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<document id="5789C84DC56DE77AFA5F5E542DD2CC45" ID-DOI="10.1649/685" ID-ISSN="1938-4394" ID-Zenodo-Dep="10104682" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630169721737" checkinUser="tatiana" docAuthor="Ocampo, Federico C." docDate="2004" docId="9F2D3E5A6A29FFF7FED5FA647D70BF4E" docLanguage="en" docName="685.pdf.imf" docOrigin="The Coleopterists Bulletin 58 (2)" docSource="http://dx.doi.org/10.1649/685" docStyle="DocumentStyle:07711681861DB98305E87147CAB6DAD5.4:TheColeopBull.2003-2009.journal_article.0cover" docStyleId="07711681861DB98305E87147CAB6DAD5" docStyleName="TheColeopBull.2003-2009.journal_article.0cover" docStyleVersion="4" docTitle="Glyphoderus sterquilinus" docType="treatment" docVersion="4" lastPageNumber="304" masterDocId="631446226A21FFFEFFA4FFD47F21BD05" masterDocTitle="Food Relocation Behavior and Synopsis of the Southern South American Genus Glyphoderus Westwood (Scarabaeidae: Scarabaeinae: Eucraniini)" masterLastPageNumber="305" masterPageNumber="295" pageNumber="303" updateTime="1699630397919" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D69C5D868E50FAA79CCFD19FFE32B0E0">Food Relocation Behavior and Synopsis of the Southern South American Genus Glyphoderus Westwood (Scarabaeidae: Scarabaeinae: Eucraniini)</mods:title>
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<mods:namePart id="BCDA9D29E1E7CBC5066A4C81BA6A0315">Ocampo, Federico C.</mods:namePart>
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<mods:title id="CAEA83FB2D8BC5069C6618507FE58B31">The Coleopterists Bulletin</mods:title>
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<mods:date id="BF84D93C582AC285E861A7452A4966FC">2004</mods:date>
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<mods:identifier id="279C0D8FBEB96917E3E9E846B4952DC8" type="ISSN">1938-4394</mods:identifier>
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<treatment id="9F2D3E5A6A29FFF7FED5FA647D70BF4E" ID-DOI="http://doi.org/10.5281/zenodo.5399096" ID-Zenodo-Dep="5399096" LSID="urn:lsid:plazi:treatment:9F2D3E5A6A29FFF7FED5FA647D70BF4E" httpUri="http://treatment.plazi.org/id/9F2D3E5A6A29FFF7FED5FA647D70BF4E" lastPageId="9" lastPageNumber="304" pageId="8" pageNumber="303">
<subSubSection id="5F9EDCC76A29FFF6FED5FA647C0AB8C0" box="[369,811,1456,1477]" pageId="8" pageNumber="303" type="nomenclature">
<paragraph id="173B8F4C6A29FFF6FED5FA647C0AB8C0" blockId="8.[369,811,1456,1504]" box="[369,811,1456,1477]" pageId="8" pageNumber="303">
<heading id="4C7338206A29FFF6FED5FA647C0AB8C0" box="[369,811,1456,1477]" centered="true" fontSize="9" level="2" pageId="8" pageNumber="303" reason="2">
<taxonomicName id="D084F4CF6A29FFF6FED5FA647C0AB8C0" ID-CoL="3GGNV" authority="(Westwood, 1837)" baseAuthorityName="Westwood" baseAuthorityYear="1838" box="[369,811,1456,1477]" class="Insecta" family="Scarabaeidae" genus="Glyphoderus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="303" phylum="Arthropoda" rank="species" species="sterquilinus">
<emphasis id="25F0535E6A29FFF6FED5FA647C0AB8C0" bold="true" box="[369,811,1456,1477]" pageId="8" pageNumber="303">
