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<mods:title id="3DCBF78A42571E1665E6C58DE651A839">Review of the genus Signoretia (Hemiptera: Cicadellidae: Signoretiinae) of the Oriental region with description of nine new species</mods:title>
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<mods:namePart id="588665FAAA6D33D5A9896AE1B44F1AD6">Viraktamath, C. A.</mods:namePart>
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<mods:namePart id="404691F815639A4249120A437F9CFA18">Webb, M. D.</mods:namePart>
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<treatment id="9C0F87C41875FFE2FF65F8AE48E8FDE9" ID-DOI="http://doi.org/10.5281/zenodo.3511113" ID-GBIF-Taxon="124770270" ID-Zenodo-Dep="3511113" LSID="urn:lsid:plazi:treatment:9C0F87C41875FFE2FF65F8AE48E8FDE9" httpUri="http://treatment.plazi.org/id/9C0F87C41875FFE2FF65F8AE48E8FDE9" lastPageId="3" lastPageNumber="489" pageId="2" pageNumber="488">
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<heading id="4F5181BE1875FFE3FF65F8AE49E0F885" bold="true" box="[151,475,1849,1875]" fontSize="11" level="1" pageId="2" pageNumber="488" reason="1">
<taxonomicName id="D3A64D511875FFE3FF65F8AE49E0F885" ID-CoL="4XB9F" authority="Distant" authorityName="Distant" box="[151,475,1849,1875]" class="Insecta" family="Cicadellidae" genus="Signoretia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="488" phylum="Arthropoda" rank="species" species="aureola">
<emphasis id="26D2EAC01875FFE3FF65F8AE49E0F885" bold="true" box="[151,475,1849,1875]" pageId="2" pageNumber="488">
<emphasis id="26D2EAC01875FFE3FF65F8AE4942F885" bold="true" box="[151,377,1849,1875]" italics="true" pageId="2" pageNumber="488">Signoretia aureola</emphasis>
Distant
</emphasis>
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</heading>
</paragraph>
<paragraph id="141936D21875FFE3FF65F8CB498DF8A2" blockId="2.[151,475,1849,1909]" box="[151,438,1884,1909]" pageId="2" pageNumber="488">
<figureCitation id="8C9D2A571875FFE3FF65F8CB48C1F8A2" box="[151,250,1884,1909]" captionStart="FIGURES 1 15" captionStartId="13.[151,264,1896,1919]" captionTargetBox="[208,1379,202,1866]" captionTargetId="figure@13.[194,1391,195,1879]" captionTargetPageId="13" captionText="FIGURES 1 15. Habitus of species of Signoretia. 1 4. S. aureola Distant: 1 2. Type, female; 3 4, Male from Hunli; 5. S. benguetensis Baker, syntype; 6. S. bilineata Baker, syntype; 7. S. carinata Baker, syntype; 8 11. S. dulitensis sp. nov., holotype and paratype; 12 13. S. greeni Distant, holotype; 14 15. S. lunglei sp. nov., holotype. Figs 1, 3, 6 8, 10, 12, 14. Dorsal aspect; Figs 2, 4, 5, 9, 11, 13, 15, lateral aspect." httpUri="https://zenodo.org/record/167192/files/figure.png" pageId="2" pageNumber="488">Figs 14</figureCitation>
,
<figureCitation id="8C9D2A571875FFE3FEF4F8CB491FF8A2" box="[262,292,1884,1908]" captionStart="FIGURES 47 65" captionStartId="16.[151,264,1836,1859]" captionTargetBox="[171,1423,199,1809]" captionTargetId="figure@16.[151,1436,193,1815]" captionTargetPageId="16" captionText="FIGURES 47 65. Habitus and faces of species of Signoretia. 47 48. S. takiyae sp. nov. holotype male; 49. S. aureola Distant; 50 51. S. dulitensis sp. nov., male and female; 52 53. S. lunglei sp. nov., male and female; 54. S. maculata, female; 55. S. mismiensis sp. nov.; 56 57. S. quoinensis, male and female; 58. S. rubra sp. nov.; 59 60. S. sahyadrica sp. nov. male and female; 61. S. similaris sp. nov. male; 62 63. S. sinuata, male and female; 64. S. sumatrana Schmidt; 65. S. takiyae sp. nov., male. Fig. 47. Dorsal aspect. Fig. 48. Lateral aspect. Figs 49 65. Ventral aspect." httpUri="https://zenodo.org/record/167195/files/figure.png" pageId="2" pageNumber="488">49</figureCitation>
