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<document id="865450D88CAD375987C87DC5DFB918B7" ID-DOI="10.1080/00222933.2019.1606953" ID-GBIF-Dataset="d2615e10-8703-400b-b571-6c431b0ed9a7" ID-ISSN="1464-5262" ID-Zenodo-Dep="3675571" ID-ZooBank="09AA5C86-5138-42A7-9646-9FF2A7F46767" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582134181895" checkinUser="valdenar" docAuthor="Frolov, Andrey V., Akhmetova, Lilia A. &amp; Vaz-de-Mello, Fernando Z." docDate="2019" docId="9F138799E55AB0639B7F5BBAD121FEF3" docLanguage="en" docName="JNATHIST.53.1112.725747.pdf.imf" docOrigin="Journal of Natural History 53 (11)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Aegidinus candezei" docType="treatment" docVersion="17" lastPageNumber="737" masterDocId="632AFFE1E556B06D9A1D5E52D024FFC2" masterDocTitle="Contribution to the knowledge of Aegidinus Arrow (Coleoptera: Scarabaeidae: Orphninae): new species and comments on the classification and nomenclature" masterLastPageNumber="747" masterPageNumber="725" pageNumber="736" updateTime="1698719875536" updateUser="plazi">
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<mods:title id="ED24FCD2FA3097C2C54361E6E3C8D71B">Contribution to the knowledge of Aegidinus Arrow (Coleoptera: Scarabaeidae: Orphninae): new species and comments on the classification and nomenclature</mods:title>
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<mods:namePart id="9C76C90979534B7A170801E72357EEC5">Frolov, Andrey V.</mods:namePart>
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<mods:namePart id="7319B1DC6F376877DFB76467825B514A">Akhmetova, Lilia A.</mods:namePart>
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<treatment id="9F138799E55AB0639B7F5BBAD121FEF3" ID-DOI="http://doi.org/10.5281/zenodo.3680498" ID-GBIF-Taxon="162173917" ID-Zenodo-Dep="3680498" LSID="urn:lsid:plazi:treatment:9F138799E55AB0639B7F5BBAD121FEF3" httpUri="http://treatment.plazi.org/id/9F138799E55AB0639B7F5BBAD121FEF3" lastPageId="14" lastPageNumber="737" pageId="12" pageNumber="736">
<subSubSection id="5FA06504E55AB0619B7F5BBAD3E5F9C3" box="[354,961,1512,1537]" pageId="12" pageNumber="736" type="nomenclature">
<paragraph id="1705368FE55AB0619B7F5BBAD3E5F9C3" blockId="12.[354,961,1512,1571]" box="[354,961,1512,1537]" pageId="12" pageNumber="736">
<heading id="4C4D81E3E55AB0619B7F5BBAD3E5F9C3" box="[354,961,1512,1537]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="736" reason="2">
<taxonomicName id="D0BA4D0CE55AB0619B7F5BBAD3E5F9C3" authority="(Preudhomme de Borre, 1886)" baseAuthorityName="Preudhomme de Borre" baseAuthorityYear="1886" box="[354,961,1512,1537]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="736" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="25CEEA9DE55AB0619B7F5BBAD26BF9C3" bold="true" box="[354,591,1512,1537]" italics="true" pageId="12" pageNumber="736">Aegidinus candezei</emphasis>
(Preudhomme de Borre, 1886)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5FA06504E55AB061980A5858D328F9E1" box="[535,780,1546,1571]" pageId="12" pageNumber="736" type="description">
<paragraph id="1705368FE55AB061980A5858D328F9E1" blockId="12.[354,961,1512,1571]" box="[535,780,1546,1571]" pageId="12" pageNumber="736">
<heading id="4C4D81E3E55AB061980A5858D328F9E1" box="[535,780,1546,1571]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="736" reason="5">
(
<figureCitation id="8F812A0AE55AB06198025858D2A4F9E1" box="[543,640,1546,1571]" captionStart="Figure 3" captionStartId="6.[160,225,1495,1518]" captionTargetBox="[167,1148,143,1463]" captionTargetId="figure@6.[167,1149,142,1464]" captionTargetPageId="6" captionText="Figure 3. Aegidinus spp. Aegidinus candezei, lectotype, male (a, e), male (bd), female (h, k), female, holotype of Ae. sunidigea (g, j); Aegidinus sp., paralectotype of Ae. candezei (f, i, l). Habitus in dorsal view (a, g, h, i); labels (e, f); female external genitalia (jl); aedeagus in lateral (b) and ventral view (c); parameres in dorsal view (d)." figureDoi="http://doi.org/10.5281/zenodo.3675577" httpUri="https://zenodo.org/record/3675577/files/figure.png" pageId="12" pageNumber="736">Figure 3</figureCitation>
(ae,g,h,j,k))
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5FA06504E55AB0619ABD586DD258F97D" pageId="12" pageNumber="736" type="reference_group">
<paragraph id="1705368FE55AB0619ABD586DD2C9F995" blockId="12.[160,1155,1599,1727]" box="[160,749,1599,1623]" pageId="12" pageNumber="736">
<taxonomicName id="D0BA4D0CE55AB0619ABD586DD2C9F995" authority="Preudhomme de Borre, 1886: 116" authorityName="Preudhomme de Borre" authorityPageNumber="116" authorityYear="1886" box="[160,749,1599,1623]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="736" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="25CEEA9DE55AB0619ABD586DD141F995" box="[160,357,1599,1623]" italics="true" pageId="12" pageNumber="736">Orphnus candezei</emphasis>
Preudhomme de Borre, 1886: 116
</taxonomicName>
</paragraph>
<paragraph id="1705368FE55AB0619ABD5830D1EEF95F" blockId="12.[160,1155,1599,1727]" pageId="12" pageNumber="736">
<treatmentCitationGroup id="37AA11A1E55AB0619ABD5830D1EEF95F" pageId="12" pageNumber="736">
<taxonomicName id="D0BA4D0CE55AB0619ABD5830D2A9F9B8" authority="(Preudhomme de Borre)" baseAuthorityName="Preudhomme de Borre" baseAuthorityYear="1886" box="[160,653,1634,1658]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="736" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="25CEEA9DE55AB0619ABD5830D155F9B8" box="[160,369,1634,1658]" italics="true" pageId="12" pageNumber="736">Aegidinus candezei</emphasis>
(Preudhomme de Borre)
</taxonomicName>
; Arrow, 1912: 31;
<bibRefCitation id="732B4B7EE55AB06199445830D3DEF9B8" box="[857,1018,1634,1658]" pageId="12" pageNumber="736" refId="ref9652" type="book">Schmidt, 1913</bibRefCitation>
: 76; Paulian 184: 89;
<treatmentCitation id="961B109EE55AB0619B3C58D7D1EEF95F" ID-DOI="http://doi.org/10.5281/zenodo.5450531" ID-Zenodo-Dep="5450531" author="Colby J" box="[289,458,1669,1693]" httpUri="http://treatment.plazi.org/id/03BBFE490B348748FF361BF5B268F842" page="17" pageId="12" pageNumber="736" year="2009">
<bibRefCitation id="732B4B7EE55AB0619B3C58D7D1EEF95F" author="Colby J" box="[289,458,1669,1693]" pageId="12" pageNumber="736" pagination="1 - 41" refId="ref9012" refString="Colby J. 2009. Monographic revision of the genus Aegidinus Arrow (1904) and generic phylogeny of the world Orphninae (Coleoptera: Scarabaeidae: Orphninae). Insecta Mundi. 76: 1 - 41." type="journal article" year="2009">Colby 2009: 17</bibRefCitation>
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</paragraph>
<paragraph id="1705368FE55AB0619ABD58F5D258F97D" blockId="12.[160,1155,1599,1727]" box="[160,636,1703,1727]" pageId="12" pageNumber="736">
<taxonomicName id="D0BA4D0CE55AB0619ABD58F5D22DF97D" authority="Colby, 2009" authorityName="Colby" authorityYear="2009" box="[160,521,1703,1727]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="736" phylum="Arthropoda" rank="species" species="sunidigea">
<emphasis id="25CEEA9DE55AB0619ABD58F5D15EF97D" box="[160,378,1703,1727]" italics="true" pageId="12" pageNumber="736">Aegidinus sunidigea</emphasis>
Colby, 2009
</taxonomicName>
, syn. nov.
