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<document id="54AD77ADF3EA4A136197117153BCFA9F" ID-CLB-Dataset="29151" ID-DOI="10.11646/zootaxa.4422.1.5" ID-GBIF-Dataset="ea160d3a-63f4-4a24-b4b1-042ef1508b5c" ID-ISSN="1175-5326" ID-Zenodo-Dep="1455505" ID-ZooBank="D167DDAD-17D0-4F35-9873-85B4BC7E8FEB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539184953362" checkinUser="plazi" docAuthor="Jaschhof, Mathias &amp; Spungis, Voldemars" docDate="2018" docId="9E1D954CFFE0966CFF51FA30DF0DFE40" docLanguage="en" docName="zootaxa.4422.1.5.pdf" docOrigin="Zootaxa 4422 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dicerura yezoensis Jaschhof &amp; Spungis 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="91" masterDocId="6224ED34FFE5966AFFC6FF8ADA0CFFEC" masterDocTitle="Towards reliable identification of male Dicerura: descriptions of three new and seven poorly known species in the Palearctic region (Diptera: Cecidomyiidae, Porricondylinae)" masterLastPageNumber="103" masterPageNumber="85" pageNumber="90" updateTime="1698573658146" updateUser="ExternalLinkService">
<mods:mods id="48ACC628CA4849C7A70D1D55DF5A5998" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="733DC6799A47B67FA7EB9414C742E218">Towards reliable identification of male Dicerura: descriptions of three new and seven poorly known species in the Palearctic region (Diptera: Cecidomyiidae, Porricondylinae)</mods:title>
</mods:titleInfo>
<mods:name id="B79BCF99B4018AF02C350EAEDC746C57" type="personal">
<mods:role id="2E11A73ACE2DF3E96AFA7BDDF259716C">
<mods:roleTerm id="A2B3E0A0EC3D0B0EE20FF9DFB5B9960A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F3FDF0D1198FD46413C013ED9AB9FE83">Jaschhof, Mathias</mods:namePart>
</mods:name>
<mods:name id="60E21EFE3DDF3F6C5B1685DA97653C26" type="personal">
<mods:role id="D59967865ABC7F22661A5C1DE7DBCF0A">
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<mods:namePart id="03BEF27776DF7ED9B583E1ACD18A995F">Spungis, Voldemars</mods:namePart>
</mods:name>
<mods:typeOfResource id="F7AC9D212CDB20D69CE50F4F54A2DBFB">text</mods:typeOfResource>
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<mods:title id="C2FF2F741FB0884D4B9E47D0A978D880">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0BA36B730FBD045E5244E56ECECCF250">
<mods:date id="BCFB2CBB49AB2E5B15B1793BA0380BE3">2018</mods:date>
<mods:detail id="0749B0A161617A07C89F3DCA4FD8CDA4" type="volume">
<mods:number id="2BEC20CDE91B9DFCF28DC94A1EC3E688">4422</mods:number>
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<mods:number id="D5E416156FC02EB169019C69FE79B384">1</mods:number>
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<mods:classification id="78E46ACDE1DB87AB50494176FD3D7AE3">journal article</mods:classification>
<mods:identifier id="F7528EE6948ED9D2404CD0CB937685AE" type="CLB-Dataset">29151</mods:identifier>
<mods:identifier id="4240D04E913966A8C56187BE15745E3E" type="DOI">10.11646/zootaxa.4422.1.5</mods:identifier>
<mods:identifier id="E65D51BE14D7991B53AAFC0A5B881824" type="GBIF-Dataset">ea160d3a-63f4-4a24-b4b1-042ef1508b5c</mods:identifier>
<mods:identifier id="4E7FC92C2744A0467C9A69CA27ACE0D0" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F3A4D9C164F90D338A1C94C7AD8EE93F" type="Zenodo-Dep">1455505</mods:identifier>
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</mods:mods>
<treatment id="9E1D954CFFE0966CFF51FA30DF0DFE40" ID-DOI="http://doi.org/10.5281/zenodo.6489287" ID-GBIF-Taxon="148671193" ID-Zenodo-Dep="6489287" LSID="urn:lsid:plazi:treatment:9E1D954CFFE0966CFF51FA30DF0DFE40" httpUri="http://treatment.plazi.org/id/9E1D954CFFE0966CFF51FA30DF0DFE40" lastPageId="6" lastPageNumber="91" pageId="5" pageNumber="90">