,
<figureCitation id="8C9D2A571875FFE3FEC2F8CB4975F8A3" box="[304,334,1884,1909]" captionStart="FIGURES 66 79" captionStartId="17.[151,264,1923,1946]" captionTargetBox="[159,1410,217,1893]" captionTargetId="figure@17.[151,1436,193,1902]" captionTargetPageId="17" captionText="FIGURES 66 79. Profile of head and thorax of species of Signoretia. 66. S. aureola Distant; 67. S. dulitensis sp. nov.; 68. S. greeni Distant; 69. S. lunglei sp. nov.; 70 71. S. maculata Baker; 72. S. mishmiensis sp. nov.; 73. S. quoinensis sp. nov.; 74. S. rubra sp. nov.; 75 76. S. sahyadrica sp. nov.; 77. S. similaris sp. nov. 78. S. sinuata sp. nov.; 79. S. takiyae sp. nov." httpUri="https://zenodo.org/record/257129/files/figure.png" pageId="2" pageNumber="488">66</figureCitation>
,
<figureCitation id="8C9D2A571875FFE3FEA8F8CB4989F8A2" box="[346,434,1884,1908]" captionStart="FIGURES 98 104" captionStartId="20.[151,264,1946,1969]" captionTargetBox="[180,1401,208,1907]" captionTargetId="figure@20.[165,1412,193,1920]" captionTargetPageId="20" captionText="FIGURES 98 104. Signoretia aureola Distant. 98. Male genital capsule except subgenital plates, lateral view; 99. Process of segment X; 100. Connective, dorsal view; 101. Style, dorsal view; 102. Aedeagus, lateral view; 103. Ventral process of aedeagus, posterior view; 104. Adeagus, posterior view." httpUri="https://zenodo.org/record/167199/files/figure.png" pageId="2" pageNumber="488">98104</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5CBC65591875FFE3FF65F8334D59F86C" box="[151,1378,1956,1978]" pageId="2" pageNumber="488" type="reference_group">
<paragraph id="141936D21875FFE3FF65F8334D59F86C" blockId="2.[151,1378,1956,1978]" box="[151,1378,1956,1978]" pageId="2" pageNumber="488">
<treatmentCitationGroup id="34B611FC1875FFE3FF65F8334D59F86C" box="[151,1378,1956,1978]" pageId="2" pageNumber="488">
<taxonomicName id="D3A64D511875FFE3FF65F8334A2BF86C" ID-CoL="4XB9F" authority="Distant 1908: 232" authorityName="Distant" authorityPageNumber="232" authorityYear="1908" box="[151,528,1956,1978]" class="Insecta" family="Cicadellidae" genus="Signoretia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="488" phylum="Arthropoda" rank="species" species="aureola">
<emphasis id="26D2EAC01875FFE3FF65F8334969F86C" box="[151,338,1956,1978]" italics="true" pageId="2" pageNumber="488">Signoretia aureola</emphasis>
<treatmentCitation id="950710C31875FFE3FEA8F8334A2BF86C" author="Distant" box="[346,528,1956,1978]" page="232" pageId="2" pageNumber="488" year="1908">
<bibRefCitation id="70374B231875FFE3FEA8F83349E4F86C" author="Distant" box="[346,479,1956,1978]" pageId="2" pageNumber="488" refString="Distant, W. L. (1908) Rhynchota, vol. IV. Homoptera and Appendix (Pt). In: Bingham, C. T. (Ed.), The Fauna of British India including Ceylon and Burma, 501 pp. [Taylor and Francis, London.]" type="book" year="1908">Distant 1908</bibRefCitation>
: 232
</treatmentCitation>
</taxonomicName>
233, fig. 148;
