405 lines
42 KiB
XML
405 lines
42 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.413.7172" ID-GBIF-Dataset="f45c1ebe-039d-4180-bfd0-a9d772f72281" ID-PMC="PMC4086027" ID-Pensoft-Pub="1313-2970-413-1" ID-PubMed="25009414" ID-ZBK="5791CE9C1CC0472095838A585DA79446" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-413-1" ModsDocOrigin="ZooKeys 413" ModsDocTitle="The hyper-diverse ant genus Tetramorium Mayr (Hymenoptera, Formicidae) in the Malagasy region taxonomic revision of the T. naganum, T. plesiarum, T. schaufussii, and T. severini species groups" checkinTime="1451245833965" checkinUser="pensoft" docAuthor="Hita Garcia, Francisco & Fisher, Brian L." docDate="2014" docId="9D033737280D28C2D907093017DEAC40" docLanguage="en" docName="ZooKeys 413: 1-170" docOrigin="ZooKeys 413" docSource="http://dx.doi.org/10.3897/zookeys.413.7172" docTitle="Tetramorium tenuinode Hita Garcia & Fisher, sp. n." docType="treatment" docUuid="4856B8A4-FBBA-4629-8619-21F62101BAA7" docUuidSource="ZooBank" docVersion="4" lastPageNumber="78" masterDocId="FFFAFFE7FFB3C93AB161C804401EFFC2" masterDocTitle="The hyper-diverse ant genus Tetramorium Mayr (Hymenoptera, Formicidae) in the Malagasy region taxonomic revision of the T. naganum, T. plesiarum, T. schaufussii, and T. severini species groups" masterLastPageNumber="170" masterPageNumber="1" pageNumber="74" updateTime="1668158573061" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The hyper-diverse ant genus Tetramorium Mayr (Hymenoptera, Formicidae) in the Malagasy region taxonomic revision of the T. naganum, T. plesiarum, T. schaufussii, and T. severini species groups</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hita Garcia, Francisco</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>413</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>170</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.413.7172</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.413.7172</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-413-1</mods:identifier>
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<mods:identifier type="ZBK">5791CE9C1CC0472095838A585DA79446</mods:identifier>
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<mods:identifier type="ZooBank">5791CE9C1CC0472095838A585DA79446</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152053296" LSID="urn:lsid:zoobank.org:act:4856B8A4-FBBA-4629-8619-21F62101BAA7" httpUri="http://treatment.plazi.org/id/9D033737280D28C2D907093017DEAC40" lastPageId="77" lastPageNumber="78" pageId="73" pageNumber="74">
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<subSubSection pageId="73" pageNumber="74" type="nomenclature">
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<paragraph pageId="73" pageNumber="74">
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<taxonomicName LSID="http://zoobank.org/4856B8A4-FBBA-4629-8619-21F62101BAA7" authority="Hita Garcia & Fisher" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode Hita Garcia & Fisher</taxonomicName>
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<taxonomicNameLabel pageId="73" pageNumber="74">sp. n.</taxonomicNameLabel>
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Figs 24B, 25C, 26A, 26B, 39, 64
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74" type="type material">
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<paragraph pageId="73" pageNumber="74">Type material.</paragraph>
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<paragraph lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74">
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Holotype, pinned worker, MADAGASCAR, Fianarantsoa, Parc National de Ranomafana, Vatoharanana River, 4.1 km 231° SW Ranomafana,
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<geoCoordinate direction="south" orientation="latitude" precision="555" value="-21.29">21.29°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="47.43333">47.43333°E</geoCoordinate>
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<pageBreakToken pageId="74" pageNumber="75" start="start">,</pageBreakToken>
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1100 m, montane rainforest, sifted litter (leaf mold, rotten wood), collection code BLF08400, 27.-31.III.2003 (B.L. Fisher et al.) (CAS: CASENT0040115). Paratypes, 41 pinned workers with same data as holotype (CAS: CASENT0039644; CASENT0039646; CASENT0039651; CASENT0039655; CASENT0039660; CASENT0039664; CASENT0039668; CASENT0039671; CASENT0039673; CASENT0039737; CASENT0039739; CASENT0039745; CASENT0039749; CASENT0039750; CASENT0039754; CASENT0039759; CASENT0039761; CASENT0039809; CASENT0040000; CASENT0040020; CASENT0040023; CASENT0040035; CASENT0040036; CASENT0040090; CASENT0040092; CASENT0040096; CASENT0040099; CASENT0040105; CASENT0040106; CASENT0040112; CASENT0040123; CASENT0040181; CASENT0040279; CASENT0040284; CASENT0040285; CASENT0040293; CASENT0040296; CASENT0040304; CASENT0040306; CASENT0040311; CASENT0040317); and nine pinned workers with same data as holotype except collected ex rotten log and collection code BLF08488 (BMNH: CASENT0497623; CAS: CASENT0497621; MCZ: CASENT0497622).
