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<mods:title id="82F0CEE6DDEFD63BD636CEB86545E081">The identity of Hamiltons Ticto Barb, Pethia ticto (Teleostei: Cyprinidae)</mods:title>
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<treatment id="9D112B52814E4947FF46BDEBFE3800ED" ID-DOI="http://doi.org/10.5281/zenodo.5664390" ID-GBIF-Taxon="119617815" ID-Zenodo-Dep="5664390" LSID="urn:lsid:plazi:treatment:9D112B52814E4947FF46BDEBFE3800ED" httpUri="http://treatment.plazi.org/id/9D112B52814E4947FF46BDEBFE3800ED" lastPageId="13" lastPageNumber="413" pageId="2" pageNumber="403">
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<heading id="4E4F2D28814E4948FF46BDEBFE6300E2" bold="true" box="[151,507,220,247]" fontSize="11" level="1" pageId="2" pageNumber="403" reason="1">
<taxonomicName id="D2B8E1C7814E4948FF46BDEBFE6300E2" ID-CoL="4FDDM" authority="Hamilton, 1822" authorityName="Hamilton" authorityYear="1822" box="[151,507,220,247]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="403" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC4656814E4948FF46BDEBFE6300E2" bold="true" box="[151,507,220,247]" pageId="2" pageNumber="403">
<emphasis id="27CC4656814E4948FF46BDEBFEB800E3" bold="true" box="[151,288,220,246]" italics="true" pageId="2" pageNumber="403">Pethia ticto</emphasis>
(
<bibRefCitation id="7129E7B5814E4948FEE1BDEAFE6B00E2" author="Hamilton" box="[304,499,221,247]" pageId="2" pageNumber="403" refString="Hamilton, F. (1822) An Account of the Fishes of River Ganges and its Branches. George Ramsay and Co., London, vii + 405 pp, 39 pls." type="book" year="1822">Hamilton, 1822</bibRefCitation>
)
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(
<figureCitation id="8D8386C1814E4948FF4FBDC8FE9D010D" box="[158,261,255,280]" captionStart="FIGURE 1" captionStartId="1.[151,250,1163,1185]" captionTargetBox="[194,1393,623,1141]" captionTargetId="figure@1.[194,1394,623,1141]" captionTargetPageId="1" captionText="FIGURE 1. Illustration of Pethia ticto based on topotype, male, BNHS FWF 127, 35.9 mm SL, showing distribution of prominent body pigmentation and banding pattern and lateral and transverse scale rows." httpUri="https://zenodo.org/record/233765/files/figure.png" pageId="2" pageNumber="403">Figures 1</figureCitation>
A, B, 2, 3)
</paragraph>
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<subSubSection id="5DA2C9CF814E4948FF46BC7EFD7D01CD" pageId="2" pageNumber="403" type="reference_group">
<paragraph id="15079A44814E4948FF46BC7EFD2A014B" blockId="2.[151,1162,328,472]" box="[151,690,328,350]" pageId="2" pageNumber="403">
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<taxonomicName id="D2B8E1C7814E4948FF46BC7EFD99014B" ID-CoL="33HZK" authority="Hamilton, 1822: 314" authorityName="Hamilton" authorityPageNumber="314" authorityYear="1822" box="[151,513,328,350]" class="Actinopterygii" family="Cyprinidae" genus="Cyprinus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="403" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC4656814E4948FF46BC7EFEBF014B" box="[151,295,329,350]" italics="true" pageId="2" pageNumber="403">Cyprinus ticto</emphasis>
<treatmentCitation id="9419BC55814E4948FEFCBC7FFD99014B" author="Hamilton" box="[301,513,328,350]" page="314" pageId="2" pageNumber="403" year="1822">
<bibRefCitation id="7129E7B5814E4948FEFCBC7FFE57014B" author="Hamilton" box="[301,463,328,350]" pageId="2" pageNumber="403" refString="Hamilton, F. (1822) An Account of the Fishes of River Ganges and its Branches. George Ramsay and Co., London, vii + 405 pp, 39 pls." type="book" year="1822">Hamilton, 1822</bibRefCitation>
: 314
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, plate VIII fig.87
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<treatmentCitationGroup id="35A8BD6A814E4948FF46BC50FD7D01CD" pageId="2" pageNumber="403">
<taxonomicName id="D2B8E1C7814E4948FF46BC50FE7B0169" ID-CoL="5456F" authority="Hamilton, 1822" authorityName="Hamilton" authorityYear="1822" box="[151,483,358,381]" class="Actinopterygii" family="Cyprinidae" genus="Systomus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="403" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC4656814E4948FF46BC50FEB00168" box="[151,296,359,381]" italics="true" pageId="2" pageNumber="403">Systomus ticto</emphasis>
(
<bibRefCitation id="7129E7B5814E4948FEE7BC51FE420169" author="Hamilton" box="[310,474,358,380]" pageId="2" pageNumber="403" refString="Hamilton, F. (1822) An Account of the Fishes of River Ganges and its Branches. George Ramsay and Co., London, vii + 405 pp, 39 pls." type="book" year="1822">Hamilton, 1822</bibRefCitation>
)
</taxonomicName>
: MClelland 1839: 382, plate
<specimenCount id="03BE51CD814E4948FCC0BC51FCAB0169" box="[785,819,358,380]" pageId="2" pageNumber="403" type="generic">8 f.</specimenCount>
87;
<treatmentCitation id="9419BC55814E4948FCB1BC51FB830169" author="Bleeker" box="[864,1051,358,380]" page="260" pageId="2" pageNumber="403" year="1844">
<bibRefCitation id="7129E7B5814E4948FCB1BC51FC730169" author="Bleeker" box="[864,1003,358,380]" pageId="2" pageNumber="403" refString="Bleeker (1844) Enumeratio specierum piscium hucusque in Archipelago Indico observatarum, adjectis habitationibus citationibusque ubi descriptions earum recentiores reperiuntur. Weber and de Beaufort, 15, 1 - 276." type="journal article" year="1844">Bleeker 1844</bibRefCitation>
: 260
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<taxonomicName id="D2B8E1C7814E4948FF46BCB2FE55018E" ID-CoL="KNY4" authority="Hamilton, 1822" authorityName="Hamilton" authorityYear="1822" box="[151,461,389,411]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="403" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC4656814E4948FF46BCB2FE8B018E" box="[151,275,389,411]" italics="true" pageId="2" pageNumber="403">Barbus ticto</emphasis>
(
<bibRefCitation id="7129E7B5814E4948FEF1BCB2FE5D018E" author="Hamilton" box="[288,453,389,411]" pageId="2" pageNumber="403" refString="Hamilton, F. (1822) An Account of the Fishes of River Ganges and its Branches. George Ramsay and Co., London, vii + 405 pp, 39 pls." type="book" year="1822">Hamilton, 1822</bibRefCitation>
)
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:
<treatmentCitation id="9419BC55814E4948FE06BCB2FD01018E" author="Gunther" box="[471,665,389,411]" page="153" pageId="2" pageNumber="403" year="1868">
<bibRefCitation id="7129E7B5814E4948FE06BCB2FDFE018E" author="Gunther" box="[471,614,389,411]" pageId="2" pageNumber="403" refString="Gunther, A. (1868) Catalogue of the fishes in the British Museum. Catalogue of the Physostomi, containing the families Heteropygii, Cyprinidae, Gonorhynchidae, Hyodontidae, Osteoglossidae, Clupeidae, Chirocenridae, Alepocephalidae, Notopteridae, Halosauridae, in the collection of the British Museum. British Museum, London, xx + 512 pp." type="book" year="1868">Günther 1868</bibRefCitation>
: 153
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;
<treatmentCitation id="9419BC55814E4948FD72BCB1FCA5018E" author="Day" box="[675,829,389,411]" page="576" pageId="2" pageNumber="403" year="1878">
<bibRefCitation id="7129E7B5814E4948FD72BCB1FC92018E" author="Day" box="[675,778,389,411]" pageId="2" pageNumber="403" refString="Day, F. (1878) The Fishes of India; Being a Natural History of the Fishes Known to Inhabit the Seas and Freshwaters of India, Burma and Ceylon. William Dawson &amp; Sons, London, xx + 778 pp, 196 pls." type="book" year="1878">Day 1878</bibRefCitation>
: 576
</treatmentCitation>
;
<treatmentCitation id="9419BC55814E4948FC96BCB1FC79018E" author="Day" box="[839,993,389,411]" page="290" pageId="2" pageNumber="403" year="1871">
<bibRefCitation id="7129E7B5814E4948FC96BCB1FC36018E" author="Day" box="[839,942,389,411]" pageId="2" pageNumber="403" refString="Day, F. (1871) Monograph of Indian Cyprinidae. Part 2. Journal and Proceedings of the Asiatic Society of Bengal, 40, 277 - 336." type="journal article" year="1871">Day 1871</bibRefCitation>
: 290
</treatmentCitation>
;
<treatmentCitation id="9419BC55814E4948FC3ABCB1FB12018E" author="Hora" box="[1003,1162,389,411]" page="263" pageId="2" pageNumber="403" year="1939">
<bibRefCitation id="7129E7B5814E4948FC3ABCB1FBC2018E" author="Hora" box="[1003,1114,389,411]" pageId="2" pageNumber="403" refString="Hora, S. L., Misra, K. S. &amp; Malik, G. M. (1939) A study of variations in Barbus (Puntius) ticto (Hamilton). Records of the Indian Museum, 41, 263 - 279." type="journal article" year="1939">Hora 1939</bibRefCitation>
: 263
</treatmentCitation>
<taxonomicName id="D2B8E1C7814E4948FF46BC92FE4A01AF" ID-CoL="6WPFT" authority="Hamilton, 1822" authorityName="Hamilton" authorityYear="1822" box="[151,466,420,442]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="403" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC4656814E4948FF46BC92FE8001AF" box="[151,280,421,442]" italics="true" pageId="2" pageNumber="403">Puntius ticto</emphasis>
(
<bibRefCitation id="7129E7B5814E4948FEF4BC93FE5201AF" author="Hamilton" box="[293,458,420,442]" pageId="2" pageNumber="403" refString="Hamilton, F. (1822) An Account of the Fishes of River Ganges and its Branches. George Ramsay and Co., London, vii + 405 pp, 39 pls." type="book" year="1822">Hamilton, 1822</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="9419BC55814E4948FE0ABC93FD6E01AF" author="Motwani" box="[475,758,420,442]" page="11" pageId="2" pageNumber="403" year="1957">
<bibRefCitation id="7129E7B5814E4948FE0ABC93FD4901AF" author="Motwani" box="[475,721,420,442]" pageId="2" pageNumber="403" refString="Motwani, M. P. &amp; Darvid, A. (1957) Fishes of the river Sone with observation on the zoo-geographical significance. Journal of Zoological Society India, 9, 9 - 15." type="journal article" year="1957">Motwani &amp; David 1957</bibRefCitation>
: 11
</treatmentCitation>
;
<treatmentCitation id="9419BC55814E4948FCD0BC92FC2001AF" author="Menon" box="[769,952,420,442]" page="268" pageId="2" pageNumber="403" year="1952">
<bibRefCitation id="7129E7B5814E4948FCD0BC92FC1D01AF" author="Menon" box="[769,901,420,442]" pageId="2" pageNumber="403" refString="Menon, M. A. S. (1952) On a collection of fish from Manipur, Assam. Records of the Indian Museum, 50, 265 - 270." type="journal article" year="1952">Menon 1952</bibRefCitation>
: 268
</treatmentCitation>
;
<treatmentCitation id="9419BC55814E4948FC13BC92FBEE01AF" author="Menon" box="[962,1142,420,442]" page="103" pageId="2" pageNumber="403" year="1999">
<bibRefCitation id="7129E7B5814E4948FC13BC92FBDE01AF" author="Menon" box="[962,1094,420,442]" pageId="2" pageNumber="403" refString="Menon, A. G. K. (1999) Check list-fresh water fishes of India. Records of the Zoological Survey of India, Miscellaneous Publication, Occasional Paper, 175, 366." type="journal article" year="1999">Menon 1999</bibRefCitation>
: 103
</treatmentCitation>
<taxonomicName id="D2B8E1C7814E4948FF46BCF5FE5E01CD" ID-CoL="4FDDM" authority="Hamilton, 1822" authorityName="Hamilton" authorityYear="1822" box="[151,454,450,472]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="403" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC4656814E4948FF46BCF5FE9401CD" box="[151,268,450,472]" italics="true" pageId="2" pageNumber="403">Pethia ticto</emphasis>
(
<bibRefCitation id="7129E7B5814E4948FEC8BCF5FE2601CD" author="Hamilton" box="[281,446,450,472]" pageId="2" pageNumber="403" refString="Hamilton, F. (1822) An Account of the Fishes of River Ganges and its Branches. George Ramsay and Co., London, vii + 405 pp, 39 pls." type="book" year="1822">Hamilton, 1822</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="9419BC55814E4948FE01BCF5FD7D01CD" author="Pethiyagoda" box="[464,741,450,472]" page="81" pageId="2" pageNumber="403" year="2012">
<bibRefCitation id="7129E7B5814E4948FE01BCF5FD5B01CD" author="Pethiyagoda" box="[464,707,450,472]" pageId="2" pageNumber="403" refString="Pethiyagoda, R., Meegaskumbura, M. &amp; Maduwage K. (2012) A synopsis of the South Asian fishes referred to Puntius (Pisces: Cyprinidae). Ichthyological Exploration of Freshwaters, 23, 69 - 95." type="journal article" year="2012">
Pethiyagoda
<emphasis id="27CC4656814E4948FD82BCF4FD1E01CD" box="[595,646,450,472]" italics="true" pageId="2" pageNumber="403">et al.</emphasis>
2012
</bibRefCitation>
: 81
</treatmentCitation>
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</paragraph>
</subSubSection>
<subSubSection id="5DA2C9CF814E4948FF46BF32FB82030F" pageId="2" pageNumber="403" type="materials_examined">
<paragraph id="15079A44814E4948FF46BF32FB82030F" blockId="2.[151,1437,517,1946]" pageId="2" pageNumber="403">
<emphasis id="27CC4656814E4948FF46BF32FE1A020B" bold="true" box="[151,386,517,542]" pageId="2" pageNumber="403">Material examined.</emphasis>
5 ex.,
<collectionCode id="73A90281814E4948FE05BF31FD87020B" box="[468,543,518,542]" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="2" pageNumber="403">BNHS</collectionCode>
<collectionCode id="73A90281814E4948FDF6BF31FDF90208" box="[551,609,518,541]" pageId="2" pageNumber="403">FWF</collectionCode>
127 to 131, Beri Baor, Ramnagar, Kolkata, West Bengal,
<collectingCountry id="6DAFDAD4814E4948FB2EBF32FAA1020B" box="[1279,1337,517,542]" name="India" pageId="2" pageNumber="403">India</collectingCountry>
(
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a.s.l.), coll. U. Katwate, R. Raghavan and N. Dahanukar,
<date id="6106BC84814E4948FB97BF1EFB4A0254" box="[1094,1234,553,578]" pageId="2" pageNumber="403" value="2014-06-06">6 June 2014</date>
; 4 ex., WILD-15- PIS-145 to 148, Beri Baor, Ramnagar, Kolkata, West Bengal,
<collectingCountry id="6DAFDAD4814E4948FC97BF7AFC180273" box="[838,896,589,614]" name="India" pageId="2" pageNumber="403">India</collectingCountry>
(
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a.s.l.), coll. U. Katwate, R. Raghavan and N. Dahanukar,
<date id="6106BC84814E4948FDAFBF46FC94029F" box="[638,780,625,650]" pageId="2" pageNumber="403" value="2014-06-06">6 June 2014</date>
; 2 ex., ZSI-WRC P 4360, Beri Baor, Ramnagar, Kolkata, West Bengal,
<collectingCountry id="6DAFDAD4814E4948FEE4BFA2FEF702BB" box="[309,367,661,686]" name="India" pageId="2" pageNumber="403">India</collectingCountry>
(
<geoCoordinate id="708CFC83814E4948FEAEBFA1FD8F02BB" box="[383,535,661,686]" direction="north" orientation="latitude" pageId="2" pageNumber="403" precision="15" value="22.908888">22° 54' 32&quot;N</geoCoordinate>
,
<geoCoordinate id="708CFC83814E4948FDF5BFA1FD2702B8" box="[548,703,661,686]" direction="east" orientation="longitude" pageId="2" pageNumber="403" precision="15" value="88.85389">88° 51' 14&quot; E</geoCoordinate>
,
<quantity id="D24037A1814E4948FD1CBFA2FD6102B8" box="[717,761,661,686]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="403" unit="m" value="5.0">5 m</quantity>
a.s.l.), coll. U. Katwate, R. Raghavan and N. Dahanukar,
<date id="6106BC84814E4948FA5FBFA2FE9502C7" pageId="2" pageNumber="403" value="2014-06-06">6 June 2014</date>
; 2 ex., DABFUK/FI/223, Beri Baor, Ramnagar, Kolkata, West Bengal,
<collectingCountry id="6DAFDAD4814E4948FBE3BF8EFBF402C7" box="[1074,1132,697,722]" name="India" pageId="2" pageNumber="403">India</collectingCountry>
(
<geoCoordinate id="708CFC83814E4948FBAABF8DFA8A02C7" box="[1147,1298,697,722]" direction="north" orientation="latitude" pageId="2" pageNumber="403" precision="15" value="22.908888">22° 54' 32&quot;N</geoCoordinate>
,
<geoCoordinate id="708CFC83814E4948FACCBF8DFF3202E0" direction="east" orientation="longitude" pageId="2" pageNumber="403" precision="15" value="88.85389">88° 51' 14&quot; E</geoCoordinate>
,
<quantity id="D24037A1814E4948FF6DBFEAFF7502E0" box="[188,237,733,758]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="403" unit="m" value="5.0">5 m</quantity>
a.s.l.), coll. U. Katwate, R. Raghavan and N. Dahanukar,
<date id="6106BC84814E4948FC65BFEAFBD402E3" box="[948,1100,733,758]" pageId="2" pageNumber="403" value="2014-06-06">6 June 2014</date>
; 1 ex., unregistered, Dhaka,
<collectingCountry id="6DAFDAD4814E4948FF46BE36FE85030F" box="[151,285,769,794]" name="Bangladesh" pageId="2" pageNumber="403">Bangladesh</collectingCountry>
, coll. Muntasir Akash,
<date id="6106BC84814E4948FDF0BE35FD54030F" box="[545,716,769,794]" pageId="2" pageNumber="403" value="2015-03-24">24 March 2015</date>
(only photograph examined).
</paragraph>
</subSubSection>
<subSubSection id="5DA2C9CF814E4948FF16BE12FDDE0400" pageId="2" pageNumber="403" type="diagnosis">
<paragraph id="15079A44814E4948FF16BE12FDDE0400" blockId="2.[151,1437,517,1946]" pageId="2" pageNumber="403">
<emphasis id="27CC4656814E4948FF16BE12FED9032B" bold="true" box="[199,321,805,830]" pageId="2" pageNumber="403">Diagnosis.</emphasis>
<taxonomicName id="D2B8E1C7814E4948FE9CBE12FE4D0328" box="[333,469,805,829]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="403" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC4656814E4948FE9CBE12FE4D0328" box="[333,469,805,829]" italics="true" pageId="2" pageNumber="403">Pethia ticto</emphasis>
</taxonomicName>
can be distinguished from all other species of
<taxonomicName id="D2B8E1C7814E4948FBC1BE12FBC10328" box="[1040,1113,805,829]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="403" phylum="Chordata" rank="genus">
<emphasis id="27CC4656814E4948FBC1BE12FBC10328" box="[1040,1113,805,829]" italics="true" pageId="2" pageNumber="403">Pethia</emphasis>
</taxonomicName>
by the combination of the following characters: barbels absent; lateral line incomplete, with 612 pored scales; 2326 scales in lateral-scale row; predorsal scales 9; scales in transverse series ½4/1/3½4; dorsal fin originating after pelvic-fin origin; gill rakers 7 on first ceratobranchial; 17 abdominal including 4 weberian and 13 caudal vertebrae including posteriormost compound centrum (i.e. 4+13+13); banding pattern includes a small black humeral spot covering third and fourth lateral-line scales; a prominent spot on the caudal peduncle on scales 1619 of lateral scale row; and dorsal fin with 2 oblique rows of black spots.