<subSubSection id="5EAE77D1FFE0966FFF51FA30DAF5FA1A" pageId="5" pageNumber="90" type="nomenclature">
<paragraph id="160B245AFFE0966FFF51FA30D8EDFA39" blockId="5.[151,737,1466,1526]" box="[151,737,1466,1493]" pageId="5" pageNumber="90">
<heading id="4D439336FFE0966FFF51FA30D8EDFA39" bold="true" box="[151,737,1466,1493]" fontSize="11" level="1" pageId="5" pageNumber="90" reason="1">
<emphasis id="24C0F848FFE0966FFF51FA30D8EDFA39" bold="true" box="[151,737,1466,1493]" pageId="5" pageNumber="90">
<taxonomicName id="D1B45FD9FFE0966FFF51FA30D870FA39" authority="Jaschhof &amp; Spungis" authorityName="Jaschhof &amp; Spungis" authorityYear="2018" box="[151,636,1466,1493]" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="yezoensis" status="sp. nov.">
<emphasis id="24C0F848FFE0966FFF51FA30DB77FA38" bold="true" box="[151,379,1466,1492]" italics="true" pageId="5" pageNumber="90">Dicerura yezoensis</emphasis>
Jaschhof &amp; Spungis
</taxonomicName>
<taxonomicNameLabel id="3FF34533FFE0966FFD45FA31D8EDFA39" box="[643,737,1467,1493]" pageId="5" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="160B245AFFE0966FFF51FA57DAF5FA1A" blockId="5.[151,737,1466,1526]" box="[151,249,1501,1526]" pageId="5" pageNumber="90">
<figureCitation id="8E8F38DFFFE0966FFF51FA57DAF5FA1A" box="[151,249,1501,1526]" captionStart="FIGURES 6–8" captionStartId="6.[151,264,1554,1576]" captionTargetBox="[185,1388,518,1492]" captionTargetId="figure@6.[151,1436,483,1532]" captionTargetPageId="6" captionText="FIGURES 6–8. Dicerura yezoenSiS, male, holotype. 6: Genitalia, ventral. 7: Ninth tergite, dorsal. 8: Tegmen (apex slightly distorted) and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455511/files/figure.png" pageId="5" pageNumber="90">Figs 6–8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="5EAE77D1FFE0966CFF51F9AFDED0FEF0" lastPageId="6" lastPageNumber="91" pageId="5" pageNumber="90" type="diagnosis">
<paragraph id="160B245AFFE0966FFF51F9AFDBF4F95E" blockId="5.[151,1437,1573,2005]" pageId="5" pageNumber="90">
<emphasis id="24C0F848FFE0966FFF51F9AFDB1DF9D2" bold="true" box="[151,273,1573,1598]" pageId="5" pageNumber="90">Diagnosis.</emphasis>
<taxonomicName id="D1B45FD9FFE0966FFEDFF9ACDBFEF9D1" authorityName="Jaschhof &amp; Spungis" authorityYear="2018" box="[281,498,1574,1597]" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="yezoensis">
<emphasis id="24C0F848FFE0966FFEDFF9ACDBFEF9D1" box="[281,498,1574,1597]" italics="true" pageId="5" pageNumber="90">Dicerura yezoensis</emphasis>
</taxonomicName>
, a member of the
<taxonomicName id="D1B45FD9FFE0966FFD02F9ACD92CF9D2" authorityName="Mamaev" authorityYear="1998" box="[708,800,1574,1598]" class="Insecta" family="Cecidomyiidae" genus="Dicerura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="formosa">
<emphasis id="24C0F848FFE0966FFD02F9ACD92CF9D2" box="[708,800,1574,1598]" italics="true" pageId="5" pageNumber="90">formosa</emphasis>
</taxonomicName>
group, is distinguished as follows: the gonostylus has a broadly rounded, not pointed, apex (
<figureCitation id="8E8F38DFFFE0966FFD86F9C6D884F989" box="[576,648,1612,1637]" captionStart="FIGURES 6–8" captionStartId="6.[151,264,1554,1576]" captionTargetBox="[185,1388,518,1492]" captionTargetId="figure@6.[151,1436,483,1532]" captionTargetPageId="6" captionText="FIGURES 6–8. Dicerura yezoenSiS, male, holotype. 6: Genitalia, ventral. 7: Ninth tergite, dorsal. 8: Tegmen (apex slightly distorted) and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455511/files/figure.png" pageId="5" pageNumber="90">Fig. 6</figureCitation>