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</paragraph>
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<caption pageId="74" pageNumber="75">
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<paragraph pageId="74" pageNumber="75">
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Figure 39.
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<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode</taxonomicName>
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holotype worker (CASENT0040115). A Body in profile B Body in dorsal view C Head in full-face view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="75" lastPageNumber="76" pageId="74" pageNumber="75" type="non-type material">
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<paragraph pageId="74" pageNumber="75">Non-type material.</paragraph>
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<paragraph lastPageId="75" lastPageNumber="76" pageId="74" pageNumber="75">
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MADAGASCAR: Antsiranana,
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
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Ambanitaza, 26.1 km 347° Antalaha,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-14.67933">14.67933°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="50.18367">50.18367°E</geoCoordinate>
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, 240 m, rainforest, 27.XI.1994 (
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<pageBreakToken pageId="75" pageNumber="76" start="start">B</pageBreakToken>
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.L. Fisher); Antsiranana, 1 km W Andampibe, Cap Masoala,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-15.69361">15.69361°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="50.18139">50.18139°E</geoCoordinate>
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, 125 m, lowland rainforest, 1.XII.1993 (G.D. Alpert); Fianarantsoa, 45 km S Ambalavao,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-22.21667">22.21667°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="47.01667">47.01667°E</geoCoordinate>
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, 785 m, rainforest, 25.IX.1993 (B.L. Fisher); Fianarantsoa, 45 km S Ambalavao,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-22.21667">22.21667°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="47.01667">47.01667°E</geoCoordinate>
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, 720 m, rainforest edge, 31.X.1993 (B.L. Fisher); Fianarantsoa, 2 km W Andrambovato, along river Tatamaly,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-21.51167">21.51167°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="555" value="47.41">47.41°E</geoCoordinate>
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, 1075 m, montane rainforest, 3.-5.VI.2005 (B.L. Fisher et al.); Fianarantsoa,
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<normalizedToken originalValue="Rés">Res</normalizedToken>
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. Andringitra, 43 km S Ambalavao,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-22.23333">22.23333°S</geoCoordinate>
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, 47°E, 825 m, rainforest, 5.X.1993 (B.L. Fisher); Fianarantsoa, R.S. Ivohibe, 7.5 km ENE Ivohibe,
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<geoCoordinate direction="south" orientation="latitude" precision="555" value="-22.47">22.47°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="555" value="46.96">46.96°E</geoCoordinate>
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, 900 m, rainforest, 7.-12.X.1997 (B.L. Fisher); Fianarantsoa, R.S. Ivohibe, 8.0 km E Ivohibe,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-22.48333">22.48333°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="46.96833">46.96833°E</geoCoordinate>
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, 1200 m, montane rainforest, 15.-21.X.1997 (B.L. Fisher); Fianarantsoa, 9.0 km NE Ivohibe,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-22.42667">22.42667°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="46.93833">46.93833°E</geoCoordinate>
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, 900 m, rainforest, 17.XI.1997 (B.L. Fisher); Fianarantsoa,
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
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Speciale Manombo 24.5 km 228° Farafangana,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-23.01583">23.01583°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="55" value="47.719">47.719°E</geoCoordinate>
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, 30 m, lowland rainforest, 20.IV.2006 (B.L. Fisher et al.); Fianarantsoa, Ranomafana Nat. Park, Maharira forest, 21°S, 47°E, 1190 m, montane forest, 11.X.1992 (E. Rajeriarison); Fianarantsoa, Parc National de Ranomafana, Vatoharanana River, 4.1 km 231° SW Ranomafana,
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<geoCoordinate direction="south" orientation="latitude" precision="555" value="-21.29">21.29°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="47.43333">47.43333°E</geoCoordinate>