<emphasis id="25F0535E6A29FFF6FED5FA647D4DB8C0" bold="true" box="[369,620,1456,1477]" italics="true" pageId="8" pageNumber="303">Glyphoderus sterquilinus</emphasis>
(
<bibRefCitation id="7315F2BD6A29FFF6FDDEFA647C05B8C0" author="Westwood" box="[634,804,1456,1477]" pageId="8" pageNumber="303" pagination="12 - 13" refId="ref6276" refString="Westwood, J. O. 1837. Illustrations of exotic entomology, containing upwards of six hundred and fifty figures and descriptions of foreign insects, interspersed with remarks and reflections on their nature and properties. Proceedings of the Zoological Society of London 5: 12 - 13." type="journal article" year="1837">Westwood, 1837</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5F9EDCC76A29FFF6FDBCFA1F7DA5B8E5" box="[536,644,1483,1504]" pageId="8" pageNumber="303" type="description">
<paragraph id="173B8F4C6A29FFF6FDBCFA1F7DA5B8E5" blockId="8.[369,811,1456,1504]" box="[536,644,1483,1504]" pageId="8" pageNumber="303">
(
<figureCitation id="8FBF93C96A29FFF6FDBBFA1F7D42B8E5" box="[543,611,1483,1504]" captionStart="Fig" captionStartId="6.[197,228,744,763]" captionTargetBox="[176,1007,194,730]" captionTargetId="figure-374@6.[175,1007,194,730]" captionTargetPageId="6" captionText="Fig. 5. Dorsal view of the pronotum of Glyphoderus sterquilinus (Westwood)." figureDoi="http://doi.org/10.5281/zenodo.10104693" httpUri="https://zenodo.org/record/10104693/files/figure.png" pageId="8" pageNumber="303">Figs. 5</figureCitation>
,
<figureCitation id="8FBF93C96A29FFF6FDD4FA1F7D5DB8E5" box="[624,636,1483,1504]" captionStart="Fig" captionStartId="7.[169,200,1157,1176]" captionTargetBox="[148,980,195,1143]" captionTargetId="figure-300@7.[148,980,194,1144]" captionTargetPageId="7" captionText="Fig. 6. Map showing distribution of Glyphoderus species." figureDoi="http://doi.org/10.5281/zenodo.10104695" httpUri="https://zenodo.org/record/10104695/files/figure.png" pageId="8" pageNumber="303">6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5F9EDCC76A29FFF6FF0BFA277C63BB27" pageId="8" pageNumber="303" type="reference_group">
<paragraph id="173B8F4C6A29FFF6FF0BFA277DA3BB0D" blockId="8.[175,834,1523,1570]" box="[175,642,1523,1544]" pageId="8" pageNumber="303">
<taxonomicName id="D084F4CF6A29FFF6FF0BFA277D43BB0D" ID-CoL="3GGNV" authority="(Westwood, 1837)" baseAuthorityName="Westwood" baseAuthorityYear="1838" box="[175,610,1523,1544]" class="Insecta" family="Scarabaeidae" genus="Glyphoderus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="303" phylum="Arthropoda" rank="species" species="sterquilinus">
<emphasis id="25F0535E6A29FFF6FF0BFA277E83BB0D" box="[175,418,1523,1544]" italics="true" pageId="8" pageNumber="303">Glyphoderus sterquilinus</emphasis>
(
<bibRefCitation id="7315F2BD6A29FFF6FE14FA277D7ABB0D" author="Westwood" box="[432,603,1523,1544]" pageId="8" pageNumber="303" pagination="12 - 13" refId="ref6276" refString="Westwood, J. O. 1837. Illustrations of exotic entomology, containing upwards of six hundred and fifty figures and descriptions of foreign insects, interspersed with remarks and reflections on their nature and properties. Proceedings of the Zoological Society of London 5: 12 - 13." type="journal article" year="1837">Westwood, 1837</bibRefCitation>
)
</taxonomicName>
:13.
