245 lines
31 KiB
XML
245 lines
31 KiB
XML
<document id="2CB8B03CF54CF3AEE4337F66C00341A5" ID-DOI="10.24199/j.mmv.2006.63.17" ID-ISSN="1447-2554" ID-Zenodo-Dep="12210410" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718991714234" checkinUser="felipe" docAuthor="Sandford, Andrew C. & Holloway, David J." docDate="2006" docId="9C6887D7FF9E3F1B6536F975AA21FDAF" docLanguage="en" docName="MemMusVictoria.63.2.215-255.pdf" docOrigin="Memoirs of Museum Victoria 63 (2)" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-63-issue-2-2006/pages-215-255/" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Bessazoon Curtis and Lane 1998" docType="treatment" docVersion="2" lastPageNumber="240" masterDocId="6051FFAFFF893F02660FFF86AF6EFF93" masterDocTitle="Early Silurian phacopide trilobites from central Victoria, Australia" masterLastPageNumber="255" masterPageNumber="215" pageNumber="238" updateTime="1719002137797" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="DCEF476EB6560AF8CE56613B6B165D6E">Early Silurian phacopide trilobites from central Victoria, Australia</mods:title>
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<mods:namePart id="16C905A2B6F9F9C620232A11B97DF716">Sandford, Andrew C.</mods:namePart>
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<mods:namePart id="2B13E5551DBFBE6A1397BB64E99C7CF5">Holloway, David J.</mods:namePart>
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<mods:title id="91C2BE27438F0A73A60221663AC5D7D6">Memoirs of Museum Victoria</mods:title>
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<mods:date id="91AE4F51B29A351C8E39AD287CC53D18">2006</mods:date>
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<treatment id="9C6887D7FF9E3F1B6536F975AA21FDAF" LSID="urn:lsid:plazi:treatment:9C6887D7FF9E3F1B6536F975AA21FDAF" httpUri="http://treatment.plazi.org/id/9C6887D7FF9E3F1B6536F975AA21FDAF" lastPageId="25" lastPageNumber="240" pageId="23" pageNumber="238">
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<subSubSection id="5CDB654AFF9E3F156536F975ABE0F89B" box="[825,1166,1779,1801]" pageId="23" pageNumber="238" type="nomenclature">
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<paragraph id="147E36C1FF9E3F156536F975ABE0F89B" blockId="23.[825,1166,1779,1801]" box="[825,1166,1779,1801]" pageId="23" pageNumber="238">
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<heading id="4F3681ADFF9E3F156536F975ABE0F89B" box="[825,1166,1779,1801]" fontSize="9" level="2" pageId="23" pageNumber="238" reason="5">
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<taxonomicName id="D3C14D42FF9E3F156536F975ABE0F89B" ID-CoL="62GKX" authority="Curtis and Lane, 1998" authorityName="Curtis and Lane" authorityYear="1998" box="[825,1166,1779,1801]" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="23" pageNumber="238" phylum="Arthropoda" rank="genus">
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<emphasis id="26B5EAD3FF9E3F156536F975ACC9F89B" bold="true" box="[825,935,1779,1800]" italics="true" pageId="23" pageNumber="238">Bessazoon</emphasis>
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||
<bibRefCitation id="70504B30FF9E3F1565A2F975ABE0F89B" author="Curtis, N. J. & Lane, P. D." box="[941,1166,1779,1801]" pageId="23" pageNumber="238" pagination="51 - 101" refId="ref29505" refString="Curtis, N. J. and Lane, P. D. 1998. The Llandovery trilobites of England and Wales. Part 2. Monograph of the Palaeontographical Society London 152 (608): 51 - 101, pls 4 - 12." type="journal article" year="1998">Curtis and Lane, 1998</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="5CDB654AFF9E3F1B6536F8A6AA21FDAF" lastPageId="25" lastPageNumber="240" pageId="23" pageNumber="238" type="materials_examined">
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<paragraph id="147E36C1FF9E3F156536F8A6AA6AF8FC" blockId="23.[825,1456,1823,1903]" pageId="23" pageNumber="238">
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<materialsCitation id="A4A93C9CFF9E3F156536F8A6AA6EF8FC" country="United Kingdom" location="Hughley Shales" municipality="upper Llandovery" pageId="23" pageNumber="238" specimenCount="1" stateProvince="Shropshire" typeStatus="holotype">
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<emphasis id="26B5EAD3FF9E3F156536F8A6ACA0F8A6" box="[825,974,1824,1845]" italics="true" pageId="23" pageNumber="238">
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||
<typeStatus id="CB7A8863FF9E3F156536F8A6AC07F8A6" box="[825,873,1824,1845]" pageId="23" pageNumber="238">Type</typeStatus>
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species.