</paragraph>
</subSubSection>
<subSubSection id="5DA2C9CF814E4941FF16B916FBF604DC" lastPageId="11" lastPageNumber="412" pageId="2" pageNumber="403" type="description">
<paragraph id="15079A44814E4948FF16B916FDFF044B" blockId="2.[151,1437,517,1946]" pageId="2" pageNumber="403">
<emphasis id="27CC4656814E4948FF16B916FEC1042F" bold="true" box="[199,345,1057,1082]" pageId="2" pageNumber="403">Description.</emphasis>
Illustration and photograph of topotypes are provided in
<figureCitation id="8D8386C1814E4948FC0EB916FBD0042F" box="[991,1096,1057,1082]" captionStart="FIGURE 1" captionStartId="1.[151,250,1163,1185]" captionTargetBox="[194,1393,623,1141]" captionTargetId="figure@1.[194,1394,623,1141]" captionTargetPageId="1" captionText="FIGURE 1. Illustration of Pethia ticto based on topotype, male, BNHS FWF 127, 35.9 mm SL, showing distribution of prominent body pigmentation and banding pattern and lateral and transverse scale rows." httpUri="https://zenodo.org/record/233765/files/figure.png" pageId="2" pageNumber="403">Figures 1</figureCitation>
,
<figureCitation id="8D8386C1814E4948FB86B915FBFD042F" box="[1111,1125,1058,1082]" captionStart="FIGURE 2" captionStartId="4.[151,250,1439,1461]" captionTargetBox="[272,1313,194,1403]" captionTargetId="figure@4.[271,1315,193,1418]" captionTargetPageId="4" captionText="FIGURE 2. Pethia ticto. (A) Topotype, BNHS FWF, male, 127, 35.9 mm SL, from West Bengal, India. (B) Cyprinus tic to (iconotype) from Hamiltons (1822) illustration in the collection of the Linnean Society of London (courtesy: Linnean Society of London)." httpUri="https://zenodo.org/record/233766/files/figure.png" pageId="2" pageNumber="403">2</figureCitation>
AB, 3AB. Morphometric and meristic data are provided in
<tableCitation id="583AAFFF814E4948FDDFB972FDF9044B" box="[526,609,1093,1118]" captionStart="TABLE 1" captionStartId="3.[151,239,152,175]" captionTargetBox="[159,1168,234,1951]" captionTargetPageId="3" captionText="TABLE 1. Morphometric data of topotypic Pethia ticto (n = 13, BNHS FWF 127 to 131, WILD- 15 - PIS- 145 to 148, ZSI- WRC P 4360 and DABFUK / FI / 223." httpUri="http://table.plazi.org/id/41C7CACC814F4949FF46BDAFFD8400DA" pageId="2" pageNumber="403" tableUuid="41C7CACC814F4949FF46BDAFFD8400DA">Table 1</tableCitation>
.
</paragraph>
<paragraph id="15079A44814E4948FF16B95EFAAC04FB" blockId="2.[151,1437,517,1946]" pageId="2" pageNumber="403">Body short, deep, its length 2.8 to 3.2 times depth; compressed laterally; predorsal contour slightly convex, rising gradually up to dorsal-fin origin, thereafter sloping down towards hypural notch in a sharp concave slope. Ventral profile convex up to base of pectoral fin, then almost straight up to anterior extremity of anal-fin base, sloping gradually towards hypural notch. Caudal peduncle broad, short, its length 1.21.6 times its depth.</paragraph>
<paragraph id="15079A44814E4948FF16B9CEFE6A056B" blockId="2.[151,1437,517,1946]" pageId="2" pageNumber="403">Head small, laterally compressed. Snout pointed, smooth, its length less than eye diameter. Eyes large, midlaterally positioned, closer to snout tip than margin of operculum, diameter 0.81.3 interorbital width. Mouth small, terminal, ventrally U shaped, angle of gape not reaching to vertical from anterior margin of eye. Upper and lower lips thin. Barbels absent.</paragraph>
<paragraph id="15079A44814E4948FF16B8BEFB5D06D7" blockId="2.[151,1437,517,1946]" pageId="2" pageNumber="403">
Dorsal fin originating posterior to pelvic-fin origin, closer to tip of snout than to base of caudal fin, its distal margin concave, height 0.91.1 times head length. Dorsal fin with 4 unbranched (including 3 supernumerary rays) and 8 branched rays, last unbranched ray strong, its proximal half smooth, densely serrated posteriorly with 11 (1), 13 (3), 14 (1), 15 (1), 16 (2), 17 (1), 18 (1), 19 (2) or 21 (1) serrae on its distal half, 2 serrae on apex. Pectoral fin with one unbranched and 12 (3), 13 (3), 14 (1) or 15 (6) branched rays, its tip rounded, extending beyond pelvic-fin origin (
<figureCitation id="8D8386C1814E4948FF38BB0AFED20643" box="[233,330,1597,1622]" captionStart="FIGURE 1" captionStartId="1.[151,250,1163,1185]" captionTargetBox="[194,1393,623,1141]" captionTargetId="figure@1.[194,1394,623,1141]" captionTargetPageId="1" captionText="FIGURE 1. Illustration of Pethia ticto based on topotype, male, BNHS FWF 127, 35.9 mm SL, showing distribution of prominent body pigmentation and banding pattern and lateral and transverse scale rows." httpUri="https://zenodo.org/record/233765/files/figure.png" pageId="2" pageNumber="403">Figure 1</figureCitation>
,
<figureCitation id="8D8386C1814E4948FE8BBB09FEF00643" box="[346,360,1598,1622]" captionStart="FIGURE 2" captionStartId="4.[151,250,1439,1461]" captionTargetBox="[272,1313,194,1403]" captionTargetId="figure@4.[271,1315,193,1418]" captionTargetPageId="4" captionText="FIGURE 2. Pethia ticto. (A) Topotype, BNHS FWF, male, 127, 35.9 mm SL, from West Bengal, India. (B) Cyprinus tic to (iconotype) from Hamiltons (1822) illustration in the collection of the Linnean Society of London (courtesy: Linnean Society of London)." httpUri="https://zenodo.org/record/233766/files/figure.png" pageId="2" pageNumber="403">2</figureCitation>
A, 3A &amp; B). Pelvic fin with one unbranched and 7 branched rays, its tip rounded, not reaching vent when adpressed. Anal fin with 3 unbranched (including 2 supernumerary) and 5 branched rays, its distal margin concave, with rounded angles. Caudal fin forked, lobes representing more than half of fin length, tips pointed. Principal caudal-fin rays dorsally 9, ventrally 8; procurrent rays dorsally 7, ventrally 6.
</paragraph>
<paragraph id="15079A44814E4948FF16BBFAFCF50747" blockId="2.[151,1437,517,1946]" pageId="2" pageNumber="403">Lateral line incomplete with 6 (1), 7 (3), 9 (5), 11 (3) or 12 (1) pored scales. Lateral-scale row with 23 (3), 24 (1), 25 (6) or 26 (3) scales. Scales in transverse series ½4/1/3½ (3) or ½4/1/4 (10); predorsal scales 9; prepelvic scales 10 (9), 11 (2) or 12 (2); preanal scales 16 (4), 17 (8) or 19 (1); circumpeduncular scales 12. Pelvic axillary scale present, reaching to one-fourth adpressed pelvic-fin length.</paragraph>
<paragraph id="15079A44814E4948FF16BA6AFDED078F" blockId="2.[151,1437,517,1946]" pageId="2" pageNumber="403">
<emphasis id="27CC4656814E4948FF16BA6AFEDB0763" bold="true" box="[199,323,1885,1910]" pageId="2" pageNumber="403">Osteology.</emphasis>
Complete lateral view of the cleared and stained specimens of
<emphasis id="27CC4656814E4948FBC3BA69FBC50760" box="[1042,1117,1886,1909]" italics="true" pageId="2" pageNumber="403">P. t i c t o</emphasis>
(BNHS FWF 130, 131) are provided in
<figureCitation id="8D8386C1814E4948FECFBAB6FEE4078F" box="[286,380,1921,1946]" captionStart="FIGURE 4" captionStartId="6.[151,249,1233,1255]" captionTargetBox="[151,1375,197,1197]" captionTargetId="figure@6.[151,1435,175,1225]" captionTargetPageId="6" captionText="FIGURE 4. Lateral view of entire skeleton of topotypic Pethia ticto (A) male, BNHS FWF 130, 32.2 mm SL; (B) male, BNHS FWF 131, 30.7 mm SL. AF, anal fin; CF, caudal fin; DF, dorsal fin; En, epineural intermusculars; Ep, epipleural intermusculars; Nc, neurocranium; PcF, pectoral fin; PvF, pelvic fin; WeV, Weberian vertebrae." httpUri="https://zenodo.org/record/233768/files/figure.png" pageId="2" pageNumber="403">Figure 4</figureCitation>
A and B, respectively.
</paragraph>
<caption id="41C7CACC814F4949FF46BDAFFD8400DA" ID-Table-UUID="41C7CACC814F4949FF46BDAFFD8400DA" httpUri="http://table.plazi.org/id/41C7CACC814F4949FF46BDAFFD8400DA" pageId="3" pageNumber="404" targetBox="[159,1168,234,1951]" targetIsTable="true" targetPageId="3">
<paragraph id="15079A44814F4949FF46BDAFFD8400DA" blockId="3.[151,1436,151,207]" pageId="3" pageNumber="404">
<emphasis id="27CC4656814F4949FF46BDAFFE9000BA" bold="true" box="[151,264,152,175]" pageId="3" pageNumber="404">TABLE 1.</emphasis>
Morphometric data of topotypic
<taxonomicName id="D2B8E1C7814F4949FDB8BDA0FD7D00BA" box="[617,741,151,175]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="404" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC4656814F4949FDB8BDA0FD7D00BA" box="[617,741,151,175]" italics="true" pageId="3" pageNumber="404">Pethia ticto</emphasis>
</taxonomicName>
(n = 13, BNHS FWF 127 to 131, WILD-15-PIS-145 to 148, ZSI- WRC P 4360 and DABFUK/FI/223.
</paragraph>
</caption>
<paragraph id="15079A44814F4949FF4EBDDDFBEF078A" pageId="3" pageNumber="404">
<table id="67B868E4814FB6B5FF4EBDDDFB08078A" box="[159,1168,234,1951]" gridcols="3" gridrows="42" pageId="3" pageNumber="404">
<tr id="AB889806814FB6B5FF4EBDDDFB080115" box="[159,1168,234,256]" gridrow="0" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EBDDDFE450115" box="[159,477,234,256]" gridcol="0" gridrow="0" pageId="3" pageNumber="404">Character</th>
<th id="E859F17A814FB6B5FD0CBDDDFCDD0115" box="[733,837,234,256]" gridcol="1" gridrow="0" pageId="3" pageNumber="404">Mean (sd)</th>
<th id="E859F17A814FB6B5FBCCBDDDFB080115" box="[1053,1168,234,256]" gridcol="2" gridrow="0" pageId="3" pageNumber="404">Range</th>
</tr>
<tr id="AB889806814FB6B5FF4EBC23FB08013F" box="[159,1168,276,298]" gridrow="1" pageId="3" pageNumber="404" rowspan-1="1" rowspan-2="1">
<th id="E859F17A814FB6B5FF4EBC23FE45013F" box="[159,477,276,298]" gridcol="0" gridrow="1" pageId="3" pageNumber="404">Morphometrics</th>
</tr>
<tr id="AB889806814FB6B5FF4EBC0AFB080146" box="[159,1168,317,339]" gridrow="2" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EBC0AFE450146" box="[159,477,317,339]" gridcol="0" gridrow="2" pageId="3" pageNumber="404">Total length (mm)</th>
<td id="E859F17A814FB6B5FD0CBC0AFCDD0146" box="[733,837,317,339]" gridcol="1" gridrow="2" pageId="3" pageNumber="404">37.1 (4.4)</td>
<td id="E859F17A814FB6B5FBCCBC0AFB080146" box="[1053,1168,317,339]" gridcol="2" gridrow="2" pageId="3" pageNumber="404">32.047.2</td>
</tr>
<tr id="AB889806814FB6B5FF4EBC51FB080169" box="[159,1168,358,380]" gridrow="3" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EBC51FE450169" box="[159,477,358,380]" gridcol="0" gridrow="3" pageId="3" pageNumber="404">Standard length (mm, SL)</th>
<td id="E859F17A814FB6B5FD0CBC51FCDD0169" box="[733,837,358,380]" gridcol="1" gridrow="3" pageId="3" pageNumber="404">28.6 (3.0)</td>
<td id="E859F17A814FB6B5FBCCBC51FB080169" box="[1053,1168,358,380]" gridcol="2" gridrow="3" pageId="3" pageNumber="404">25.436.0</td>
</tr>
<tr id="AB889806814FB6B5FF4EBCA7FB0801B3" box="[159,1168,400,422]" gridrow="4" pageId="3" pageNumber="404" rowspan-1="1" rowspan-2="1">
<th id="E859F17A814FB6B5FF4EBCA7FE4501B3" box="[159,477,400,422]" gridcol="0" gridrow="4" pageId="3" pageNumber="404">% SL</th>
</tr>
<tr id="AB889806814FB6B5FF4EBC8EFB0801DA" box="[159,1168,441,463]" gridrow="5" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EBC8EFE4501DA" box="[159,477,441,463]" gridcol="0" gridrow="5" pageId="3" pageNumber="404">Head length (HL)</th>
<td id="E859F17A814FB6B5FD0CBC8EFCDD01DA" box="[733,837,441,463]" gridcol="1" gridrow="5" pageId="3" pageNumber="404">30.4 (1.7)</td>
<td id="E859F17A814FB6B5FBCCBC8EFB0801DA" box="[1053,1168,441,463]" gridcol="2" gridrow="5" pageId="3" pageNumber="404">27.633.7</td>
</tr>
<tr id="AB889806814FB6B5FF4EBCD5FB0801ED" box="[159,1168,482,504]" gridrow="6" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EBCD5FE4501ED" box="[159,477,482,504]" gridcol="0" gridrow="6" pageId="3" pageNumber="404">Body depth</th>
<td id="E859F17A814FB6B5FD0CBCD5FCDD01ED" box="[733,837,482,504]" gridcol="1" gridrow="6" pageId="3" pageNumber="404">33.6 (1.3)</td>
<td id="E859F17A814FB6B5FBCCBCD5FB0801ED" box="[1053,1168,482,504]" gridcol="2" gridrow="6" pageId="3" pageNumber="404">31.535.8</td>
</tr>
<tr id="AB889806814FB6B5FF4EBF3BFB080237" box="[159,1168,524,546]" gridrow="7" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EBF3BFE450237" box="[159,477,524,546]" gridcol="0" gridrow="7" pageId="3" pageNumber="404">Body width at Dorsal fin origin</th>
<td id="E859F17A814FB6B5FD0CBF3BFCDD0237" box="[733,837,524,546]" gridcol="1" gridrow="7" pageId="3" pageNumber="404">12.9 (1.4)</td>
<td id="E859F17A814FB6B5FBCCBF3BFB080237" box="[1053,1168,524,546]" gridcol="2" gridrow="7" pageId="3" pageNumber="404">10.715.4</td>
</tr>
<tr id="AB889806814FB6B5FF4EBF02FB08025E" box="[159,1168,565,587]" gridrow="8" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EBF02FE45025E" box="[159,477,565,587]" gridcol="0" gridrow="8" pageId="3" pageNumber="404">Body width at anal fin origin</th>
<td id="E859F17A814FB6B5FD0CBF02FCDD025E" box="[733,837,565,587]" gridcol="1" gridrow="8" pageId="3" pageNumber="404">8.8 (1.9)</td>
<td id="E859F17A814FB6B5FBCCBF02FB08025E" box="[1053,1168,565,587]" gridcol="2" gridrow="8" pageId="3" pageNumber="404">5.311.1</td>
</tr>
<tr id="AB889806814FB6B5FF4EBF69FB080261" box="[159,1168,606,628]" gridrow="9" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EBF69FE450261" box="[159,477,606,628]" gridcol="0" gridrow="9" pageId="3" pageNumber="404">Predorsal distance</th>
<td id="E859F17A814FB6B5FD0CBF69FCDD0261" box="[733,837,606,628]" gridcol="1" gridrow="9" pageId="3" pageNumber="404">49.6 (1.3)</td>
<td id="E859F17A814FB6B5FBCCBF69FB080261" box="[1053,1168,606,628]" gridcol="2" gridrow="9" pageId="3" pageNumber="404">47.251.7</td>
</tr>
<tr id="AB889806814FB6B5FF4EBFBFFB08028B" box="[159,1168,648,670]" gridrow="10" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EBFBFFE45028B" box="[159,477,648,670]" gridcol="0" gridrow="10" pageId="3" pageNumber="404">Dorsal to hypural distance</th>
<td id="E859F17A814FB6B5FD0CBFBFFCDD028B" box="[733,837,648,670]" gridcol="1" gridrow="10" pageId="3" pageNumber="404">53.8 (1.4)</td>
<td id="E859F17A814FB6B5FBCCBFBFFB08028B" box="[1053,1168,648,670]" gridcol="2" gridrow="10" pageId="3" pageNumber="404">52.157.3</td>
</tr>
<tr id="AB889806814FB6B5FF4EBF86FB0802D2" box="[159,1168,689,711]" gridrow="11" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EBF86FE4502D2" box="[159,477,689,711]" gridcol="0" gridrow="11" pageId="3" pageNumber="404">Prepelvic distance</th>
<td id="E859F17A814FB6B5FD0CBF86FCDD02D2" box="[733,837,689,711]" gridcol="1" gridrow="11" pageId="3" pageNumber="404">48.5 (1.8)</td>
<td id="E859F17A814FB6B5FBCCBF86FB0802D2" box="[1053,1168,689,711]" gridcol="2" gridrow="11" pageId="3" pageNumber="404">45.551.6</td>
</tr>
<tr id="AB889806814FB6B5FF4EBFEDFB0802E5" box="[159,1168,730,752]" gridrow="12" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EBFEDFE4502E5" box="[159,477,730,752]" gridcol="0" gridrow="12" pageId="3" pageNumber="404">Preanal distance</th>
<td id="E859F17A814FB6B5FD0CBFEDFCDD02E5" box="[733,837,730,752]" gridcol="1" gridrow="12" pageId="3" pageNumber="404">68.9 (1.2)</td>
<td id="E859F17A814FB6B5FBCCBFEDFB0802E5" box="[1053,1168,730,752]" gridcol="2" gridrow="12" pageId="3" pageNumber="404">67.671.5</td>
</tr>
<tr id="AB889806814FB6B5FF4EBE33FB08030F" box="[159,1168,772,794]" gridrow="13" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EBE33FE45030F" box="[159,477,772,794]" gridcol="0" gridrow="13" pageId="3" pageNumber="404">Prepectoral distance</th>
<td id="E859F17A814FB6B5FD0CBE33FCDD030F" box="[733,837,772,794]" gridcol="1" gridrow="13" pageId="3" pageNumber="404">29.9 (1.8)</td>
<td id="E859F17A814FB6B5FBCCBE33FB08030F" box="[1053,1168,772,794]" gridcol="2" gridrow="13" pageId="3" pageNumber="404">27.733.3</td>
</tr>
<tr id="AB889806814FB6B5FF4EBE1AFB080356" box="[159,1168,813,835]" gridrow="14" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EBE1AFE450356" box="[159,477,813,835]" gridcol="0" gridrow="14" pageId="3" pageNumber="404">Dorsal fin length</th>
<td id="E859F17A814FB6B5FD0CBE1AFCDD0356" box="[733,837,813,835]" gridcol="1" gridrow="14" pageId="3" pageNumber="404">29.6 (3.3)</td>
<td id="E859F17A814FB6B5FBCCBE1AFB080356" box="[1053,1168,813,835]" gridcol="2" gridrow="14" pageId="3" pageNumber="404">24.334.7</td>
</tr>
<tr id="AB889806814FB6B5FF4EBE61FB080379" box="[159,1168,854,876]" gridrow="15" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EBE61FE450379" box="[159,477,854,876]" gridcol="0" gridrow="15" pageId="3" pageNumber="404">Dorsal fin spine length</th>
<td id="E859F17A814FB6B5FD0CBE61FCDD0379" box="[733,837,854,876]" gridcol="1" gridrow="15" pageId="3" pageNumber="404">22.1 (3.6)</td>
<td id="E859F17A814FB6B5FBCCBE61FB080379" box="[1053,1168,854,876]" gridcol="2" gridrow="15" pageId="3" pageNumber="404">16.629.6</td>
</tr>
<tr id="AB889806814FB6B5FF4EBEB7FB080383" box="[159,1168,896,918]" gridrow="16" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EBEB7FE450383" box="[159,477,896,918]" gridcol="0" gridrow="16" pageId="3" pageNumber="404">Length of Dorsal fin base</th>
<td id="E859F17A814FB6B5FD0CBEB7FCDD0383" box="[733,837,896,918]" gridcol="1" gridrow="16" pageId="3" pageNumber="404">18.7 (1.9)</td>
<td id="E859F17A814FB6B5FBCCBEB7FB080383" box="[1053,1168,896,918]" gridcol="2" gridrow="16" pageId="3" pageNumber="404">15.422.1</td>
</tr>
<tr id="AB889806814FB6B5FF4EBE9EFB0803AA" box="[159,1168,937,959]" gridrow="17" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EBE9EFE4503AA" box="[159,477,937,959]" gridcol="0" gridrow="17" pageId="3" pageNumber="404">Pectoralfin length</th>
<td id="E859F17A814FB6B5FD0CBE9EFCDD03AA" box="[733,837,937,959]" gridcol="1" gridrow="17" pageId="3" pageNumber="404">22.7 (2.2)</td>
<td id="E859F17A814FB6B5FBCCBE9EFB0803AA" box="[1053,1168,937,959]" gridcol="2" gridrow="17" pageId="3" pageNumber="404">19.326.3</td>
</tr>
<tr id="AB889806814FB6B5FF4EBEE5FB0803FD" box="[159,1168,978,1000]" gridrow="18" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EBEE5FE4503FD" box="[159,477,978,1000]" gridcol="0" gridrow="18" pageId="3" pageNumber="404">Analfin depth</th>
<td id="E859F17A814FB6B5FD0CBEE5FCDD03FD" box="[733,837,978,1000]" gridcol="1" gridrow="18" pageId="3" pageNumber="404">19.1 (2.1)</td>