, ↓1); the ventromedial gonocoxal processes are big and subglobular rather than ovate (
<figureCitation id="8E8F38DFFFE0966FFEB4F9F9DBB0F960" box="[370,444,1651,1676]" captionStart="FIGURES 6–8" captionStartId="6.[151,264,1554,1576]" captionTargetBox="[185,1388,518,1492]" captionTargetId="figure@6.[151,1436,483,1532]" captionTargetPageId="6" captionText="FIGURES 6–8. Dicerura yezoenSiS, male, holotype. 6: Genitalia, ventral. 7: Ninth tergite, dorsal. 8: Tegmen (apex slightly distorted) and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455511/files/figure.png" pageId="5" pageNumber="90">Fig. 6</figureCitation>
, ↓2); and the posterior edge of the ninth tergite is provided with a pair of pointed, microtrichose lobes (
<figureCitation id="8E8F38DFFFE0966FFE44F913DBCAF95D" box="[386,454,1689,1714]" captionStart="FIGURES 6–8" captionStartId="6.[151,264,1554,1576]" captionTargetBox="[185,1388,518,1492]" captionTargetId="figure@6.[151,1436,483,1532]" captionTargetPageId="6" captionText="FIGURES 6–8. Dicerura yezoenSiS, male, holotype. 6: Genitalia, ventral. 7: Ninth tergite, dorsal. 8: Tegmen (apex slightly distorted) and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455511/files/figure.png" pageId="5" pageNumber="90">Fig. 7</figureCitation>
, ↓3).
</paragraph>
<paragraph id="160B245AFFE0966FFF01F937DF90F93A" blockId="5.[151,1437,1573,2005]" box="[199,1436,1725,1751]" pageId="5" pageNumber="90">
<emphasis id="24C0F848FFE0966FFF01F937DBD6F93A" bold="true" box="[199,474,1725,1750]" pageId="5" pageNumber="90">Other male characters.</emphasis>
Body length
<quantity id="D14C89BFFFE0966FFDA9F934D8CBF93A" box="[623,711,1726,1750]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="5" pageNumber="90" unit="mm" value="3.3">3.3 mm</quantity>
.
<emphasis id="24C0F848FFE0966FFD17F937D914F93A" bold="true" box="[721,792,1725,1750]" pageId="5" pageNumber="90">Head.</emphasis>
Eye bridge 3–4 ommatidia long dorsally. Antenna clearly
</paragraph>
<paragraph id="160B245AFFE0966FFF51F96BDF90F916" blockId="5.[151,1437,1573,2005]" box="[151,1436,1761,1786]" pageId="5" pageNumber="90">shorter than body; scape and pedicel yellowish, lighter than flagellum. Circumfila with 1, seldom 2 short</paragraph>
<paragraph id="160B245AFFE0966FFF51F88FDF91F8F2" blockId="5.[151,1437,1573,2005]" box="[151,1437,1797,1822]" pageId="5" pageNumber="90">extensions. Neck of fourth flagellomere slightly shorter than node. Palpus 4-segmented, 0.7 times the head height.</paragraph>
<paragraph id="160B245AFFE0966FFF51F8A3DF90F8AE" blockId="5.[151,1437,1573,2005]" box="[151,1436,1833,1858]" pageId="5" pageNumber="90">
<emphasis id="24C0F848FFE0966FFF51F8A3DAFAF8AE" bold="true" box="[151,246,1833,1858]" pageId="5" pageNumber="90">Thorax.</emphasis>
Scutum with dark stripes dorsally and laterally. Anepisternum with 10 setae.
<emphasis id="24C0F848FFE0966FFBA3F8A3DEAAF8AE" bold="true" box="[1125,1190,1833,1858]" pageId="5" pageNumber="90">Wing</emphasis>
as long as body, with
</paragraph>
<paragraph id="160B245AFFE0966FFF51F8C7DF90F887" blockId="5.[151,1437,1573,2005]" box="[151,1436,1869,1899]" pageId="5" pageNumber="90">brownish tinge. Veins thick, distinct; Rs long, ¼ the length of R1, r-m slightly longer than Rs, apical portion of M1+2</paragraph>
<paragraph id="160B245AFFE0966FFF51F8FEDF90F861" blockId="5.[151,1437,1573,2005]" box="[151,1436,1908,1933]" pageId="5" pageNumber="90">
distinct, basal portion vanished.