</paragraph>
<paragraph id="173B8F4C6A29FFF6FF0BF9D97C63BB27" blockId="8.[175,834,1523,1570]" box="[175,834,1549,1570]" pageId="8" pageNumber="303">
<treatmentCitationGroup id="3794A8626A29FFF6FF0BF9D97C63BB27" box="[175,834,1549,1570]" pageId="8" pageNumber="303">
<treatmentCitation id="9625A95D6A29FFF6FF0BF9D97D43BB27" author="Westwood" box="[175,610,1549,1570]" page="13" pageId="8" pageNumber="303" year="1837">
<taxonomicName id="D084F4CF6A29FFF6FF0BF9D97D43BB27" ID-CoL="5V6LJ" authority="Westwood, 1837: 13" authorityName="Westwood" authorityPageNumber="13" authorityYear="1837" box="[175,610,1549,1570]" class="Insecta" family="Scarabaeidae" genus="Anomiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="303" phylum="Arthropoda" rank="species" species="sterquilinus">
<emphasis id="25F0535E6A29FFF6FF0BF9D97EB7BB27" box="[175,406,1549,1570]" italics="true" pageId="8" pageNumber="303">Anomiopsis sterquilinus</emphasis>
<bibRefCitation id="7315F2BD6A29FFF6FE3AF9D97D43BB27" author="Westwood" box="[414,610,1549,1570]" pageId="8" pageNumber="303" pagination="12 - 13" refId="ref6276" refString="Westwood, J. O. 1837. Illustrations of exotic entomology, containing upwards of six hundred and fifty figures and descriptions of foreign insects, interspersed with remarks and reflections on their nature and properties. Proceedings of the Zoological Society of London 5: 12 - 13." type="journal article" year="1837">Westwood, 1837:13</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Original combination.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="5F9EDCC76A28FFF7FF08FF167C88BCC3" pageId="9" pageNumber="304" type="diagnosis">
<paragraph id="173B8F4C6A28FFF7FF08FF167C88BCC3" blockId="9.[148,977,194,587]" pageId="9" pageNumber="304">
<emphasis id="25F0535E6A28FFF7FF08FF167E35BDD2" bold="true" box="[172,276,194,215]" pageId="9" pageNumber="304">Diagnosis.</emphasis>
Males. This species is separated from other
<taxonomicName id="D084F4CF6A28FFF7FD63FF167CACBDD2" box="[711,909,194,215]" class="Insecta" family="Scarabaeidae" genus="Glyphoderus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="304" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="25F0535E6A28FFF7FD63FF167C63BDD2" box="[711,834,194,215]" italics="true" pageId="9" pageNumber="304">Glyphoderus</emphasis>
species
</taxonomicName>
by the shape and placement of the pronotal horns and tubercles (
<figureCitation id="8FBF93C96A28FFF7FD5FFF097C1DBDF7" box="[763,828,221,242]" captionStart="Fig" captionStartId="6.[197,228,744,763]" captionTargetBox="[176,1007,194,730]" captionTargetId="figure-374@6.[175,1007,194,730]" captionTargetPageId="6" captionText="Fig. 5. Dorsal view of the pronotum of Glyphoderus sterquilinus (Westwood)." figureDoi="http://doi.org/10.5281/zenodo.10104693" httpUri="https://zenodo.org/record/10104693/files/figure.png" pageId="9" pageNumber="304">Fig. 5</figureCitation>
).