</paragraph>
</subSubSection>
<subSubSection id="5FA06504E55BB0609ABD5EDCD40FFD10" pageId="13" pageNumber="737" type="materials_examined">
<paragraph id="1705368FE55BB0609ABD5EDCD232FF65" blockId="13.[160,534,142,167]" box="[160,534,142,167]" pageId="13" pageNumber="737">
<emphasis id="25CEEA9DE55BB0609ABD5EDCD11AFF65" bold="true" box="[160,318,142,167]" italics="true" pageId="13" pageNumber="737">Type locality.</emphasis>
Colombia,
<collectingRegion id="D57EF86DE55BB0609BDA5EDDD235FF65" box="[455,529,143,167]" country="Colombia" name="Choco" pageId="13" pageNumber="737">Chocó</collectingRegion>
.
</paragraph>
<paragraph id="1705368FE55BB0609ABD5E86D1E6FF2F" blockId="13.[160,1156,212,479]" box="[160,450,212,237]" pageId="13" pageNumber="737">
<emphasis id="25CEEA9DE55BB0609ABD5E86D1E6FF2F" bold="true" box="[160,450,212,237]" italics="true" pageId="13" pageNumber="737">Type material examined</emphasis>
</paragraph>
<paragraph id="1705368FE55BB0609ABD5EA5D351FE96" blockId="13.[160,1156,212,479]" pageId="13" pageNumber="737">
<materialsCitation id="A7D23CD2E55BB0609ABD5EA5D351FE96" ID-GBIF-Occurrence="2569344914" collectionCode="IRSNB" collectorName="Coll. R. I. Sc. N. B. &amp; Coll. R. I. Sc." country="Colombia" isTailEnumeration="true" location="Choco" pageId="13" pageNumber="737" specimenCount="1" specimenCount-male="1" stateProvince="Choco" typeStatus="lectotype">
<emphasis id="25CEEA9DE55BB0609ABD5EA5D133FECD" bold="true" box="[160,279,247,271]" pageId="13" pageNumber="737">
<typeStatus id="C801882DE55BB0609ABD5EA5D133FECD" box="[160,279,247,271]" pageId="13" pageNumber="737" type="lectotype">Lectotype</typeStatus>
</emphasis>
(designated here,
<figureCitation id="8F812A0AE55BB0609BE95EA5D270FECD" box="[500,596,247,271]" captionStart="Figure 3" captionStartId="6.[160,225,1495,1518]" captionTargetBox="[167,1148,143,1463]" captionTargetId="figure@6.[167,1149,142,1464]" captionTargetPageId="6" captionText="Figure 3. Aegidinus spp. Aegidinus candezei, lectotype, male (a, e), male (bd), female (h, k), female, holotype of Ae. sunidigea (g, j); Aegidinus sp., paralectotype of Ae. candezei (f, i, l). Habitus in dorsal view (a, g, h, i); labels (e, f); female external genitalia (jl); aedeagus in lateral (b) and ventral view (c); parameres in dorsal view (d)." figureDoi="http://doi.org/10.5281/zenodo.3675577" httpUri="https://zenodo.org/record/3675577/files/figure.png" pageId="13" pageNumber="737">Figure 3</figureCitation>
(a,e)), male at
<collectionCode id="71ABAE4AE55BB06098E05EA5D31BFECD" box="[765,831,247,271]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="13" pageNumber="737">IRSNB</collectionCode>
labelled
<emphasis id="25CEEA9DE55BB06099B05EA5D396FECD" box="[941,946,247,271]" italics="true" pageId="13" pageNumber="737"></emphasis>
<collectorName id="BA4F5359E55BB06099AE5EA5D4A0FECD" box="[947,1156,247,271]" pageId="13" pageNumber="737">Coll. R. I. Sc. N. B.</collectorName>
<collectingCountry id="6FAD761FE55BB0609ABD5F48D128FEF0" box="[160,268,282,306]" name="Colombia" pageId="13" pageNumber="737">Colombie</collectingCountry>
ex coll Candeze
<collectingRegion id="D57EF86DE55BB0609BD45F48D234FEF0" box="[457,528,282,306]" country="Colombia" name="Choco" pageId="13" pageNumber="737">Choco</collectingRegion>
<collectingCountry id="6FAD761FE55BB06098055F48D27AFEF0" box="[536,606,282,306]" name="United Kingdom" pageId="13" pageNumber="737">Wallis</collectingCountry>
/det. P.