<emphasis id="24C0F848FFE0966FFDE2F8FED86FF861" bold="true" box="[548,611,1908,1933]" pageId="5" pageNumber="90">Legs.</emphasis>
Empodia rudimentary. Claws slightly bent, 4 basal teeth gradually
</paragraph>
<paragraph id="160B245AFFE0966FFF51F812DF90F85D" blockId="5.[151,1437,1573,2005]" box="[151,1436,1944,1969]" pageId="5" pageNumber="90">
decreasing in size.
<emphasis id="24C0F848FFE0966FFEBAF812DBFDF85D" bold="true" box="[380,497,1944,1969]" pageId="5" pageNumber="90">Genitalia.</emphasis>
Ninth tergite subtrapezoid, with setae of various lengths except along median;
</paragraph>
<paragraph id="160B245AFFE0966CFF51F836DED0FEF0" blockId="5.[151,1437,1573,2005]" lastBlockId="6.[151,1437,151,429]" lastPageId="6" lastPageNumber="91" pageId="5" pageNumber="90">
anterior edge straight, faintly contoured (
<figureCitation id="8E8F38DFFFE0966FFDA4F836D8AAF839" box="[610,678,1980,2005]" captionStart="FIGURES 6–8" captionStartId="6.[151,264,1554,1576]" captionTargetBox="[185,1388,518,1492]" captionTargetId="figure@6.[151,1436,483,1532]" captionTargetPageId="6" captionText="FIGURES 6–8. Dicerura yezoenSiS, male, holotype. 6: Genitalia, ventral. 7: Ninth tergite, dorsal. 8: Tegmen (apex slightly distorted) and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455511/files/figure.png" pageId="5" pageNumber="90">Fig. 7</figureCitation>
). Gonocoxites: ventral surface densely setose; ventral emargination broadly rounded, sclerotized basally; dorsolateral processes larger than ventromedial processes, both with fine setae; dorsal portions angular-shaped posteriorly; dorsal apodemes moderately long (
<figureCitation id="8E8F38DFFFE3966CFBB5FF36DEB7FF39" box="[1139,1211,188,213]" captionStart="FIGURES 6–8" captionStartId="6.[151,264,1554,1576]" captionTargetBox="[185,1388,518,1492]" captionTargetId="figure@6.[151,1436,483,1532]" captionTargetPageId="6" captionText="FIGURES 6–8. Dicerura yezoenSiS, male, holotype. 6: Genitalia, ventral. 7: Ninth tergite, dorsal. 8: Tegmen (apex slightly distorted) and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455511/files/figure.png" pageId="6" pageNumber="91">Fig. 6</figureCitation>
). Gonostylus twice longer than broad, slightly bent, slightly tapered towards apex, large setae laterally, both small setae and dense, short microtrichia medially (
<figureCitation id="8E8F38DFFFE3966CFE11FE8ED817FEF1" box="[471,539,260,285]" captionStart="FIGURES 6–8" captionStartId="6.[151,264,1554,1576]" captionTargetBox="[185,1388,518,1492]" captionTargetId="figure@6.[151,1436,483,1532]" captionTargetPageId="6" captionText="FIGURES 6–8. Dicerura yezoenSiS, male, holotype. 6: Genitalia, ventral. 7: Ninth tergite, dorsal. 8: Tegmen (apex slightly distorted) and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455511/files/figure.png" pageId="6" pageNumber="91">Fig. 6</figureCitation>
). Tegminal apex with collar of 5 inconspicuous knobs (
<figureCitation id="8E8F38DFFFE3966CFB4FFE8EDEC2FEF0" box="[1161,1230,260,285]" captionStart="FIGURES 6–8" captionStartId="6.[151,264,1554,1576]" captionTargetBox="[185,1388,518,1492]" captionTargetId="figure@6.[151,1436,483,1532]" captionTargetPageId="6" captionText="FIGURES 6–8. Dicerura yezoenSiS, male, holotype. 6: Genitalia, ventral. 7: Ninth tergite, dorsal. 8: Tegmen (apex slightly distorted) and ejaculatory apodeme, ventral. Scales 0.05 mm. Arrows refer to characters described in the diagnosis." httpUri="https://zenodo.org/record/1455511/files/figure.png" pageId="6" pageNumber="91">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5EAE77D1FFE3966CFF01FEADDBD0FE89" pageId="6" pageNumber="91" type="etymology">
<paragraph id="160B245AFFE3966CFF01FEADDBD0FE89" blockId="6.[151,1437,151,429]" pageId="6" pageNumber="91">
<emphasis id="24C0F848FFE3966CFF01FEADDB42FEAC" bold="true" box="[199,334,295,320]" pageId="6" pageNumber="91">Etymology.</emphasis>
The species epithet is derived from Yezo, a now historical name of
<collectingRegion id="D470EAB8FFE3966CFB8EFEADDEB5FEAC" box="[1096,1209,295,320]" country="Japan" name="Hokkaido" pageId="6" pageNumber="91">Hokkaido</collectingRegion>
, where the
<typeStatus id="C90F9AF8FFE3966CFAFCFEADDF90FEAC" box="[1338,1436,295,320]" pageId="6" pageNumber="91" type="holotype">holotype</typeStatus>
of this species was collected.