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, 1100 m, montane rainforest, 27.-31.III.2003 (B.L. Fisher et al.); Fianarantsoa,
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
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de Vevembe, 66.6 km 293° Farafangana,
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<geoCoordinate direction="south" orientation="latitude" precision="55" value="-22.791">22.791°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="47.18183">47.18183°E</geoCoordinate>
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, 600 m, rainforest, transition to montane forest, 23.IV.2006 (B.L. Fisher et al.); Toamasina, Montagne
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<normalizedToken originalValue="d’Akirindro">d'Akirindro</normalizedToken>
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7.6 km 341° NNW Ambinanitelo,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-15.28833">15.28833°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.54833">49.54833°E</geoCoordinate>
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, 600 m, rainforest, 17.-21.III.2003 (B.L. Fisher et al.); Toamasina, 6.3 km S Ambanizana, Andranobe,
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<geoCoordinate direction="south" orientation="latitude" precision="5" value="-15.6813">15.6813°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="55" value="49.958">49.958°E</geoCoordinate>
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, 25 m, rainforest, 14.XI.1993 (B.L. Fisher); Toamasina,
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
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Ambatovaky, Sandrangato river,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-16.77274">16.77274°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.26551">49.26551°E</geoCoordinate>
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, 450 m, rainforest, 20.-22.II.2010 (B.L. Fisher et al.); Toamasina,
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
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Ambatovaky, Sandrangato river,
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<geoCoordinate direction="south" orientation="latitude" precision="5" value="-16.7633">16.7633°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.26692">49.26692°E</geoCoordinate>
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, 520 m, rainforest, 22.-24.II.2010 (B.L. Fisher et al.); Toamasina,
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
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Ambatovaky, Sandrangato river,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-16.81753">16.81753°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.29498">49.29498°E</geoCoordinate>
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, 360 m, rainforest, 25.-27.II.2010 (B.L. Fisher et al.); Toamasina,
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
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Ambatovy, 14.3 km 57° Moramanga,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.85083">18.85083°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="555" value="48.32">48.32°E</geoCoordinate>
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, 1075 m, montane rainforest, 21.III.2004 (Malagasy ant team); Toamasina, Ambatovy, 12.4 km NE Moramanga,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.83937">18.83937°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.30842">48.30842°E</geoCoordinate>
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, 1080 m, montane rainforest, 4.-8.III.2007 (B.L. Fisher et al.); Toamasina, Analamay,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.80623">18.80623°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.33707">48.33707°E</geoCoordinate>
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, 1068 m, montane rainforest, 21.III.2004 (Malagasy ant team); Toamasina, Corridor Forestier Analamay-Mantadia, Ambatoharanana,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.80388">18.80388°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.40506">48.40506°E</geoCoordinate>
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, 1013 m, rainforest, 12.-19.XII.2012 (B.L.Fisher et al.); Toamasina, Corridor Forestier Analamay-Mantadia, Ambohibolakely,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.76087">18.76087°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.37128">48.37128°E</geoCoordinate>
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, 1044 m, rainforest, 23.-28.XI.2012 (B.L.Fisher et al.); Toamasina, Corridor Forestier Analamay-Mantadia, Ambohibolakely,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.77898">18.77898°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.36375">48.36375°E</geoCoordinate>
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, 918 m, rainforest, 23.-28.XI.2012 (B.L.Fisher et al.); Toamasina, Corridor Forestier Analamay-Mantadia, Ambohibolakely,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.76131">18.76131°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.36437">48.36437°E</geoCoordinate>
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, 983 m, rainforest, 26.XI.2012 (B.L.Fisher et al.); Toamasina, Corridor Forestier Analamay-Mantadia, Tsaravoniana,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.76124">18.76124°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.42134">48.42134°E</geoCoordinate>