<bibRefCitation id="732B4B7EE55BB06098B25F48D368FEF0" author="Preudhomme de Borre A" box="[687,844,282,306]" pageId="13" pageNumber="737" pagination="103 - 120" refId="ref9549" refString="Preudhomme de Borre A. 1886. Liste des Lamellicornes Laparostictiques recueillis par feu Camille Van Volxem pendant son voyage au Bresil et a la Plata en 1872 i suivie de la description de dix huit especes nouvelles et un genre nouveau. Annales de la Societe entomologique de Belgique. 30: 103 - 120." type="journal article" year="1886">de Borre 1886</bibRefCitation>
<taxonomicName id="D0BA4D0CE55BB060994E5F48D406FEF0" authority="Candezei" authorityName="Candezei" box="[851,1058,282,306]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="737" phylum="Arthropoda" rank="genus">Orphnus Candezei</taxonomicName>
<specimenCount id="01BCFD06E55BB0609E345F48D419FEF0" box="[1065,1085,282,306]" pageId="13" pageNumber="737" type="male"></specimenCount>
Type/
<typeStatus id="C801882DE55BB0609ABD5F6ED121FE96" box="[160,261,316,340]" pageId="13" pageNumber="737" type="holotype">Holotype</typeStatus>
/
<typeStatus id="C801882DE55BB0609B105F6ED1AFFE96" box="[269,395,316,340]" pageId="13" pageNumber="737" type="lectotype">Lectotypus</typeStatus>
<taxonomicName id="D0BA4D0CE55BB0609B895F6ED245FE96" authorityName="Preudhomme de Borre" authorityYear="1886" box="[404,609,316,340]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="737" phylum="Arthropoda" rank="species" species="candezei">Orphnus candezei</taxonomicName>
P.d.B. Frolov des. 2018
<emphasis id="25CEEA9DE55BB06099775F6ED354FE96" box="[874,880,316,340]" italics="true" pageId="13" pageNumber="737"></emphasis>
.
</materialsCitation>
</paragraph>
<paragraph id="1705368FE55BB0609ADD5F0DD30BFE1D" blockId="13.[160,1156,212,479]" pageId="13" pageNumber="737">
<materialsCitation id="A7D23CD2E55BB0609ADD5F0DD30BFE1D" ID-GBIF-Occurrence="2569344942" collectionCode="IRSNB" collectorName="Coll. R. I. Sc. N. B. &amp; Coll. R. I. Sc." country="Colombia" isTailEnumeration="true" location="Choco" pageId="13" pageNumber="737" specimenCount="1" specimenCount-male="1" stateProvince="Choco" typeStatus="paralectotype">
<emphasis id="25CEEA9DE55BB0609ADD5F0DD142FEB5" bold="true" box="[192,358,351,375]" pageId="13" pageNumber="737">
<typeStatus id="C801882DE55BB0609ADD5F0DD142FEB5" box="[192,358,351,375]" pageId="13" pageNumber="737" type="paralectotype">Paralectotype</typeStatus>
</emphasis>
(
<figureCitation id="8F812A0AE55BB0609B6A5F0DD1F2FEB5" box="[375,470,351,375]" captionStart="Figure 3" captionStartId="6.[160,225,1495,1518]" captionTargetBox="[167,1148,143,1463]" captionTargetId="figure@6.[167,1149,142,1464]" captionTargetPageId="6" captionText="Figure 3. Aegidinus spp. Aegidinus candezei, lectotype, male (a, e), male (bd), female (h, k), female, holotype of Ae. sunidigea (g, j); Aegidinus sp., paralectotype of Ae. candezei (f, i, l). Habitus in dorsal view (a, g, h, i); labels (e, f); female external genitalia (jl); aedeagus in lateral (b) and ventral view (c); parameres in dorsal view (d)." figureDoi="http://doi.org/10.5281/zenodo.3675577" httpUri="https://zenodo.org/record/3675577/files/figure.png" pageId="13" pageNumber="737">Figure 3</figureCitation>
(f,i,l)), female at
<collectionCode id="71ABAE4AE55BB06098935F0DD2F4FEB5" box="[654,720,351,375]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="13" pageNumber="737">IRSNB</collectionCode>
labelled
<emphasis id="25CEEA9DE55BB06099265F0DD365FEB5" box="[827,833,351,375]" italics="true" pageId="13" pageNumber="737"></emphasis>
<collectorName id="BA4F5359E55BB060995D5F0DD349FEB5" box="[832,877,351,375]" pageId="13" pageNumber="737">Coll</collectorName>
. R. I. Sc. N. B.
<collectingCountry id="6FAD761FE55BB0609E0F5F0DD4A5FEB5" box="[1042,1153,351,375]" name="Colombia" pageId="13" pageNumber="737">Colombie</collectingCountry>
:
<collectingRegion id="D57EF86DE55BB0609ABD5FD0D0C3FE58" box="[160,231,386,410]" country="Colombia" name="Choco" pageId="13" pageNumber="737">Choco</collectingRegion>
ex
<collectingCountry id="6FAD761FE55BB0609B065FD0D178FE58" box="[283,348,386,410]" name="United Kingdom" pageId="13" pageNumber="737">Wallis</collectingCountry>
ex coll.
<location id="12656054E55BB0609BD45FD0D20BFE58" LSID="urn:lsid:plazi:treatment:9F138799E55AB0639B7F5BBAD121FEF3:12656054E55BB0609BD45FD0D20BFE58" box="[457,559,386,410]" country="Colombia" name="Candeze" pageId="13" pageNumber="737" stateProvince="Choco">Candeze</location>
/det. P.
<bibRefCitation id="732B4B7EE55BB06098965FD0D310FE58" author="Preudhomme de Borre A" box="[651,820,386,410]" pageId="13" pageNumber="737" pagination="103 - 120" refId="ref9549" refString="Preudhomme de Borre A. 1886. Liste des Lamellicornes Laparostictiques recueillis par feu Camille Van Volxem pendant son voyage au Bresil et a la Plata en 1872 i suivie de la description de dix huit especes nouvelles et un genre nouveau. Annales de la Societe entomologique de Belgique. 30: 103 - 120." type="journal article" year="1886">de Borre 1886</bibRefCitation>
<taxonomicName id="D0BA4D0CE55BB060995C5FD0D432FE58" authority="Candezei" authorityName="Candezei" box="[833,1046,386,410]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="737" phylum="Arthropoda" rank="genus">Orphnus Candezei</taxonomicName>
<specimenCount id="01BCFD06E55BB0609E3E5FD0D413FE58" box="[1059,1079,386,410]" pageId="13" pageNumber="737" type="female"></specimenCount>
Type/
<typeStatus id="C801882DE55BB0609ABD5FF6D0D8FE7E" box="[160,252,420,444]" pageId="13" pageNumber="737" type="allotype">Allotype</typeStatus>
/
<taxonomicName id="D0BA4D0CE55BB0609B195FF6D37CFE7E" authority="(Preudhomme de Borre)" baseAuthorityName="Preudhomme de Borre" baseAuthorityYear="1886" box="[260,856,420,444]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="737" phylum="Arthropoda" rank="species" species="candezei">Aegidinus candezei (Preudhomme de Borre)</taxonomicName>
det. J.B.