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</emphasis>
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||
<taxonomicName id="D3C14D42FF9E3F1565EDF899ACBCF8C2" authority="Whittard, 1938" authorityName="Whittard" authorityYear="1938" class="Trilobita" family="Dalmanitidae" genus="Dalmanites" kingdom="Animalia" order="Phacopida" pageId="23" pageNumber="238" phylum="Arthropoda" rank="variety" species="weaveri" variety="tenuimucronata">
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<emphasis id="26B5EAD3FF9E3F1565EDF899ABD0F8A6" box="[994,1214,1823,1845]" italics="true" pageId="23" pageNumber="238">Dalmanites weaveri</emphasis>
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var.
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<emphasis id="26B5EAD3FF9E3F156301F8A6AADEF8A6" box="[1294,1456,1824,1845]" italics="true" pageId="23" pageNumber="238">tenuimucronata</emphasis>
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||
<bibRefCitation id="70504B30FF9E3F156536F8BAACBCF8C2" author="Whittard, W. F." box="[825,978,1852,1874]" pageId="23" pageNumber="238" pagination="85 - 140" refId="ref33225" refString="Whittard, W. F. 1938. The upper Valentian trilobite fauna of Shropshire. Annals and Magazine of Natural History, Series 11, 1: 85 - 140." type="journal article" year="1938">Whittard, 1938</bibRefCitation>
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</taxonomicName>
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from the
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<location id="111E601AFF9E3F15623DF8BAABBEF8C1" LSID="urn:lsid:plazi:treatment:9C6887D7FF9E3F1B6536F975AA21FDAF:111E601AFF9E3F15623DF8BAABBEF8C1" box="[1074,1232,1852,1874]" country="United Kingdom" municipality="upper Llandovery" name="Hughley Shales" pageId="23" pageNumber="238" stateProvince="Shropshire">Hughley Shales</location>
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(
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<collectingMunicipality id="F41AACBBFF9E3F1562D4F8BAAAE1F8C1" box="[1243,1423,1852,1874]" pageId="23" pageNumber="238">upper Llandovery</collectingMunicipality>
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) of
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<collectingRegion id="D605F823FF9E3F156536F8DFACC4F8FC" box="[825,938,1881,1903]" country="United Kingdom" name="Shropshire" pageId="23" pageNumber="238">Shropshire</collectingRegion>
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,
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<collectingCountry id="6CD67651FF9E3F1565BBF8DFAB65F8FC" box="[948,1035,1881,1903]" name="United Kingdom" pageId="23" pageNumber="238">England</collectingCountry>
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, by original designation
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</materialsCitation>
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.