</paragraph>
</subSubSection>
<subSubSection id="5EAE77D1FFE3966CFF01FEE5DF0DFE40" pageId="6" pageNumber="91" type="materials_examined">
<paragraph id="160B245AFFE3966CFF01FEE5DF0DFE40" blockId="6.[151,1437,151,429]" pageId="6" pageNumber="91">
<materialsCitation id="A6DC2E07FFE3966CFF01FEE5DEF1FE40" ID-GBIF-Occurrence="1915497322" collectingDate="1999-06-30" collectingDateMax="1999-07-26" collectingDateMin="1999-06-30" collectionCode="KUEC" collectorName="Malaisetrap, M." country="Japan" county="Male" location="Hokkaido University Experimental Forest" municipality="Tomakomai" pageId="6" pageNumber="91" specimenCount="2" stateProvince="Hokkaido" typeStatus="holotype">
<emphasis id="24C0F848FFE3966CFF01FEE5DB74FE64" bold="true" box="[199,376,367,392]" pageId="6" pageNumber="91">
<typeStatus id="C90F9AF8FFE3966CFF01FEE5DB0EFE64" box="[199,258,367,392]" pageId="6" pageNumber="91">Type</typeStatus>
material.
</emphasis>
<typeStatus id="C90F9AF8FFE3966CFE45FEE5DBE3FE64" box="[387,495,367,392]" pageId="6" pageNumber="91" type="holotype">Holotype</typeStatus>
.
<collectingCounty id="FF6A5CD6FFE3966CFE3BFEE5D835FE64" box="[509,569,367,392]" pageId="6" pageNumber="91">Male</collectingCounty>
,
<collectingCountry id="6EA364CAFFE3966CFD8FFEFAD880FE6B" box="[585,652,368,391]" name="Japan" pageId="6" pageNumber="91">Japan</collectingCountry>
,
<collectingRegion id="D470EAB8FFE3966CFD5DFEE5D900FE64" box="[667,780,367,392]" country="Japan" name="Hokkaido" pageId="6" pageNumber="91">Hokkaido</collectingRegion>
,
<collectingMunicipality id="F66FBE20FFE3966CFCDDFEE5D9A8FE64" box="[795,932,367,392]" pageId="6" pageNumber="91">Tomakomai</collectingMunicipality>
,
<location id="136B7281FFE3966CFC74FEE5DF94FE6B" LSID="urn:lsid:plazi:treatment:9E1D954CFFE0966CFF51FA30DF0DFE40:136B7281FFE3966CFC74FEE5DF94FE6B" box="[946,1432,367,392]" country="Japan" county="Male" municipality="Tomakomai" name="Hokkaido University Experimental Forest" pageId="6" pageNumber="91" stateProvince="Hokkaido">Hokkaido University Experimental Forest</location>
, mixed broadleaf deciduous forest,
<collectingDate id="724EFB72FFE3966CFDDDFE1ED91FFE40" box="[539,787,404,429]" pageId="6" pageNumber="91" value="1999-06-30" valueMax="1999-07-26" valueMin="1999-06-30">
<date id="620A029AFFE3966CFDDDFE1ED91FFE40" box="[539,787,404,429]" pageId="6" pageNumber="91" value="1999-06-30" valueMax="1999-07-26" valueMin="1999-06-30">30 June–26 July 1999</date>
</collectingDate>
,
<collectorName id="BB41418CFFE3966CFCD8FE1ED9DCFE40" box="[798,976,404,429]" pageId="6" pageNumber="91">Malaisetrap, M.</collectorName>
&amp; C. Jaschhof (in
<collectionCode id="70A5BC9FFFE3966CFB65FE1EDEF9FE40" box="[1187,1269,404,428]" country="Japan" httpUri="http://grbio.org/cool/txbq-e9hf" name="Kyushu University Entomology Collection" pageId="6" pageNumber="91">KUEC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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