<bibRefCitation id="732B4B7EE55BB06099FC5FF6D45EFE7E" author="Colby J" box="[993,1146,420,444]" pageId="13" pageNumber="737" pagination="1 - 41" refId="ref9012" refString="Colby J. 2009. Monographic revision of the genus Aegidinus Arrow (1904) and generic phylogeny of the world Orphninae (Coleoptera: Scarabaeidae: Orphninae). Insecta Mundi. 76: 1 - 41." type="journal article" year="2009">Colby 2009</bibRefCitation>
/
<typeStatus id="C801882DE55BB0609ABD5F95D161FE1D" box="[160,325,455,479]" pageId="13" pageNumber="737" type="paralectotype">Paralectotypus</typeStatus>
<taxonomicName id="D0BA4D0CE55BB0609B535F95D23FFE1D" authorityName="Preudhomme de Borre" authorityYear="1886" box="[334,539,455,479]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="737" phylum="Arthropoda" rank="species" species="candezei">Orphnus candezei</taxonomicName>
P.d.B. Frolov des. 2018
<emphasis id="25CEEA9DE55BB06099395F95D30DFE1D" box="[804,809,455,479]" italics="true" pageId="13" pageNumber="737"></emphasis>
.
</materialsCitation>
</paragraph>
<paragraph id="1705368FE55BB0609ABD5C5ED221FDE7" blockId="13.[160,1157,524,722]" box="[160,517,524,549]" pageId="13" pageNumber="737">
<emphasis id="25CEEA9DE55BB0609ABD5C5ED221FDE7" bold="true" box="[160,517,524,549]" italics="true" pageId="13" pageNumber="737">Additional material examined</emphasis>
</paragraph>
<paragraph id="1705368FE55BB0609ABD5C7DD40FFD10" blockId="13.[160,1157,524,722]" pageId="13" pageNumber="737">
<materialsCitation id="A7D23CD2E55BB0609ABD5C7DD4A0FD85" ID-GBIF-Occurrence="2569344947" box="[160,1156,559,583]" collectingDate="1995-01" collectionCode="CEMT" collectorName="Arriagada" country="Colombia" isTailEnumeration="true" location="May" municipality="Otanchei" pageId="13" pageNumber="737" specimenCount="1" stateProvince="Boyaca">
<emphasis id="25CEEA9DE55BB0609ABD5C7DD1B6FD85" bold="true" box="[160,402,559,583]" pageId="13" pageNumber="737">
<collectingCountry id="6FAD761FE55BB0609ABD5C7DD103FD85" box="[160,295,559,583]" name="Colombia" pageId="13" pageNumber="737">COLOMBIA</collectingCountry>
.
<collectingRegion id="D57EF86DE55BB0609B215C7DD1B6FD85" box="[316,402,559,583]" country="Colombia" name="Boyaca" pageId="13" pageNumber="737">Boyacá</collectingRegion>
</emphasis>
:
<collectingMunicipality id="F761ACF5E55BB0609BB75C7DD22AFD85" box="[426,526,559,583]" pageId="13" pageNumber="737">Otanche,</collectingMunicipality>
5
<location id="12656054E55BB060985D5C7DD24BFD85" LSID="urn:lsid:plazi:treatment:9F138799E55AB0639B7F5BBAD121FEF3:12656054E55BB060985D5C7DD24BFD85" box="[576,623,559,583]" country="Colombia" municipality="Otanchei" name="May" pageId="13" pageNumber="737" stateProvince="Boyaca">May</location>
<date id="6304104FE55BB060989D5C7DD29AFD85" box="[640,702,559,583]" pageId="13" pageNumber="737" value="1995-01">
<collectingDate id="7340E9A7E55BB060989D5C7DD29AFD85" box="[640,702,559,583]" pageId="13" pageNumber="737" value="1995-01">1995,</collectingDate>
</date>
<collectorName id="BA4F5359E55BB06098D25C7DD318FD85" box="[719,828,559,583]" pageId="13" pageNumber="737">Arriágada</collectorName>
leg., five females (
<collectionCode id="71ABAE4AE55BB0609E285C7DD45FFD85" box="[1077,1147,559,583]" pageId="13" pageNumber="737">CEMT</collectionCode>
).
</materialsCitation>
<materialsCitation id="A7D23CD2E55BB0609ABD5C00D4A1FD4E" ID-GBIF-Occurrence="2569344940" collectingDate="1995-01" collectionCode="CEMT" collectorName="Arriagada" country="Colombia" isTailEnumeration="true" location="May" municipality="Otanchei" pageId="13" pageNumber="737" specimenCount="1" stateProvince="Boyaca">
<emphasis id="25CEEA9DE55BB0609ABD5C00D138FDA8" bold="true" box="[160,284,594,618]" pageId="13" pageNumber="737">
<collectingRegion id="D57EF86DE55BB0609ABD5C00D138FDA8" box="[160,284,594,618]" country="Colombia" name="Santander" pageId="13" pageNumber="737">Santander</collectingRegion>
</emphasis>
:
<collectingMunicipality id="F761ACF5E55BB0609B365C00D2C4FDA8" box="[299,736,594,618]" pageId="13" pageNumber="737">Puerto Araújo, Los Manantiales Ranch,</collectingMunicipality>
<location id="12656054E55BB06098F45C00D33CFDA8" LSID="urn:lsid:plazi:treatment:9F138799E55AB0639B7F5BBAD121FEF3:12656054E55BB06098F45C00D33CFDA8" box="[745,792,594,618]" country="Colombia" municipality="Puerto Araujoi Los Manantiales Ranchi" name="May" pageId="13" pageNumber="737" stateProvince="Santander">May</location>
<date id="6304104FE55BB060993F5C00D344FDA8" box="[802,864,594,618]" pageId="13" pageNumber="737" value="2000-01">
<collectingDate id="7340E9A7E55BB060993F5C00D344FDA8" box="[802,864,594,618]" pageId="13" pageNumber="737" value="2000-01">2000,</collectingDate>
</date>
J.
<collectorName id="BA4F5359E55BB060999D5C00D3FCFDA8" box="[896,984,594,618]" pageId="13" pageNumber="737">Noriega</collectorName>
leg., one male (
<collectionCode id="71ABAE4AE55BB0609AB85C26D0CFFD4E" box="[165,235,628,652]" pageId="13" pageNumber="737">CEMT</collectionCode>
);
<location id="12656054E55BB0609AE15C26D205FD4E" LSID="urn:lsid:plazi:treatment:9F138799E55AB0639B7F5BBAD121FEF3:12656054E55BB0609AE15C26D205FD4E" box="[252,545,628,652]" country="Colombia" name="Puerto Araujoi Las Mariasi" pageId="13" pageNumber="737" stateProvince="Santander">Puerto Araújo, Las Marias,</location>
<locationDeviation id="06B72FADE55BB06098345C26D289FD4E" box="[553,685,628,652]" location="May" pageId="13" pageNumber="737">
<quantity id="D0429B6AE55BB06098345C26D252FD4E" box="[553,630,628,652]" metricMagnitude="5" metricUnit="m" metricValue="2.0921472" pageId="13" pageNumber="737" unit="mi" value="130.0">130 m,</quantity>
May
</locationDeviation>
<date id="6304104FE55BB06098A85C26D2D7FD4E" box="[693,755,628,652]" pageId="13" pageNumber="737" value="2000-01">
<collectingDate id="7340E9A7E55BB06098A85C26D2D7FD4E" box="[693,755,628,652]" pageId="13" pageNumber="737" value="2000-01">2000,</collectingDate>
</date>
J.