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</paragraph>
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<caption id="40BE6649FF913F1A6683F938AB5CF8FD" ID-DOI="http://doi.org/10.5281/zenodo.12210444" ID-Zenodo-Dep="12210444" httpUri="https://zenodo.org/record/12210444/files/figure.png" pageId="24" pageNumber="239" startId="24.[140,198,1726,1745]" targetBox="[262,1320,326,1692]" targetPageId="24" targetType="figure">
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<paragraph id="147E36C1FF913F1A6683F938AB5CF8FD" blockId="24.[140,1441,1726,1903]" pageId="24" pageNumber="239">
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Figure 13. A–K,
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<taxonomicName id="D3C14D42FF913F1A672EF938AD28F942" authority="Opik, 1953" authorityName="Opik" authorityYear="1953" box="[289,582,1726,1746]" class="Trilobita" family="Dalmanitidae" genus="Dalmanites" kingdom="Animalia" order="Phacopida" pageId="24" pageNumber="239" phylum="Arthropoda" rank="species" species="athamas">
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<emphasis id="26B5EAD3FF913F1A672EF938AEB5F942" box="[289,475,1726,1745]" italics="true" pageId="24" pageNumber="239">Dalmanites athamas</emphasis>
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<bibRefCitation id="70504B30FF913F1A67EEF938AD28F942" author="Opik, A. A." box="[481,582,1726,1746]" pageId="24" pageNumber="239" pagination="1 - 42" refId="ref31184" refString="Opik, A. A. 1953. Lower Silurian fossils from the ʻ Illaenus Bandʼ Heathcote, Victoria. Memoirs of the Geological Survey of Victoria 19: 1 - 42, pls 1 - 13." type="journal article" year="1953">Öpik, 1953</bibRefCitation>
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</taxonomicName>
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. A, NMV P138215, thoracopygon, × 1.5, from the ʻ
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<taxonomicName id="D3C14D42FF913F1A6206F938AB3CF942" authorityName="Dalman" authorityYear="1827" box="[1033,1106,1726,1745]" class="Trilobita" family="Illaenidae" genus="Illaenus" kingdom="Animalia" order="Corynexochida" pageId="24" pageNumber="239" phylum="Arthropoda" rank="genus">
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<emphasis id="26B5EAD3FF913F1A6206F938AB3CF942" box="[1033,1106,1726,1745]" italics="true" pageId="24" pageNumber="239">Illaenus</emphasis>
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||
</taxonomicName>
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bandʼ, Costerfield. B, NMV P138219, pygidium, × 3.5, from PL2269, Costerfield. C, paratype NMV P52485, cranidium, × 2, from PL2262, Costerfield. D, G, NMV P138217, pygidium, × 2, from PL389, Costerfield. E, paratype NMV P52486, cranidium, × 2, from PL2269, Costerfield. F, NMV P138218 pygidium, × 2, from PL2269, Costerfield. H, holotype NMV P52484, pygidium, × 1.5, from PL2262, Costerfield. I, NMV P138223, thorax, enlargement showing granulation, from PL2269, Costerfield. J, NMV P139805, teratological pygidium, × 2, from PL386, Costerfield. K, NMV P52482, incomplete fixigena, holotype of ʻ
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<taxonomicName id="D3C14D42FF913F1A669DF8C7AD05F8C7" authority="ʼ Opik, 1953" authorityName="ʼ Opik" authorityYear="1953" box="[146,619,1856,1876]" class="Trilobita" family="Dalmanitidae" genus="Dalmanitina" kingdom="Animalia" order="Phacopida" pageId="24" pageNumber="239" phylum="Arthropoda" rank="species" species="aborigenum" subGenus="Eudolatites">
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<emphasis id="26B5EAD3FF913F1A669DF8C7AE6BF8C7" box="[146,261,1857,1876]" italics="true" pageId="24" pageNumber="239">Dalmanitina</emphasis>
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(
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<emphasis id="26B5EAD3FF913F1A671CF8C7AE17F8C7" box="[275,377,1857,1876]" italics="true" pageId="24" pageNumber="239">Eudolatites</emphasis>
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)
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<emphasis id="26B5EAD3FF913F1A6786F8C7AE9AF8C7" box="[393,500,1857,1876]" italics="true" pageId="24" pageNumber="239">aborigenum</emphasis>