<taxonomicName id="D084F4CF6A28FFF7FCF2FF097E25BC08" baseAuthorityName="Westwood" baseAuthorityYear="1838" class="Insecta" family="Scarabaeidae" genus="Glyphoderus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="304" phylum="Arthropoda" rank="species" species="sterquilinus">
<emphasis id="25F0535E6A28FFF7FCF2FF097E25BC08" italics="true" pageId="9" pageNumber="304">Glyphoderus sterquilinus</emphasis>
</taxonomicName>
has the pronotum with two horns in middle and two transversally elongated tubercles (sometimes developed as horns) on basal half; horns on margin of central depression; tubercles of basal half included in central depression; apical medial horn with arms robust, with anteromedial notch. Some specimens of
<taxonomicName id="D084F4CF6A28FFF7FCB4FE937CBFBC59" baseAuthorityName="Westwood" baseAuthorityYear="1838" box="[784,926,327,348]" class="Insecta" family="Scarabaeidae" genus="Glyphoderus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="304" phylum="Arthropoda" rank="species" species="sterquilinus">
<emphasis id="25F0535E6A28FFF7FCB4FE937CBFBC59" box="[784,926,327,348]" italics="true" pageId="9" pageNumber="304">G. sterquilinus</emphasis>
</taxonomicName>
have the posterior pronotal tubercles on the central depression margin (observed particularly in minor males), in this case, the shape of the medial horn will separate this species from
<taxonomicName id="D084F4CF6A28FFF7FF6EFE437E69BCA9" box="[202,328,407,428]" class="Insecta" family="Scarabaeidae" genus="Glyphoderus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="304" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="25F0535E6A28FFF7FF6EFE437E69BCA9" box="[202,328,407,428]" italics="true" pageId="9" pageNumber="304">G. monticola</emphasis>
</taxonomicName>
(
<figureCitation id="8FBF93C96A28FFF7FEFCFE437EBCBCA9" box="[344,413,407,428]" captionStart="Fig" captionStartId="5.[169,200,1175,1194]" captionTargetBox="[151,980,199,1162]" captionTargetId="figure-166@5.[148,980,194,1162]" captionTargetPageId="5" captionText="Fig. 4. Dorsal habitus of Glyphoderus monticola Burmeister." figureDoi="http://doi.org/10.5281/zenodo.10104691" httpUri="https://zenodo.org/record/10104691/files/figure.png" pageId="9" pageNumber="304">Figs. 4</figureCitation>
,
<figureCitation id="8FBF93C96A28FFF7FE0EFE437E96BCA9" box="[426,439,407,428]" captionStart="Fig" captionStartId="6.[197,228,744,763]" captionTargetBox="[176,1007,194,730]" captionTargetId="figure-374@6.[175,1007,194,730]" captionTargetPageId="6" captionText="Fig. 5. Dorsal view of the pronotum of Glyphoderus sterquilinus (Westwood)." figureDoi="http://doi.org/10.5281/zenodo.10104693" httpUri="https://zenodo.org/record/10104693/files/figure.png" pageId="9" pageNumber="304">5</figureCitation>
). No morphological characters were found to diagnose females of this species, females can be identified only by association with males.
</paragraph>
</subSubSection>
<subSubSection id="5F9EDCC76A28FFF7FF08FE187D70BF4E" pageId="9" pageNumber="304" type="distribution">
<paragraph id="173B8F4C6A28FFF7FF08FE187CF1BF13" blockId="9.[148,977,194,587]" pageId="9" pageNumber="304">
<emphasis id="25F0535E6A28FFF7FF08FE187E08BCE4" bold="true" box="[172,297,460,481]" pageId="9" pageNumber="304">Distribution</emphasis>
(
<figureCitation id="8FBF93C96A28FFF7FE93FE187E52BCE4" box="[311,371,460,481]" captionStart="Fig" captionStartId="7.[169,200,1157,1176]" captionTargetBox="[148,980,195,1143]" captionTargetId="figure-300@7.[148,980,194,1144]" captionTargetPageId="7" captionText="Fig. 6. Map showing distribution of Glyphoderus species." figureDoi="http://doi.org/10.5281/zenodo.10104695" httpUri="https://zenodo.org/record/10104695/files/figure.png" pageId="9" pageNumber="304">Fig. 6</figureCitation>
).
<emphasis id="25F0535E6A28FFF7FE2FFE187DB2BCE4" bold="true" box="[395,659,460,481]" pageId="9" pageNumber="304">
<collectingCountry id="6F93CFDC6A28FFF7FE2FFE187D01BCE4" box="[395,544,460,481]" name="Argentina" pageId="9" pageNumber="304">ARGENTINA</collectingCountry>
:
<collectingRegion id="D54041AE6A28FFF7FD88FE187DAFBCE4" box="[556,654,460,481]" country="Argentina" name="Mendoza" pageId="9" pageNumber="304">Mendoza</collectingRegion>
:
</emphasis>
Las Catitias (
<quantity id="D07C22A96A28FFF7FCBFFE187C6FBCE4" box="[795,846,460,481]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="304" unit="km" value="7.0">7 km</quantity>
S) (6); Santa Rosa (1); Villavicencio (1).
<emphasis id="25F0535E6A28FFF7FE06FE327D24BCFE" bold="true" box="[418,517,486,507]" pageId="9" pageNumber="304">
<collectingRegion id="D54041AE6A28FFF7FE06FE327D21BCFE" box="[418,512,486,507]" country="Argentina" name="La Rioja" pageId="9" pageNumber="304">La Rioja</collectingRegion>
:
</emphasis>
Chepes (2); Ulapes (
<quantity id="D07C22A96A28FFF7FD76FE337C31BCF9" box="[722,784,487,508]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="304" unit="km" value="17.0">17 km</quantity>
S) (28); Ulapes (
<quantity id="D07C22A96A28FFF7FC1CFE337F93BF13" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="304" unit="km" value="10.0">10 km</quantity>
N) (8).
<emphasis id="25F0535E6A28FFF7FEA5FDD57E49BF13" bold="true" box="[257,360,513,534]" pageId="9" pageNumber="304">
<collectingRegion id="D54041AE6A28FFF7FEA5FDD57E45BF13" box="[257,356,513,534]" country="Argentina" name="San Juan" pageId="9" pageNumber="304">San Juan</collectingRegion>
:
</emphasis>
Encón (
<quantity id="D07C22A96A28FFF7FE1EFDD57ED9BF13" box="[442,504,513,534]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="304" unit="km" value="12.0">12 km</quantity>
E) (1); Nikizanga (2); Morayes Astica road (1).
</paragraph>
<paragraph id="173B8F4C6A28FFF7FF30FDCF7D70BF4E" blockId="9.[148,977,194,587]" pageId="9" pageNumber="304">
<materialsCitation id="A7EC85116A28FFF7FF30FDCF7F9CBF4E" collectorName="La Chanarienta" location="San Luis" pageId="9" pageNumber="304" specimenCount="1" stateProvince="San Luis">
<emphasis id="25F0535E6A28FFF7FF30FDCF7FD9BF35" bold="true" box="[148,248,539,560]" pageId="9" pageNumber="304">
<collectingRegion id="D54041AE6A28FFF7FF30FDCF7FD5BF35" box="[148,244,539,560]" country="Argentina" name="San Luis" pageId="9" pageNumber="304">San Luis</collectingRegion>
:
</emphasis>
Jarilla (1);
<collectorName id="BA71EA9A6A28FFF7FECAFDC87D23BF34" box="[366,514,540,561]" pageId="9" pageNumber="304">La Chañarienta</collectorName>
(2); San Antonio (5); Talita (8); no more data (11)
</materialsCitation>
.
<emphasis id="25F0535E6A28FFF7FF6DFDE27EBBBF4E" bold="true" box="[201,410,566,587]" pageId="9" pageNumber="304">
<collectingRegion id="D54041AE6A28FFF7FF6DFDE27EB7BF4E" box="[201,406,566,587]" country="Argentina" name="Santiago del Estero" pageId="9" pageNumber="304">Santiago del Estero</collectingRegion>
:
</emphasis>
No more data (6).
</paragraph>
</subSubSection>
</treatment>
</document>