<td id="E859F17A814FB6B5FBCCBEE5FB0803FD" box="[1053,1168,978,1000]" gridcol="2" gridrow="18" pageId="3" pageNumber="404">14.321.7</td>
</tr>
<tr id="AB889806814FB6B5FF4EBECBFB080407" box="[159,1168,1020,1042]" gridrow="19" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EBECBFE450407" box="[159,477,1020,1042]" gridcol="0" gridrow="19" pageId="3" pageNumber="404">Caudalpeduncle length</th>
<td id="E859F17A814FB6B5FD0CBECBFCDD0407" box="[733,837,1020,1042]" gridcol="1" gridrow="19" pageId="3" pageNumber="404">20.4 (1.3)</td>
<td id="E859F17A814FB6B5FBCCBECBFB080407" box="[1053,1168,1020,1042]" gridcol="2" gridrow="19" pageId="3" pageNumber="404">17.022.2</td>
</tr>
<tr id="AB889806814FB6B5FF4EB912FB08042E" box="[159,1168,1061,1083]" gridrow="20" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EB912FE45042E" box="[159,477,1061,1083]" gridcol="0" gridrow="20" pageId="3" pageNumber="404">Caudalpeduncle depth</th>
<td id="E859F17A814FB6B5FD0CB912FCDD042E" box="[733,837,1061,1083]" gridcol="1" gridrow="20" pageId="3" pageNumber="404">14.0 (0.7)</td>
<td id="E859F17A814FB6B5FBCCB912FB08042E" box="[1053,1168,1061,1083]" gridcol="2" gridrow="20" pageId="3" pageNumber="404">12.715.1</td>
</tr>
<tr id="AB889806814FB6B5FF4EB979FB080471" box="[159,1168,1102,1124]" gridrow="21" pageId="3" pageNumber="404" rowspan-1="1" rowspan-2="1">
<th id="E859F17A814FB6B5FF4EB979FE450471" box="[159,477,1102,1124]" gridcol="0" gridrow="21" pageId="3" pageNumber="404">% HL</th>
</tr>
<tr id="AB889806814FB6B5FF4EB94FFB08049B" box="[159,1168,1144,1166]" gridrow="22" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EB94FFE45049B" box="[159,477,1144,1166]" gridcol="0" gridrow="22" pageId="3" pageNumber="404">Head depth</th>
<td id="E859F17A814FB6B5FD0CB94FFCDD049B" box="[733,837,1144,1166]" gridcol="1" gridrow="22" pageId="3" pageNumber="404">77.1 (4.8)</td>
<td id="E859F17A814FB6B5FBCCB94FFB08049B" box="[1053,1168,1144,1166]" gridcol="2" gridrow="22" pageId="3" pageNumber="404">69.987.0</td>
</tr>
<tr id="AB889806814FB6B5FF4EB996FB0804A2" box="[159,1168,1185,1207]" gridrow="23" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EB996FE4504A2" box="[159,477,1185,1207]" gridcol="0" gridrow="23" pageId="3" pageNumber="404">Head width</th>
<td id="E859F17A814FB6B5FD0CB996FCDD04A2" box="[733,837,1185,1207]" gridcol="1" gridrow="23" pageId="3" pageNumber="404">49.6 (2.3)</td>
<td id="E859F17A814FB6B5FBCCB996FB0804A2" box="[1053,1168,1185,1207]" gridcol="2" gridrow="23" pageId="3" pageNumber="404">45.954.4</td>
</tr>
<tr id="AB889806814FB6B5FF4EB9FDFB0804F5" box="[159,1168,1226,1248]" gridrow="24" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EB9FDFE4504F5" box="[159,477,1226,1248]" gridcol="0" gridrow="24" pageId="3" pageNumber="404">Snout length</th>
<td id="E859F17A814FB6B5FD0CB9FDFCDD04F5" box="[733,837,1226,1248]" gridcol="1" gridrow="24" pageId="3" pageNumber="404">24.5 (1.7)</td>
<td id="E859F17A814FB6B5FBCCB9FDFB0804F5" box="[1053,1168,1226,1248]" gridcol="2" gridrow="24" pageId="3" pageNumber="404">22.227.6</td>
</tr>
<tr id="AB889806814FB6B5FF4EB9C3FB08051F" box="[159,1168,1268,1290]" gridrow="25" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EB9C3FE45051F" box="[159,477,1268,1290]" gridcol="0" gridrow="25" pageId="3" pageNumber="404">Eye diameter</th>
<td id="E859F17A814FB6B5FD0CB9C3FCDD051F" box="[733,837,1268,1290]" gridcol="1" gridrow="25" pageId="3" pageNumber="404">34.3 (2.3)</td>
<td id="E859F17A814FB6B5FBCCB9C3FB08051F" box="[1053,1168,1268,1290]" gridcol="2" gridrow="25" pageId="3" pageNumber="404">30.837.1</td>
</tr>
<tr id="AB889806814FB6B5FF4EB82AFB080526" box="[159,1168,1309,1331]" gridrow="26" pageId="3" pageNumber="404">
<th id="E859F17A814FB6B5FF4EB82AFE450526" box="[159,477,1309,1331]" gridcol="0" gridrow="26" pageId="3" pageNumber="404">Inter orbital width</th>
<td id="E859F17A814FB6B5FD0CB82AFCDD0526" box="[733,837,1309,1331]" gridcol="1" gridrow="26" pageId="3" pageNumber="404">35.3 (3.3)</td>
<td id="E859F17A814FB6B5FBCCB82AFB080526" box="[1053,1168,1309,1331]" gridcol="2" gridrow="26" pageId="3" pageNumber="404">29.539.4</td>
</tr>
<tr id="AB889806814FB6B5FF4EB871FB080549" box="[159,1168,1350,1372]" gridrow="27" pageId="3" pageNumber="404" rowspan-1="1" rowspan-2="1">
<th id="E859F17A814FB6B5FF4EB871FE450549" box="[159,477,1350,1372]" gridcol="0" gridrow="27" pageId="3" pageNumber="404">Meristics</th>
</tr>
<tr id="AB889806814FB6B5FF4EB847FB080593" box="[159,1168,1392,1414]" gridrow="28" pageId="3" pageNumber="404" rowspan-1="1">
<th id="E859F17A814FB6B5FF4EB847FE450593" box="[159,477,1392,1414]" gridcol="0" gridrow="28" pageId="3" pageNumber="404">Lateral row scales</th>
<td id="E859F17A814FB6B5FBCCB847FB080593" box="[1053,1168,1392,1414]" gridcol="2" gridrow="28" pageId="3" pageNumber="404">2326</td>
</tr>
<tr id="AB889806814FB6B5FF4EB8AEFB0805BA" box="[159,1168,1433,1455]" gridrow="29" pageId="3" pageNumber="404" rowspan-1="1">
<th id="E859F17A814FB6B5FF4EB8AEFE4505BA" box="[159,477,1433,1455]" gridcol="0" gridrow="29" pageId="3" pageNumber="404">Number of lateral line pores</th>
<td id="E859F17A814FB6B5FBCCB8AEFB0805BA" box="[1053,1168,1433,1455]" gridcol="2" gridrow="29" pageId="3" pageNumber="404">612</td>
</tr>
<tr id="AB889806814FB6B5FF4EB8F5FB0805CD" box="[159,1168,1474,1496]" gridrow="30" pageId="3" pageNumber="404" rowspan-1="1">
<th id="E859F17A814FB6B5FF4EB8F5FE4505CD" box="[159,477,1474,1496]" gridcol="0" gridrow="30" pageId="3" pageNumber="404">Dorsalfin spine serrae</th>
<td id="E859F17A814FB6B5FBCCB8F5FB0805CD" box="[1053,1168,1474,1496]" gridcol="2" gridrow="30" pageId="3" pageNumber="404">1121</td>
</tr>
<tr id="AB889806814FB6B5FF4EB8DBFB080617" box="[159,1168,1516,1538]" gridrow="31" pageId="3" pageNumber="404" rowspan-1="1">
<th id="E859F17A814FB6B5FF4EB8DBFE450617" box="[159,477,1516,1538]" gridcol="0" gridrow="31" pageId="3" pageNumber="404">Transverse scale rows</th>
<td id="E859F17A814FB6B5FBCCB8DBFB080617" box="[1053,1168,1516,1538]" gridcol="2" gridrow="31" pageId="3" pageNumber="404">½4/1/3½4</td>
</tr>
<tr id="AB889806814FB6B5FF4EBB22FB08063E" box="[159,1168,1557,1579]" gridrow="32" pageId="3" pageNumber="404" rowspan-1="1">
<th id="E859F17A814FB6B5FF4EBB22FE45063E" box="[159,477,1557,1579]" gridcol="0" gridrow="32" pageId="3" pageNumber="404">Predorsal scales</th>
<td id="E859F17A814FB6B5FBCCBB22FB08063E" box="[1053,1168,1557,1579]" gridcol="2" gridrow="32" pageId="3" pageNumber="404">9</td>
</tr>
<tr id="AB889806814FB6B5FF4EBB09FB080641" box="[159,1168,1598,1620]" gridrow="33" pageId="3" pageNumber="404" rowspan-1="1">
<th id="E859F17A814FB6B5FF4EBB09FE450641" box="[159,477,1598,1620]" gridcol="0" gridrow="33" pageId="3" pageNumber="404">Prepelvic scales</th>
<td id="E859F17A814FB6B5FBCCBB09FB080641" box="[1053,1168,1598,1620]" gridcol="2" gridrow="33" pageId="3" pageNumber="404">1012</td>
</tr>
<tr id="AB889806814FB6B5FF4EBB5FFB08066B" box="[159,1168,1640,1662]" gridrow="34" pageId="3" pageNumber="404" rowspan-1="1">
<th id="E859F17A814FB6B5FF4EBB5FFE45066B" box="[159,477,1640,1662]" gridcol="0" gridrow="34" pageId="3" pageNumber="404">Preanal scales</th>
<td id="E859F17A814FB6B5FBCCBB5FFB08066B" box="[1053,1168,1640,1662]" gridcol="2" gridrow="34" pageId="3" pageNumber="404">1619</td>
</tr>
<tr id="AB889806814FB6B5FF4EBBA6FB0806B2" box="[159,1168,1681,1703]" gridrow="35" pageId="3" pageNumber="404" rowspan-1="1">
<th id="E859F17A814FB6B5FF4EBBA6FE4506B2" box="[159,477,1681,1703]" gridcol="0" gridrow="35" pageId="3" pageNumber="404">Circumpeduncular scales</th>
<td id="E859F17A814FB6B5FBCCBBA6FB0806B2" box="[1053,1168,1681,1703]" gridcol="2" gridrow="35" pageId="3" pageNumber="404">12</td>
</tr>
<tr id="AB889806814FB6B5FF4EBB8DFB0806C5" box="[159,1168,1722,1744]" gridrow="36" pageId="3" pageNumber="404" rowspan-1="1">
<th id="E859F17A814FB6B5FF4EBB8DFE4506C5" box="[159,477,1722,1744]" gridcol="0" gridrow="36" pageId="3" pageNumber="404">Dorsalfin rays</th>
<td id="E859F17A814FB6B5FBCCBB8DFB0806C5" box="[1053,1168,1722,1744]" gridcol="2" gridrow="36" pageId="3" pageNumber="404">iii8</td>
</tr>
<tr id="AB889806814FB6B5FF4EBBD3FB0806EF" box="[159,1168,1764,1786]" gridrow="37" pageId="3" pageNumber="404" rowspan-1="1">
<th id="E859F17A814FB6B5FF4EBBD3FE4506EF" box="[159,477,1764,1786]" gridcol="0" gridrow="37" pageId="3" pageNumber="404">Pectoralfin rays</th>
<td id="E859F17A814FB6B5FBCCBBD3FB0806EF" box="[1053,1168,1764,1786]" gridcol="2" gridrow="37" pageId="3" pageNumber="404">i1215</td>
</tr>
<tr id="AB889806814FB6B5FF4EBA3AFB080736" box="[159,1168,1805,1827]" gridrow="38" pageId="3" pageNumber="404" rowspan-1="1">
<th id="E859F17A814FB6B5FF4EBA3AFE450736" box="[159,477,1805,1827]" gridcol="0" gridrow="38" pageId="3" pageNumber="404">Pelvicfin rays</th>
<td id="E859F17A814FB6B5FBCCBA3AFB080736" box="[1053,1168,1805,1827]" gridcol="2" gridrow="38" pageId="3" pageNumber="404">i7</td>
</tr>
<tr id="AB889806814FB6B5FF4EBA01FB080759" box="[159,1168,1846,1868]" gridrow="39" pageId="3" pageNumber="404" rowspan-1="1">
<th id="E859F17A814FB6B5FF4EBA01FE450759" box="[159,477,1846,1868]" gridcol="0" gridrow="39" pageId="3" pageNumber="404">Analfin rays</th>
<td id="E859F17A814FB6B5FBCCBA01FB080759" box="[1053,1168,1846,1868]" gridcol="2" gridrow="39" pageId="3" pageNumber="404">iii5</td>
</tr>
<tr id="AB889806814FB6B5FF4EBA57FB080763" box="[159,1168,1888,1910]" gridrow="40" pageId="3" pageNumber="404" rowspan-1="1">
<th id="E859F17A814FB6B5FF4EBA57FE450763" box="[159,477,1888,1910]" gridcol="0" gridrow="40" pageId="3" pageNumber="404">Caudalfin rays (procurrent)</th>
<td id="E859F17A814FB6B5FBCCBA57FB080763" box="[1053,1168,1888,1910]" gridcol="2" gridrow="40" pageId="3" pageNumber="404">46+46</td>
</tr>
<tr id="AB889806814FB6B5FF4EBABEFB08078A" box="[159,1168,1929,1951]" gridrow="41" pageId="3" pageNumber="404" rowspan-1="1">
<th id="E859F17A814FB6B5FF4EBABEFE45078A" box="[159,477,1929,1951]" gridcol="0" gridrow="41" pageId="3" pageNumber="404">Caudalfin rays (principal)</th>
<td id="E859F17A814FB6B5FBCCBABEFB08078A" box="[1053,1168,1929,1951]" gridcol="2" gridrow="41" pageId="3" pageNumber="404">89+89</td>
</tr>
</table>
</paragraph>
<caption id="41C7CACC8148494EFF46B8A8FE8805E7" httpUri="https://zenodo.org/record/233766/files/figure.png" pageId="4" pageNumber="405" targetBox="[272,1313,194,1403]" targetPageId="4">
<paragraph id="15079A448148494EFF46B8A8FE8805E7" blockId="4.[151,1436,1439,1522]" pageId="4" pageNumber="405">
<emphasis id="27CC46568148494EFF46B8A8FE8D05A1" bold="true" box="[151,277,1439,1461]" pageId="4" pageNumber="405">FIGURE 2.</emphasis>
<taxonomicName id="D2B8E1C78148494EFECCB8A8FE0D05A3" box="[285,405,1439,1462]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="405" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC46568148494EFECCB8A8FE0D05A3" box="[285,405,1439,1462]" italics="true" pageId="4" pageNumber="405">Pethia ticto</emphasis>
</taxonomicName>
. (A) Topotype, BNHS FWF, male, 127, 35.9 mm SL, from West Bengal, India. (B)
<taxonomicName id="D2B8E1C78148494EFAD8B897FA1F05A3" box="[1289,1415,1440,1462]" class="Actinopterygii" family="Cyprinidae" genus="Cyprinus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="405" phylum="Chordata" rank="species" species="tic">
<emphasis id="27CC46568148494EFAD8B897FA1F05A3" box="[1289,1415,1440,1462]" italics="true" pageId="4" pageNumber="405">Cyprinus tic</emphasis>
</taxonomicName>
to (iconotype) from Hamiltons (1822) illustration in the collection of the Linnean Society of London (courtesy: Linnean Society of London).
</paragraph>
</caption>
<paragraph id="15079A448148494FFF16BB2DFEF001E0" blockId="4.[151,1437,1562,2019]" lastBlockId="5.[151,1436,151,824]" lastPageId="5" lastPageNumber="406" pageId="4" pageNumber="405">
<emphasis id="27CC46568148494EFF16BB2DFEE30626" bold="true" box="[199,379,1562,1587]" pageId="4" pageNumber="405">
<emphasis id="27CC46568148494EFF16BB2DFEEA0626" bold="true" box="[199,370,1562,1587]" italics="true" pageId="4" pageNumber="405">Neurocranium</emphasis>
:
</emphasis>
Dorsal, lateral and ventral view of neurocranium provided in Figure 5AC. In general appearance, it is broad and wide at otic region, and narrow and slopes sloping dorsally towards ethmoid region. Ethmoid region is composed of mesethmoid, prethmoid, ethmoid, kinethmoid (Figure 5A), vomer and large winged lateral ethmoid. Ethmoid region slopes down anteroventrally, meeting the triangular mesethmoid, which is placed between lateral wings of considerably cartilaginous ethmoid portion. Lateral ethmoid contributes large portion in the ethmoid complex, clearly visible, lateral processes well developed, tapers and slopes down anterolaterally, covers anterior most portion of orbital section. Prethmoid closely attached with anterior most tips of vomer. Nasals are in the form of small bony plate, anterior down to the lateral ethmoid. Frontals large and contributes as the largest anterior portion of neurocranium, with a clear supraorbital lateral-line canal running along its lateral margin. Parietals large, with a sensory canal running on lateroposterior side forming the distal margin of neurocranium. Orbitosphenoid large, distinct, attached dorsally with fontanel and ventrally with dorsovertical flap of parasphenoid. Pterosphenoid pierced, anteriorly attached with distal margin of orbitosphenoid. Parasphenoid long, elongated with dorsovertical flap, attaches anteriorly with vomer and cartilaginous trabecula communis, runs posteriorly till otic region. Prootic with large auditory foramen seems to be the largest bone of the ventral surface of cranium, attached posterolaterally with anterior most part of the subtemporal fossa. Autopterotic forms the posterolateral corner of the neurocranium, ventrally articulates with hyomandibular. On the ventral side of cranium, autopterotic forms the posterolateral wall of the deep subtemporal fossa. Supraoccipital attaches with the parietal on the anterior side, dorsally concave, spine well developed but does not supersedes weberian complex. Exoccipital large, well ossified with little cartilaginous remains, forms posterior wall of the otic capsule and the posterodorsal part of the otic bulla. Basioccipital large, covers ventral portion of the otic bulla, and the articulation between the neurocranium and the first vertebra, it is a posteroventral most point of the occipital region. Basioccipital process well developed, elongated with round concave masticatory plate on the anterior side and it reaches half of weberian complex ventrally.
</paragraph>
<paragraph id="15079A448149494FFF16BCC8FB6C032D" blockId="5.[151,1436,151,824]" pageId="5" pageNumber="406">
<emphasis id="27CC46568149494FFF16BCC8FD12020D" bold="true" box="[199,650,511,536]" pageId="5" pageNumber="406">
<emphasis id="27CC46568149494FFF16BCC8FD19020D" bold="true" box="[199,641,511,536]" italics="true" pageId="5" pageNumber="406">Hyopalatine arch and opercular series</emphasis>
:
</emphasis>
See Figure 5D. The dorsal edge of the hyomandibular articulates with the autosphenotic, the autopterotic, and the prootic cranium bones and unites with the opercle in anterodorsal portion. Laterally, the hyomandibular overlaps with the posterodorsal margin of the preopercle. Opercle overlaps with interopercle and subopercle anteroventrally. The quadrate well ossified, zone of cartilage, a remnant of the palatoquadrate cartilage is distinctly visible. Its large posteroventral process extends laterally and overlaps the anterior most point of the preopercle. On the anterior side, articular condyle of the quadrate articulates strongly with the anguloarticular. The metapterygoid is well ossified, ventrally attaches with the quadrate on anterior, preopercle and hyomandibular on posterior and overlaps laterally with symplectic. Endopterygoid and ectopterygoid well ossified like metapterygoid and overlaps on dorsal and anterior face of quadrate.
</paragraph>
<caption id="41C7CACC8149494FFF46BAADFB5807DB" httpUri="https://zenodo.org/record/233767/files/figure.png" pageId="5" pageNumber="406" targetBox="[382,1204,871,1923]" targetPageId="5">
<paragraph id="15079A448149494FFF46BAADFB5807DB" blockId="5.[151,1436,1946,1998]" pageId="5" pageNumber="406">
<emphasis id="27CC46568149494FFF46BAADFE8B07BA" bold="true" box="[151,275,1946,1968]" pageId="5" pageNumber="406">FIGURE 3.</emphasis>
Topotype of
<taxonomicName id="D2B8E1C78149494FFE4DBAADFD8907A5" box="[412,529,1946,1968]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="406" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC46568149494FFE4DBAADFD8907A5" box="[412,529,1946,1968]" italics="true" pageId="5" pageNumber="406">Pethia ticto</emphasis>
</taxonomicName>
in life (A) from Beri Baor, West Bangal, India (BNHS FWF 127, 35.9 mm SL), humeral and caudal spots disappear immediately after collection, and (B) from Dhaka, Bangladesh (unregistered).
</paragraph>
</caption>
<caption id="41C7CACC814A494CFF46B9E6FC200531" httpUri="https://zenodo.org/record/233768/files/figure.png" pageId="6" pageNumber="407" targetBox="[151,1375,197,1197]" targetPageId="6">
<paragraph id="15079A44814A494CFF46B9E6FC200531" blockId="6.[151,1436,1233,1316]" pageId="6" pageNumber="407">
<emphasis id="27CC4656814A494CFF46B9E6FE8904F3" bold="true" box="[151,273,1233,1255]" pageId="6" pageNumber="407">FIGURE 4.</emphasis>
Lateral view of entire skeleton of topotypic
<taxonomicName id="D2B8E1C7814A494CFD03B9E6FCDF04F2" box="[722,839,1233,1255]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC4656814A494CFD03B9E6FCDF04F2" box="[722,839,1233,1255]" italics="true" pageId="6" pageNumber="407">Pethia ticto</emphasis>
</taxonomicName>
(A) male, BNHS FWF 130, 32.2 mm SL; (B) male, BNHS FWF 131, 30.7 mm SL. AF, anal fin; CF, caudal fin; DF, dorsal fin; En, epineural intermusculars; Ep, epipleural intermusculars; Nc, neurocranium; PcF, pectoral fin; PvF, pelvic fin; WeV, Weberian vertebrae.