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ʼ
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<bibRefCitation id="70504B30FF913F1A640BF8C6AD05F8C7" author="Opik, A. A." box="[516,619,1856,1876]" pageId="24" pageNumber="239" pagination="1 - 42" refId="ref31184" refString="Opik, A. A. 1953. Lower Silurian fossils from the ʻ Illaenus Bandʼ Heathcote, Victoria. Memoirs of the Geological Survey of Victoria 19: 1 - 42, pls 1 - 13." type="journal article" year="1953">Öpik, 1953</bibRefCitation>
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</taxonomicName>
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, × 2, from PL2269, Costerfield. L,
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<taxonomicName id="D3C14D42FF913F1A65BBF8C7AB53F8C7" box="[948,1085,1857,1876]" class="Trilobita" family="Dalmanitidae" genus="Struveria" kingdom="Animalia" order="Phacopida" pageId="24" pageNumber="239" phylum="Arthropoda" rank="species" species="undefined-2">
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<emphasis id="26B5EAD3FF913F1A65BBF8C7AB66F8C7" box="[948,1032,1857,1876]" italics="true" pageId="24" pageNumber="239">Struveria</emphasis>
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sp. 2
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</taxonomicName>
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, NMV P52483, pygidium, paratype of ʻ
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<taxonomicName id="D3C14D42FF913F1A669DF8DDAD31F8FD" authority="ʼ Opik, 1953" authorityName="ʼ Opik" authorityYear="1953" box="[146,607,1883,1903]" class="Trilobita" family="Dalmanitidae" genus="Dalmanitina" kingdom="Animalia" order="Phacopida" pageId="24" pageNumber="239" phylum="Arthropoda" rank="species" species="aborigenum" subGenus="Eudolatites">
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||
<emphasis id="26B5EAD3FF913F1A669DF8DDAE6BF8FD" box="[146,261,1883,1902]" italics="true" pageId="24" pageNumber="239">Dalmanitina</emphasis>
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||
(
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<emphasis id="26B5EAD3FF913F1A671FF8DDAE18F8FD" box="[272,374,1883,1902]" italics="true" pageId="24" pageNumber="239">Eudolatites</emphasis>
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)
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<emphasis id="26B5EAD3FF913F1A678CF8DDAE80F8FD" box="[387,494,1883,1902]" italics="true" pageId="24" pageNumber="239">aborigenum</emphasis>
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||
ʼ
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<bibRefCitation id="70504B30FF913F1A67F4F8DDAD31F8FD" author="Opik, A. A." box="[507,607,1883,1903]" pageId="24" pageNumber="239" pagination="1 - 42" refId="ref31184" refString="Opik, A. A. 1953. Lower Silurian fossils from the ʻ Illaenus Bandʼ Heathcote, Victoria. Memoirs of the Geological Survey of Victoria 19: 1 - 42, pls 1 - 13." type="journal article" year="1953">Öpik, 1953</bibRefCitation>
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</taxonomicName>
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, × 3, from PL2269, Costerfield. (B, D are latex casts)
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</paragraph>
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</caption>
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<paragraph id="147E36C1FF903F1B6685FF63AD2EFA65" blockId="25.[138,770,229,1903]" pageId="25" pageNumber="240">
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<emphasis id="26B5EAD3FF903F1B6685FF63AF8DFF69" box="[138,227,229,250]" italics="true" pageId="25" pageNumber="240">Remarks</emphasis>
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. Of the characters regarded by
|
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<bibRefCitation id="70504B30FF903F1B6414FF63AFC5FE8B" author="Curtis, N. J. & Lane, P. D." pageId="25" pageNumber="240" pagination="51 - 101" refId="ref29505" refString="Curtis, N. J. and Lane, P. D. 1998. The Llandovery trilobites of England and Wales. Part 2. Monograph of the Palaeontographical Society London 152 (608): 51 - 101, pls 4 - 12." type="journal article" year="1998">Curtis and Lane (1998: 62)</bibRefCitation>