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, 939 m, rainforest, 2.-7.XII.2012 (B.L.Fisher et al.); Toamasina, Corridor Forestier Analamay-Mantadia, Tsaravoniana,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.76465">18.76465°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.41938">48.41938°E</geoCoordinate>
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, 1039 m, rainforest, 2.-7.XII.2012 (B.L.Fisher et al.); Toamasina, Station
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<normalizedToken originalValue="Forestière">Forestiere</normalizedToken>
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Analamazaotra, Analamazaotra 1.3 km S Andasibe,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.38466">18.38466°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.41271">48.41271°E</geoCoordinate>
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, 980 m, montane rainforest, 11.-13.XII.2007 (B.L. Fisher et al.); Toamasina, F.C. Andriantantely,
|
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<geoCoordinate direction="south" orientation="latitude" precision="55" value="-18.695">18.695°S</geoCoordinate>
|
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.81333">48.81333°E</geoCoordinate>
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, 530 m, rainforest, 4.-7.XII.1998 (H.J. Ratsirarson); Toamasina, Montagne
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<normalizedToken originalValue="d’Anjanaharibe">d'Anjanaharibe</normalizedToken>
|
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, 18.0 km 21° NNE Ambinanitelo,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-15.18833">15.18833°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="55" value="49.615">49.615°E</geoCoordinate>
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, 470 m, rainforest, 8.-12.III.2003 (B.L. Fisher et al.); Toamasina, Ankerana,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.40829">18.40829°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.82107">48.82107°E</geoCoordinate>
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, 750 m, rainforest, 21.-26.I.2012 (B.L. Fisher et al.); Toamasina, Ankerana,
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<geoCoordinate direction="south" orientation="latitude" precision="5" value="-18.4104">18.4104°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="5" value="48.8189">48.8189°E</geoCoordinate>
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, 855 m, rainforest, 22.-27.I.2012 (B.L. Fisher et al.); Toamasina, Reserve Betampona, Camp Vohitsivalana, 37.1 km 338° Toamasina,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-17.88667">17.88667°S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="5" value="49.2025">49.2025°E</geoCoordinate>
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, 520 m, rainforest, 1.-3.XII.2005 (B.L.Fisher et al.); Toamasina, Reserve Betampona, Camp Rendrirendry, 34.1 km 332° Toamasina,
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<geoCoordinate direction="south" orientation="latitude" precision="55" value="-17.924">17.924°S</geoCoordinate>
|
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.19967">49.19967°E</geoCoordinate>
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, 390 m, rainforest, 28.XI.2005 (B.L.Fisher et al.); Toamasina, F.C. Didy,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.19833">18.19833°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.57833">48.57833°E</geoCoordinate>
|
||
, 960 m, rainforest, 16.-23.XII.1998 (H.J. Ratsirarson); Toamasina, Parc National Mananara-Nord, 7.1 km 261° Antanambe,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="55" value="-16.455">16.455°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="5" value="49.7875">49.7875°E</geoCoordinate>
|
||
, 225 m, rainforest, 14.XI.2005 (B.L. Fisher et al.); Toamasina, 19 km ESE Maroantsetra,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-15.48333">15.48333°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="5555" value="49.9">49.9°E</geoCoordinate>
|
||
, 350 m, rainforest, 22.IV.1989 (P.S. Ward); Toamasina, 16 km S Moramanga,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-19.08333">19.08333°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.23333">48.23333°E</geoCoordinate>
|
||
, 950 m, rainforest, 18.XI.1990 (P.S. Ward); Toamasina, Moramanga,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.94417">18.94417°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.23067">48.23067°E</geoCoordinate>
|
||
, 922 m, urban/garden, 14.II.2007 (B.L. Fisher et al.); Toamasina, F.C. Sandranantitra,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.04833">18.04833°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.09167">49.09167°E</geoCoordinate>
|
||
, 450 m, rainforest, 18.-24.I.1999 (H.J. Ratsirarson); Toamasina, Parc National de Zahamena, Onibe River,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-17.75908">17.75908°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.85468">48.85468°E</geoCoordinate>
|
||