</paragraph>
</caption>
<paragraph id="15079A44814A494CFF16B865FB3C0657" blockId="6.[151,1437,1361,2034]" pageId="6" pageNumber="407">Upper jaw consists of premaxilla and maxilla, relatively large and well ossified. Premaxilla proportionately smaller than maxilla curved anteriorly and free of any joints with maxilla and dentary, while only attached with strong ligaments. Premaxilla articulates with the dentary on posteroventral surface. The lower jaw comprises Meckels cartilage, dentary, anguloarticular and the coronomeckelian. Dentary articulates with maxilla on dorsolateral side, broad on posterior edge and pointed towards anterior edge. Coronomeckelian bone presents on dorsal surface of remnant of rod shape Meckels cartilage. Kinethmoid a small bone contributing to the ethmoid complex is present in association with maxillae and premaxilla, bound with small ligaments.</paragraph>
<paragraph id="15079A44814A494CFF16BB7BFE2F072B" blockId="6.[151,1437,1361,2034]" pageId="6" pageNumber="407">
<emphasis id="27CC4656814A494CFF16BB7BFD070673" bold="true" box="[199,671,1612,1638]" pageId="6" pageNumber="407">
<emphasis id="27CC4656814A494CFF16BB7BFD0F0670" bold="true" box="[199,663,1612,1637]" italics="true" pageId="6" pageNumber="407">Infraorbital series and scleral skeleton</emphasis>
:
</emphasis>
See Figure 5E &amp; F. Five infraorbital bones present, bearing the laterosensory canal which encircles the orbital margin. Infraorbital 1 well developed, wider than infraorbital 2. Infraorbital 2 with laterosensory canal appears like a thin tube, encircles the orbital margin. Infraorbital 3, largest among all and overlaps preopercle. Infraorbital 4 well developed but less wide than infraorbital 1 and 3, infraorbital 5 very small. Supraorbital large and well ossified, covers eye dorsally. Infraorbital 6 absent. Scleral bones well ossified, covers proximal and distal perimeter of eye sclera in lateral view, dorsal and ventral perimeter of eye is cartilaginous (Figure 5E).
</paragraph>
<paragraph id="15079A44814A494CFF16BA7FFD0507E7" blockId="6.[151,1437,1361,2034]" pageId="6" pageNumber="407">
<emphasis id="27CC4656814A494CFF16BA7FFECA0774" bold="true" box="[199,338,1864,1889]" italics="true" pageId="6" pageNumber="407">Hyoid arch:</emphasis>
See Figure 5G &amp; H. Anterior ceratohyal distinct, broad posteriorly, narrows down anteriorly. It is attached anteriorly with dorsal hypohyal and ventral hypohyal and posteriorly with the posterior ceratohyal. Dorsal and ventral hypohyal are small and well ossified with little or no cartilaginous remnants. Large urohyal, with three well spread lateral flanges, attached anteriorly to ventral hypohyal with ligaments. Three well developed thin branchiostegal rays in each half of hyoid arch.
</paragraph>
<paragraph id="15079A44814B494DFF46B804FED106AE" blockId="7.[151,1436,1331,1723]" pageId="7" pageNumber="408">
<emphasis id="27CC4656814B494DFF46B804FE8B055D" bold="true" box="[151,275,1331,1353]" pageId="7" pageNumber="408">FIGURE 5.</emphasis>
<taxonomicName id="D2B8E1C7814B494DFECAB804FE09055C" box="[283,401,1331,1353]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="408" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC4656814B494DFECAB804FE09055C" box="[283,401,1331,1353]" italics="true" pageId="7" pageNumber="408">Pethia ticto</emphasis>
</taxonomicName>
, topotype, male, BNHS FWF 130,
<quantity id="D24037A1814B494DFD2EB804FCC2055C" box="[767,858,1331,1353]" metricMagnitude="-2" metricUnit="m" metricValue="3.22" pageId="7" pageNumber="408" unit="mm" value="32.2">32.2 mm</quantity>
SL; (A) dorsal neurocranium, (B) lateral neurocranium, (C) ventral neurocranium, (D) hyopalatine arch, (E) scleral cartilage and bone; (F) hyoid arch, dorsal view; (G) infraorbital series, lateral view; (H) urohyal bone, (I) Gill arch dorsal view, (J) ventral gill arches dorsal view, (K) dorsal gill arches ventral view and (L-M) 5th ceratobranchial showing pharyngeal teeth in two different angles. Apto, autopterotic; Asph, autosphenotic; Boc, basioccipital; BocPr, basioccipital process; E, ethmoid; Exoc, exoccipital; Epo, epioccipital; Fr, frontal; LE, lateral ethmoid; MaPl, masticatory plate of the basioccipital; Me, mesethmoid; N, nasal; Orsph, orbitosphenoid; Pa, parietal; Pe, prethmoid;
<collectingCountry id="6DAFDAD4814B494DFEDDB8D9FEAA0616" box="[268,306,1518,1539]" name="Palau" pageId="7" pageNumber="408">Pro</collectingCountry>
, prootic; Psph, parasphenoid; Ptsph, pterosphenoid; Soc, supraoccipital; STF, subtemporal fossa; TrC, trabecula communis; Vo, vomer; Ana, anguloarticular; Apal, autopalatine; Cm, coronomeckelian;
<collectingCountry id="6DAFDAD4814B494DFBC1BB3AFBA80637" box="[1040,1072,1549,1570]" name="South Georgia and the South Sandwich Islands" pageId="7" pageNumber="408">De</collectingCountry>
, dentary; Ecpt, ectopterygoid; Enpt, endopterygoid; Hy, hyomandibular; Iop, interopercle; KE, kinethmoid; MC, Meckels cartilage; Mpt, metapterygoid;
<collectingCountry id="6DAFDAD4814B494DFAA3BB1CFA000655" box="[1394,1432,1579,1600]" name="Mexico" pageId="7" pageNumber="408">Mx</collectingCountry>
, maxilla; Op, opercle; Pmx, premaxilla; Pop, preopercle; PvPQ, posteroventral process of quadrate; Q, quadrate; Ra, retroarticular; Sop, subopercle; Sy, symplectic; ACh, anterior ceratohyal; BR, branchiostegal ray; DHh, dorsal hypohyal; IO1- 5, infraorbital 15; PCh, posterior ceratohyal; Scl B, scleral bone; Scl C, scleral cartilage; So, supraorbital; Uh, urohyal; VHh, ventral hypohyal.
</paragraph>
<paragraph id="15079A44814B494DFF16BBD0FA0507CC" blockId="7.[151,1437,1767,2009]" pageId="7" pageNumber="408">
<emphasis id="27CC4656814B494DFF16BBD0FE010715" bold="true" box="[199,409,1767,1792]" italics="true" pageId="7" pageNumber="408">Branchial arches:</emphasis>
See Figure
<date id="6106BC84814B494DFDF2BBDFFDA10714" box="[547,569,1768,1793]" pageId="7" pageNumber="408">5I</date>
M. Three well ossified cylindrical basibranchial, connected by congregated cartilage; first one is smaller than second and third. Three pairs of small, round, ossified hypobranchials, covered with cartilage, attaches to the respective basibranchial and ceratobranchial. First basibranchial attaches to the basihyal by ligaments; no cartilaginous connection was seen between basihyal and first basibranchial. Fourth and fifth ceratobranchial connects with basibranchial 3 by strong ligaments and basibranchial 4 cartilage remnants. Five pairs of well developed ceratobranchials present. Ceratobranchial 1 with 7 gill rakers on anterior side and 12 14 on posterior. Ceratobranchial 2 with 1415 gill rakers along their anterior margins and 1516 on posterior.
</paragraph>
<paragraph id="15079A4481444942FF46BDA0FAEA019D" blockId="8.[151,1437,151,2016]" pageId="8" pageNumber="409">Ceratobranchial 3 with 20 gill rakers on anterior and 18 on posterior margin. Ceratobranchial 4 with 19 gill rakers on anterior and 16 on posterior margin. Ceratobranchial 5 well ossified, toothed having 14 gill rakers on anterior margin. Proximal half of ceratobranchial 5 marked with 3 rows of well ossified conical teeth. Anterior most teeth row comprises 2, middle 3 and posterior most comprises 4 conical and one small tooth (Figure 5LM). Upper gill arch constitutes by four pairs of well-developed epibranchials. Epibranchial 1 broad and wing shaped followed by relatively less wide epibranchial 2 and 3. Epibranchial 4 narrow bodied. Pharyngobranchial 1 was not located, pharyngobranchial 2 small whereas pharyngobranchial 3 is well developed, pharyngobranchial 4 cartilaginous.</paragraph>
<paragraph id="15079A4481444942FF16BCA4FABE02C5" blockId="8.[151,1437,151,2016]" pageId="8" pageNumber="409">
<emphasis id="27CC465681444942FF16BCA4FE1C01B9" bold="true" box="[199,388,403,428]" italics="true" pageId="8" pageNumber="409">Pectoral Girdle:</emphasis>
See
<figureCitation id="8D8386C181444942FE12BCA3FDBE01B8" box="[451,550,404,429]" captionStart="FIGURE 6" captionStartId="9.[151,249,1673,1695]" captionTargetBox="[249,1332,197,1640]" captionTargetId="figure@9.[249,1333,193,1641]" captionTargetPageId="9" captionText="FIGURE 6. Pethia ticto, topotype, male, BNHS FWF 130, 32.2 mm SL; (A) pectoral girdle lateral view; (B) pelvic girdle dorsal view, (C) pelvic girdle ventral view, (D) dorsal fin, lateral view, (E) anal fin lateral view, (F) weberian apparatus lateral view and (G) Caudal fin. Cl, cleithrum; Co, coracoid; DRC, distal radial cartilage; Pcl, postcleithrum; PeR 1 4, pectoral radial 1 4; Sc, scapula; Scl, supracleithrum; Bp, basipterygium; PS, pelvic splint; PR, pelvic radial; AFS, anal fin stay; DFS, dorsal fin stay; DR, distal radial; MR, middle radial; P MR, proximal middle radial; PR, proximal radial; Sn 5 - 8, supraneural 5 8; Apto, autopterotic; Boc, basioccipital; C, claustrum; Epo, epioccipital; Exoc, exoccipital; Ios, inner arm of the os suspensorium; Lp 1, lateral process of the first vertebral centrum; Lp 2, lateral process of the second vertebral centrum; MaPl, masticatory plate of the basioccipital process; NA, neural arch; NS, neural spine; Oos, outer arm of the os suspensorium; Ri, rib; S, scaphium; Sn, supraneural; Soc, supraoccipital; T, tripus; V, vertebral centrum (numbers after abbreviations refer to the vertebral centrum to which the structure is associated); CC, compound centrum; CIHPU 4, inter-hemal spine cartilage of preural centrum 4; Ep, epural; H 1 - 6, hypurals 1 6; HS, hemal spine; NS, neural spine; Ph, parhypural; Pls, pleurostyle; PU 2 - 3, preural centra 2 3." httpUri="https://zenodo.org/record/233769/files/figure.png" pageId="8" pageNumber="409">Figure 6</figureCitation>
A. Pectoral girdle positioned most anteriorly and does not overlap weberian apparatus laterally; articulate with posterolateral surface of neurocranium through vertically aligned supracleithrum and cleithrum. Supracleithrum small, laterally overlaps with cleithrum by articulating more than 3/ 4th area. Cleithrum large, narrow and pointed dorsally, articulates with postcleithrum lateroventrally. Postcleithrum single, elongated and well developed, supports pectoral girdle posteriorly. Coracoid large, rectangular, articulates posteriorly with scapula and pectoral radials, dorsally overlaps with cleithrum. Four large well ossified pectoral radials present. Three distal radials less ossified, with three medial elements predominantly cartilaginous. Eleven distal radial present, first eight distal radial appear to be well ossified whereas remaining three are cartilaginous and abbreviated as distal radial cartilage, supports one unbranched and 12 branched (i.
<date id="6106BC8481444942FBFCBF81FBC002DA" box="[1069,1112,694,719]" pageId="8" pageNumber="409">11.i</date>
) pectoral fin rays.
</paragraph>
<paragraph id="15079A4481444942FF16BFEDFADB0375" blockId="8.[151,1437,151,2016]" pageId="8" pageNumber="409">
<emphasis id="27CC465681444942FF16BFEDFEF802E6" bold="true" box="[199,352,730,755]" italics="true" pageId="8" pageNumber="409">Pelvic girdle:</emphasis>
See
<figureCitation id="8D8386C181444942FE46BFEDFE6D02E6" box="[407,501,730,755]" captionStart="FIGURE 6" captionStartId="9.[151,249,1673,1695]" captionTargetBox="[249,1332,197,1640]" captionTargetId="figure@9.[249,1333,193,1641]" captionTargetPageId="9" captionText="FIGURE 6. Pethia ticto, topotype, male, BNHS FWF 130, 32.2 mm SL; (A) pectoral girdle lateral view; (B) pelvic girdle dorsal view, (C) pelvic girdle ventral view, (D) dorsal fin, lateral view, (E) anal fin lateral view, (F) weberian apparatus lateral view and (G) Caudal fin. Cl, cleithrum; Co, coracoid; DRC, distal radial cartilage; Pcl, postcleithrum; PeR 1 4, pectoral radial 1 4; Sc, scapula; Scl, supracleithrum; Bp, basipterygium; PS, pelvic splint; PR, pelvic radial; AFS, anal fin stay; DFS, dorsal fin stay; DR, distal radial; MR, middle radial; P MR, proximal middle radial; PR, proximal radial; Sn 5 - 8, supraneural 5 8; Apto, autopterotic; Boc, basioccipital; C, claustrum; Epo, epioccipital; Exoc, exoccipital; Ios, inner arm of the os suspensorium; Lp 1, lateral process of the first vertebral centrum; Lp 2, lateral process of the second vertebral centrum; MaPl, masticatory plate of the basioccipital process; NA, neural arch; NS, neural spine; Oos, outer arm of the os suspensorium; Ri, rib; S, scaphium; Sn, supraneural; Soc, supraoccipital; T, tripus; V, vertebral centrum (numbers after abbreviations refer to the vertebral centrum to which the structure is associated); CC, compound centrum; CIHPU 4, inter-hemal spine cartilage of preural centrum 4; Ep, epural; H 1 - 6, hypurals 1 6; HS, hemal spine; NS, neural spine; Ph, parhypural; Pls, pleurostyle; PU 2 - 3, preural centra 2 3." httpUri="https://zenodo.org/record/233769/files/figure.png" pageId="8" pageNumber="409">Figure 6</figureCitation>
BC. Pelvic girdle consists of a pair of thin, flat and elongated basipterygium, each supporting one unbranched and eight branched fin rays (i.
<date id="6106BC8481444942FCF0BFC9FCD80302" box="[801,832,766,791]" pageId="8" pageNumber="409">7.i</date>
). On the anterior side basipterygium bears two spinelike processes, external and internal. External process slightly elongated than internal. Five well ossified pelvic radials present, pelvic splint is well developed and laterally associated with first unbranched pelvic-fin ray.
</paragraph>
<paragraph id="15079A4481444942FF16BE5DFE2104B6" blockId="8.[151,1437,151,2016]" pageId="8" pageNumber="409">
<emphasis id="27CC465681444942FF16BE5DFEDE0396" bold="true" box="[199,326,874,899]" italics="true" pageId="8" pageNumber="409">Dorsal fin:</emphasis>
See
<figureCitation id="8D8386C181444942FEAFBE5DFE460396" box="[382,478,874,899]" captionStart="FIGURE 6" captionStartId="9.[151,249,1673,1695]" captionTargetBox="[249,1332,197,1640]" captionTargetId="figure@9.[249,1333,193,1641]" captionTargetPageId="9" captionText="FIGURE 6. Pethia ticto, topotype, male, BNHS FWF 130, 32.2 mm SL; (A) pectoral girdle lateral view; (B) pelvic girdle dorsal view, (C) pelvic girdle ventral view, (D) dorsal fin, lateral view, (E) anal fin lateral view, (F) weberian apparatus lateral view and (G) Caudal fin. Cl, cleithrum; Co, coracoid; DRC, distal radial cartilage; Pcl, postcleithrum; PeR 1 4, pectoral radial 1 4; Sc, scapula; Scl, supracleithrum; Bp, basipterygium; PS, pelvic splint; PR, pelvic radial; AFS, anal fin stay; DFS, dorsal fin stay; DR, distal radial; MR, middle radial; P MR, proximal middle radial; PR, proximal radial; Sn 5 - 8, supraneural 5 8; Apto, autopterotic; Boc, basioccipital; C, claustrum; Epo, epioccipital; Exoc, exoccipital; Ios, inner arm of the os suspensorium; Lp 1, lateral process of the first vertebral centrum; Lp 2, lateral process of the second vertebral centrum; MaPl, masticatory plate of the basioccipital process; NA, neural arch; NS, neural spine; Oos, outer arm of the os suspensorium; Ri, rib; S, scaphium; Sn, supraneural; Soc, supraoccipital; T, tripus; V, vertebral centrum (numbers after abbreviations refer to the vertebral centrum to which the structure is associated); CC, compound centrum; CIHPU 4, inter-hemal spine cartilage of preural centrum 4; Ep, epural; H 1 - 6, hypurals 1 6; HS, hemal spine; NS, neural spine; Ph, parhypural; Pls, pleurostyle; PU 2 - 3, preural centra 2 3." httpUri="https://zenodo.org/record/233769/files/figure.png" pageId="8" pageNumber="409">Figure 6</figureCitation>
D. Dorsal fin constitutes thirteen rays (iv.
<date id="6106BC8481444942FC65BE5DFC560396" box="[948,974,874,899]" pageId="8" pageNumber="409">8.i</date>
) supported by a series of pterygiophores placed between neural spines of vertebrae
<date id="6106BC8481444942FDBEBEB9FD1703B2" box="[623,655,910,935]" pageId="8" pageNumber="409">V8</date>
/V9V
<date id="6106BC8481444942FD0EBEB9FCA903B2" box="[735,817,910,935]" pageId="8" pageNumber="409" value="1915-05-14">14/V15</date>
(
<figureCitation id="8D8386C181444942FCEEBEB9FC0603B2" box="[831,926,910,935]" captionStart="FIGURE 4" captionStartId="6.[151,249,1233,1255]" captionTargetBox="[151,1375,197,1197]" captionTargetId="figure@6.[151,1435,175,1225]" captionTargetPageId="6" captionText="FIGURE 4. Lateral view of entire skeleton of topotypic Pethia ticto (A) male, BNHS FWF 130, 32.2 mm SL; (B) male, BNHS FWF 131, 30.7 mm SL. AF, anal fin; CF, caudal fin; DF, dorsal fin; En, epineural intermusculars; Ep, epipleural intermusculars; Nc, neurocranium; PcF, pectoral fin; PvF, pelvic fin; WeV, Weberian vertebrae." httpUri="https://zenodo.org/record/233768/files/figure.png" pageId="8" pageNumber="409">Figure 4</figureCitation>
A &amp; B). First three pterygiophores constitutes a large proximalmiddle radial, articulates with three unbranched supernumerary and one serrated last unbranched dorsal-fin ray, distal radial absent. Posterior to proximalmiddle radial first three pterygiophores articulates with respective branched fin rays by having only distal radial, middle radials untraceable in this articulation. Middle radial appears in articulation with distal radial and dorsal base of pterygiophore on fourth proximal radial. All middle and distal radials are well ossified. Last thirteenth fin ray found to be closely united with twelfth branched fin ray, whereas articulated separately with last pterygiophore. 3 (1)4 (1) free supraneurals, Sn58 present on anterior side of dorsal fin.