|
||
as diagnostic of
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<taxonomicName id="D3C14D42FF903F1B6760FE85AF98FEA7" authority=", Chatterton and Ludvigsen (2004: 47)" authorityName=", Chatterton and Ludvigsen" authorityPageNumber="47" authorityYear="2004" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="genus">
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||
<emphasis id="26B5EAD3FF903F1B6760FE85AEB4FE8B" box="[367,474,259,280]" italics="true" pageId="25" pageNumber="240">Bessazoon</emphasis>
|
||
,
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||
<bibRefCitation id="70504B30FF903F1B67E2FE84AF98FEA7" author="Chatterton, B. D. E. & Ludvigsen, R." pageId="25" pageNumber="240" pagination="1 - 264" refId="ref29332" refString="Chatterton, B. D. E. and Ludvigsen, R. 2004. Early Silurian trilobites of Anticosti Island, Quebec, Canada. Palaeontographica Canadiana 22: 1 - 264." type="journal article" year="2004">Chatterton and Ludvigsen (2004: 47)</bibRefCitation>
|
||
</taxonomicName>
|
||
questioned the taxonomic value of the size of the eyes and the loss of the pygidial mucro in large specimens. In the
|
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<typeStatus id="CB7A8863FF903F1B66BBFEDCAF8EFEFC" box="[180,224,346,367]" pageId="25" pageNumber="240">type</typeStatus>
|
||
species and
|
||
<taxonomicName id="D3C14D42FF903F1B677FFEDCAFADFE18" authority="(Holloway and Sandford, 1993)" baseAuthorityName="Holloway and Sandford" baseAuthorityYear="1993" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="tigerense">
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||
<emphasis id="26B5EAD3FF903F1B677FFEDCAE83FEFC" box="[368,493,346,367]" italics="true" pageId="25" pageNumber="240">B. tigerense</emphasis>
|
||
(
|
||
<bibRefCitation id="70504B30FF903F1B640FFEDFAFD2FE18" author="Holloway, D. J. & Sandford, A. C." pageId="25" pageNumber="240" pagination="85 - 102" refId="ref30463" refString="Holloway, D. J. and Sandford, A. C. 1993. An early Silurian trilobite fauna from Tasmania. Memoirs of the Association of Australasian Palaeontologists 15: 85 - 102." type="journal article" year="1993">Holloway and Sandford, 1993</bibRefCitation>
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||
)
|
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</taxonomicName>
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the eyes are very large and occupy almost the entire length of the genal field, but in Curtis and Laneʼs species
|
||
<taxonomicName id="D3C14D42FF903F1B64E4FE12AE4FFE56" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="buttingtonense">
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<emphasis id="26B5EAD3FF903F1B64E4FE12AE4FFE56" italics="true" pageId="25" pageNumber="240">B. buttingtonense</emphasis>
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</taxonomicName>
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and the cephalon they illustrated as
|
||
<emphasis id="26B5EAD3FF903F1B64A6FE37ADD1FE55" box="[681,703,433,454]" italics="true" pageId="25" pageNumber="240">B.</emphasis>
|
||
cf.
|
||
<taxonomicName id="D3C14D42FF903F1B64E4FE37AE51FE70" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="tenuimucronatum">
|
||
<emphasis id="26B5EAD3FF903F1B64E4FE37AE51FE70" italics="true" pageId="25" pageNumber="240">B. tenuimucronatum</emphasis>
|
||
</taxonomicName>
|
||
(their pl. 9, fig. 1) the eyes are smaller and do not extend very close to the border furrows anteriorly and/or posteriorly. In regard to the loss of the mucro, Curtis and Lane illustrated three pygidia assigned to
|
||
<taxonomicName id="D3C14D42FF903F1B67F4FDA3ADA3FDA9" box="[507,717,549,570]" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="tenuimucronatum">
|
||
<emphasis id="26B5EAD3FF903F1B67F4FDA3ADA3FDA9" box="[507,717,549,570]" italics="true" pageId="25" pageNumber="240">B. tenuimucronatum</emphasis>
|
||
</taxonomicName>
|
||