<treatmentCitation id="950710C31875FFE3FD59F8334B4DF86C" author="Anufriev" box="[683,886,1956,1978]" page="725" pageId="2" pageNumber="488" year="1971">
<bibRefCitation id="70374B231875FFE3FD59F8334B79F86C" author="Anufriev" box="[683,834,1956,1978]" pageId="2" pageNumber="488" refString="Anufriev, G. A. (1971) Study of leafhoppers from the tribe Signoretiini (Homoptera, Aunchenorrhyncha) with description of two new species of Signoretia Stal from western Africa. Bulletin de l'Akademie Plonoaise des Sciences, Serie des Sciences Biologiques, 19, 721 - 726." type="journal article" year="1971">Anufriev 1971</bibRefCitation>
: 725
</treatmentCitation>
: figs 1219;
<treatmentCitation id="950710C31875FFE3FC0BF8334CDFF86C" ID-DOI="http://doi.org/10.5281/zenodo.3507017" ID-Zenodo-Dep="3507017" author="Takiya" box="[1017,1252,1956,1978]" httpUri="http://treatment.plazi.org/id/25BB1FD858A15D49B2E0BE20E8D501A5" page="315" pageId="2" pageNumber="488" year="2013">
<bibRefCitation id="70374B231875FFE3FC0BF8334C88F86C" author="Takiya" box="[1017,1203,1956,1978]" pageId="2" pageNumber="488" refString="Takiya, D. M., Dietrich, C. H. &amp; Viraktamath, C. A. (2013) The unusual Afrotropical and Oriental leafhopper subfamily Signoretiinae (Hemiptera, Cicadellidae): taxonomic notes, new distributional records, and description of two new Signoretia species. Zookeys, 319, 303 - 323." type="journal article" year="2013">
Takiya
<emphasis id="26D2EAC01875FFE3FBB1F8324C4BF86C" box="[1091,1136,1956,1978]" italics="true" pageId="2" pageNumber="488">et al</emphasis>
. 2013
</bibRefCitation>
: 315
</treatmentCitation>
, figs 1920.
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</paragraph>
</subSubSection>
<subSubSection id="5CBC65591874FFE2FF65FF004C98FF2D" pageId="3" pageNumber="489" type="materials_examined">
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<location id="117960091874FFE2FF65FF0048C4FF66" LSID="urn:lsid:plazi:treatment:9C0F87C41875FFE2FF65F8AE48E8FDE9:117960091874FFE2FF65FF0048C4FF66" box="[151,255,151,176]" country="Myanmar" name="Material" pageId="3" pageNumber="489">Material</location>
examined.
</emphasis>
<collectingCountry id="6CB176421874FFE2FE67FF0F4A17FF66" box="[405,556,152,176]" name="Myanmar" pageId="3" pageNumber="489">MYANMAR</collectingCountry>
:
<typeStatus id="CB1D88701874FFE2FDCCFF0F4A95FF66" box="[574,686,152,176]" pageId="3" pageNumber="489" type="lectotype">Lectotype</typeStatus>
<specimenCount id="02A0FD5B1874FFE2FD4EFF0F4AEAFF79" box="[700,721,152,175]" pageId="3" pageNumber="489" type="female"></specimenCount>
(here designated)
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,
<materialsCitation id="A4CE3C8F1874FFE2FC4FFF0F4DA2FF66" ID-GBIF-Occurrence="1326058673" box="[957,1433,151,176]" collectionCode="BMNH" collectorName="Ruby Mines &amp; Doherty" country="Myanmar" location="Burma" pageId="3" pageNumber="489" specimenCount="1" typeStatus="lectotype">
<collectingCountry id="6CB176421874FFE2FC4FFF0F4C37FF79" box="[957,1036,152,175]" name="Myanmar" pageId="3" pageNumber="489">Burma</collectingCountry>
,
<collectorName id="B95353041874FFE2FBEDFF004C89FF66" box="[1055,1202,151,176]" pageId="3" pageNumber="489">Ruby Mines</collectorName>
,
<collectorName id="B95353041874FFE2FB36FF004D1BFF66" box="[1220,1312,151,176]" pageId="3" pageNumber="489">Doherty</collectorName>
(
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)
</materialsCitation>
.