</paragraph>
<paragraph id="15079A4481444942FF16B999FD00058A" blockId="8.[151,1437,151,2016]" pageId="8" pageNumber="409">
<emphasis id="27CC465681444942FF16B999FEB704D2" bold="true" box="[199,303,1198,1223]" italics="true" pageId="8" pageNumber="409">Anal fin:</emphasis>
See
<figureCitation id="8D8386C181444942FEB4B999FE5C04D2" box="[357,452,1198,1223]" captionStart="FIGURE 6" captionStartId="9.[151,249,1673,1695]" captionTargetBox="[249,1332,197,1640]" captionTargetId="figure@9.[249,1333,193,1641]" captionTargetPageId="9" captionText="FIGURE 6. Pethia ticto, topotype, male, BNHS FWF 130, 32.2 mm SL; (A) pectoral girdle lateral view; (B) pelvic girdle dorsal view, (C) pelvic girdle ventral view, (D) dorsal fin, lateral view, (E) anal fin lateral view, (F) weberian apparatus lateral view and (G) Caudal fin. Cl, cleithrum; Co, coracoid; DRC, distal radial cartilage; Pcl, postcleithrum; PeR 1 4, pectoral radial 1 4; Sc, scapula; Scl, supracleithrum; Bp, basipterygium; PS, pelvic splint; PR, pelvic radial; AFS, anal fin stay; DFS, dorsal fin stay; DR, distal radial; MR, middle radial; P MR, proximal middle radial; PR, proximal radial; Sn 5 - 8, supraneural 5 8; Apto, autopterotic; Boc, basioccipital; C, claustrum; Epo, epioccipital; Exoc, exoccipital; Ios, inner arm of the os suspensorium; Lp 1, lateral process of the first vertebral centrum; Lp 2, lateral process of the second vertebral centrum; MaPl, masticatory plate of the basioccipital process; NA, neural arch; NS, neural spine; Oos, outer arm of the os suspensorium; Ri, rib; S, scaphium; Sn, supraneural; Soc, supraoccipital; T, tripus; V, vertebral centrum (numbers after abbreviations refer to the vertebral centrum to which the structure is associated); CC, compound centrum; CIHPU 4, inter-hemal spine cartilage of preural centrum 4; Ep, epural; H 1 - 6, hypurals 1 6; HS, hemal spine; NS, neural spine; Ph, parhypural; Pls, pleurostyle; PU 2 - 3, preural centra 2 3." httpUri="https://zenodo.org/record/233769/files/figure.png" pageId="8" pageNumber="409">Figure 6</figureCitation>
E. Anal fin constitutes 9 rays (iii.
<date id="6106BC8481444942FCE7B999FCC904D2" box="[822,849,1198,1223]" pageId="8" pageNumber="409">5.i</date>
) supported by a series of anal pterygiophores placed between hemal spines of vertebrae
<date id="6106BC8481444942FDEDB9E4FDF304FE" box="[572,619,1235,1259]" pageId="8" pageNumber="409" value="1917-05">V17</date>
/V18 (2)
<date id="6106BC8481444942FD0CB9E4FC9704FE" box="[733,783,1235,1259]" pageId="8" pageNumber="409" value="1920-05">V20</date>
/V21 (1) or
<date id="6106BC8481444942FC75B9E4FC4A04FE" box="[932,978,1235,1259]" pageId="8" pageNumber="409" value="1919-05">V19</date>
/V20 (1) (
<figureCitation id="8D8386C181444942FB9CB9E5FB2A04FE" box="[1101,1202,1234,1259]" captionStart="FIGURE 4" captionStartId="6.[151,249,1233,1255]" captionTargetBox="[151,1375,197,1197]" captionTargetId="figure@6.[151,1435,175,1225]" captionTargetPageId="6" captionText="FIGURE 4. Lateral view of entire skeleton of topotypic Pethia ticto (A) male, BNHS FWF 130, 32.2 mm SL; (B) male, BNHS FWF 131, 30.7 mm SL. AF, anal fin; CF, caudal fin; DF, dorsal fin; En, epineural intermusculars; Ep, epipleural intermusculars; Nc, neurocranium; PcF, pectoral fin; PvF, pelvic fin; WeV, Weberian vertebrae." httpUri="https://zenodo.org/record/233768/files/figure.png" pageId="8" pageNumber="409">Figure 4</figureCitation>
A &amp; B). First three pterygiophores constitutes long, elongated proximalmiddle radial, articulates with two unbranched supernumerary and last unbranched anal-fin ray, distal radial present with no traces of middle radial. Middle radial appears in articulation with distal radial and dorsal base of pterygiophore on third proximal radial. All middle and distal radials are well ossified. Last sixth branched fin ray found to be closely united with fifth anal-fin ray, whereas articulated separately with last pterygiophore.
</paragraph>
<paragraph id="15079A4481444942FF16B89DFD0E06AA" blockId="8.[151,1437,151,2016]" pageId="8" pageNumber="409">
<emphasis id="27CC465681444942FF16B89DFED405D6" bold="true" box="[199,332,1450,1475]" italics="true" pageId="8" pageNumber="409">Caudal fin:</emphasis>
See
<figureCitation id="8D8386C181444942FE55B89DFE7905D6" box="[388,481,1450,1475]" captionStart="FIGURE 6" captionStartId="9.[151,249,1673,1695]" captionTargetBox="[249,1332,197,1640]" captionTargetId="figure@9.[249,1333,193,1641]" captionTargetPageId="9" captionText="FIGURE 6. Pethia ticto, topotype, male, BNHS FWF 130, 32.2 mm SL; (A) pectoral girdle lateral view; (B) pelvic girdle dorsal view, (C) pelvic girdle ventral view, (D) dorsal fin, lateral view, (E) anal fin lateral view, (F) weberian apparatus lateral view and (G) Caudal fin. Cl, cleithrum; Co, coracoid; DRC, distal radial cartilage; Pcl, postcleithrum; PeR 1 4, pectoral radial 1 4; Sc, scapula; Scl, supracleithrum; Bp, basipterygium; PS, pelvic splint; PR, pelvic radial; AFS, anal fin stay; DFS, dorsal fin stay; DR, distal radial; MR, middle radial; P MR, proximal middle radial; PR, proximal radial; Sn 5 - 8, supraneural 5 8; Apto, autopterotic; Boc, basioccipital; C, claustrum; Epo, epioccipital; Exoc, exoccipital; Ios, inner arm of the os suspensorium; Lp 1, lateral process of the first vertebral centrum; Lp 2, lateral process of the second vertebral centrum; MaPl, masticatory plate of the basioccipital process; NA, neural arch; NS, neural spine; Oos, outer arm of the os suspensorium; Ri, rib; S, scaphium; Sn, supraneural; Soc, supraoccipital; T, tripus; V, vertebral centrum (numbers after abbreviations refer to the vertebral centrum to which the structure is associated); CC, compound centrum; CIHPU 4, inter-hemal spine cartilage of preural centrum 4; Ep, epural; H 1 - 6, hypurals 1 6; HS, hemal spine; NS, neural spine; Ph, parhypural; Pls, pleurostyle; PU 2 - 3, preural centra 2 3." httpUri="https://zenodo.org/record/233769/files/figure.png" pageId="8" pageNumber="409">Figure 6</figureCitation>
G. Seven dorsal procurrent caudal-fin rays and six ventral procurrent caudal-fin rays. In cleared and stained specimens, 9 dorsal and 8 ventral principal caudal-fin rays present. Principal caudal-fin rays supported by neural and hemal spines of third preural centrum, epural, pleurostyle, six hypurals and a parhypural. A single interhemal spine cartilage of preural centrum 4 present, placed anterior to ventral tip of hemal spine of preural centrum
<quantity id="D24037A181444942FE80BB0CFEE40646" box="[337,380,1594,1619]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="8" pageNumber="409" unit="in" value="3.0">3 in</quantity>
male. Single epural runs laterally with anterior margin of pleurostyle on dorsal half, distal tip of pleurostyle cartilaginous. Parhypural and hypural 16 well developed, broad and ossified. In male, pleural centra 2 supports single neural and hemal spine, pleural centra 3 supports well developed neural and hemal spine which support peduncle arch. Free uroneural absent.
</paragraph>
<paragraph id="15079A4481444941FF16BBFDFC0900ED" blockId="8.[151,1437,151,2016]" lastBlockId="11.[151,1437,151,2018]" lastPageId="11" lastPageNumber="412" pageId="8" pageNumber="409">
<emphasis id="27CC465681444942FF16BBFDFE2506F7" bold="true" box="[199,445,1738,1763]" italics="true" pageId="8" pageNumber="409">Weberian apparatus:</emphasis>
See
<figureCitation id="8D8386C181444942FE2BBBFDFDC406F6" box="[506,604,1738,1763]" captionStart="FIGURE 6" captionStartId="9.[151,249,1673,1695]" captionTargetBox="[249,1332,197,1640]" captionTargetId="figure@9.[249,1333,193,1641]" captionTargetPageId="9" captionText="FIGURE 6. Pethia ticto, topotype, male, BNHS FWF 130, 32.2 mm SL; (A) pectoral girdle lateral view; (B) pelvic girdle dorsal view, (C) pelvic girdle ventral view, (D) dorsal fin, lateral view, (E) anal fin lateral view, (F) weberian apparatus lateral view and (G) Caudal fin. Cl, cleithrum; Co, coracoid; DRC, distal radial cartilage; Pcl, postcleithrum; PeR 1 4, pectoral radial 1 4; Sc, scapula; Scl, supracleithrum; Bp, basipterygium; PS, pelvic splint; PR, pelvic radial; AFS, anal fin stay; DFS, dorsal fin stay; DR, distal radial; MR, middle radial; P MR, proximal middle radial; PR, proximal radial; Sn 5 - 8, supraneural 5 8; Apto, autopterotic; Boc, basioccipital; C, claustrum; Epo, epioccipital; Exoc, exoccipital; Ios, inner arm of the os suspensorium; Lp 1, lateral process of the first vertebral centrum; Lp 2, lateral process of the second vertebral centrum; MaPl, masticatory plate of the basioccipital process; NA, neural arch; NS, neural spine; Oos, outer arm of the os suspensorium; Ri, rib; S, scaphium; Sn, supraneural; Soc, supraoccipital; T, tripus; V, vertebral centrum (numbers after abbreviations refer to the vertebral centrum to which the structure is associated); CC, compound centrum; CIHPU 4, inter-hemal spine cartilage of preural centrum 4; Ep, epural; H 1 - 6, hypurals 1 6; HS, hemal spine; NS, neural spine; Ph, parhypural; Pls, pleurostyle; PU 2 - 3, preural centra 2 3." httpUri="https://zenodo.org/record/233769/files/figure.png" pageId="8" pageNumber="409">
Figure
<specimenCount id="03BE51CD81444942FD9FBBFDFDC406F6" box="[590,604,1738,1763]" pageId="8" pageNumber="409" type="generic">6</specimenCount>
</figureCitation>
F. A complex structure made up by first four vertebral centra and their associated elements, covers the anterodorsal portion of swim bladder. The first centrum relatively small but distinct, possesses lateral process 1 (Lp1). Second centra equivalent in size of first, possesses large lateral process of second vertebra (Lp2). Lp2 is more developed and distinct than LP1 and grown laterally in wing shaped processes. Third and fourth centrum is much larger and each equivalent to size of first two. Scaphium and claustrum small, rounded and well ossified, present on dorsal in association with first centra. Tripus well developed, triangular grows posteriorly and overlaps inner arm of the os suspensorium, associated with third vertebral centrum. Outer arms of the os suspensorium much grown, long and reaches behind the postcleithrum bone of pectoral girdle. Neural arch 3 and neural arch 4 well developed and ossified. Supraneural 3 seems to be a second largest bone of weberian apparatus after outer arm of the os suspensorium. Supraneural 2, 3 and neural spine 4 is associated with neural arch 3 and 4. Supraneural 4 absent.
</paragraph>
<caption id="41C7CACC81454943FF46BBBEFE9007E4" httpUri="https://zenodo.org/record/233769/files/figure.png" pageId="9" pageNumber="410" targetBox="[249,1332,197,1640]" targetPageId="9">
<paragraph id="15079A4481454943FF46BBBEFE9007E4" blockId="9.[151,1436,1673,2033]" pageId="9" pageNumber="410">
<emphasis id="27CC465681454943FF46BBBEFE8C068B" bold="true" box="[151,276,1673,1695]" pageId="9" pageNumber="410">FIGURE 6.</emphasis>
<taxonomicName id="D2B8E1C781454943FECCBBBEFE0D068A" box="[285,405,1673,1695]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="410" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC465681454943FECCBBBEFE0D068A" box="[285,405,1673,1695]" italics="true" pageId="9" pageNumber="410">Pethia ticto</emphasis>
</taxonomicName>
, topotype, male, BNHS FWF 130, 32.2 mm SL; (A) pectoral girdle lateral view; (B) pelvic girdle dorsal view, (C) pelvic girdle ventral view, (D) dorsal fin, lateral view, (E) anal fin lateral view, (F) weberian apparatus lateral view and (G) Caudal fin. Cl, cleithrum; Co, coracoid; DRC, distal radial cartilage; Pcl, postcleithrum; PeR14, pectoral radial 14; Sc, scapula; Scl, supracleithrum; Bp, basipterygium; PS, pelvic splint; PR, pelvic radial; AFS, anal fin stay; DFS, dorsal fin stay; DR, distal radial; MR, middle radial; PMR, proximalmiddle radial; PR, proximal radial; Sn5-8, supraneural 58; Apto, autopterotic; Boc, basioccipital; C, claustrum; Epo, epioccipital; Exoc, exoccipital; Ios, inner arm of the os suspensorium; Lp1, lateral process of the first vertebral centrum; Lp2, lateral process of the second vertebral centrum; MaPl, masticatory plate of the basioccipital process; NA, neural arch; NS, neural spine; Oos, outer arm of the os suspensorium; Ri, rib; S, scaphium; Sn, supraneural; Soc, supraoccipital; T, tripus; V, vertebral centrum (numbers after abbreviations refer to the vertebral centrum to which the structure is associated); CC, compound centrum; CIHPU4, inter-hemal spine cartilage of preural centrum 4; Ep, epural; H1-6, hypurals 16; HS, hemal spine; NS, neural spine; Ph, parhypural; Pls, pleurostyle; PU2-3, preural centra 23.
</paragraph>
</caption>
<caption id="41C7CACC81464940FF46BA63FB1507FA" httpUri="https://zenodo.org/record/233770/files/figure.png" pageId="10" pageNumber="411" targetBox="[284,1293,201,1815]" targetPageId="10">
<paragraph id="15079A4481464940FF46BA63FB1507FA" blockId="10.[151,1437,1876,2031]" pageId="10" pageNumber="411">
<emphasis id="27CC465681464940FF46BA63FE8C077C" bold="true" box="[151,276,1876,1898]" pageId="10" pageNumber="411">FIGURE 7.</emphasis>
Maximum likelihood tree for
<taxonomicName id="D2B8E1C781464940FD87BA63FD00077F" box="[598,664,1876,1898]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="411" phylum="Chordata" rank="genus">
<emphasis id="27CC465681464940FD87BA63FD00077F" box="[598,664,1876,1898]" italics="true" pageId="10" pageNumber="411">Pethia</emphasis>
</taxonomicName>
species based on cytochrome b sequences. Nucleotide substitution model used was Tamura &amp; Nei (1993) with invariant sites and gamma distribution (TN93+G+I, BIC = 15662.99, lnL = -6933.45, I = 0.42, G = 0.78). Values along the nodes are percent bootstraps for 1000 iterations.
<taxonomicName id="D2B8E1C781464940FBC8BAA0FBC007B9" box="[1049,1112,1943,1964]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="411" phylum="Chordata" rank="genus">
<emphasis id="27CC465681464940FBC8BAA0FBC007B9" box="[1049,1112,1943,1964]" italics="true" pageId="10" pageNumber="411">Garra</emphasis>
</taxonomicName>
species are used as outgroup. Sequences in red are from current study and indicate topotypic
<taxonomicName id="D2B8E1C781464940FC81BA80FC5307DB" box="[848,971,1975,1998]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="411" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC465681464940FC81BA80FC5307DB" box="[848,971,1975,1998]" italics="true" pageId="10" pageNumber="411">Pethia ticto</emphasis>
</taxonomicName>
. Sequence in blue from NCBI database is conspecific with topotypic material. Sequences in green from NCBI database are misidentifications.
</paragraph>
</caption>
<paragraph id="15079A4481474941FF16BC34FCF8019D" blockId="11.[151,1437,151,2018]" pageId="11" pageNumber="412">
<emphasis id="27CC465681474941FF16BC34FE2E0109" bold="true" box="[199,438,259,284]" italics="true" pageId="11" pageNumber="412">Intermuscular bone:</emphasis>
See
<figureCitation id="8D8386C181474941FE3FBC33FDD50109" box="[494,589,260,285]" captionStart="FIGURE 4" captionStartId="6.[151,249,1233,1255]" captionTargetBox="[151,1375,197,1197]" captionTargetId="figure@6.[151,1435,175,1225]" captionTargetPageId="6" captionText="FIGURE 4. Lateral view of entire skeleton of topotypic Pethia ticto (A) male, BNHS FWF 130, 32.2 mm SL; (B) male, BNHS FWF 131, 30.7 mm SL. AF, anal fin; CF, caudal fin; DF, dorsal fin; En, epineural intermusculars; Ep, epipleural intermusculars; Nc, neurocranium; PcF, pectoral fin; PvF, pelvic fin; WeV, Weberian vertebrae." httpUri="https://zenodo.org/record/233768/files/figure.png" pageId="11" pageNumber="412">Figure 4</figureCitation>
AB. Two series of intermuscular bones, epineural and epipleural. Total 24 epineural and 11 epipleural intermuscular bones present; first epineural appears before neural spine of sixth vertebra whereas first epipleural appears before hemal spine of seventieth vertebra. All epipleural and epineural bifurcated at tip and becomes wider in caudal peduncle portion.
</paragraph>
<paragraph id="15079A4481474941FF16BCA4FC140257" blockId="11.[151,1437,151,2018]" pageId="11" pageNumber="412">
<emphasis id="27CC465681474941FF16BCA4FE0001B9" bold="true" box="[199,408,403,428]" italics="true" pageId="11" pageNumber="412">Vertebral column:</emphasis>
See
<figureCitation id="8D8386C181474941FE01BCA3FDB701B9" box="[464,559,404,429]" captionStart="FIGURE 4" captionStartId="6.[151,249,1233,1255]" captionTargetBox="[151,1375,197,1197]" captionTargetId="figure@6.[151,1435,175,1225]" captionTargetPageId="6" captionText="FIGURE 4. Lateral view of entire skeleton of topotypic Pethia ticto (A) male, BNHS FWF 130, 32.2 mm SL; (B) male, BNHS FWF 131, 30.7 mm SL. AF, anal fin; CF, caudal fin; DF, dorsal fin; En, epineural intermusculars; Ep, epipleural intermusculars; Nc, neurocranium; PcF, pectoral fin; PvF, pelvic fin; WeV, Weberian vertebrae." httpUri="https://zenodo.org/record/233768/files/figure.png" pageId="11" pageNumber="412">Figure 4</figureCitation>
AB. 4+4 (2) predorsal vertebrae; 4+26 (2) total vertebrae including posterior most compound centrum, with 4+13 (2) abdominal and 13 (2) caudal vertebrae. Vertebral centra much elongated than deep. Neural pre and postzygopsis appears on all centra posterior to weberian complex. Hemal postzygopsis appear on all caudal vertebrae posterior to 14th (1) or 15th (1) centra. First hemal prezygopsis appears on 21st (1) or 22th (1) centra. Total 11 ribs are associated with 5th to 15th vertebrae.
</paragraph>
<paragraph id="15079A4481474941FF16BF7AFAD4030C" blockId="11.[151,1437,151,2018]" pageId="11" pageNumber="412">
<emphasis id="27CC465681474941FF16BF7AFD930273" bold="true" box="[199,523,589,614]" pageId="11" pageNumber="412">Coloration in preservative.</emphasis>
For general appearance see
<figureCitation id="8D8386C181474941FC80BF7AFC350273" box="[849,941,589,614]" captionStart="FIGURE 1" captionStartId="1.[151,250,1163,1185]" captionTargetBox="[194,1393,623,1141]" captionTargetId="figure@1.[194,1394,623,1141]" captionTargetPageId="1" captionText="FIGURE 1. Illustration of Pethia ticto based on topotype, male, BNHS FWF 127, 35.9 mm SL, showing distribution of prominent body pigmentation and banding pattern and lateral and transverse scale rows." httpUri="https://zenodo.org/record/233765/files/figure.png" pageId="11" pageNumber="412">Figure 1</figureCitation>
A; body above lateral scale row grey; head and dorsum dark brown; lower lip, cheek, opercular region below inferior border of eye cream; ventral region uniformly cream white. Body with prominent dark black humeral spot, covering third and fourth lateral line pored scales; a prominent spot on caudal peduncle covering scales 1619 of lateral scale row, encircling peduncle above. Scale pockets above lateral scale row studded with melanophores. Dorsal fin with 2 longitudinal rows of intense black spots (
<figureCitation id="8D8386C181474941FEF3BE36FE15030F" box="[290,397,769,794]" captionStart="FIGURE 1" captionStartId="1.[151,250,1163,1185]" captionTargetBox="[194,1393,623,1141]" captionTargetId="figure@1.[194,1394,623,1141]" captionTargetPageId="1" captionText="FIGURE 1. Illustration of Pethia ticto based on topotype, male, BNHS FWF 127, 35.9 mm SL, showing distribution of prominent body pigmentation and banding pattern and lateral and transverse scale rows." httpUri="https://zenodo.org/record/233765/files/figure.png" pageId="11" pageNumber="412">Figures 1</figureCitation>
,
<figureCitation id="8D8386C181474941FE4ABE35FE32030F" box="[411,426,770,794]" captionStart="FIGURE 2" captionStartId="4.[151,250,1439,1461]" captionTargetBox="[272,1313,194,1403]" captionTargetId="figure@4.[271,1315,193,1418]" captionTargetPageId="4" captionText="FIGURE 2. Pethia ticto. (A) Topotype, BNHS FWF, male, 127, 35.9 mm SL, from West Bengal, India. (B) Cyprinus tic to (iconotype) from Hamiltons (1822) illustration in the collection of the Linnean Society of London (courtesy: Linnean Society of London)." httpUri="https://zenodo.org/record/233766/files/figure.png" pageId="11" pageNumber="412">2</figureCitation>
A). Pectoral, pelvic, anal and caudal fins hyaline, without any color bands or spots.
</paragraph>
<paragraph id="15079A4481474941FF16BE12FAAE0377" blockId="11.[151,1437,151,2018]" pageId="11" pageNumber="412">
<emphasis id="27CC465681474941FF16BE12FE02032B" bold="true" box="[199,410,805,830]" pageId="11" pageNumber="412">Coloration in life.</emphasis>
See
<figureCitation id="8D8386C181474941FE00BE12FDB6032B" box="[465,558,805,830]" captionStart="FIGURE 3" captionStartId="5.[151,250,1946,1968]" captionTargetBox="[382,1204,871,1923]" captionTargetId="figure@5.[382,1204,871,1924]" captionTargetPageId="5" captionText="FIGURE 3. Topotype of Pethia ticto in life (A) from Beri Baor, West Bangal, India (BNHS FWF 127, 35.9 mm SL), humeral and caudal spots disappear immediately after collection, and (B) from Dhaka, Bangladesh (unregistered)." httpUri="https://zenodo.org/record/233767/files/figure.png" pageId="11" pageNumber="412">Figure 3</figureCitation>
A, B. Body yellow. Position of humeral and caudal spot same as described for preserved specimens, spot on caudal peduncle with a golden hoop on its anterior margin. All fins hyaline.
</paragraph>
<paragraph id="15079A4481474941FF16BE5AFBF604DC" blockId="11.[151,1437,151,2018]" pageId="11" pageNumber="412">
<emphasis id="27CC465681474941FF16BE5AFE170393" bold="true" box="[199,399,877,902]" pageId="11" pageNumber="412">Genetic analysis.</emphasis>
Nucleotide substitution rates followed
<bibRefCitation id="7129E7B581474941FC94BE59FBAF0393" author="Tamura" box="[837,1079,877,902]" pageId="11" pageNumber="412" refString="Tamura, K. &amp; Nei, M. (1993) Estimation of the number of nucleotide substitutions in the control region of mitochondrial DNA in humans and chimpanzees. Molecular Biology and Evolution, 10, 512 - 526." type="journal article" year="1993">Tamura &amp; Nei (1993)</bibRefCitation>
model with gamma distribution and invariant sites (TN93+G+I, BIC = 15828.18, lnL = -6923.37, I = 0.42, G = 0.78). Maximum likelihood tree based on the best fit model is shown in
<figureCitation id="8D8386C181474941FDA5BE82FD4003D8" box="[628,728,949,974]" captionStart="FIGURE 7" captionStartId="10.[151,249,1876,1898]" captionTargetBox="[284,1293,201,1815]" captionTargetId="figure@10.[270,1301,193,1830]" captionTargetPageId="10" captionText="FIGURE 7. Maximum likelihood tree for Pethia species based on cytochrome b sequences. Nucleotide substitution model used was Tamura &amp; Nei (1993) with invariant sites and gamma distribution (TN 93 + G + I, BIC = 15662.99, lnL = - 6933.45, I = 0.42, G = 0.78). Values along the nodes are percent bootstraps for 1000 iterations. Garra species are used as outgroup. Sequences in red are from current study and indicate topotypic Pethia ticto. Sequence in blue from NCBI database is conspecific with topotypic material. Sequences in green from NCBI database are misidentifications." httpUri="https://zenodo.org/record/233770/files/figure.png" pageId="11" pageNumber="412">Figure 7</figureCitation>
. Based on the comparison of cytb sequences from topotypic material collected in the present study, we suggest that the sequence
<accessionNumber id="0AEB07A781474941FC72BEEEFB8403E7" box="[931,1052,985,1010]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EU604679" pageId="11" pageNumber="412">EU604679</accessionNumber>
(from
<bibRefCitation id="7129E7B581474941FBBDBEEEFAE203E7" author="Pethiyagoda" box="[1132,1402,985,1010]" pageId="11" pageNumber="412" refString="Pethiyagoda, R., Meegaskumbura, M. &amp; Maduwage K. (2012) A synopsis of the South Asian fishes referred to Puntius (Pisces: Cyprinidae). Ichthyological Exploration of Freshwaters, 23, 69 - 95." type="journal article" year="2012">
Pethiyagoda
<emphasis id="27CC465681474941FB2FBEEDFAAF03E4" box="[1278,1335,985,1009]" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
2012
</bibRefCitation>
) is conspecific to
<emphasis id="27CC465681474941FEEABEC9FE1D0400" box="[315,389,1022,1045]" italics="true" pageId="11" pageNumber="412">P. t i c t o</emphasis>
with a genetic distance of just 0.4%. However, sequence
<accessionNumber id="0AEB07A781474941FBDEBECAFB110403" box="[1039,1161,1021,1046]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AY708250" pageId="11" pageNumber="412">AY708250</accessionNumber>
deposited as
<emphasis id="27CC465681474941FAF0BEC9FAF30400" box="[1313,1387,1022,1045]" italics="true" pageId="11" pageNumber="412">P. t i c t o</emphasis>
is a misidentification with a genetic distance of 10.3% and should be attributed to
<taxonomicName id="D2B8E1C781474941FBCEB915FB60042C" box="[1055,1272,1057,1081]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="melanomaculata">
<emphasis id="27CC465681474941FBCEB915FB60042C" box="[1055,1272,1057,1081]" italics="true" pageId="11" pageNumber="412">P. melanomaculata</emphasis>
</taxonomicName>
as there is no genetic distance between
<accessionNumber id="0AEB07A781474941FE69B972FDAB044B" box="[440,563,1093,1118]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AY708250" pageId="11" pageNumber="412">AY708250</accessionNumber>
and
<taxonomicName id="D2B8E1C781474941FDBBB971FCDA0448" box="[618,834,1093,1117]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="melanomaculata">
<emphasis id="27CC465681474941FDBBB971FCDA0448" box="[618,834,1093,1117]" italics="true" pageId="11" pageNumber="412">P. melanomaculata</emphasis>
</taxonomicName>
sequence
<accessionNumber id="0AEB07A781474941FC69B972FBA9044B" box="[952,1073,1093,1118]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EU604678" pageId="11" pageNumber="412">EU604678</accessionNumber>
(from
<bibRefCitation id="7129E7B581474941FB51B972FA08044B" author="Pethiyagoda" box="[1152,1424,1093,1118]" pageId="11" pageNumber="412" refString="Pethiyagoda, R., Meegaskumbura, M. &amp; Maduwage K. (2012) A synopsis of the South Asian fishes referred to Puntius (Pisces: Cyprinidae). Ichthyological Exploration of Freshwaters, 23, 69 - 95." type="journal article" year="2012">
Pethiyagoda
<emphasis id="27CC465681474941FAC2B971FAD40448" box="[1299,1356,1093,1117]" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
2012
</bibRefCitation>
). Sequences
<accessionNumber id="0AEB07A781474941FEC6B95EFE0C0497" box="[279,404,1129,1154]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KC696520" pageId="11" pageNumber="412">KC696520</accessionNumber>
,
<accessionNumber id="0AEB07A781474941FE72B95EFD800497" box="[419,536,1129,1154]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ795474" pageId="11" pageNumber="412">JQ795474</accessionNumber>
,
<accessionNumber id="0AEB07A781474941FDF6B95EFD050497" box="[551,669,1129,1154]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ795475" pageId="11" pageNumber="412">JQ795475</accessionNumber>
,
<accessionNumber id="0AEB07A781474941FD7AB95EFCB90497" box="[683,801,1129,1154]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ795476" pageId="11" pageNumber="412">JQ795476</accessionNumber>
,
<accessionNumber id="0AEB07A781474941FCFEB95EFC3A0497" box="[815,930,1129,1154]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ795477" pageId="11" pageNumber="412">JQ795477</accessionNumber>
and
<accessionNumber id="0AEB07A781474941FC0EB95EFBCA0497" box="[991,1106,1129,1154]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ795478" pageId="11" pageNumber="412">JQ795478</accessionNumber>
deposited as
<taxonomicName id="D2B8E1C781474941FB23B95DFADA0494" box="[1266,1346,1130,1153]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC465681474941FB23B95DFADA0494" box="[1266,1346,1130,1153]" italics="true" pageId="11" pageNumber="412">P. ticto</emphasis>
</taxonomicName>
are not conspecific to topotypic
<emphasis id="27CC465681474941FE7BB9B9FE6B04B0" box="[426,499,1166,1189]" italics="true" pageId="11" pageNumber="412">P. t i c t o</emphasis>
and have genetic distance ranging between 8.18.4%. We attribute these sequences (
<accessionNumber id="0AEB07A781474941FF4FB986FE8604DF" box="[158,286,1201,1226]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KC696520" pageId="11" pageNumber="412">KC696520</accessionNumber>
,
<accessionNumber id="0AEB07A781474941FEF8B986FE3804DF" box="[297,416,1201,1226]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ795474" pageId="11" pageNumber="412">JQ795474</accessionNumber>
,
<accessionNumber id="0AEB07A781474941FE7BB986FDB804DF" box="[426,544,1201,1226]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ795475" pageId="11" pageNumber="412">JQ795475</accessionNumber>
,
<accessionNumber id="0AEB07A781474941FDFAB986FD3904DF" box="[555,673,1201,1226]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ795476" pageId="11" pageNumber="412">JQ795476</accessionNumber>
,
<accessionNumber id="0AEB07A781474941FD7DB986FC8704DF" box="[684,799,1201,1226]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ795477" pageId="11" pageNumber="412">JQ795477</accessionNumber>
and
<accessionNumber id="0AEB07A781474941FC87B986FC5304DF" box="[854,971,1201,1226]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ795478" pageId="11" pageNumber="412">JQ795478</accessionNumber>
) to
<taxonomicName id="D2B8E1C781474941FC27B985FBFF04DC" box="[1014,1127,1201,1226]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC465681474941FC27B985FB9004DC" box="[1014,1032,1202,1225]" italics="true" pageId="11" pageNumber="412">P.</emphasis>
cf.
<emphasis id="27CC465681474941FBE4B985FBFF04DC" box="[1077,1127,1202,1225]" italics="true" pageId="11" pageNumber="412">ticto</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="5DA2C9CF81474941FF16B9E2FCA20627" pageId="11" pageNumber="412" type="distribution">
<paragraph id="15079A4481474941FF16B9E2FCA20627" blockId="11.[151,1437,151,2018]" pageId="11" pageNumber="412">
<emphasis id="27CC465681474941FF16B9E2FEF804FB" bold="true" box="[199,352,1237,1262]" pageId="11" pageNumber="412">Distribution.</emphasis>
In the absence of a precise
<typeStatus id="CA0324E681474941FD4AB9E1FD5304FB" box="[667,715,1238,1262]" pageId="11" pageNumber="412">type</typeStatus>
locality, we consider the specimens from Beri Baor, Ramnagar, Kolkata, West Bengal,
<collectingCountry id="6DAFDAD481474941FE71B9CEFE440507" box="[416,476,1273,1298]" name="India" pageId="11" pageNumber="412">India</collectingCountry>
, as putative topotypes, given that the area is within the general
<typeStatus id="CA0324E681474941FB6EB9CDFB770507" box="[1215,1263,1274,1298]" pageId="11" pageNumber="412">type</typeStatus>
locality south eastern parts of Bengal. Although we were not able to procure specimens from
<collectingCountry id="6DAFDAD481474941FBC2B82AFB0E0523" box="[1043,1174,1309,1334]" name="Bangladesh" pageId="11" pageNumber="412">Bangladesh</collectingCountry>
for genetic analysis for logistical reasons, the
<emphasis id="27CC465681474941FE40B875FE43054C" box="[401,475,1346,1369]" italics="true" pageId="11" pageNumber="412">P. t i c t o</emphasis>
specimen collected from Dhaka,
<collectingCountry id="6DAFDAD481474941FC89B876FC46054F" box="[856,990,1345,1370]" name="Bangladesh" pageId="11" pageNumber="412">Bangladesh</collectingCountry>
, was morphologically indistinguishable from the Beri Baor specimens (
<figureCitation id="8D8386C181474941FE29B852FDC2056B" box="[504,602,1381,1406]" captionStart="FIGURE 3" captionStartId="5.[151,250,1946,1968]" captionTargetBox="[382,1204,871,1923]" captionTargetId="figure@5.[382,1204,871,1924]" captionTargetPageId="5" captionText="FIGURE 3. Topotype of Pethia ticto in life (A) from Beri Baor, West Bangal, India (BNHS FWF 127, 35.9 mm SL), humeral and caudal spots disappear immediately after collection, and (B) from Dhaka, Bangladesh (unregistered)." httpUri="https://zenodo.org/record/233767/files/figure.png" pageId="11" pageNumber="412">Figure 3</figureCitation>
). Specimens identified as
<taxonomicName id="D2B8E1C781474941FC53B851FC480568" box="[898,976,1382,1405]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC465681474941FC53B851FC480568" box="[898,976,1382,1405]" italics="true" pageId="11" pageNumber="412">P. ticto</emphasis>
</taxonomicName>
by
<bibRefCitation id="7129E7B581474941FC2AB852FA04056B" author="Linthoingambi" box="[1019,1436,1381,1406]" pageId="11" pageNumber="412" refString="Linthoingambi, I. &amp; Vishwanath, W. (2007) Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa, 1450, 45 - 56." type="journal article" year="2007">Linthoingambi &amp; Vishwanath (2007)</bibRefCitation>
from the Brahmaputra River System in Assam and Manipur are morphologically different (see Remarks) and hence may not be conspecific with
<taxonomicName id="D2B8E1C781474941FE38B899FDA105D0" box="[489,569,1454,1477]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC465681474941FE38B899FDA105D0" box="[489,569,1454,1477]" italics="true" pageId="11" pageNumber="412">P. ticto</emphasis>
</taxonomicName>
. Further, genetic data of specimens from Lucknow (
<accessionNumber id="0AEB07A781474941FB4DB89AFA8C05D3" box="[1180,1300,1453,1478]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ795474" pageId="11" pageNumber="412">JQ795474</accessionNumber>
,
<accessionNumber id="0AEB07A781474941FAF3B89AFA0005D3" box="[1314,1432,1453,1478]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ795475" pageId="11" pageNumber="412">JQ795475</accessionNumber>
,
<accessionNumber id="0AEB07A781474941FF46B8E6FE9505FF" box="[151,269,1489,1514]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ795476" pageId="11" pageNumber="412">JQ795476</accessionNumber>
,
<accessionNumber id="0AEB07A781474941FEC9B8E6FE1305FF" box="[280,395,1489,1514]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ795477" pageId="11" pageNumber="412">JQ795477</accessionNumber>
and
<accessionNumber id="0AEB07A781474941FE10B8E6FDAE05FF" box="[449,566,1489,1514]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ795478" pageId="11" pageNumber="412">JQ795478</accessionNumber>
) are also different from
<emphasis id="27CC465681474941FC95B8E5FC1605FC" box="[836,910,1490,1513]" italics="true" pageId="11" pageNumber="412">P. t i c t o</emphasis>
topotypes. Therefore, as currently understood,
<taxonomicName id="D2B8E1C781474941FF46B8C1FF7D0618" box="[151,229,1526,1549]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" sensu="stricto" species="ticto">
<emphasis id="27CC465681474941FF46B8C1FF7D0618" box="[151,229,1526,1549]" italics="true" pageId="11" pageNumber="412">P. ticto</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3CFFFB2D81474941FF3DB8C1FEE0061B" box="[236,376,1525,1550]" pageId="11" pageNumber="412" sensu="stricto">sensu stricto</taxonomicNameLabel>
is confined to the Beri Baor, Ramnagar, Kolkata, West Bengal,
<collectingCountry id="6DAFDAD481474941FB9BB8C2FB1E061B" box="[1098,1158,1525,1550]" name="India" pageId="11" pageNumber="412">India</collectingCountry>
, and Dhaka,
<collectingCountry id="6DAFDAD481474941FACBB8C2FA05061B" box="[1306,1437,1525,1550]" name="Bangladesh" pageId="11" pageNumber="412">Bangladesh</collectingCountry>
(
<figureCitation id="8D8386C181474941FF4FBB2EFF670627" box="[158,255,1561,1586]" captionStart="FIGURE 8" captionStartId="12.[151,249,998,1020]" captionTargetBox="[151,1436,301,976]" captionTargetId="figure@12.[151,1436,301,977]" captionTargetPageId="12" captionText="FIGURE 8. Collection location of topotypes of Pethia ticto from Beri Baor, West Bengal, India, and Dhaka, Bangladesh." httpUri="https://zenodo.org/record/233771/files/figure.png" pageId="11" pageNumber="412">Figure 8</figureCitation>
), both of which lie within the Gangetic floodplain.
</paragraph>
</subSubSection>
<subSubSection id="5DA2C9CF81474947FF16BB0AFE3800ED" lastPageId="13" lastPageNumber="414" pageId="11" pageNumber="412" type="discussion">
<paragraph id="15079A4481474941FF16BB0AFDE00760" blockId="11.[151,1437,151,2018]" pageId="11" pageNumber="412">
<emphasis id="27CC465681474941FF16BB0AFEA30643" bold="true" box="[199,315,1597,1622]" pageId="11" pageNumber="412">Remarks.</emphasis>
<bibRefCitation id="7129E7B581474941FE94BB0AFD870643" author="McClelland" box="[325,543,1597,1622]" pageId="11" pageNumber="412" refString="McClelland, J. (1839) Indian Cyprinidae. Asiatic Researches, 19, 217 - 471." type="journal article" year="1839">McClelland (1839)</bibRefCitation>
mentioned two names
<taxonomicName id="D2B8E1C781474941FCE2BB09FBB50640" box="[819,1069,1597,1621]" class="Actinopterygii" family="Cyprinidae" genus="Cyprinus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="bimaculatus">
<emphasis id="27CC465681474941FCE2BB09FBB50640" box="[819,1069,1597,1621]" italics="true" pageId="11" pageNumber="412">Cyprinus bimaculatus</emphasis>
</taxonomicName>
and
<taxonomicName id="D2B8E1C781474941FBBABB09FAD50640" box="[1131,1357,1597,1621]" class="Actinopterygii" family="Cyprinidae" genus="Cyprinus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="quadrimaculatus">
<emphasis id="27CC465681474941FBBABB09FAD50640" box="[1131,1357,1597,1621]" italics="true" pageId="11" pageNumber="412">C. quadrimaculatus</emphasis>
</taxonomicName>
, when redescribing
<taxonomicName id="D2B8E1C781474941FEFCBB55FE4B066C" box="[301,467,1634,1657]" class="Actinopterygii" family="Cyprinidae" genus="Systomus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC465681474941FEFCBB55FE4B066C" box="[301,467,1634,1657]" italics="true" pageId="11" pageNumber="412">Systomus ticto</emphasis>
</taxonomicName>
. However, as these name were not accompanied by any descriptive or diagnostic information,
<bibRefCitation id="7129E7B581474941FEFDBBB1FDA9068B" box="[300,561,1669,1694]" pageId="11" pageNumber="412" refString="Katwate, U., Baby, F., Raghavan, R. &amp; Dahanukar, N. (2014 a) The identity of Pethia punctata, a senior synonym of P. muvattupuzhaensis (Teleostei: Cyprinidae). Zootaxa, 3884 (3), 201 - 221. http: // dx. doi. org / 10.11646 / zootaxa. 3884.3.1" type="journal article">
Katwate
<emphasis id="27CC465681474941FE45BBB1FE480688" box="[404,464,1669,1693]" italics="true" pageId="11" pageNumber="412">et al.</emphasis>
(2014a)
</bibRefCitation>
considered them to be
<emphasis id="27CC465681474941FC99BBB1FC460688" box="[840,990,1669,1693]" italics="true" pageId="11" pageNumber="412">nomina nuda</emphasis>
, not synonyms of
<emphasis id="27CC465681474941FB6DBBB1FA920688" box="[1212,1290,1670,1693]" italics="true" pageId="11" pageNumber="412">P. t i c t o</emphasis>
, contrary to
<bibRefCitation id="7129E7B581474941FF46BB9EFED106D7" author="Bleeker" box="[151,329,1705,1730]" pageId="11" pageNumber="412" refString="Bleeker (1844) Enumeratio specierum piscium hucusque in Archipelago Indico observatarum, adjectis habitationibus citationibusque ubi descriptions earum recentiores reperiuntur. Weber and de Beaufort, 15, 1 - 276." type="journal article" year="1844">Bleeker (1844)</bibRefCitation>
.
<bibRefCitation id="7129E7B581474941FE8ABB9DFE7906D7" author="Day" box="[347,481,1706,1730]" pageId="11" pageNumber="412" refString="Day, F. (1878) The Fishes of India; Being a Natural History of the Fishes Known to Inhabit the Seas and Freshwaters of India, Burma and Ceylon. William Dawson &amp; Sons, London, xx + 778 pp, 196 pls." type="book" year="1878">Day (1878)</bibRefCitation>
considered
<taxonomicName id="D2B8E1C781474941FDA7BB9DFB8306D7" authority="Jerdon, 1849" authorityName="Jerdon" authorityYear="1849" box="[630,1051,1705,1730]" class="Actinopterygii" family="Cyprinidae" genus="Systomus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="tripunctatus">
<emphasis id="27CC465681474941FDA7BB9DFCEB06D4" box="[630,883,1706,1729]" italics="true" pageId="11" pageNumber="412">Systomus tripunctatus</emphasis>
<bibRefCitation id="7129E7B581474941FC50BB9EFB8306D7" author="Jerdon" box="[897,1051,1705,1730]" pageId="11" pageNumber="412" refString="Jerdon, T. C. (1849) On the freshwater fishes of southern India. Madras Journal of Literature and Science, 15 (2), 302 - 346." type="journal article" year="1849">Jerdon, 1849</bibRefCitation>
</taxonomicName>
, as a synonym of
<taxonomicName id="D2B8E1C781474941FAD5BB9EFA0D06D4" box="[1284,1429,1705,1729]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC465681474941FAD5BB9EFA0D06D4" box="[1284,1429,1705,1729]" italics="true" pageId="11" pageNumber="412">Barbus ticto</emphasis>
</taxonomicName>
. However, the colour pattern of
<taxonomicName id="D2B8E1C781474941FE2FBBF9FD3B06F0" box="[510,675,1742,1765]" class="Actinopterygii" family="Cyprinidae" genus="Systomus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="tripunctatus">
<emphasis id="27CC465681474941FE2FBBF9FD3B06F0" box="[510,675,1742,1765]" italics="true" pageId="11" pageNumber="412">S. tripunctatus</emphasis>
</taxonomicName>
provided by
<bibRefCitation id="7129E7B581474941FCECBBFAFC4306F3" author="Jerdon" box="[829,987,1741,1766]" pageId="11" pageNumber="412" refString="Jerdon, T. C. (1849) On the freshwater fishes of southern India. Madras Journal of Literature and Science, 15 (2), 302 - 346." type="journal article" year="1849">Jerdon (1849)</bibRefCitation>
does not match with that of
<emphasis id="27CC465681474941FAF9BBF9FAEC06F0" box="[1320,1396,1742,1765]" italics="true" pageId="11" pageNumber="412">P. t i c t o</emphasis>
by having three spots over body, two black spots under end of the dorsal and one at the base of the tail vs. two spots over body, one humeral and one on caudal peduncle in
<taxonomicName id="D2B8E1C781474941FCDEBA21FCC60738" box="[783,862,1814,1837]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC465681474941FCDEBA21FCC60738" box="[783,862,1814,1837]" italics="true" pageId="11" pageNumber="412">P. ticto</emphasis>
</taxonomicName>
. Further,
<taxonomicName id="D2B8E1C781474941FC1FBA21FBEB0738" box="[974,1139,1814,1837]" class="Actinopterygii" family="Cyprinidae" genus="Systomus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="tripunctatus">
<emphasis id="27CC465681474941FC1FBA21FBEB0738" box="[974,1139,1814,1837]" italics="true" pageId="11" pageNumber="412">S. tripunctatus</emphasis>
</taxonomicName>
was described from near the coast in Canara, in Southern India, and is unlikely to have a range that extends to Bengal. We thus do not consider this species a synonym of
<emphasis id="27CC465681474941FDF2BA69FDF60760" box="[547,622,1886,1909]" italics="true" pageId="11" pageNumber="412">P. t icto</emphasis>
.
</paragraph>
<paragraph id="15079A4481474946FF16BAB6FD8D00ED" blockId="11.[151,1437,151,2018]" lastBlockId="12.[151,1436,151,248]" lastPageId="12" lastPageNumber="413" pageId="11" pageNumber="412">
The redescription of
<taxonomicName id="D2B8E1C781474941FE13BAB5FD8D078C" box="[450,533,1922,1945]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="412" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC465681474941FE13BAB5FD8D078C" box="[450,533,1922,1945]" italics="true" pageId="11" pageNumber="412">P. ticto</emphasis>
</taxonomicName>
by
<bibRefCitation id="7129E7B581474941FD9BBAB6FC63078F" author="Linthoingambi" box="[586,1019,1921,1946]" pageId="11" pageNumber="412" refString="Linthoingambi, I. &amp; Vishwanath, W. (2007) Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa, 1450, 45 - 56." type="journal article" year="2007">Linthoingambi &amp; Vishwanath (2007)</bibRefCitation>
was based on specimens from the Brahmaputra River System in Assam and Manipur. Topotypic material from the Ganges basin used in the present study differs from the Brahmaputra form by a lesser number of transverse scale rows ½4/1/3½4 (vs. ½5/1/5½), lesser body depth 31.535.8%SL (vs. 36.845.0%SL) and a more compressed body width at dorsal fin origin (body width 10.715.4% SL, vs. 16.518.4% SL). Further studies are required for testing the conspecificity of these Gangetic and Brahmaputra forms.
</paragraph>
<caption id="41C7CACC81404946FF46BED1FAFE03E9" box="[151,1382,998,1020]" httpUri="https://zenodo.org/record/233771/files/figure.png" pageId="12" pageNumber="413" targetBox="[151,1436,301,976]" targetPageId="12">
<paragraph id="15079A4481404946FF46BED1FAFE03E9" blockId="12.[151,1382,998,1020]" box="[151,1382,998,1020]" pageId="12" pageNumber="413">
<emphasis id="27CC465681404946FF46BED1FE8A03EE" bold="true" box="[151,274,998,1020]" pageId="12" pageNumber="413">FIGURE 8.</emphasis>
Collection location of topotypes of
<taxonomicName id="D2B8E1C781404946FD51BED1FD6D03E9" box="[640,757,998,1020]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC465681404946FD51BED1FD6D03E9" box="[640,757,998,1020]" italics="true" pageId="12" pageNumber="413">Pethia ticto</emphasis>
</taxonomicName>
from Beri Baor, West Bengal, India, and Dhaka, Bangladesh.
</paragraph>
</caption>
<paragraph id="15079A4481404946FF16B911FAA706B6" blockId="12.[151,1436,1062,2023]" pageId="12" pageNumber="413">
Other congeners of
<taxonomicName id="D2B8E1C781404946FE7DB911FDA8042B" box="[428,560,1062,1086]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC465681404946FE7DB911FDA8042B" box="[428,560,1062,1086]" italics="true" pageId="12" pageNumber="413">Pethia ticto</emphasis>
</taxonomicName>
with an incomplete lateral line include
<taxonomicName id="D2B8E1C781404946FC2FB910FAB7042A" authority="Hamilton, 1822" authorityName="Hamilton" authorityYear="1822" box="[1022,1327,1062,1087]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="gelius">
<emphasis id="27CC465681404946FC2FB910FBC7042B" box="[1022,1119,1062,1086]" italics="true" pageId="12" pageNumber="413">P. gelius</emphasis>
(
<bibRefCitation id="7129E7B581404946FBA1B911FAB0042A" author="Hamilton" box="[1136,1320,1062,1087]" pageId="12" pageNumber="413" refString="Hamilton, F. (1822) An Account of the Fishes of River Ganges and its Branches. George Ramsay and Co., London, vii + 405 pp, 39 pls." type="book" year="1822">Hamilton, 1822</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FAECB910FEB60477" authority="Knight, 2013" authorityName="Knight" authorityYear="2013" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="aurea">
<emphasis id="27CC465681404946FAECB910FA04042B" box="[1341,1436,1063,1086]" italics="true" pageId="12" pageNumber="413">P. aurea</emphasis>
<bibRefCitation id="7129E7B581404946FF46B97DFEB60477" author="Knight" box="[151,302,1098,1123]" pageId="12" pageNumber="413" refString="Knight, J. D. M. (2013) Pethia aurea (Teleostei: Cyprinidae), a new species of barb from West Bengal, India, with redescription of P. gelius and P. canius. Zootaxa, 3700 (1), 173 - 184. http: // dx. doi. org / 10.11646 / zootaxa. 3700.1.7" type="journal article" year="2013">Knight, 2013</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FEEBB97CFD1F0477" authority="Hamilton, 1822" authorityName="Hamilton" authorityYear="1822" box="[314,647,1098,1123]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="phutunio">
<emphasis id="27CC465681404946FEEBB97CFE220477" box="[314,442,1098,1122]" italics="true" pageId="12" pageNumber="413">P. phutunio</emphasis>
(
<bibRefCitation id="7129E7B581404946FE18B97DFD180477" author="Hamilton" box="[457,640,1098,1123]" pageId="12" pageNumber="413" refString="Hamilton, F. (1822) An Account of the Fishes of River Ganges and its Branches. George Ramsay and Co., London, vii + 405 pp, 39 pls." type="book" year="1822">Hamilton, 1822</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FD45B97CFC5E0477" authority="Hamilton, 1822" authorityName="Hamilton" authorityYear="1822" box="[660,966,1098,1123]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="canius">
<emphasis id="27CC465681404946FD45B97CFD610477" box="[660,761,1099,1122]" italics="true" pageId="12" pageNumber="413">P. canius</emphasis>
(
<bibRefCitation id="7129E7B581404946FCD9B97DFC260477" author="Hamilton" box="[776,958,1098,1123]" pageId="12" pageNumber="413" refString="Hamilton, F. (1822) An Account of the Fishes of River Ganges and its Branches. George Ramsay and Co., London, vii + 405 pp, 39 pls." type="book" year="1822">Hamilton, 1822</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FC02B97CFE310492" authority="Knight, Rema Devi, Indra &amp; Arunachalam, 2012" authorityName="Knight, Rema Devi, Indra &amp; Arunachalam" authorityYear="2012" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="nigripinna">
<emphasis id="27CC465681404946FC02B97CFBFD0477" box="[979,1125,1099,1122]" italics="true" pageId="12" pageNumber="413">P. nigripinna</emphasis>
(Knight, Rema
<bibRefCitation id="7129E7B581404946FACDB97DFE390492" author="Devi" pageId="12" pageNumber="413" refString="Knight, J. D. M., Devi, K. R., Indra, T. J. &amp; Arunachalam, M. (2012) A new species of barb Puntius nigripinnis (Teleostei: Cyprinidae) from southern Western Ghats, India. Journal of Threatened Taxa, 4, 2409 - 2416. http: // dx. doi. org / 10.11609 / JoTT. o 3014.2409 - 16" type="journal article" year="2012">Devi, Indra &amp; Arunachalam, 2012</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FE6BB958FCC60492" authority="Kullander, 2008" authorityName="Kullander" authorityYear="2008" box="[442,862,1134,1159]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="macrogramma">
<emphasis id="27CC465681404946FE6BB958FD180493" box="[442,640,1135,1158]" italics="true" pageId="12" pageNumber="413">P. macrogramma</emphasis>
(Kullander, 2008)
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FCBEB958FEEB04BF" authority="Meegaskumbura, Silva, Maduwage &amp; Pethiyagoda, 2008" authorityName="Meegaskumbura, Silva, Maduwage &amp; Pethiyagoda" authorityYear="2008" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="reval">
<emphasis id="27CC465681404946FCBEB958FC510493" box="[879,969,1134,1158]" italics="true" pageId="12" pageNumber="413">P. reval</emphasis>
(
<bibRefCitation id="7129E7B581404946FC0FB959FEF304BF" author="Meegaskumbura" pageId="12" pageNumber="413" refString="Meegaskumbura, M., Silva, A., Maduwage, K. &amp; Pethiyagoda, R. (2008) Puntius reval, a new barb from Sri Lanka (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 19, 141 - 152." type="journal article" year="2008">Meegaskumbura, Silva, Maduwage &amp; Pethiyagoda, 2008</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FE53B9A4FC8904BF" authority="Kullander, 2008" authorityName="Kullander" authorityYear="2008" box="[386,785,1170,1195]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="erythromycter">
<emphasis id="27CC465681404946FE53B9A4FDA504BF" box="[386,573,1170,1194]" italics="true" pageId="12" pageNumber="413">P. erythromycter</emphasis>
(Kullander, 2008)
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FCCFB9A4FBC504BE" authority="Gunther, 1868" authorityName="Gunther" authorityYear="1868" box="[798,1117,1170,1195]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="cumingii">
<emphasis id="27CC465681404946FCCFB9A4FC0504BF" box="[798,925,1171,1194]" italics="true" pageId="12" pageNumber="413">P. cumingii</emphasis>
(
<bibRefCitation id="7129E7B581404946FC7FB9A5FBCD04BE" author="Gunther" box="[942,1109,1170,1195]" pageId="12" pageNumber="413" refString="Gunther, A. (1868) Catalogue of the fishes in the British Museum. Catalogue of the Physostomi, containing the families Heteropygii, Cyprinidae, Gonorhynchidae, Hyodontidae, Osteoglossidae, Clupeidae, Chirocenridae, Alepocephalidae, Notopteridae, Halosauridae, in the collection of the British Museum. British Museum, London, xx + 512 pp." type="book" year="1868">Günther, 1868</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FBBAB9A4FF4304DA" authority="Kullander &amp; Fang, 2005" authorityName="Kullander &amp; Fang" authorityYear="2005" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="didi">
<emphasis id="27CC465681404946FBBAB9A4FB2C04BF" box="[1131,1204,1170,1194]" italics="true" pageId="12" pageNumber="413">P. didi</emphasis>
(
<bibRefCitation id="7129E7B581404946FB15B9A5FF4B04DA" author="Kullander" pageId="12" pageNumber="413" refString="Kullander, S. O. &amp; Fang, F. (2005) Two new species of Puntius from northern Myanmar (Teleostei: Cyprinidae). Copeia (2), 290 - 302. http: // dx. doi. org / 10.1643 / CI- 04 - 138 R 1" type="journal article" year="2005">Kullander &amp; Fang, 2005</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FF3FB980FD5004DA" authority="Kullander &amp; Britz, 2008" authorityName="Kullander &amp; Britz" authorityYear="2008" box="[238,712,1206,1231]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="padamya">
<emphasis id="27CC465681404946FF3FB980FEE104DB" box="[238,377,1206,1230]" italics="true" pageId="12" pageNumber="413">P. padamya</emphasis>
(
<bibRefCitation id="7129E7B581404946FE5EB981FD5804DA" author="Kullander" box="[399,704,1206,1231]" pageId="12" pageNumber="413" refString="Kullander, S. O. &amp; Britz, R. (2008) Puntius padamya, a new species of cyprinid fish from Myanmar (Teleostei: Cyprinidae). Electronic Journal of Ichthyology, 2, 56 - 66." type="journal article" year="2008">Kullander &amp; Britz, 2008</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FD0DB980FA8C04DA" authority="Arunkumar &amp; Tombi Singh, 2003" authorityName="Arunkumar &amp; Tombi Singh" authorityYear="2003" box="[732,1300,1206,1231]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="yuensis">
<emphasis id="27CC465681404946FD0DB980FCCA04DB" box="[732,850,1207,1230]" italics="true" pageId="12" pageNumber="413">P. yuensis</emphasis>
(
<bibRefCitation id="7129E7B581404946FCB8B981FA9404DA" author="Arunkumar" box="[873,1292,1206,1231]" pageId="12" pageNumber="413" refString="Arunkumar, L. &amp; Tombi Singh, H. T. (2003) Two new species of puntiid fish from the Yu River system of Manipur. Journal of the Bombay Natural History Society, 99, 481 - 487." type="journal article" year="2003">Arunkumar &amp; Tombi Singh, 2003</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FAF6B980FDCC04E7" authority="Linthoingambi &amp; Vishwanath, 2007" authorityName="Linthoingambi &amp; Vishwanath" authorityYear="2007" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="khugae">
<emphasis id="27CC465681404946FAF6B980FA0404DB" box="[1319,1436,1206,1230]" italics="true" pageId="12" pageNumber="413">P. khugae</emphasis>
(
<bibRefCitation id="7129E7B581404946FF4FB9EDFDD404E7" author="Linthoingambi" box="[158,588,1242,1267]" pageId="12" pageNumber="413" refString="Linthoingambi, I. &amp; Vishwanath, W. (2007) Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa, 1450, 45 - 56." type="journal article" year="2007">Linthoingambi &amp; Vishwanath, 2007</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="27CC465681404946FDB7B9ECFD7804E7" box="[614,736,1242,1266]" italics="true" pageId="12" pageNumber="413">
<taxonomicName id="D2B8E1C781404946FDB7B9ECFD5304E7" box="[614,715,1242,1266]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="sharm">P. sharm</taxonomicName>
ai
</emphasis>
(
<bibRefCitation id="7129E7B581404946FD29B9EDFBC804E7" author="Menon" box="[760,1104,1242,1267]" pageId="12" pageNumber="413" refString="Menon, A. G. K. &amp; Rema Devi, K. (1993) Puntius sharmai, a new cyprinid fish from Madras. Journal of the Bombay Natural History Society, 89, 353 - 354." type="journal article" year="1993">Menon &amp; Rema Devi, 1993</bibRefCitation>
),
<emphasis id="27CC465681404946FBBBB9ECFB2004E7" box="[1130,1208,1243,1266]" italics="true" pageId="12" pageNumber="413">P. a t r a</emphasis>
(
<bibRefCitation id="7129E7B581404946FB01B9EDFEFE0502" author="Linthoingambi" pageId="12" pageNumber="413" refString="Linthoingambi, I. &amp; Vishwanath, W. (2007) Two new fish species of the genus Puntius Hamilton (Cyprinidae) from Manipur, India, with notes on P. ticto (Hamilton) and P. stoliczkanus (Day). Zootaxa, 1450, 45 - 56." type="journal article" year="2007">Linthoingambi &amp; Vishwanath, 2007</bibRefCitation>
),
<taxonomicName id="D2B8E1C781404946FEABB9C8FCFA0502" authority="Kottelat &amp; Pethiyagoda, 1991" authorityName="Kottelat &amp; Pethiyagoda" authorityYear="1991" box="[378,866,1278,1303]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="bandula">
<emphasis id="27CC465681404946FEABB9C8FE6B0503" box="[378,499,1278,1302]" italics="true" pageId="12" pageNumber="413">P. bandula</emphasis>
(
<bibRefCitation id="7129E7B581404946FDD3B9C9FCC10502" author="Kottelat" box="[514,857,1278,1303]" pageId="12" pageNumber="413" refString="Kottelat, M. &amp; Pethiyagoda, R. (1991) Descriptions of three new species of cyprinid fishes from Sri Lanka. In: Pethiyagoda, R. (Ed.), Freshwater fishes of Sri Lanka. Wildlife Heritage Trust of Sri Lanka, Colombo, pp. 298 - 313" type="book chapter" year="1991">Kottelat &amp; Pethiyagoda, 1991</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FCBFB9C8FB410502" authority="Hamilton, 1822" authorityName="Hamilton" authorityYear="1822" box="[878,1241,1278,1303]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="conchonius">
<emphasis id="27CC465681404946FCBFB9C8FB930503" box="[878,1035,1278,1302]" italics="true" pageId="12" pageNumber="413">P. conchonius</emphasis>
(
<bibRefCitation id="7129E7B581404946FBCBB9C9FB490502" author="Hamilton" box="[1050,1233,1278,1303]" pageId="12" pageNumber="413" refString="Hamilton, F. (1822) An Account of the Fishes of River Ganges and its Branches. George Ramsay and Co., London, vii + 405 pp, 39 pls." type="book" year="1822">Hamilton, 1822</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FB34B9C8FEF9052F" authority="Kullander, 2008" authorityName="Kullander" authorityYear="2008" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="nankyweensis">
<emphasis id="27CC465681404946FB34B9C8FA040503" box="[1253,1436,1278,1302]" italics="true" pageId="12" pageNumber="413">P. nankyweensis</emphasis>
(Kullander, 2008)
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FEBCB814FCDF052F" authority="Vishwanath &amp; Laisram, 2004" authorityName="Vishwanath &amp; Laisram" authorityYear="2004" box="[365,839,1314,1339]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="ornatus">
<emphasis id="27CC465681404946FEBCB814FE47052F" box="[365,479,1315,1338]" italics="true" pageId="12" pageNumber="413">P. ornatus</emphasis>
(
<bibRefCitation id="7129E7B581404946FE3FB815FCA7052F" author="Vishwanath" box="[494,831,1314,1339]" pageId="12" pageNumber="413" refString="Vishwanath, W. &amp; Laisram, J. (2004) Two new species of Puntius Hamilton-Buchanan (Cypriniformes: Cyprinidae) from Manipur, India, with an account of Puntius species from the state. Journal of the Bombay Natural History Society, 101, 130 - 137." type="journal article" year="2004">Vishwanath &amp; Laisram, 2004</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FC82B814FA92052F" authority="Mercy &amp; Jacob, 2007" authorityName="Mercy &amp; Jacob" authorityYear="2007" box="[851,1290,1314,1339]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="pookodensis">
<emphasis id="27CC465681404946FC82B814FC62052F" box="[851,1018,1314,1338]" italics="true" pageId="12" pageNumber="413">P. pookodensis</emphasis>
(
<bibRefCitation id="7129E7B581404946FBD9B815FA9A052F" author="Mercy" box="[1032,1282,1314,1339]" pageId="12" pageNumber="413" refString="Mercy, T. V. A. &amp; Jacob, E. (2007) A new species of Teleostei: Puntius pookodensis (Cyprinidae) from Wayanad, Kerala, India. Journal of the Bombay Natural History Society, 104, 76 - 78." type="journal article" year="2007">Mercy &amp; Jacob, 2007</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="27CC465681404946FAC7B814FA00052F" box="[1302,1432,1314,1338]" italics="true" pageId="12" pageNumber="413">P. s ha l y n i u s</emphasis>
(
<bibRefCitation id="7129E7B581404946FF4FB871FE50054A" author="Yazdani" box="[158,456,1350,1375]" pageId="12" pageNumber="413" refString="Yazdani, G. M. &amp; Talukdar, S. (1975) A new species of Puntius (Cypriniformes: Cyprinidae) from Khasi and Jaintia Hills (Meghalaya), India. Journal of the Bombay Natural History Society, 72, 218 - 221." type="journal article" year="1975">Yazdani &amp; Talukdar, 1975</bibRefCitation>
),
<emphasis id="27CC465681404946FE0AB870FDAC054B" box="[475,564,1350,1374]" italics="true" pageId="12" pageNumber="413">P. t he l ys</emphasis>
(Kullander, 2008),
<taxonomicName id="D2B8E1C781404946FCC3B870FA00054A" authority="Menon, Rema Devi &amp; Viswanath, 2000" authorityName="Menon, Rema Devi &amp; Viswanath" authorityYear="2000" box="[786,1432,1350,1375]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="manipurensis">
<emphasis id="27CC465681404946FCC3B870FC5B054B" box="[786,963,1351,1374]" italics="true" pageId="12" pageNumber="413">P. manipurensis</emphasis>
(Menon, Rema Devi &amp; Viswanath, 2000)
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FF46B85CFC1D0597" authority="Katwate, Paingankar, Raghavan, Dahanukar, 2014" authorityName="Katwate, Paingankar, Raghavan, Dahanukar" authorityYear="2014" box="[151,901,1386,1411]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="longicauda">
<emphasis id="27CC465681404946FF46B85CFEAC0597" box="[151,308,1386,1410]" italics="true" pageId="12" pageNumber="413">P. longicauda</emphasis>
<bibRefCitation id="7129E7B581404946FEEEB85CFC1D0597" author="Katwate" box="[319,901,1386,1411]" pageId="12" pageNumber="413" refString="Katwate, U., Katwate, C., Raghavan, R., Paingankar, M. S. &amp; Dahanukar, N. (2014 c) Pethia lutea, a new species of barb (Teleostei: Cyprinidae) and new records of P. punctata from northern Western Ghats of India. Journal of Threatened Taxa, 6, 5797 - 5818. http: // dx. doi. org / 10.11609 / JoTT. o 3929.5797 - 818" type="journal article" year="2014">Katwate, Paingankar, Raghavan, Dahanukar, 2014</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FC45B85CFF4005B2" authority="Arunkumar &amp; Tombi Singh, 2003" authorityName="Arunkumar &amp; Tombi Singh" authorityYear="2003" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="meingangbii">
<emphasis id="27CC465681404946FC45B85CFBA70597" box="[916,1087,1386,1410]" italics="true" pageId="12" pageNumber="413">P. meingangbii</emphasis>
(
<bibRefCitation id="7129E7B581404946FB80B85DFF4905B2" author="Arunkumar" pageId="12" pageNumber="413" refString="Arunkumar, L. &amp; Tombi Singh, H. T. (2003) Two new species of puntiid fish from the Yu River system of Manipur. Journal of the Bombay Natural History Society, 99, 481 - 487." type="journal article" year="2003">Arunkumar &amp; Tombi Singh, 2003</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="D2B8E1C781404946FEC4B8B8FD7605B2" authority="Deraniyagala, 1956" authorityName="Deraniyagala" authorityYear="1956" box="[277,750,1422,1447]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="melanomaculata">
<emphasis id="27CC465681404946FEC4B8B8FE7705B3" box="[277,495,1422,1446]" italics="true" pageId="12" pageNumber="413">P. melanomaculata</emphasis>
(
<bibRefCitation id="7129E7B581404946FDD0B8B9FD7D05B2" author="Deraniyagala" box="[513,741,1422,1447]" pageId="12" pageNumber="413" refString="Deraniyagala, P. E. P. (1956) Two new subspecies and one new species of cyprinoid fishes from Ceylon. Proceedings of the 12 th Annual Sessions of the Ceylon Association for the Advancement of Science, 1, 34 - 35." type="journal article" year="1956">Deraniyagala, 1956</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="D2B8E1C781404946FD2DB8B9FC1905B3" box="[764,897,1422,1446]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC465681404946FD2DB8B9FC1905B3" box="[764,897,1422,1446]" italics="true" pageId="12" pageNumber="413">Pethia ticto</emphasis>
</taxonomicName>
differs from these by having 612 lateral-line pored scales (vs.
<quantity id="D24037A181404946FE8EB885FE3005DE" box="[351,424,1458,1483]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="12" pageNumber="413" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">34 in</quantity>
<emphasis id="27CC465681404946FE60B884FD9605DF" box="[433,526,1458,1482]" italics="true" pageId="12" pageNumber="413">P. g e l i us</emphasis>
,
<taxonomicName id="D2B8E1C781404946FDF3B884FD1905DF" box="[546,641,1459,1482]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="aurea">
<emphasis id="27CC465681404946FDF3B884FD1905DF" box="[546,641,1459,1482]" italics="true" pageId="12" pageNumber="413">P. aurea</emphasis>
</taxonomicName>
and
<taxonomicName id="D2B8E1C781404946FD6AB884FCA505DF" box="[699,829,1458,1482]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="phutunio">
<emphasis id="27CC465681404946FD6AB884FCA505DF" box="[699,829,1458,1482]" italics="true" pageId="12" pageNumber="413">P. phutunio</emphasis>
</taxonomicName>
;
<quantity id="D24037A181404946FC9FB885FC0F05DE" box="[846,919,1458,1483]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" pageId="12" pageNumber="413" unit="in" value="4.0" valueMax="5.0" valueMin="3.0">35 in</quantity>
<taxonomicName id="D2B8E1C781404946FC71B884FB9F05DF" box="[928,1031,1459,1482]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="canius">
<emphasis id="27CC465681404946FC71B884FB9F05DF" box="[928,1031,1459,1482]" italics="true" pageId="12" pageNumber="413">P. canius</emphasis>
</taxonomicName>
and
<taxonomicName id="D2B8E1C781404946FB93B884FB4E05DF" box="[1090,1238,1459,1482]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="nigripinna">
<emphasis id="27CC465681404946FB93B884FB4E05DF" box="[1090,1238,1459,1482]" italics="true" pageId="12" pageNumber="413">P. nigripinna</emphasis>
</taxonomicName>
; and
<quantity id="D24037A181404946FAC8B885FAE605DE" box="[1305,1406,1458,1483]" metricMagnitude="-1" metricUnit="m" metricValue="4.699" metricValueMax="5.842" metricValueMin="3.556" pageId="12" pageNumber="413" unit="in" value="18.5" valueMax="23.0" valueMin="14.0">1423 in</quantity>
<taxonomicName id="D2B8E1C781404946FA56B884FEA305FB" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="macrogramma">
<emphasis id="27CC465681404946FA56B884FEA305FB" italics="true" pageId="12" pageNumber="413">P. macrogramma</emphasis>
</taxonomicName>
), 2326 scales in lateral-scale row (vs.
<quantity id="D24037A181404946FCD6B8E0FCF505FA" box="[775,877,1494,1519]" metricMagnitude="-1" metricUnit="m" metricValue="4.953" metricValueMax="5.334" metricValueMin="4.572" pageId="12" pageNumber="413" unit="in" value="19.5" valueMax="21.0" valueMin="18.0">1821 in</quantity>
<taxonomicName id="D2B8E1C781404946FCA7B8E0FC5405FB" box="[886,972,1494,1518]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="reval">
<emphasis id="27CC465681404946FCA7B8E0FC5405FB" box="[886,972,1494,1518]" italics="true" pageId="12" pageNumber="413">P. reval</emphasis>
</taxonomicName>
;
<quantity id="D24037A181404946FC0CB8E0FBDB05FA" box="[989,1091,1494,1519]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" metricValueMax="5.08" metricValueMin="4.572" pageId="12" pageNumber="413" unit="in" value="19.0" valueMax="20.0" valueMin="18.0">1820 in</quantity>
<taxonomicName id="D2B8E1C781404946FB9DB8E0FA9005FB" box="[1100,1288,1494,1518]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="erythromycter">
<emphasis id="27CC465681404946FB9DB8E0FA9005FB" box="[1100,1288,1494,1518]" italics="true" pageId="12" pageNumber="413">P. erythromycter</emphasis>
</taxonomicName>
;
<quantity id="D24037A181404946FAC8B8E0FAE605FA" box="[1305,1406,1494,1519]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" metricValueMax="5.334" metricValueMin="4.826" pageId="12" pageNumber="413" unit="in" value="20.0" valueMax="21.0" valueMin="19.0">1921 in</quantity>
<taxonomicName id="D2B8E1C781404946FA56B8E0FF610607" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="cumingii">
<emphasis id="27CC465681404946FA56B8E0FF610607" italics="true" pageId="12" pageNumber="413">P. cumingii</emphasis>
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FED7B8CCFED50607" box="[262,333,1530,1554]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="didi">
<emphasis id="27CC465681404946FED7B8CCFED50607" box="[262,333,1530,1554]" italics="true" pageId="12" pageNumber="413">P. didi</emphasis>
</taxonomicName>
and
<taxonomicName id="D2B8E1C781404946FE53B8CCFD9B0607" box="[386,515,1530,1554]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="padamya">
<emphasis id="27CC465681404946FE53B8CCFD9B0607" box="[386,515,1530,1554]" italics="true" pageId="12" pageNumber="413">P. padamya</emphasis>
</taxonomicName>
;
<quantity id="D24037A181404946FDC8B8CDFDE30606" box="[537,635,1530,1555]" metricMagnitude="-1" metricUnit="m" metricValue="5.461" metricValueMax="5.588" metricValueMin="5.334" pageId="12" pageNumber="413" unit="in" value="21.5" valueMax="22.0" valueMin="21.0">2122 in</quantity>
<taxonomicName id="D2B8E1C781404946FD53B8CCFD770607" box="[642,751,1531,1554]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="yuensis">
<emphasis id="27CC465681404946FD53B8CCFD770607" box="[642,751,1531,1554]" italics="true" pageId="12" pageNumber="413">P. yuensis</emphasis>
</taxonomicName>
;
<quantity id="D24037A181404946FD2CB8CDFCF80606" box="[765,864,1530,1555]" metricMagnitude="-1" metricUnit="m" metricValue="7.366" metricValueMax="7.62" metricValueMin="7.112" pageId="12" pageNumber="413" unit="in" value="29.0" valueMax="30.0" valueMin="28.0">2830 in</quantity>
<taxonomicName id="D2B8E1C781404946FCB7B8CCFC4A0607" box="[870,978,1530,1554]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="khugae">
<emphasis id="27CC465681404946FCB7B8CCFC4A0607" box="[870,978,1530,1554]" italics="true" pageId="12" pageNumber="413">P. khugae</emphasis>
</taxonomicName>
and
<quantity id="D24037A181404946FBD6B8CDFBD80606" box="[1031,1088,1530,1555]" metricMagnitude="0" metricUnit="m" metricValue="1.0668" pageId="12" pageNumber="413" unit="in" value="42.0">42 in</quantity>
<taxonomicName id="D2B8E1C781404946FB97B8CCFB250607" box="[1094,1213,1530,1554]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="sharmai">
<emphasis id="27CC465681404946FB97B8CCFB250607" box="[1094,1213,1530,1554]" italics="true" pageId="12" pageNumber="413">P. sharmai</emphasis>
</taxonomicName>
), two bands of dark spots on dorsal fin (vs. no bands in
<emphasis id="27CC465681404946FDF3BB28FDF10623" box="[546,617,1567,1590]" italics="true" pageId="12" pageNumber="413">P. at r a</emphasis>
,
<taxonomicName id="D2B8E1C781404946FD51BB28FD600623" box="[640,760,1566,1590]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="bandula">
<emphasis id="27CC465681404946FD51BB28FD600623" box="[640,760,1566,1590]" italics="true" pageId="12" pageNumber="413">P. bandula</emphasis>
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FCDDBB28FC3E0623" box="[780,934,1566,1590]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="conchonius">
<emphasis id="27CC465681404946FCDDBB28FC3E0623" box="[780,934,1566,1590]" italics="true" pageId="12" pageNumber="413">P. conchonius</emphasis>
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FC6ABB28FBE80623" box="[955,1136,1566,1590]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="nankyweensis">
<emphasis id="27CC465681404946FC6ABB28FBE80623" box="[955,1136,1566,1590]" italics="true" pageId="12" pageNumber="413">P. nankyweensis</emphasis>
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FBAFBB28FB770623" box="[1150,1263,1567,1590]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="ornatus">
<emphasis id="27CC465681404946FBAFBB28FB770623" box="[1150,1263,1567,1590]" italics="true" pageId="12" pageNumber="413">P. ornatus</emphasis>
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FAD5BB28FA0D0623" box="[1284,1429,1567,1590]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="nigripinna">
<emphasis id="27CC465681404946FAD5BB28FA0D0623" box="[1284,1429,1567,1590]" italics="true" pageId="12" pageNumber="413">P. nigripinna</emphasis>
</taxonomicName>
,
<taxonomicName id="D2B8E1C781404946FF46BB74FEA6064F" box="[151,318,1602,1626]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="pookodensis">
<emphasis id="27CC465681404946FF46BB74FEA6064F" box="[151,318,1602,1626]" italics="true" pageId="12" pageNumber="413">P. pookodensis</emphasis>
</taxonomicName>
and
<emphasis id="27CC465681404946FEA9BB74FE63064F" box="[376,507,1602,1626]" italics="true" pageId="12" pageNumber="413">P. s h al y ni us</emphasis>
; and one band in
<taxonomicName id="D2B8E1C781404946FD1FBB74FCB4064F" box="[718,812,1602,1626]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="thelys">
<emphasis id="27CC465681404946FD1FBB74FCB4064F" box="[718,812,1602,1626]" italics="true" pageId="12" pageNumber="413">P. thelys</emphasis>
</taxonomicName>
), ½4 transverse series scales between dorsal fin origin and lateral line (vs. ½
<quantity id="D24037A181404946FE5FBB51FE21066A" box="[398,441,1638,1663]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="12" pageNumber="413" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="D2B8E1C781404946FE11BB50FDEA066B" box="[448,626,1639,1662]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="manipurensis">
<emphasis id="27CC465681404946FE11BB50FDEA066B" box="[448,626,1639,1662]" italics="true" pageId="12" pageNumber="413">P. manipurensis</emphasis>
</taxonomicName>
and
<taxonomicName id="D2B8E1C781404946FD7BBB50FCDC066B" box="[682,836,1638,1662]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="longicauda">
<emphasis id="27CC465681404946FD7BBB50FCDC066B" box="[682,836,1638,1662]" italics="true" pageId="12" pageNumber="413">P. longicauda</emphasis>
</taxonomicName>
and ½
<quantity id="D24037A181404946FC42BB51FC25066A" box="[915,957,1638,1663]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="12" pageNumber="413" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="D2B8E1C781404946FC14BB50FBF5066B" box="[965,1133,1638,1662]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="meingangbii">
<emphasis id="27CC465681404946FC14BB50FBF5066B" box="[965,1133,1638,1662]" italics="true" pageId="12" pageNumber="413">P. meingangbii</emphasis>
</taxonomicName>
), dorsal fin with 2 oblique rows of black spots (vs. dorsal fin without any distinct marks in
<taxonomicName id="D2B8E1C781404946FCB7BBBCFBA506B7" box="[870,1085,1674,1698]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="melanomaculata">
<emphasis id="27CC465681404946FCB7BBBCFBA506B7" box="[870,1085,1674,1698]" italics="true" pageId="12" pageNumber="413">P. melanomaculata</emphasis>
</taxonomicName>
;
<bibRefCitation id="7129E7B581404946FB9ABBBDFAAB06B6" author="Batuwita" box="[1099,1331,1674,1699]" pageId="12" pageNumber="413" refString="Batuwita, S., Maduwage, K. &amp; Sudasinghe, H. (2015) Redescription of Pethia melanomaculata (Teleostei: Cyprinidae) from Sri Lanka. Zootaxa, 3936 (4), 575 - 583. http: // dx. doi. org / 10.11646 / zootaxa. 3936.4.7" type="journal article" year="2015">
Batuwita
<emphasis id="27CC465681404946FB66BBBCFB7706B7" box="[1207,1263,1674,1698]" italics="true" pageId="12" pageNumber="413">et al.</emphasis>
2015
</bibRefCitation>
).
</paragraph>
<paragraph id="15079A4481404946FF16BB99FEBE076E" blockId="12.[151,1436,1062,2023]" pageId="12" pageNumber="413">
<taxonomicName id="D2B8E1C781404946FF16BB99FED106D3" box="[199,329,1710,1734]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="ticto">
<emphasis id="27CC465681404946FF16BB99FED106D3" box="[199,329,1710,1734]" italics="true" pageId="12" pageNumber="413">Pethia ticto</emphasis>
</taxonomicName>
differs from
<taxonomicName id="D2B8E1C781404946FE0EBB98FDC506D3" box="[479,605,1711,1734]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="punctata">
<emphasis id="27CC465681404946FE0EBB98FDC506D3" box="[479,605,1711,1734]" italics="true" pageId="12" pageNumber="413">P. punctata</emphasis>
</taxonomicName>
in having an incomplete lateral line with 612 pored scales (vs. complete lateral line with 2325 pored scales), 9 predorsal scales (vs. 8), dorsal fin originates behind pelvic fin origin (vs. dorsal fin originates opposite to or slightly in advance of pelvic fin origin) and dark black humeral spot that covers third and fourth lateral line pored scales (vs. humeral spot covers anterior half of forth scale of the row below the lateral line row). Further, the genetic distance in cytb sequences among topotypes of both the species falls between 13.113.5%.
</paragraph>
<paragraph id="15079A4481404947FF16BAB1FE3800ED" blockId="12.[151,1436,1062,2023]" lastBlockId="13.[151,1436,151,248]" lastPageId="13" lastPageNumber="414" pageId="12" pageNumber="413">
While describing
<taxonomicName id="D2B8E1C781404946FE5EBAB1FCF4078A" authority="Katwate et al." authorityName="Katwate et al." authorityYear="2014" box="[399,876,1926,1951]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="longicauda">
<emphasis id="27CC465681404946FE5EBAB1FDC6078B" box="[399,606,1926,1950]" italics="true" pageId="12" pageNumber="413">Pethia longicauda</emphasis>
,
<bibRefCitation id="7129E7B581404946FDBCBAB0FCF4078A" box="[621,876,1926,1951]" pageId="12" pageNumber="413" refString="Katwate, U., Paingankar, M. S., Raghavan, R. &amp; Dahanukar, N. (2014 b) Pethia longicauda, a new species of barb (Teleostei: Cyprinidae) from the northern Western Ghats, India. Zootaxa, 3846 (2), 235 - 248. http: // dx. doi. org / 10.11646 / zootaxa. 3846.2.4" type="journal article">
Katwate
<emphasis id="27CC465681404946FD03BAB0FC94078B" box="[722,780,1926,1950]" italics="true" pageId="12" pageNumber="413">et al.</emphasis>
(2014b)
</bibRefCitation>
</taxonomicName>
attributed sequences
<accessionNumber id="0AEB07A781404946FBB1BAB1FB4B078A" box="[1120,1235,1926,1951]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ795476" pageId="12" pageNumber="413">JQ795476</accessionNumber>
and
<accessionNumber id="0AEB07A781404946FADABAB1FAE6078A" box="[1291,1406,1926,1951]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ795475" pageId="12" pageNumber="413">JQ795475</accessionNumber>
to topotypic
<emphasis id="27CC465681404946FEDFBA9CFEC707D7" box="[270,351,1963,1986]" italics="true" pageId="12" pageNumber="413">P. t i c t o</emphasis>
. However, this was in error as the voucher specimens used for sequences
<accessionNumber id="0AEB07A781404946FB22BA9DFAFE07D6" box="[1267,1382,1962,1987]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ795476" pageId="12" pageNumber="413">JQ795476</accessionNumber>
and
<accessionNumber id="0AEB07A781404946FF46BAF9FE9207F2" box="[151,266,1998,2023]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JQ795475" pageId="12" pageNumber="413">JQ795475</accessionNumber>
came from the River Ganges at Lucknow (
<geoCoordinate id="708CFC8381404946FD3EBAF9FCCC07F3" box="[751,852,1998,2023]" direction="north" orientation="latitude" pageId="12" pageNumber="413" precision="925" value="26.85">26º51' N</geoCoordinate>
,
<geoCoordinate id="708CFC8381404946FC8EBAF9FC5807F2" box="[863,960,1998,2023]" direction="east" orientation="longitude" pageId="12" pageNumber="413" precision="925" value="80.95">80º57' E</geoCoordinate>
). Nevertheless,
<taxonomicName id="D2B8E1C781404946FBA2BAF8FA9407F3" box="[1139,1292,1998,2022]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="413" phylum="Chordata" rank="species" species="longicauda">
<emphasis id="27CC465681404946FBA2BAF8FA9407F3" box="[1139,1292,1998,2022]" italics="true" pageId="12" pageNumber="413">P. longicauda</emphasis>
</taxonomicName>
has a 17.2% difference in the cytb gene sequence compared to
<emphasis id="27CC465681414947FD14BDAEFC9700BA" box="[709,783,152,176]" italics="true" pageId="13" pageNumber="414">P. t icto</emphasis>
.
<taxonomicName id="D2B8E1C781414947FCCEBDAFFEC500C1" authority="Katwate, Katwate, Raghavan, Paingankar &amp; Dahanukar, 2014" authorityName="Katwate, Katwate, Raghavan, Paingankar &amp; Dahanukar" authorityYear="2014" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="414" phylum="Chordata" rank="species" species="lutea">
<emphasis id="27CC465681414947FCCEBDAFFC3F00A5" box="[799,935,152,176]" italics="true" pageId="13" pageNumber="414">Pethia lutea</emphasis>
<bibRefCitation id="7129E7B581414947FC7FBDAFFEC500C1" author="Katwate" pageId="13" pageNumber="414" refString="Katwate, U., Katwate, C., Raghavan, R., Paingankar, M. S. &amp; Dahanukar, N. (2014 c) Pethia lutea, a new species of barb (Teleostei: Cyprinidae) and new records of P. punctata from northern Western Ghats of India. Journal of Threatened Taxa, 6, 5797 - 5818. http: // dx. doi. org / 10.11609 / JoTT. o 3929.5797 - 818" type="journal article" year="2014">Katwate, Katwate, Raghavan, Paingankar &amp; Dahanukar, 2014</bibRefCitation>
</taxonomicName>
, another species recently described from the
<emphasis id="27CC465681414947FCB8BD8AFC2F00C1" box="[873,951,189,212]" italics="true" pageId="13" pageNumber="414">P. t i c t o</emphasis>
complex, differs by a genetic distance of 18.4% in the cytb gene.
</paragraph>
</subSubSection>
</treatment>
</document>