with the posterior termination preserved. Two of these (Curtis and Laneʼs pl. 8, figs 6, 8) have a mucro, that on the larger specimen being considerably shorter than the one on the smaller specimen. The third pygidium, which is very much larger that the others (Curtis and Laneʼs ʻtype 2ʼ pygidium, pl. 8, fig. 2), does not have ʻa shorter mucral spineʼ as stated by
|
||
<bibRefCitation id="70504B30FF903F1B646CFD54AE20FC97" author="Chatterton, B. D. E. & Ludvigsen, R." pageId="25" pageNumber="240" pagination="1 - 264" refId="ref29332" refString="Chatterton, B. D. E. and Ludvigsen, R. 2004. Early Silurian trilobites of Anticosti Island, Quebec, Canada. Palaeontographica Canadiana 22: 1 - 264." type="journal article" year="2004">Chatterton and Ludvigsen (2004)</bibRefCitation>
|
||
but an embayment in the margin posteromedially. Because of this difference, as well as the differences in size and the much greater number of axial rings and pleural furrows, it is not possible to be confident that this pygidium is correctly assigned to
|
||
<taxonomicName id="D3C14D42FF903F1B6412FCE5AD94FCEB" box="[541,762,867,888]" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="tenuimucronatum">
|
||
<emphasis id="26B5EAD3FF903F1B6412FCE5AD94FCEB" box="[541,762,867,888]" italics="true" pageId="25" pageNumber="240">B. tenuimucronatum</emphasis>
|
||
</taxonomicName>
|
||
. However, we note that it has a distinctive pleural structure in which the posterior bands expand distally and the anterior bands are pinched out, and that this structure appears to be shared by the smaller pygidium in pl. 8, fig. 6 (the structure is not clear in the third pygidium as the dorsal surface is broken away distally to reveal the doublure). Although we believe that these pygidia are correctly assigned to the
|
||
<taxonomicName id="D3C14D42FF903F1B6434FBA8ADA9FBD7" authorityName="Vodges" authorityYear="1890" box="[571,711,1070,1092]" class="Trilobita" family="Dalmanitidae" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="family">Dalmanitidae</taxonomicName>
|
||
, their pleural structure is unusual for Silurian and Devonian representatives of the family, in most of which it is the anterior rather than the posterior pleural band that is dominant distally (
|
||
<bibRefCitation id="70504B30FF903F1B669EFB24AE0CFB24" author="Holloway, D. J." box="[145,354,1186,1208]" pageId="25" pageNumber="240" pagination="695 - 731" refId="ref30324" refString="Holloway, D. J. 1981. Silurian dalmanitacean trilobites from North America and the origins of the Dalmanitinae and Synphoriinae. Palaeontology 24: 695 - 731." type="journal article" year="1981">Holloway, 1981: 710</bibRefCitation>
|
||
), as is also the case in
|
||
<taxonomicName id="D3C14D42FF903F1B6442FB24AC6FFB24" box="[589,769,1186,1207]" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="buttingtonense">
|
||
<emphasis id="26B5EAD3FF903F1B6442FB24AC6FFB24" box="[589,769,1186,1207]" italics="true" pageId="25" pageNumber="240">B. buttingtonense</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="70504B30FF903F1B669EFB38AE01FB47" author="Curtis, N. J. & Lane, P. D." box="[145,367,1214,1236]" pageId="25" pageNumber="240" pagination="51 - 101" refId="ref29505" refString="Curtis, N. J. and Lane, P. D. 1998. The Llandovery trilobites of England and Wales. Part 2. Monograph of the Palaeontographical Society London 152 (608): 51 - 101, pls 4 - 12." type="journal article" year="1998">Curtis and Lane, 1998</bibRefCitation>
|
||
, pl. 9, figs 2, 4, 7),
|
||
<taxonomicName id="D3C14D42FF903F1B6412FB39ADFDFB47" baseAuthorityName="Holloway and Sandford" baseAuthorityYear="1993" box="[541,659,1215,1236]" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="tigerense">
|
||
<emphasis id="26B5EAD3FF903F1B6412FB39ADFDFB47" box="[541,659,1215,1236]" italics="true" pageId="25" pageNumber="240">B. tigerense</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="70504B30FF903F1B6492FB38AE3BFB62" author="Holloway, D. J. & Sandford, A. C." pageId="25" pageNumber="240" pagination="85 - 102" refId="ref30463" refString="Holloway, D. J. and Sandford, A. C. 1993. An early Silurian trilobite fauna from Tasmania. Memoirs of the Association of Australasian Palaeontologists 15: 85 - 102." type="journal article" year="1993">Holloway and Sandford, 1993</bibRefCitation>