<materialsCitation id="A4CE3C8F1874FFE2FF65FF284CA4FF2D" ID-GBIF-Occurrence="1326058676" collectingDate="2014-09-12" collectionCode="UASB" collectorName="Km S. &amp; Yeshwanth, H. M. &amp; C. A. Viraktamath" country="India" elevation="1597" location="Hunli" pageId="3" pageNumber="489" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="lectotype">
<collectingCountry id="6CB176421874FFE2FF65FF2848D0FF01" box="[151,235,191,215]" name="India" pageId="3" pageNumber="489">INDIA</collectingCountry>
:
<collectingRegion id="D662F8301874FFE2FF0AFF2949EFFF01" box="[248,468,190,215]" country="India" name="Arunachal Pradesh" pageId="3" pageNumber="489">Arunachal Pradesh</collectingRegion>
: 2 Ƌ, 10
<collectorName id="B95353041874FFE2FDB4FF574AABFF01" box="[582,656,191,215]" pageId="3" pageNumber="489">Km S.</collectorName>
of
<location id="117960091874FFE2FD4EFF294AC4FF01" LSID="urn:lsid:plazi:treatment:9C0F87C41875FFE2FF65F8AE48E8FDE9:117960091874FFE2FD4EFF294AC4FF01" box="[700,767,190,215]" country="India" name="Hunli" pageId="3" pageNumber="489" stateProvince="Arunachal Pradesh">Hunli</location>
, 28o, 19 05.1” N, 95o 56 015.2” E,
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<quantity id="D35E9B371874FFE2FB46FF294D35FF01" box="[1204,1294,190,215]" metricMagnitude="3" metricUnit="m" metricValue="1.597" pageId="3" pageNumber="489" unit="m" value="1597.0">1597 m</quantity>
</elevation>
,
<collectingDate id="705CE9FA1874FFE2FAEFFF294DACFF01" box="[1309,1431,190,215]" pageId="3" pageNumber="489" value="2014-09-12">
<date id="601810121874FFE2FAEFFF294DACFF01" box="[1309,1431,190,215]" pageId="3" pageNumber="489" value="2014-09-12">12.ix.2014</date>
</collectingDate>
,
<collectingMethod id="CDE74EC51874FFE2FF65FF744936FF2C" box="[151,269,227,251]" pageId="3" pageNumber="489">Sweep net</collectingMethod>
,
<collectorName id="B95353041874FFE2FEEBFF7549E4FF2C" box="[281,479,226,251]" pageId="3" pageNumber="489">Yeshwanth, H.M.</collectorName>
, 1 Ƌ, same data except collected by
<collectorName id="B95353041874FFE2FC8BFF754C0CFF2D" box="[889,1079,226,251]" pageId="3" pageNumber="489">C.A.Viraktamath</collectorName>
(
<collectionCode id="72B7AE171874FFE2FBB5FF744CACFF2D" box="[1095,1175,227,251]" country="India" httpUri="http://grbio.org/cool/yr7i-jnjm" name="University of Agricultural Sciences" pageId="3" pageNumber="489">UASB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5CBC65591874FFE2FF35FE9148E8FDE9" pageId="3" pageNumber="489" type="discussion">
<paragraph id="141936D21874FFE2FF35FE9148E8FDE9" blockId="3.[151,1437,151,575]" pageId="3" pageNumber="489">
<emphasis id="26D2EAC01874FFE2FF35FE91490FFEC9" bold="true" box="[199,308,262,287]" pageId="3" pageNumber="489">Remarks</emphasis>
.
<bibRefCitation id="70374B231874FFE2FEB7FE9149D0FEC9" author="Distant" box="[325,491,262,287]" pageId="3" pageNumber="489" refString="Distant, W. L. (1908) Rhynchota, vol. IV. Homoptera and Appendix (Pt). In: Bingham, C. T. (Ed.), The Fauna of British India including Ceylon and Burma, 501 pp. [Taylor and Francis, London.]" type="book" year="1908">Distant (1908)</bibRefCitation>
described this species based on unspecified number of specimens collected from “
<collectingCountry id="6CB176421874FFE2FF53FEBC48C9FE95" box="[161,242,299,323]" name="Myanmar" pageId="3" pageNumber="489">Burma</collectingCountry>
, Ruby Mines&quot; (presently Mogok). This species appears to vary in coloration with respect to the dark markings on the crown and the pronotum as already noted by
<bibRefCitation id="70374B231874FFE2FC81FED94C26FEB1" author="Distant" box="[883,1053,334,359]" pageId="3" pageNumber="489" refString="Distant, W. L. (1908) Rhynchota, vol. IV. Homoptera and Appendix (Pt). In: Bingham, C. T. (Ed.), The Fauna of British India including Ceylon and Burma, 501 pp. [Taylor and Francis, London.]" type="book" year="1908">Distant (1908)</bibRefCitation>
. The male specimens from
<collectingCountry id="6CB176421874FFE2FA90FED94DA7FEB1" box="[1378,1436,334,359]" name="India" pageId="3" pageNumber="489">India</collectingCountry>
identified here have a fuscous stripe along the claval suture, costal margin and claval commissure, and the apical veins are fuscous and the forewings lack the golden yellow coloration of the female
<typeStatus id="CB1D88701874FFE2FBBEFE0F4C47FE66" box="[1100,1148,408,432]" pageId="3" pageNumber="489">type</typeStatus>
(
<figureCitation id="8C9D2A571874FFE2FB79FE014CCAFE79" box="[1163,1265,406,431]" captionStart="FIGURES 1 15" captionStartId="13.[151,264,1896,1919]" captionTargetBox="[208,1379,202,1866]" captionTargetId="figure@13.[194,1391,195,1879]" captionTargetPageId="13" captionText="FIGURES 1 15. Habitus of species of Signoretia. 1 4. S. aureola Distant: 1 2. Type, female; 3 4, Male from Hunli; 5. S. benguetensis Baker, syntype; 6. S. bilineata Baker, syntype; 7. S. carinata Baker, syntype; 8 11. S. dulitensis sp. nov., holotype and paratype; 12 13. S. greeni Distant, holotype; 14 15. S. lunglei sp. nov., holotype. Figs 1, 3, 6 8, 10, 12, 14. Dorsal aspect; Figs 2, 4, 5, 9, 11, 13, 15, lateral aspect." httpUri="https://zenodo.org/record/167192/files/figure.png" pageId="3" pageNumber="489">Figs 12</figureCitation>
). However, the male genitalia are similar to those illustrated by
<bibRefCitation id="70374B231874FFE2FD35FE2D4BB8FE05" author="Anufriev" box="[711,899,442,467]" pageId="3" pageNumber="489" refString="Anufriev, G. A. (1971) Study of leafhoppers from the tribe Signoretiini (Homoptera, Aunchenorrhyncha) with description of two new species of Signoretia Stal from western Africa. Bulletin de l'Akademie Plonoaise des Sciences, Serie des Sciences Biologiques, 19, 721 - 726." type="journal article" year="1971">Anufriev (1971)</bibRefCitation>
. This species can readily be recognised by the greatly elongated ventral process of the pygofer similar to that of
<taxonomicName id="D3A64D511874FFE2FC9DFE484C69FE21" authority="Baker" authorityName="Baker" box="[879,1106,478,503]" class="Insecta" family="Cicadellidae" genus="Signoretia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="489" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="26D2EAC01874FFE2FC9DFE484BC9FE21" box="[879,1010,479,503]" italics="true" pageId="3" pageNumber="489">S. maculata</emphasis>
(Baker)
</taxonomicName>
. However, both species differ in the shape of the atrium of the aedeagus which is much narrower in
<taxonomicName id="D3A64D511874FFE2FC53FD944C29FDCD" box="[929,1042,515,539]" class="Insecta" family="Cicadellidae" genus="Signoretia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="489" phylum="Arthropoda" rank="species" species="aureola">
<emphasis id="26D2EAC01874FFE2FC53FD944B94FDCC" box="[929,943,515,538]" italics="true" pageId="3" pageNumber="489">S</emphasis>
.
<emphasis id="26D2EAC01874FFE2FC4EFD944C29FDCD" box="[956,1042,515,539]" italics="true" pageId="3" pageNumber="489">aureola</emphasis>
</taxonomicName>
compared to
<taxonomicName id="D3A64D511874FFE2FB59FD944D14FDCD" box="[1195,1327,515,539]" class="Insecta" family="Cicadellidae" genus="Signoretia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="489" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="26D2EAC01874FFE2FB59FD944C82FDCC" box="[1195,1209,515,538]" italics="true" pageId="3" pageNumber="489">S</emphasis>
.
<emphasis id="26D2EAC01874FFE2FB35FD944D14FDCD" box="[1223,1327,515,539]" italics="true" pageId="3" pageNumber="489">maculata</emphasis>
</taxonomicName>
in caudal view.
</paragraph>
</subSubSection>
</treatment>
</document>