<collectorName id="BA4F5359E55BB060990F5C26D34EFD4E" box="[786,874,628,652]" pageId="13" pageNumber="737">Noriega</collectorName>
leg., one female (
<collectionCode id="71ABAE4AE55BB0609E285C26D45FFD4E" box="[1077,1147,628,652]" pageId="13" pageNumber="737">CEMT</collectionCode>
).
</materialsCitation>
<materialsCitation id="A7D23CD2E55BB0609ABD5CC5D4A1FD6D" ID-GBIF-Occurrence="2569344902" box="[160,1157,663,687]" collectingDate="1995-01" collectionCode="CEMT" collectorName="Arriagada" country="Colombia" isTailEnumeration="true" location="May" municipality="Otanchei" pageId="13" pageNumber="737" specimenCount="1" stateProvince="Boyaca">
<emphasis id="25CEEA9DE55BB0609ABD5CC5D0D6FD6D" bold="true" box="[160,242,663,687]" pageId="13" pageNumber="737">
<collectingRegion id="D57EF86DE55BB0609ABD5CC5D0D6FD6D" box="[160,242,663,687]" country="Colombia" name="Tolima" pageId="13" pageNumber="737">Tolima</collectingRegion>
</emphasis>
:
<location id="12656054E55BB0609AE25CC5D1C2FD6D" LSID="urn:lsid:plazi:treatment:9F138799E55AB0639B7F5BBAD121FEF3:12656054E55BB0609AE25CC5D1C2FD6D" box="[255,486,663,687]" country="Colombia" name="Mariquitai Municipal" pageId="13" pageNumber="737" stateProvince="Tolima">Mariquita, Municipal</location>
forest,
<locationDeviation id="06B72FADE55BB06098265CC5D2E1FD6D" box="[571,709,663,687]" location="April" pageId="13" pageNumber="737">
<quantity id="D0429B6AE55BB06098265CC5D2ACFD6D" box="[571,648,663,687]" metricMagnitude="6" metricUnit="m" metricValue="1.09435392" pageId="13" pageNumber="737" unit="mi" value="680.0">680 m,</quantity>
April
</locationDeviation>
<date id="6304104FE55BB06098D15CC5D32EFD6D" box="[716,778,663,687]" pageId="13" pageNumber="737" value="2000-01">
<collectingDate id="7340E9A7E55BB06098D15CC5D32EFD6D" box="[716,778,663,687]" pageId="13" pageNumber="737" value="2000-01">2000,</collectingDate>
</date>
J.
<collectorName id="BA4F5359E55BB06099355CC5D3A4FD6D" box="[808,896,663,687]" pageId="13" pageNumber="737">Noriega</collectorName>
leg., one male (
<collectionCode id="71ABAE4AE55BB0609E285CC5D45FFD6D" box="[1077,1147,663,687]" pageId="13" pageNumber="737">CEMT</collectionCode>
).
</materialsCitation>
<materialsCitation id="A7D23CD2E55BB0609ABD5CE8D40FFD10" ID-GBIF-Occurrence="2569344903" box="[160,1067,698,722]" collectingDate="1995-01" collectionCode="CEMT" collectorName="Arriagada" country="Colombia" isTailEnumeration="true" location="May" municipality="Otanchei" pageId="13" pageNumber="737" specimenCount="1" stateProvince="Boyaca">
<emphasis id="25CEEA9DE55BB0609ABD5CE8D132FD10" bold="true" box="[160,278,698,722]" pageId="13" pageNumber="737">
<collectingRegion id="D57EF86DE55BB0609ABD5CE8D132FD10" box="[160,278,698,722]" country="Colombia" name="Antioquia" pageId="13" pageNumber="737">Antioquia</collectingRegion>
</emphasis>
:
<collectingMunicipality id="F761ACF5E55BB0609B385CE8D1AFFD10" box="[293,395,698,722]" pageId="13" pageNumber="737">Medellín,</collectingMunicipality>
forest, pitfall,
<location id="12656054E55BB060982D5CE8D241FD10" LSID="urn:lsid:plazi:treatment:9F138799E55AB0639B7F5BBAD121FEF3:12656054E55BB060982D5CE8D241FD10" box="[560,613,698,722]" country="Colombia" municipality="Medellini" name="April" pageId="13" pageNumber="737" stateProvince="Antioquia">April</location>
<date id="6304104FE55BB06098705CE8D28FFD10" box="[621,683,698,722]" pageId="13" pageNumber="737" value="1998-01">
<collectingDate id="7340E9A7E55BB06098705CE8D28FFD10" box="[621,683,698,722]" pageId="13" pageNumber="737" value="1998-01">1998,</collectingDate>
</date>
J.
<collectorName id="BA4F5359E55BB06098D65CE8D307FD10" box="[715,803,698,722]" pageId="13" pageNumber="737">Noriega</collectorName>
leg., one male (
<collectionCode id="71ABAE4AE55BB06099C65CE8D405FD10" box="[987,1057,698,722]" pageId="13" pageNumber="737">CEMT</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="5FA06504E55BB0609ABD5CACD355FCBD" pageId="13" pageNumber="737" type="diagnosis">
<paragraph id="1705368FE55BB0609ABD5CACD130FCD5" blockId="13.[160,1155,766,895]" box="[160,276,766,791]" pageId="13" pageNumber="737">
<emphasis id="25CEEA9DE55BB0609ABD5CACD130FCD5" bold="true" box="[160,276,766,791]" italics="true" pageId="13" pageNumber="737">Diagnosis</emphasis>
</paragraph>
<paragraph id="1705368FE55BB0609ABD5D70D355FCBD" blockId="13.[160,1155,766,895]" pageId="13" pageNumber="737">
<taxonomicName id="D0BA4D0CE55BB0609ABD5D70D10DFCF8" authorityName="Preudhomme de Borre" authorityYear="1886" box="[160,297,802,826]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="737" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="25CEEA9DE55BB0609ABD5D70D10DFCF8" box="[160,297,802,826]" italics="true" pageId="13" pageNumber="737">Ae. candezei</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="D0BA4D0CE55BB0609BE25D70D2A6FCF8" authority="Akhmetova &amp; Vaz-de-Mello, 2019" authorityName="Frolov &amp; Akhmetova &amp; Vaz-de-Mello" authorityYear="2019" box="[511,642,802,826]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="737" phylum="Arthropoda" rank="species" species="noriegai" status="sp. nov.">
<emphasis id="25CEEA9DE55BB0609BE25D70D2A6FCF8" box="[511,642,802,826]" italics="true" pageId="13" pageNumber="737">Ae. noriegai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3EFD57E6E55BB06098975D70D2C7FCF8" box="[650,739,802,826]" pageId="13" pageNumber="737" rank="species">sp. nov.</taxonomicNameLabel>
but can be distinguished from it by the shorter and less asymmetrical parameres, ventroapical plate of the phallobase wider than long (
<figureCitation id="8F812A0AE55BB0609B025D35D15AFCBD" box="[287,382,871,895]" captionStart="Figure 3" captionStartId="6.[160,225,1495,1518]" captionTargetBox="[167,1148,143,1463]" captionTargetId="figure@6.[167,1149,142,1464]" captionTargetPageId="6" captionText="Figure 3. Aegidinus spp. Aegidinus candezei, lectotype, male (a, e), male (bd), female (h, k), female, holotype of Ae. sunidigea (g, j); Aegidinus sp., paralectotype of Ae. candezei (f, i, l). Habitus in dorsal view (a, g, h, i); labels (e, f); female external genitalia (jl); aedeagus in lateral (b) and ventral view (c); parameres in dorsal view (d)." figureDoi="http://doi.org/10.5281/zenodo.3675577" httpUri="https://zenodo.org/record/3675577/files/figure.png" pageId="13" pageNumber="737">Figure 3</figureCitation>
(b
<emphasis id="25CEEA9DE55BB0609B8E5D35D184FCBD" box="[403,416,871,895]" italics="true" pageId="13" pageNumber="737"></emphasis>
d)), and protibia with medioapical tooth.