, 780 m, rainforest, 21.-23.II.2009 (B.L. Fisher et al.); Toamasina, Parc National de Zahamena, Tetezambatana forest, near junction of Nosivola and Manakambahiny rivers,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-17.74298">17.74298°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.72936">48.72936°E</geoCoordinate>
|
||
, 860 m, rainforest, 18.-19.II.2009 (B.L. Fisher et al.); Toliara,
|
||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||
. Andohahela, 10 km NW Enakara,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-24.56667">24.56667°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="1" value="46.81667">46.81667°E</geoCoordinate>
|
||
, 430 m, rainforest, 22.XI.1992 (B.L. Fisher); Toliara,
|
||
<normalizedToken originalValue="Rés">Res</normalizedToken>
|
||
. Andohahela, 6 km SSW Eminiminy,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-24.73333">24.73333°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="5555" value="46.8">46.8°E</geoCoordinate>
|
||
, 330 m, rainforest, 4.II.1993 (G.D. Alpert & P.S. Ward); Toliara, Parc National
|
||
<normalizedToken originalValue="d’Andohahela">d'Andohahela</normalizedToken>
|
||
, Col du Sedro, 3.8 km 113° ESE Mahamavo, 37.6 km 341° NNW Tolagnaro,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-24.76389">24.76389°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="1" value="46.75167">46.75167°E</geoCoordinate>
|
||
, 900 m, montane rainforest, 21.-25.I.2002 (B.L. Fisher et al.); Toliara, Andohahela,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-24.77639">24.77639°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="1" value="46.70528">46.70528°E</geoCoordinate>
|
||
, 320 m, 9.XII.2007 (A. Ballerio); Toliara, Grand Lavasoa, 25.9 km W Tolagnaro,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-25.08767">25.08767°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="55" value="46.749">46.749°E</geoCoordinate>
|
||
, 450 m, rainforest, 30.XI.-2.XII.2006 (B.L. Fisher et al.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="75" pageNumber="76" type="diagnosis">
|
||
<paragraph pageId="75" pageNumber="76">Diagnosis.</paragraph>
|
||
<paragraph pageId="75" pageNumber="76">
|
||
The following character set clearly distinguishes
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode</taxonomicName>
|
||
from the remainder of the
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium cognatum" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="cognatum">Tetramorium cognatum</taxonomicName>
|
||
species complex: eyes relatively large (OI 25-27); antennal scapes very short (SI 66-70); frontal carinae well developed, noticeably raised, and approaching or ending at posterior head margin; petiolar node high rounded nodiform, in profile around 1.8 to 2.2 times higher than long (LPeI 45-54), in dorsal view around 1.3 to 1.4 times wider than long (DPeI 125-143); mesosoma with only two pairs of long, standing hairs, one on anterior pronotum and one on anterior mesonotum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="75" pageNumber="76" type="worker measurements">
|
||
<paragraph pageId="75" pageNumber="76">Worker measurements</paragraph>
|
||
<paragraph pageId="75" pageNumber="76">(N=12). HL 0.62-0.75 (0.71); HW 0.58-0.70 (0.66); SL 0.40-0.47 (0.45); EL 0.16-0.18 (0.17); PH 0.30-0.36 (0.34); PW 0.42-0.54 (0.50); WL 0.76-0.94 (0.88); PSL 0.11-0.18 (0.14); PTL 0.13-0.17 (0.15); PTH 0.26-0.32 (0.30); PTW 0.16-0.21 (0.20); PPL 0.15-0.22 (0.20); PPH 0.25-0.32 (0.30); PPW 0.24-0.33 (0.30); CI 91-93 (92); SI 66-70 (68); OI 25-27 (26); DMI 55-59 (57); LMI 38-40 (39); PSLI 17-24 (20); PeNI 36-42 (39); LPeI 45-54 (49); DPeI 125-143 (133); PpNI 57-64 (61); LPpI 60-72 (67); DPpI 143-168 (153); PPI 148-167 (155).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76" type="worker description">
|
||
<paragraph pageId="75" pageNumber="76">Worker description.</paragraph>
|
||
<paragraph lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76">
|
||
Head longer than wide (CI 91-93); in full-face view posterior head margin weakly to moderately concave. Anterior clypeal margin with distinct median impression. Frontal carinae well developed, noticeably raised, diverging posteriorly, approaching or ending at posterior head margin. Antennal scrobes weakly developed, shallow and without clear and distinct posterior and ventral margins. Antennal scapes very short, not reaching posterior head margin (SI 66-70). Eyes relatively large (OI 25-27). Mesosomal outline in profile flat to weakly convex, moderately low and long (LMI 38-40), moderately marginate from lateral to dorsal mesosoma; promesonotal suture absent; metanotal groove very weak to absent. Propodeal spines short to moderately long, usually elongate-triangular, and acute (PSLI 17-24), propodeal lobes short and triangular, always much shorter than propodeal spines. Petiolar node in profile high rounded nodiform and relatively thin, around 1.8 to 2.2 times higher than long (LPeI 45-54), anterior and posterior faces approximately parallel, anterodorsal and posterodorsal margins usually situated at about same height and moderately rounded, petiolar dorsum distinctly convex; petiolar node in dorsal view around 1.2 to 1.4 times wider than long (DPeI 125-143), in dorsal view pronotum between 2.4 to 2.8 times wider than petiolar node (PeNI 36-42). Postpetiole in profile subglobular and weakly anteroposteriorly compressed, around 1.4 to 1.7 times higher than long (LPpI 60-72); in dorsal view between 1.4 to 1.7 times wider than long (DPpI 143-168), pronotum around 1.6 to 1.7 times wider than postpetiole (PpNI 57-64). Postpetiole in profile appearing more voluminous than petiolar node, postpetiole in dorsal view around 1.5 to 1.7 times wider