|
||
, fig. 6) and in the unnamed species from
|
||
<collectingRegion id="D605F823FF903F1B6685FB7EAFB2FA9D" box="[138,220,1272,1294]" country="Australia" name="Victoria" pageId="25" pageNumber="240">Victoria</collectingRegion>
|
||
described below. The only pygidium of
|
||
<taxonomicName id="D3C14D42FF903F1B64E4FB7FAE4FFAB9" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="buttingtonense">
|
||
<emphasis id="26B5EAD3FF903F1B64E4FB7FAE4FFAB9" italics="true" pageId="25" pageNumber="240">B. buttingtonense</emphasis>
|
||
</taxonomicName>
|
||
with the posterior termination preserved (Curtis and Lane, pl. 9, fig. 2a, b) is of about the same size as the ʻtype 2ʼ pygidium of
|
||
<taxonomicName id="D3C14D42FF903F1B6722FAD6AD6FFAF6" box="[301,513,1360,1381]" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="tenuimucronatum">
|
||
<emphasis id="26B5EAD3FF903F1B6722FAD6AD6FFAF6" box="[301,513,1360,1381]" italics="true" pageId="25" pageNumber="240">B. tenuimucronatum</emphasis>
|
||
</taxonomicName>
|
||
but has neither a mucro nor a posteromedian embayment; instead the margin is rather truncated in dorsal view and arched upwards in posterior view. There is no evidence that the mucro is lost in
|
||
<taxonomicName id="D3C14D42FF903F1B646FFA21ADB7FA2F" baseAuthorityName="Holloway and Sandford" baseAuthorityYear="1993" box="[608,729,1447,1468]" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="tigerense">
|
||
<emphasis id="26B5EAD3FF903F1B646FFA21ADB7FA2F" box="[608,729,1447,1468]" italics="true" pageId="25" pageNumber="240">B. tigerense</emphasis>
|
||
</taxonomicName>
|
||
but all known pygidia are smaller than that of
|
||
<taxonomicName id="D3C14D42FF903F1B6431FA45ADBBFA4B" box="[574,725,1475,1496]" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="buttingtonense">
|
||
<emphasis id="26B5EAD3FF903F1B6431FA45ADBBFA4B" box="[574,725,1475,1496]" italics="true" pageId="25" pageNumber="240">buttingtonense</emphasis>
|
||
</taxonomicName>
|
||
and the ʻtype 2ʼ pygidium of
|
||
<taxonomicName id="D3C14D42FF903F1B678AFA67AD54FA65" box="[389,570,1505,1526]" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="tenuimucronatum">
|
||
<emphasis id="26B5EAD3FF903F1B678AFA67AD54FA65" box="[389,570,1505,1526]" italics="true" pageId="25" pageNumber="240">tenuimucronatum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="147E36C1FF903F1B66BFFA7BAA21FDAF" blockId="25.[138,770,229,1903]" lastBlockId="25.[806,1437,229,572]" pageId="25" pageNumber="240">
|
||
Some of the other characters listed by Curtis and Lane as diagnostic of
|
||
<taxonomicName id="D3C14D42FF903F1B672AF99DAEFFF9A3" authorityName=", Chatterton and Ludvigsen" authorityYear="2004" box="[293,401,1563,1584]" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26B5EAD3FF903F1B672AF99DAEFFF9A3" box="[293,401,1563,1584]" italics="true" pageId="25" pageNumber="240">Bessazoon</emphasis>
|
||
</taxonomicName>
|
||
cannot in our view be used to distinguish
|
||
<taxonomicName id="D3C14D42FF903F1B670EF9BEAE03F9DE" authorityName=", Chatterton and Ludvigsen" authorityYear="2004" box="[257,365,1592,1613]" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26B5EAD3FF903F1B670EF9BEAE03F9DE" box="[257,365,1592,1613]" italics="true" pageId="25" pageNumber="240">Bessazoon</emphasis>
|
||
</taxonomicName>
|
||
either. The size of the palpebral area is determined by the size of the eye, and thus in comparison with other Silurian and Devonian dalmanitids is not particularly large in
|
||
<taxonomicName id="D3C14D42FF903F1B66D1F909AEFCF930" box="[222,402,1678,1700]" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="buttingtonense">
|
||
<emphasis id="26B5EAD3FF903F1B66D1F909AEFCF930" box="[222,402,1678,1700]" italics="true" pageId="25" pageNumber="240">B. buttingtonense</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="26B5EAD3FF903F1B67CBF909AEB4F937" box="[452,474,1679,1700]" italics="true" pageId="25" pageNumber="240">B.</emphasis>
|
||
cf.