</paragraph>
</subSubSection>
<subSubSection id="5FA06504E55BB0639ABD5DFED121FEF3" lastPageId="14" lastPageNumber="738" pageId="13" pageNumber="737" type="discussion">
<paragraph id="1705368FE55BB0609ABD5DFED122FC07" blockId="13.[160,1157,940,1727]" box="[160,262,940,965]" pageId="13" pageNumber="737">
<emphasis id="25CEEA9DE55BB0609ABD5DFED122FC07" bold="true" box="[160,262,940,965]" italics="true" pageId="13" pageNumber="737">Remarks</emphasis>
</paragraph>
<paragraph id="1705368FE55BB0609ABD5D9DD11CF9B8" blockId="13.[160,1157,940,1727]" pageId="13" pageNumber="737">
<taxonomicName id="D0BA4D0CE55BB0609ABD5D9DD155FC25" baseAuthorityName="Preudhomme de Borre" baseAuthorityYear="1886" box="[160,369,975,999]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="737" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="25CEEA9DE55BB0609ABD5D9DD155FC25" box="[160,369,975,999]" italics="true" pageId="13" pageNumber="737">Aegidinus candezei</emphasis>
</taxonomicName>
was described with no explicit information about the number of the examined specimens although it was indicated that both sexes were studied. There are two specimens (
<figureCitation id="8F812A0AE55BB0609B795A47D1E2FBEF" box="[356,454,1045,1069]" captionStart="Figure 3" captionStartId="6.[160,225,1495,1518]" captionTargetBox="[167,1148,143,1463]" captionTargetId="figure@6.[167,1149,142,1464]" captionTargetPageId="6" captionText="Figure 3. Aegidinus spp. Aegidinus candezei, lectotype, male (a, e), male (bd), female (h, k), female, holotype of Ae. sunidigea (g, j); Aegidinus sp., paralectotype of Ae. candezei (f, i, l). Habitus in dorsal view (a, g, h, i); labels (e, f); female external genitalia (jl); aedeagus in lateral (b) and ventral view (c); parameres in dorsal view (d)." figureDoi="http://doi.org/10.5281/zenodo.3675577" httpUri="https://zenodo.org/record/3675577/files/figure.png" pageId="13" pageNumber="737">Figure 3</figureCitation>
(a,i)) in the IRSNB collection which agree with the original description and might be the only specimens studied by Preudhomme de Borre. These specimens should be considered
<typeStatus id="C801882DE55BB060980E5A08D251FBB0" box="[531,629,1114,1138]" pageId="13" pageNumber="737" type="syntype">syntypes</typeStatus>
since no
<typeStatus id="C801882DE55BB06098C25A08D36CFBB0" box="[735,840,1114,1138]" pageId="13" pageNumber="737" type="lectotype">lectotype</typeStatus>
had been designated so far. The male specimen bears a red printed label that reads
<emphasis id="25CEEA9DE55BB060993D5A2ED302FB56" box="[800,806,1148,1172]" italics="true" pageId="13" pageNumber="737"></emphasis>
<typeStatus id="C801882DE55BB060993B5A2FD3A9FB57" box="[806,909,1149,1173]" pageId="13" pageNumber="737" type="holotype">Holotype</typeStatus>
<emphasis id="25CEEA9DE55BB06099915A2ED3B6FB56" box="[908,914,1148,1172]" italics="true" pageId="13" pageNumber="737"></emphasis>
(
<figureCitation id="8F812A0AE55BB06099BF5A2FD3DBFB57" box="[930,1023,1149,1173]" captionStart="Figure 3" captionStartId="6.[160,225,1495,1518]" captionTargetBox="[167,1148,143,1463]" captionTargetId="figure@6.[167,1149,142,1464]" captionTargetPageId="6" captionText="Figure 3. Aegidinus spp. Aegidinus candezei, lectotype, male (a, e), male (bd), female (h, k), female, holotype of Ae. sunidigea (g, j); Aegidinus sp., paralectotype of Ae. candezei (f, i, l). Habitus in dorsal view (a, g, h, i); labels (e, f); female external genitalia (jl); aedeagus in lateral (b) and ventral view (c); parameres in dorsal view (d)." figureDoi="http://doi.org/10.5281/zenodo.3675577" httpUri="https://zenodo.org/record/3675577/files/figure.png" pageId="13" pageNumber="737">Figure 3</figureCitation>
(e)) and the female specimen the label
<emphasis id="25CEEA9DE55BB0609BCC5ACDD1F3FB75" box="[465,471,1183,1207]" italics="true" pageId="13" pageNumber="737"></emphasis>
<typeStatus id="C801882DE55BB0609BCB5ACDD217FB75" box="[470,563,1183,1207]" pageId="13" pageNumber="737" type="allotype">Allotype</typeStatus>
<emphasis id="25CEEA9DE55BB06098295ACDD21EFB75" box="[564,570,1183,1207]" italics="true" pageId="13" pageNumber="737"></emphasis>
(
<figureCitation id="8F812A0AE55BB06098555ACDD282FB75" box="[584,678,1183,1207]" captionStart="Figure 3" captionStartId="6.[160,225,1495,1518]" captionTargetBox="[167,1148,143,1463]" captionTargetId="figure@6.[167,1149,142,1464]" captionTargetPageId="6" captionText="Figure 3. Aegidinus spp. Aegidinus candezei, lectotype, male (a, e), male (bd), female (h, k), female, holotype of Ae. sunidigea (g, j); Aegidinus sp., paralectotype of Ae. candezei (f, i, l). Habitus in dorsal view (a, g, h, i); labels (e, f); female external genitalia (jl); aedeagus in lateral (b) and ventral view (c); parameres in dorsal view (d)." figureDoi="http://doi.org/10.5281/zenodo.3675577" httpUri="https://zenodo.org/record/3675577/files/figure.png" pageId="13" pageNumber="737">Figure 3</figureCitation>
(f)). These labels were placed by the IRSNB technical staff in 1980
<emphasis id="25CEEA9DE55BB0609B885A90D186FB18" box="[405,418,1218,1242]" italics="true" pageId="13" pageNumber="737"></emphasis>
2000 (Alain Drumont, pers. comm.).
<bibRefCitation id="732B4B7EE55BB060992B5A90D3F1FB18" author="Paulian R" box="[822,981,1218,1242]" pageId="13" pageNumber="737" pagination="65 - 92" refId="ref9518" refString="Paulian R. 1984. Les Orphnidae Americains (Coleopteresi Scarabaeoidea). Annales de la Societe Entomologique de France (N S). 20: 65 - 92." type="journal article" year="1984">Paulian (1984)</bibRefCitation>
mentioned the
<emphasis id="25CEEA9DE55BB0609ABD5AB6D082FB3E" box="[160,166,1252,1276]" italics="true" pageId="13" pageNumber="737"></emphasis>
<typeStatus id="C801882DE55BB0609ABB5AB7D12EFB3F" box="[166,266,1253,1277]" pageId="13" pageNumber="737" type="holotype">holotype</typeStatus>
<emphasis id="25CEEA9DE55BB0609B175AB6D134FB3E" box="[266,272,1252,1276]" italics="true" pageId="13" pageNumber="737"></emphasis>
apparently based on these labels. The identity and conspecificity of the two specimens were not questioned by
<bibRefCitation id="732B4B7EE55BB06098335B55D2E8FADD" author="Paulian R" box="[558,716,1287,1311]" pageId="13" pageNumber="737" pagination="65 - 92" refId="ref9518" refString="Paulian R. 1984. Les Orphnidae Americains (Coleopteresi Scarabaeoidea). Annales de la Societe Entomologique de France (N S). 20: 65 - 92." type="journal article" year="1984">Paulian (1984)</bibRefCitation>
and
<bibRefCitation id="732B4B7EE55BB060991E5B55D3B5FADD" author="Colby J" box="[771,913,1287,1311]" pageId="13" pageNumber="737" pagination="1 - 41" refId="ref9012" refString="Colby J. 2009. Monographic revision of the genus Aegidinus Arrow (1904) and generic phylogeny of the world Orphninae (Coleoptera: Scarabaeidae: Orphninae). Insecta Mundi. 76: 1 - 41." type="journal article" year="2009">Colby (2009)</bibRefCitation>
, and the latter author drew the female characters of
<taxonomicName id="D0BA4D0CE55BB0609BE65B78D2A7FA80" baseAuthorityName="Preudhomme de Borre" baseAuthorityYear="1886" box="[507,643,1322,1346]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="737" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="25CEEA9DE55BB0609BE65B78D2A7FA80" box="[507,643,1322,1346]" italics="true" pageId="13" pageNumber="737">Ae. candezei</emphasis>
</taxonomicName>
from the female
<typeStatus id="C801882DE55BB06099575B78D380FA80" box="[842,932,1322,1346]" pageId="13" pageNumber="737" type="syntype">syntype</typeStatus>
. However, we think that the two
<typeStatus id="C801882DE55BB0609B215B1FD1BAFAA7" box="[316,414,1357,1381]" pageId="13" pageNumber="737" type="syntype">syntypes</typeStatus>
are not conspecific. The body punctation and the shape of the genitalia of the female
<typeStatus id="C801882DE55BB0609B835B3DD1D2FA45" box="[414,502,1391,1415]" pageId="13" pageNumber="737" type="syntype">syntype</typeStatus>
(
<figureCitation id="8F812A0AE55BB060981E5B3DD27AFA45" box="[515,606,1391,1415]" captionStart="Figure 3" captionStartId="6.[160,225,1495,1518]" captionTargetBox="[167,1148,143,1463]" captionTargetId="figure@6.[167,1149,142,1464]" captionTargetPageId="6" captionText="Figure 3. Aegidinus spp. Aegidinus candezei, lectotype, male (a, e), male (bd), female (h, k), female, holotype of Ae. sunidigea (g, j); Aegidinus sp., paralectotype of Ae. candezei (f, i, l). Habitus in dorsal view (a, g, h, i); labels (e, f); female external genitalia (jl); aedeagus in lateral (b) and ventral view (c); parameres in dorsal view (d)." figureDoi="http://doi.org/10.5281/zenodo.3675577" httpUri="https://zenodo.org/record/3675577/files/figure.png" pageId="13" pageNumber="737">Figure 3</figureCitation>
(i,l)) do not agree with the female specimens from San Carlos (
<figureCitation id="8F812A0AE55BB0609B385BC0D1A5FA68" box="[293,385,1426,1450]" captionStart="Figure 3" captionStartId="6.[160,225,1495,1518]" captionTargetBox="[167,1148,143,1463]" captionTargetId="figure@6.[167,1149,142,1464]" captionTargetPageId="6" captionText="Figure 3. Aegidinus spp. Aegidinus candezei, lectotype, male (a, e), male (bd), female (h, k), female, holotype of Ae. sunidigea (g, j); Aegidinus sp., paralectotype of Ae. candezei (f, i, l). Habitus in dorsal view (a, g, h, i); labels (e, f); female external genitalia (jl); aedeagus in lateral (b) and ventral view (c); parameres in dorsal view (d)." figureDoi="http://doi.org/10.5281/zenodo.3675577" httpUri="https://zenodo.org/record/3675577/files/figure.png" pageId="13" pageNumber="737">Figure 3</figureCitation>
(h,k)) belonging to the series of both sexes with trustworthy locality data. Only the male
<typeStatus id="C801882DE55BB0609B965BE7D1C7FA0F" box="[395,483,1461,1485]" pageId="13" pageNumber="737" type="syntype">syntype</typeStatus>
bears the original locality label
<emphasis id="25CEEA9DE55BB060994B5BE6D378FA0E" box="[854,860,1460,1484]" italics="true" pageId="13" pageNumber="737"></emphasis>
Choco
<collectingCountry id="6FAD761FE55BB06099B15BE7D3C9FA0F" box="[940,1005,1461,1485]" name="United Kingdom" pageId="13" pageNumber="737">Wallis</collectingCountry>
<emphasis id="25CEEA9DE55BB06099F05BE6D3D7FA0E" box="[1005,1011,1460,1484]" italics="true" pageId="13" pageNumber="737"></emphasis>
. The locality label of the female
<typeStatus id="C801882DE55BB0609B9C5B85D1FDFA2D" box="[385,473,1495,1519]" pageId="13" pageNumber="737" type="syntype">syntype</typeStatus>
was added by IRSNB technical staff later and put under an unlabelled specimen. Although it seems probable that both specimens originate from Candeze
<emphasis id="25CEEA9DE55BB0609B1D584ED122F9F6" box="[256,262,1564,1588]" italics="true" pageId="13" pageNumber="737"></emphasis>
s collection and might have been collected by Wallis, the locality of the female
<typeStatus id="C801882DE55BB0609ABD586DD0DCF995" box="[160,248,1599,1623]" pageId="13" pageNumber="737" type="syntype">syntype</typeStatus>
is unclear since
<collectingCountry id="6FAD761FE55BB0609BA1586DD1D9F995" box="[444,509,1599,1623]" name="United Kingdom" pageId="13" pageNumber="737">Wallis</collectingCountry>
made extensive expeditions throughout South America (
<bibRefCitation id="732B4B7EE55BB0609ABB5830D108F9B8" author="Veitch JH" box="[166,300,1634,1658]" pageId="13" pageNumber="737" refId="ref9685" refString="Veitch JH. 1906. Hortus Veitchii: A history of the rise and progress of the nurseries of Messrs. James Veitch and Sonsi together with an account of the botanical collectors and hybridists employed by them and a list of the most remarkable of their introductions. London: J. Veitch and Sons; p. 684 p." type="book" year="1906">Veitch 1906</bibRefCitation>
).
</paragraph>
<paragraph id="1705368FE55BB0639ADD58D7D121FEF3" blockId="13.[160,1157,940,1727]" lastBlockId="14.[160,1156,142,305]" lastPageId="14" lastPageNumber="738" pageId="13" pageNumber="737">
<taxonomicName id="D0BA4D0CE55BB0609ADD58D7D1C4F95F" authority="Colby" authorityName="Colby" authorityYear="2009" box="[192,480,1669,1693]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="737" phylum="Arthropoda" rank="species" species="sunidigea">
<emphasis id="25CEEA9DE55BB0609ADD58D7D1BDF95F" box="[192,409,1669,1693]" italics="true" pageId="13" pageNumber="737">Aegidinus sunidigea</emphasis>
Colby
</taxonomicName>
was described from a female (
<figureCitation id="8F812A0AE55BB060992458D7D3B1F95F" box="[825,917,1669,1693]" captionStart="Figure 3" captionStartId="6.[160,225,1495,1518]" captionTargetBox="[167,1148,143,1463]" captionTargetId="figure@6.[167,1149,142,1464]" captionTargetPageId="6" captionText="Figure 3. Aegidinus spp. Aegidinus candezei, lectotype, male (a, e), male (bd), female (h, k), female, holotype of Ae. sunidigea (g, j); Aegidinus sp., paralectotype of Ae. candezei (f, i, l). Habitus in dorsal view (a, g, h, i); labels (e, f); female external genitalia (jl); aedeagus in lateral (b) and ventral view (c); parameres in dorsal view (d)." figureDoi="http://doi.org/10.5281/zenodo.3675577" httpUri="https://zenodo.org/record/3675577/files/figure.png" pageId="13" pageNumber="737">Figure 3</figureCitation>
(g,j)) from San Carlos. As discussed above, the series from San Carlos includes two species represented by both sexes, so the female characters of
<taxonomicName id="D0BA4D0CE558B06398285EDCD29BFF64" baseAuthorityName="Preudhomme de Borre" baseAuthorityYear="1886" box="[565,703,142,166]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="738" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="25CEEA9DE558B06398285EDCD29BFF64" box="[565,703,142,166]" italics="true" pageId="14" pageNumber="738">Ae. candezei</emphasis>
</taxonomicName>
can be inferred from these specimens with sparse punctation, including the type specimen of
<taxonomicName id="D0BA4D0CE558B063990A5EE3D38CFF0B" authorityName="Colby" authorityYear="2009" box="[791,936,177,201]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="738" phylum="Arthropoda" rank="species" species="sunidigea">
<emphasis id="25CEEA9DE558B063990A5EE3D38CFF0B" box="[791,936,177,201]" italics="true" pageId="14" pageNumber="738">Ae. sunidigea</emphasis>
</taxonomicName>
. To ensure stability of the nomenclature, we here designate the male
<typeStatus id="C801882DE558B06398C05E86D311FF2E" box="[733,821,212,236]" pageId="14" pageNumber="738" type="syntype">syntype</typeStatus>
(
<figureCitation id="8F812A0AE558B06399595E86D385FF2E" box="[836,929,212,236]" captionStart="Figure 3" captionStartId="6.[160,225,1495,1518]" captionTargetBox="[167,1148,143,1463]" captionTargetId="figure@6.[167,1149,142,1464]" captionTargetPageId="6" captionText="Figure 3. Aegidinus spp. Aegidinus candezei, lectotype, male (a, e), male (bd), female (h, k), female, holotype of Ae. sunidigea (g, j); Aegidinus sp., paralectotype of Ae. candezei (f, i, l). Habitus in dorsal view (a, g, h, i); labels (e, f); female external genitalia (jl); aedeagus in lateral (b) and ventral view (c); parameres in dorsal view (d)." figureDoi="http://doi.org/10.5281/zenodo.3675577" httpUri="https://zenodo.org/record/3675577/files/figure.png" pageId="14" pageNumber="738">Figure 3</figureCitation>
(a)) as the
<typeStatus id="C801882DE558B0639E075E86D4A7FF2E" box="[1050,1155,212,236]" pageId="14" pageNumber="738" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="D0BA4D0CE558B0639ADE5EA4D1AFFECC" authorityName="Preudhomme de Borre" authorityYear="1886" box="[195,395,246,270]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="738" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="25CEEA9DE558B0639ADE5EA4D1AFFECC" box="[195,395,246,270]" italics="true" pageId="14" pageNumber="738">Orphnus candezei</emphasis>
</taxonomicName>
and propose that
<taxonomicName id="D0BA4D0CE558B063986D5EA4D36AFECC" authorityName="Colby" authorityYear="2009" box="[624,846,246,270]" class="Insecta" family="Hybosoridae" genus="Aegidinus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="738" phylum="Arthropoda" rank="species" species="sunidigea">
<emphasis id="25CEEA9DE558B063986D5EA4D36AFECC" box="[624,846,246,270]" italics="true" pageId="14" pageNumber="738">Aegidinus sunidigea</emphasis>
</taxonomicName>
is a synonym of
<taxonomicName id="D0BA4D0CE558B0639E3B5EA4D0DBFEF3" authorityName="Preudhomme de Borre" authorityYear="1886" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="738" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="25CEEA9DE558B0639E3B5EA4D0DBFEF3" italics="true" pageId="14" pageNumber="738">Orphnus candezei</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>