|
||
<pageBreakToken pageId="76" pageNumber="77" start="start">than</pageBreakToken>
|
||
petiolar node (PPI 148-167). Mandibles completely unsculptured, smooth, and shiny; clypeus irregularly longitudinally rugose/rugulose with two to seven rugae/rugulae, median ruga rarely fully developed, usually broken, most other rugulae usually broken, sometimes merging with each other; cephalic dorsum between frontal carinae longitudinally rugose, usually with six to seven, rarely eight or nine rugae, rugae running from posterior clypeal margin to posterior head margin, often interrupted, splitting up or with cross-meshes, especially posteriorly; scrobal area partly unsculptured, but mostly merging with surrounding longitudinally rugose to reticulate-rugose sculpture present on lateral head. Ground sculpture on head weakly to moderately punctate. Dorsum and sides of mesosoma reticulate-rugose/rugulose to irregularly longitudinally rugose/rugulose, lateral pronotum sometimes only weakly sculptured and relatively smooth and shining. Forecoxae mostly unsculptured, smooth and shining, sometimes with traces of ground sculpture. Ground sculpture on mesosoma usually weak to absent. Both waist segments and gaster fully unsculptured, smooth, and shining. Dorsum of head with several pairs of long, fine, standing hairs; dorsum of mesosoma with two pairs only, one on anterior pronotum and one on anterior mesonotum; propodeum, waist segments and first gastral tergite without any standing hairs at all; first gastral tergite with very short, scarce, appressed pubescence. Anterior edges of antennal scapes and dorsal (outer) surfaces of hind tibiae usually with appressed, rarely decumbent, hairs. Head, mesosoma, waist segments and gaster usually uniformly chestnut brown to very dark brown, sometimes of lighter reddish brown; appendages usually of slightly lighter brown than remainder of body.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="76" pageNumber="77" type="etymology">
|
||
<paragraph pageId="76" pageNumber="77">Etymology.</paragraph>
|
||
<paragraph pageId="76" pageNumber="77">
|
||
The name of the new species is a combination of the Latin adjective
|
||
<normalizedToken originalValue="“tenuis”">"tenuis"</normalizedToken>
|
||
, meaning thin, and the Latin noun
|
||
<normalizedToken originalValue="“nodus”">"nodus"</normalizedToken>
|
||
, meaning node.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="76" pageNumber="77" type="distribution and biology">
|
||
<paragraph pageId="76" pageNumber="77">Distribution and biology.</paragraph>
|
||
<paragraph pageId="76" pageNumber="77">
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode</taxonomicName>
|
||
is widely distributed in eastern Madagascar (Fig. 64). It is found with few interruptions in an almost straight line from Grand Lavasoa and Andohahela in the southeast to Ambanitaza in the northeast. The new species clearly prefers rainforests and montane rainforests at elevations from 25 to 1200 m, even though it was also collected several times from guava forest and urban gardens. In addition,
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode</taxonomicName>
|
||
appears to be a leaf litter inhabitant since almost all of the material was collected from this microhabitat.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77" type="discussion">
|
||
<paragraph pageId="76" pageNumber="77">Discussion.</paragraph>
|
||
<paragraph pageId="76" pageNumber="77">
|
||
This new species is easily identifiable within its species complex. The presence of well developed, raised, and long frontal carinae separates
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode</taxonomicName>
|
||
from the species
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium aspis" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="aspis">Tetramorium aspis</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium camelliae" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="camelliae">Tetramorium camelliae</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium cognatum" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="cognatum">Tetramorium cognatum</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium karthala" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="karthala">Tetramorium karthala</taxonomicName>
|
||
, and
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium rumo" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rumo">Tetramorium rumo</taxonomicName>
|
||
. These five species are generally also much smaller (WL 0.56-0.81) than
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode</taxonomicName>
|
||
(WL 0.76-0.94). Also,
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode</taxonomicName>
|
||
possesses much larger eyes (OI 25-27) than