|
||
<taxonomicName id="D3C14D42FF903F1B640FF909ADBCF937" box="[512,722,1679,1700]" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="tenuimucronatum">
|
||
<emphasis id="26B5EAD3FF903F1B640FF909AD61F937" box="[512,527,1679,1700]" italics="true" pageId="25" pageNumber="240">B</emphasis>
|
||
.
|
||
<emphasis id="26B5EAD3FF903F1B6414F909ADBCF937" box="[539,722,1679,1700]" italics="true" pageId="25" pageNumber="240">tenuimucronatum</emphasis>
|
||
</taxonomicName>
|
||
. We can see no difference from
|
||
<taxonomicName id="D3C14D42FF903F1B67CDF92DAD54F953" authorityName="Barrande" authorityYear="1852" box="[450,570,1707,1728]" class="Trilobita" family="Dalmanitidae" genus="Dalmanites" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26B5EAD3FF903F1B67CDF92DAD54F953" box="[450,570,1707,1728]" italics="true" pageId="25" pageNumber="240">Dalmanites</emphasis>
|
||
</taxonomicName>
|
||
and other closely related genera in the form of the palpebral lobe, which rises steeply from the palpebral furrow and becomes flat towards the outer margin. Finally, a posterior cephalic border furrow that fails to meet the lateral border furrow distally, an epiborder furrow on the lateral cephalic borders and genal spine, and a straight-sided pygidial axis are not unusual features for dalmanitids but are present in most Silurian and Devonian representatives. The discrimination and composition of
|
||
<taxonomicName id="D3C14D42FF903F1B6529FEA6ACFCFEA6" authorityName=", Chatterton and Ludvigsen" authorityYear="2004" box="[806,914,288,309]" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26B5EAD3FF903F1B6529FEA6ACFCFEA6" box="[806,914,288,309]" italics="true" pageId="25" pageNumber="240">Bessazoon</emphasis>
|
||
</taxonomicName>
|
||
are in need of review, but in the meantime we apply the name here to dalmanitids differing from species commonly assigned to
|
||
<taxonomicName id="D3C14D42FF903F1B6591FEDCAB78FEFC" authorityName="Barrande" authorityYear="1852" box="[926,1046,346,367]" class="Trilobita" family="Dalmanitidae" genus="Dalmanites" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26B5EAD3FF903F1B6591FEDCAB78FEFC" box="[926,1046,346,367]" italics="true" pageId="25" pageNumber="240">Dalmanites</emphasis>
|
||
</taxonomicName>
|
||
(e.g. see
|
||
<bibRefCitation id="70504B30FF903F1B627CFEDCAA74FEE3" author="Ramskold, L." box="[1139,1306,346,368]" pageId="25" pageNumber="240" pagination="1 - 62" refId="ref31314" refString="Ramskold, L. 1985. Silurian phacopid and dalmanitid trilobites from Gotland. Stockholm Contributions in Geolog y 40: 1 - 62." type="journal article" year="1985">Ramsköld, 1985</bibRefCitation>
|
||
) in lacking a well-developed anterior cephalic process (although in
|
||
<taxonomicName id="D3C14D42FF903F1B6388FEFEACB3FE39" class="Trilobita" family="Dalmanitidae" genus="Bessazoon" kingdom="Animalia" order="Phacopida" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="tenuimucronatum">
|
||
<emphasis id="26B5EAD3FF903F1B6388FEFEACB3FE39" italics="true" pageId="25" pageNumber="240">B. tenuimucronatum</emphasis>
|
||
</taxonomicName>
|
||
the cephalic margin is deflected slightly forwards medially) and tubercles on the glabella, and in having a pygidium with a curved posterolateral outline, a slender, narrow-based mucro merging anteriorly with a strong postaxial ridge, and a very wide doublure extending adaxially beyond the distal ends of the pleural and interpleural furrows.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |