259E3E655F7EFF82FF38FA4EFEE9FA09" box="[151,322,1443,1465]" italics="true" pageId="4" pageNumber="587">Eleodes binotata</emphasis>
<bibRefCitation id="737B9F865F7EFF82FEE7FA4EFE7AFA09" author="Walker" box="[328,465,1443,1465]" pageId="4" pageNumber="587" refString="Walker, F. (1866) List of Coleoptera. In: Lord, J. K. (Ed.), The Naturalist in Vancouver Island and British Columbia. Vol. 2. R. Bentley, London, pp. 309 - 334." type="book" year="1866">Walker, 1866</bibRefCitation>
</taxonomicName>
, vol. 2, p. 329.
</paragraph>
</subSubSection>
<subSubSection id="5FF0B1FC5F7EFF82FF38FA0BFEECF9DB" pageId="4" pageNumber="587" type="description">
<paragraph id="1755E2775F7EFF82FF38FA0BFEECF9DB" blockId="4.[151,1436,1510,2003]" pageId="4" pageNumber="587">
<emphasis id="259E3E655F7EFF82FF38FA0BFE3BFA4F" bold="true" box="[151,400,1510,1535]" pageId="4" pageNumber="587">Original Description</emphasis>
: “Black, slightly shining. Head and thorax extremely minutely punctured. Thorax with a slight impression on each side of the middle of the disk; sides convex before the middle; fore angles prominent, acute. Elytra subfusiform, with slight punctured striae, nearly four times the length of the thorax. Length of the body, 10 lines.”
</paragraph>
</subSubSection>
<subSubSection id="5FF0B1FC5F7EFF83FF68F99BFD36FE38" lastPageId="5" lastPageNumber="588" pageId="4" pageNumber="587" type="materials_examined">
<paragraph id="1755E2775F7EFF82FF68F99BFD1FF93F" blockId="4.[151,1436,1510,2003]" box="[199,692,1654,1679]" pageId="4" pageNumber="587">
<emphasis id="259E3E655F7EFF82FF68F99BFED3F93F" bold="true" box="[199,376,1654,1679]" pageId="4" pageNumber="587">
<typeStatus id="C8515CD55F7EFF82FF68F99BFEA9F93F" box="[199,258,1654,1679]" pageId="4" pageNumber="587">Type</typeStatus>
Material.
</emphasis>
<typeStatus id="C8515CD55F7EFF82FED1F99AFDB9F93F" box="[382,530,1655,1679]" pageId="4" pageNumber="587" type="holotype">HOLOTYPE</typeStatus>
by monotypy.
</paragraph>
<paragraph id="1755E2775F7EFF82FF68F977FEC8F967" blockId="4.[151,1436,1510,2003]" pageId="4" pageNumber="587">
<emphasis id="259E3E655F7EFF82FF68F977FEBDF903" bold="true" box="[199,278,1690,1715]" pageId="4" pageNumber="587">Labels</emphasis>
: (a) [circular, margined in red] “
<typeStatus id="C8515CD55F7EFF82FD24F976FD6EF903" box="[651,709,1691,1715]" pageId="4" pageNumber="587">Type</typeStatus>
H.T.” (b) [circular, blue, hand-script] “Brit Colum,” (c) [handscript] “
<taxonomicName id="D0EA99F45F7EFF82FF5DF953FEE4F966" box="[242,335,1726,1750]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="587" phylum="Arthropoda" rank="species" species="binotata">
<emphasis id="259E3E655F7EFF82FF5DF953FEE4F966" box="[242,335,1726,1750]" italics="true" pageId="4" pageNumber="587">binotata</emphasis>
</taxonomicName>
.”
</paragraph>
<paragraph id="1755E2775F7EFF83FF68F90FFD36FE38" blockId="4.[151,1436,1510,2003]" lastBlockId="5.[151,1436,151,392]" lastPageId="5" lastPageNumber="588" pageId="4" pageNumber="587">
<bibRefCitation id="737B9F865F7EFF82FF68F90FFEC1F94B" author="Gebien" box="[199,362,1762,1787]" pageId="4" pageNumber="587" refString="Gebien, H. (1910) Coleopterorum Catalogus, Pt. 22, Tenebrionidae II. W. Junk, Berlin, pp. 167 - 354." type="book chapter" year="1910">Gebien (1910)</bibRefCitation>
listed Walkers name as a synonym of
<taxonomicName id="D0EA99F45F7EFF82FC84F90FFB07F94B" authority="LeConte (1858)" authorityName="LeConte" authorityYear="1858" box="[811,1196,1762,1787]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="587" phylum="Arthropoda" rank="species" species="sponsus">
<emphasis id="259E3E655F7EFF82FC84F90FFC4EF94A" box="[811,997,1762,1786]" italics="true" pageId="4" pageNumber="587">Eleodes sponsus</emphasis>
<bibRefCitation id="737B9F865F7EFF82FC43F90EFB07F94B" author="LeConte" box="[1004,1196,1762,1787]" pageId="4" pageNumber="587" refString="LeConte, J. L. (1858) Note on the species of Eleodes found within the United States. Proceedings of the Academy of Natural Sciences, Philadelphia, 10, 180 - 188." type="journal article" year="1858">LeConte (1858)</bibRefCitation>
</taxonomicName>
. But
<taxonomicName id="D0EA99F45F7EFF82FB48F90EFACAF94A" box="[1255,1377,1763,1786]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="587" phylum="Arthropoda" rank="species" species="sponsus">
<emphasis id="259E3E655F7EFF82FB48F90EFB53F94A" box="[1255,1272,1763,1786]" italics="true" pageId="4" pageNumber="587">E</emphasis>
.
<emphasis id="259E3E655F7EFF82FAA7F90EFACAF94A" box="[1288,1377,1763,1786]" italics="true" pageId="4" pageNumber="587">sponsus</emphasis>
</taxonomicName>
does not occur in or near British
<collectingCountry id="6FFDA2E75F7EFF82FE60F8EBFDEBF8AF" box="[463,576,1798,1823]" name="United States of America" pageId="4" pageNumber="587">Columbia</collectingCountry>
, being native to the four-corners area of New
<collectingCountry id="6FFDA2E75F7EFF82FBE8F8EBFB37F8AF" box="[1095,1180,1798,1823]" name="Mexico" pageId="4" pageNumber="587">Mexico</collectingCountry>
(the
<typeStatus id="C8515CD55F7EFF82FB78F8EAFAACF8AF" box="[1239,1287,1799,1823]" pageId="4" pageNumber="587">type</typeStatus>
locality) and adjacent parts of Colorado, Utah and Arizona. On examination we find that Walkers beetle is actually conspecific with
<taxonomicName id="D0EA99F45F7EFF82FF7BF8A3FD94F8D7" authority="Say 1824" authorityName="Say" authorityYear="1824" box="[212,575,1870,1895]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="587" phylum="Arthropoda" rank="species" species="hispilabris">
<emphasis id="259E3E655F7EFF82FF7BF8A3FE1BF8D6" box="[212,432,1870,1894]" italics="true" pageId="4" pageNumber="587">Eleodes hispilabris</emphasis>
(Say 1824)
</taxonomicName>
.
<taxonomicName id="D0EA99F45F7EFF82FDE0F8A3FC80F8D6" box="[591,811,1870,1894]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="587" phylum="Arthropoda" rank="species" species="hispilabris">
<emphasis id="259E3E655F7EFF82FDE0F8A3FC80F8D6" box="[591,811,1870,1894]" italics="true" pageId="4" pageNumber="587">Eleodes hispilabris</emphasis>
</taxonomicName>
is a widespread and variable species occurring from southern
<collectingCountry id="6FFDA2E75F7EFF82FEACF89FFEFCF83B" box="[259,343,1906,1931]" name="Canada" pageId="4" pageNumber="587">Canada</collectingCountry>
to northern
<collectingCountry id="6FFDA2E75F7EFF82FE42F89FFDEDF83B" box="[493,582,1906,1931]" name="Mexico" pageId="4" pageNumber="587">Mexico</collectingCountry>
.
<bibRefCitation id="737B9F865F7EFF82FDFAF89FFD5EF83B" author="Boddy" box="[597,757,1906,1931]" pageId="4" pageNumber="587" refString="Boddy, D. W. (1965) Family Tenebrionidae. In, Hatch, M. H. [Ed.], The Beetles of the Pacific Northwest, Pt. IV. Macrodactyles, Palpicornes, and Heteromera. University of Washington Publications in Entomology, Vol. 16. University of Washington Press, Seattle, WA, pp. 130 - 189." type="book" year="1965">Boddy (1965)</bibRefCitation>
astutely recognized Walkers species as a synonym of
<taxonomicName id="D0EA99F45F7EFF82FA2BF89EFE4CF81F" authority="LeConte" authorityName="LeConte" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="587" phylum="Arthropoda" rank="subSpecies" species="hispilabris" subSpecies="connexa">
<emphasis id="259E3E655F7EFF82FA2BF89EFA3EF83A" box="[1412,1429,1907,1930]" italics="true" pageId="4" pageNumber="587">E</emphasis>
.
<emphasis id="259E3E655F7EFF82FF38F87BFED3F81E" box="[151,376,1942,1966]" italics="true" pageId="4" pageNumber="587">hispilabris connexa</emphasis>
LeConte
</taxonomicName>
, a variant found in New
<collectingCountry id="6FFDA2E75F7EFF82FCB6F87BFCC5F81F" box="[793,878,1942,1967]" name="Mexico" pageId="4" pageNumber="587">Mexico</collectingCountry>
and west Texas. The population in the Pacific northwest was characterized as “forma
<emphasis id="259E3E655F7EFF82FDCAF857FD0DF862" box="[613,678,1978,2002]" italics="true" pageId="4" pageNumber="587">laevis</emphasis>
” by
<bibRefCitation id="737B9F865F7EFF82FD4AF857FC0AF863" author="Blaisdell" box="[741,929,1978,2003]" pageId="4" pageNumber="587" refString="Blaisdell, F. E. (1909) A monographic revision of the Coleoptera belonging to the tenebrionid tribe Eleodiini inhabiting the United States, Lower California, and adjacent islands. Smithsonian Institution, United States National Museum Bulletin, 63, 1 - 524. http: // dx. doi. org / 10.5479 / si. 03629236.63.1" type="journal article" year="1909">Blaisdell (1909)</bibRefCitation>
, later formally described by him (
<bibRefCitation id="737B9F865F7EFF83FA98F857FF7BFF00" author="Blaisdell" lastPageId="5" lastPageNumber="588" pageId="4" pageNumber="587" refString="Blaisdell, F. E. (1918) Studies in the Tenebrionid tribe Eleodiini No. 3 (Coleop). Entomological News, 29, 162 - 169." type="journal article" year="1918">Blaisdell 1918</bibRefCitation>
) as “variety
<emphasis id="259E3E655F7FFF83FEF2FF75FE6CFF00" box="[349,455,152,176]" italics="true" pageId="5" pageNumber="588">imitabilis</emphasis>
,” which he later (
<bibRefCitation id="737B9F865F7FFF83FD21FF7AFC98FF00" author="Blaisdell" box="[654,819,151,176]" pageId="5" pageNumber="588" refString="Blaisdell, F. E. (1925) Revised checklist of the species of Eleodes inhabiting America, north of Mexico, including Lower California and adjacent islands. Pan-Pacific Entomologist, 2, 77 - 80." type="journal article" year="1925">Blaisdell 1925</bibRefCitation>
) raised to subspecific status. They have the integument dull, alutaceous, smooth; sulci very shallow, intervals feebly convex. This is the race found in eastern Washington and Oregon, eastern Idaho, northwestern Nevada, and northeastern California.
<taxonomicName id="D0EA99F45F7FFF83FB8CFF0DFA37FF48" authority="Walker 1866" authorityName="Walker" authorityYear="1866" box="[1059,1436,223,248]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="588" phylum="Arthropoda" rank="species" species="binotatus">
<emphasis id="259E3E655F7FFF83FB8CFF0DFB46FF48" box="[1059,1261,224,248]" italics="true" pageId="5" pageNumber="588">Eleodes binotatus</emphasis>
(
<bibRefCitation id="737B9F865F7FFF83FB50FF32FA3FFF48" author="Walker" box="[1279,1428,223,248]" pageId="5" pageNumber="588" refString="Walker, F. (1866) List of Coleoptera. In: Lord, J. K. (Ed.), The Naturalist in Vancouver Island and British Columbia. Vol. 2. R. Bentley, London, pp. 309 - 334." type="book" year="1866">Walker 1866</bibRefCitation>
)
</taxonomicName>
from British
<collectingCountry id="6FFDA2E75F7FFF83FE82FEE9FE30FEAD" box="[301,411,260,285]" name="United States of America" pageId="5" pageNumber="588">Columbia</collectingCountry>
belongs here as well and is thus senior as a synonym of the subspecies name
<emphasis id="259E3E655F7FFF83FA9DFEE9FA37FEAC" box="[1330,1436,260,284]" italics="true" pageId="5" pageNumber="588">imitabilis</emphasis>
<bibRefCitation id="737B9F865F7FFF83FF38FECAFEF8FEF0" author="Blaisdell" box="[151,339,295,320]" pageId="5" pageNumber="588" refString="Blaisdell, F. E. (1918) Studies in the Tenebrionid tribe Eleodiini No. 3 (Coleop). Entomological News, 29, 162 - 169." type="journal article" year="1918">Blaisdell (1918)</bibRefCitation>
. Our concept of
<taxonomicName id="D0EA99F45F7FFF83FDB7FEC4FD19FEF0" box="[536,690,296,320]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="588" phylum="Arthropoda" rank="species" species="hispilabris">
<emphasis id="259E3E655F7FFF83FDB7FEC4FD19FEF0" box="[536,690,296,320]" italics="true" pageId="5" pageNumber="588">E. hispilabris</emphasis>
</taxonomicName>
and its variants is treated in a revision of the subgenus
<taxonomicName id="D0EA99F45F7FFF83FAECFEC5FA37FEF0" box="[1347,1436,296,320]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="588" phylum="Arthropoda" rank="genus">
<emphasis id="259E3E655F7FFF83FAECFEC5FA37FEF0" box="[1347,1436,296,320]" italics="true" pageId="5" pageNumber="588">Eleodes</emphasis>
</taxonomicName>
currently in preparation. Although we do not consider the forms of
<taxonomicName id="D0EA99F45F7FFF83FC05FEA0FBEFFED4" box="[938,1092,332,356]" class="Insecta" family="Tenebrionidae" genus="Eleodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="588" phylum="Arthropoda" rank="species" species="hispilabris">
<emphasis id="259E3E655F7FFF83FC05FEA0FBEFFED4" box="[938,1092,332,356]" italics="true" pageId="5" pageNumber="588">E. hispilabris</emphasis>
</taxonomicName>
to warrant subspecific status, future revisers should be aware of the priority.
</paragraph>
</subSubSection>
</treatment>
</document>