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium gladius" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="gladius">Tetramorium gladius</taxonomicName>
|
||
(OI 19-20) and much shorter antennal scapes (SI 66-70) than
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium myrmidon" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="myrmidon">Tetramorium myrmidon</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium proximum" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="proximum">Tetramorium proximum</taxonomicName>
|
||
(SI 74-77). Lastly, the presence of two pairs of long, standing hairs on the dorsal mesosoma distinguishes
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode</taxonomicName>
|
||
from
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium freya" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="freya">Tetramorium freya</taxonomicName>
|
||
, which lacks any standing hairs on the dorsal mesosoma. The latter species also displays weaker sculpture on the head and mesosoma than
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="77" pageNumber="78">
|
||
<pageBreakToken pageId="77" pageNumber="78" start="start">It</pageBreakToken>
|
||
should be noted, however, that
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode</taxonomicName>
|
||
is morphologically still very close to
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium proximum" order="Hymenoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="proximum">Tetramorium proximum</taxonomicName>
|
||
, and on very superficial observation it is possible to confuse both. At the initial stage of the revision we considered
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode</taxonomicName>
|
||
conspecific with
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium proximum" order="Hymenoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="proximum">Tetramorium proximum</taxonomicName>
|
||
, a very variable species. However, after the detailed examination of more than five hundred pinned workers, it became apparent that the material consisted of these two fairly distinct species. Indeed, even though
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium proximum" order="Hymenoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="proximum">Tetramorium proximum</taxonomicName>
|
||
is more broadly distributed than
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode</taxonomicName>
|
||
, both co-occur in sympatry throughout most of their respective distribution ranges, and both are always very well distinguishable. The most obvious character to separate both species is pilosity on the dorsal mesosoma. In
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode</taxonomicName>
|
||
this consists of two pairs of long, standing hairs only, one on anterior pronotum and one on anterior mesonotum, which contrasts with the usually five to six (rarely four or more than six) pairs found from anterior pronotum to posterior mesonotum in
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium proximum" order="Hymenoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="proximum">Tetramorium proximum</taxonomicName>
|
||
. In addition to mesosomal pilosity and the shorter antennal scapes mentioned above,
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode</taxonomicName>
|
||
usually also has a thinner petiolar node in profile, which is around 1.8 to 2.2 times higher than long (LPeI 45-54), while the node of
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium proximum" order="Hymenoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="proximum">Tetramorium proximum</taxonomicName>
|
||
is around 1.7 to 1.9 times higher than long (LPeI 52-60). The identification key presented above might suggest that
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode</taxonomicName>
|
||
can be easily confused with
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium myrmidon" order="Hymenoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="myrmidon">Tetramorium myrmidon</taxonomicName>
|
||
, but this is not the case since the two species differ in a number of characters. Among others,
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode</taxonomicName>
|
||
has a broader head (CI 91-93), shorter antennal scapes (SI 66-70), and a higher petiolar node (LPeI 45-54) than
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium myrmidon" order="Hymenoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="myrmidon">Tetramorium myrmidon</taxonomicName>
|
||
(CI 88-91; SI 74-76; LPeI 58-60).
|
||
</paragraph>
|
||
<paragraph pageId="77" pageNumber="78">
|
||
In contrast to the more variable
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium proximum" order="Hymenoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="proximum">Tetramorium proximum</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium tenuinode" order="Hymenoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="tenuinode">Tetramorium tenuinode</taxonomicName>
|
||
has little observed intraspecific variation. Some populations vary very slightly in propodeal spine length, thickness of the petiolar node, or body colouration.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |