treatments-xml/data/9B/53/B9/9B53B917FFB0FFADFF32C9CCFBEC5E69.xml
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<document id="E9B37A5481260A0A65A8EA52F281196A" ID-CLB-Dataset="8480" ID-DOI="10.11646/zootaxa.4858.2.4" ID-GBIF-Dataset="5d0fb569-f8ca-4552-949d-6b506ec71351" ID-ISSN="1175-5326" ID-Zenodo-Dep="4411799" ID-ZooBank="04F9D9BE-70BD-410C-8BC4-DB942A0A92D0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609611275837" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2020" docId="9B53B917FFB0FFADFF32C9CCFBEC5E69" docLanguage="en" docName="zootaxa.4858.2.4.pdf" docOrigin="Zootaxa 4858 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraneseuthia libanica Jałoszyński 2020, sp. n." docType="treatment" docVersion="8" lastPageNumber="234" masterDocId="676AC16FFFB1FFAEFFA5CB1AFFD55B3C" masterDocTitle="Four new Palaearctic, Australian and Oriental species of Paraneseuthia Franz (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="240" masterPageNumber="231" pageNumber="232" updateTime="1698851499168" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F8A2BAEF62C34F181E516DE3D5C6515D">Four new Palaearctic, Australian and Oriental species of Paraneseuthia Franz (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:name id="44BC3965C1FB1707CE9E65C913159A10" type="personal">
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<mods:namePart id="ED5C51DA504D232748E5B7D829525688">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="26A27C2DB50AC17FE46ACD18A67C913D">Zootaxa</mods:title>
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<mods:part id="C5F3292681E629B666B443C31C10D5C4">
<mods:date id="5AAB3EAAEB139168AA3321E79F971E30">2020</mods:date>
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<mods:number id="464A93E7B58F9378BF6B7BBCBE78186F">2020-09-30</mods:number>
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<mods:number id="5D1F1BC075758230C993E71743EA7192">4858</mods:number>
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<mods:identifier id="94A5FEBC4BFB36366F3CDEF02D42F5B1" type="DOI">10.11646/zootaxa.4858.2.4</mods:identifier>
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<treatment id="9B53B917FFB0FFADFF32C9CCFBEC5E69" ID-DOI="http://doi.org/10.5281/zenodo.4501305" ID-GBIF-Taxon="177209810" ID-Zenodo-Dep="4501305" LSID="urn:lsid:plazi:treatment:9B53B917FFB0FFADFF32C9CCFBEC5E69" httpUri="http://treatment.plazi.org/id/9B53B917FFB0FFADFF32C9CCFBEC5E69" lastPageId="3" lastPageNumber="234" pageId="1" pageNumber="232">
<subSubSection id="5BE05B8AFFB0FFAFFF32C9CCFE2F59CD" box="[151,506,726,753]" pageId="1" pageNumber="232" type="nomenclature">
<paragraph id="13450801FFB0FFAFFF32C9CCFE2F59CD" blockId="1.[151,506,726,789]" box="[151,506,726,753]" pageId="1" pageNumber="232">
<heading id="480DBF6DFFB0FFAFFF32C9CCFE2F59CD" bold="true" box="[151,506,726,753]" fontSize="11" level="1" pageId="1" pageNumber="232" reason="1">
<emphasis id="218ED413FFB0FFAFFF32C9CCFE2F59CD" bold="true" box="[151,506,726,753]" pageId="1" pageNumber="232">
<taxonomicName id="D4FA7382FFB0FFAFFF32C9CCFE6459CD" ID-CoL="8Y7T7" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[151,433,726,753]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="species" species="libanica" status="sp. nov.">
<emphasis id="218ED413FFB0FFAFFF32C9CCFE6459CD" bold="true" box="[151,433,726,753]" italics="true" pageId="1" pageNumber="232">Paraneseuthia libanica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3ABD6968FFB0FFAFFE1DC9CDFE2F59CD" box="[440,506,727,753]" pageId="1" pageNumber="232" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5BE05B8AFFB0FFAFFF32C9E1FEDE5829" box="[151,267,763,789]" pageId="1" pageNumber="232" type="description">
<paragraph id="13450801FFB0FFAFFF32C9E1FEDE5829" blockId="1.[151,506,726,789]" box="[151,267,763,789]" pageId="1" pageNumber="232">
(
<figureCitation id="8BC11484FFB0FFAFFF3AC9E1FED75829" box="[159,258,763,789]" captionStart="FIGURES 18" captionStartId="2.[151,264,1848,1872]" captionTargetBox="[163,1430,193,1812]" captionTargetId="figure-19@2.[151,1436,179,1823]" captionTargetPageId="2" captionText="FIGURES 18. Paraneseuthia libanica sp. n., male. Dorsal habitus (1); head in ventral view (2); prothorax in ventral view (3); pterothorax in ventral view (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.4411801" httpUri="https://zenodo.org/record/4411801/files/figure.png" pageId="1" pageNumber="232">Figs 18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5BE05B8AFFB0FFAFFF32C859FE7058F5" pageId="1" pageNumber="232" type="materials_examined">
<paragraph id="13450801FFB0FFAFFF32C859FE7058F5" blockId="1.[151,1437,834,1977]" pageId="1" pageNumber="232">
<emphasis id="218ED413FFB0FFAFFF32C859FE915860" bold="true" box="[151,324,834,861]" pageId="1" pageNumber="232">Type material.</emphasis>
<materialsCitation id="A392025CFFB0FFAFFEEFC859FD2C5861" ID-GBIF-Occurrence="3029723302" box="[330,761,834,861]" country="Lebanon" location="LEBANON" pageId="1" pageNumber="232" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="218ED413FFB0FFAFFEEFC859FD8F5860" bold="true" box="[330,602,834,861]" pageId="1" pageNumber="232">
<typeStatus id="CC41B6A3FFB0FFAFFEEFC859FE6E5861" box="[330,443,835,861]" pageId="1" pageNumber="232" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="6BED4891FFB0FFAFFE62C858FD835860" box="[455,598,834,860]" name="Lebanon" pageId="1" pageNumber="232">LEBANON</collectingCountry>
:
</emphasis>
<specimenCount id="05FCC388FFB0FFAFFDC4C859FDAD5861" box="[609,632,835,861]" pageId="1" pageNumber="232" type="male"></specimenCount>
, two labels
</materialsCitation>
:
<materialsCitation id="A392025CFFB0FFAFFCA7C859FED65899" ID-GBIF-Occurrence="3029723307" collectionCode="MHNG" collectorName="P. Jaloszynski" country="Lebanon" location="LIBAN" pageId="1" pageNumber="232" specimenCount="1" typeStatus="holotype">
<collectingCountry id="6BED4891FFB0FFAFFCA9C859FCB15861" box="[780,868,835,861]" name="Lebanon" pageId="1" pageNumber="232">LIBAN</collectingCountry>
- 1975 / Damour 24 III / 1b / BESUCHET” [white, printed and handwritten]; “
<taxonomicName id="D4FA7382FFB0FFAFFE6FC87DFD7558BD" authorityName="Franz" authorityYear="1986" box="[458,672,871,897]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="genus">
<emphasis id="218ED413FFB0FFAFFE6FC87DFD7558BD" box="[458,672,871,897]" italics="true" pageId="1" pageNumber="232">PARANESEUTHIA</emphasis>
</taxonomicName>
/
<taxonomicName id="D4FA7382FFB0FFAFFD1DC87DFCC158BD" authorityName="Jałoszyński" authorityYear="2020" box="[696,788,871,897]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="232" phylum="Arthropoda" rank="species" species="libanica">
<emphasis id="218ED413FFB0FFAFFD1DC87DFCC158BD" box="[696,788,871,897]" italics="true" pageId="1" pageNumber="232">libanica</emphasis>
</taxonomicName>
m. /
<collectorName id="BE0F6DD7FFB0FFAFFCF4C87DFC2558BD" box="[849,1008,871,897]" pageId="1" pageNumber="232">P. Jałoszyński</collectorName>
, 2020 /
<typeStatus id="CC41B6A3FFB0FFAFFBE9C87DFB2358BD" box="[1100,1270,871,897]" pageId="1" pageNumber="232" type="holotype">HOLOTYPUS</typeStatus>
” [red, printed] (
<collectionCode id="75EB90C4FFB0FFAFFF3AC891FF2E5899" box="[159,251,907,933]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="232" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="A392025CFFB0FFAFFEAAC891FE7458F5" ID-GBIF-Occurrence="3029723305" collectionCode="MHNG" collectorName="P. Jaloszynski" country="Lebanon" location="LIBAN" pageId="1" pageNumber="232" specimenCount="8" specimenCount-female="5" specimenCount-male="3" typeStatus="paratype">
<emphasis id="218ED413FFB0FFAFFEAAC891FE525899" bold="true" box="[271,391,907,933]" pageId="1" pageNumber="232">
<typeStatus id="CC41B6A3FFB0FFAFFEAAC891FE525899" box="[271,391,907,933]" pageId="1" pageNumber="232" type="paratype">Paratypes</typeStatus>
</emphasis>
(17 exx., incl. 2 disarticulated):
<specimenCount id="05FCC388FFB0FFAFFD58C891FCEA5899" box="[765,831,907,933]" pageId="1" pageNumber="232" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="05FCC388FFB0FFAFFCE8C890FC5A5899" box="[845,911,906,933]" pageId="1" pageNumber="232" type="female">5 ♀♀</specimenCount>
, and 9 exx. of unknown sex, same data as for holotype (cPJ,
<collectionCode id="75EB90C4FFB0FFAFFE9BC8B5FE4C58F5" box="[318,409,943,969]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="232" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5BE05B8AFFB0FFAFFF62C8C8FC955F65" pageId="1" pageNumber="232" type="diagnosis">
<paragraph id="13450801FFB0FFAFFF62C8C8FC955F65" blockId="1.[151,1437,834,1977]" pageId="1" pageNumber="232">
<emphasis id="218ED413FFB0FFAFFF62C8C8FE9758D0" bold="true" box="[199,322,978,1004]" pageId="1" pageNumber="232">Diagnosis.</emphasis>
Body conspicuously small (BL &lt;
<quantity id="D402A5E4FFB0FFAFFD1CC8C9FCDA58D1" box="[697,783,979,1005]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="232" unit="mm" value="0.8">0.8 mm</quantity>
) and slender, covered with short and recumbent setae; eyes in both sexes small and weakly convex; head, protibia and metaventrite in male unmodified; aedeagus with asymmetrical distal half and a pair of lateral apical projections directed distodorsad, with diaphragm oblique in relation to the long axis of aedeagus and situated in subapical region.
</paragraph>
</subSubSection>
<subSubSection id="5BE05B8AFFB0FFAFFF62CF78FB0D5CA9" pageId="1" pageNumber="232" type="description">
<paragraph id="13450801FFB0FFAFFF62CF78FBDD5F9D" blockId="1.[151,1437,834,1977]" pageId="1" pageNumber="232">
<emphasis id="218ED413FFB0FFAFFF62CF78FE8C5F40" bold="true" box="[199,345,1122,1148]" pageId="1" pageNumber="232">Description.</emphasis>
Body of male (
<figureCitation id="8BC11484FFB0FFAFFDA9CF79FD9B5F41" box="[524,590,1123,1149]" captionStart="FIGURES 18" captionStartId="2.[151,264,1848,1872]" captionTargetBox="[163,1430,193,1812]" captionTargetId="figure-19@2.[151,1436,179,1823]" captionTargetPageId="2" captionText="FIGURES 18. Paraneseuthia libanica sp. n., male. Dorsal habitus (1); head in ventral view (2); prothorax in ventral view (3); pterothorax in ventral view (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.4411801" httpUri="https://zenodo.org/record/4411801/files/figure.png" pageId="1" pageNumber="232">Fig. 1</figureCitation>
) slender and moderately convex, dark brown with slightly lighter appendages, especially antennae and maxillary palps, setae light brown; BL
<quantity id="D402A5E4FFB0FFAFFC38CF9DFBD15F9D" box="[925,1028,1159,1185]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="1" pageNumber="232" unit="mm" value="0.74">0.74 mm</quantity>
.
</paragraph>
<paragraph id="13450801FFB0FFAFFF62CFB1FA4C5E45" blockId="1.[151,1437,834,1977]" pageId="1" pageNumber="232">
Head (
<figureCitation id="8BC11484FFB0FFAFFEB4CFB1FEA65FF9" box="[273,371,1195,1221]" captionStart="FIGURES 18" captionStartId="2.[151,264,1848,1872]" captionTargetBox="[163,1430,193,1812]" captionTargetId="figure-19@2.[151,1436,179,1823]" captionTargetPageId="2" captionText="FIGURES 18. Paraneseuthia libanica sp. n., male. Dorsal habitus (1); head in ventral view (2); prothorax in ventral view (3); pterothorax in ventral view (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.4411801" httpUri="https://zenodo.org/record/4411801/files/figure.png" pageId="1" pageNumber="232">Figs 12</figureCitation>
) broadest at inconspicuous, small and weakly convex eyes, HL
<quantity id="D402A5E4FFB0FFAFFB9BCFB1FB715FF9" box="[1086,1188,1195,1221]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="232" unit="mm" value="0.09">0.09 mm</quantity>
, HW
<quantity id="D402A5E4FFB0FFAFFB41CFB0FA9E5FF9" box="[1252,1355,1194,1221]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="232" unit="mm" value="0.15">0.15 mm</quantity>
; vertex and frons confluent, convex at middle and slightly impressed at sides; supraantennal tubercles barely discernible. Punctures on vertex and frons relatively large but with diffuse margins, so that cuticle appears uneven; setae short, sparse and recumbent. Antennae (
<figureCitation id="8BC11484FFB0FFAFFDBCCE0DFDAB5E0D" box="[537,638,1303,1329]" captionStart="FIGURES 18" captionStartId="2.[151,264,1848,1872]" captionTargetBox="[163,1430,193,1812]" captionTargetId="figure-19@2.[151,1436,179,1823]" captionTargetPageId="2" captionText="FIGURES 18. Paraneseuthia libanica sp. n., male. Dorsal habitus (1); head in ventral view (2); prothorax in ventral view (3); pterothorax in ventral view (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.4411801" httpUri="https://zenodo.org/record/4411801/files/figure.png" pageId="1" pageNumber="232">Figs 12</figureCitation>
) short but slender, with distinctly delimited trimerous club, AnL
<quantity id="D402A5E4FFB0FFAFFACECE0DFF105E69" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="232" unit="mm" value="0.28">0.28 mm</quantity>
, antennomeres III each elongate, III about as long as broad, IV weakly elongate, VVI each about as long as broad, VIIX each transverse, XI not broader than X, about as long as IXX together, about 1.8× as long as broad.
</paragraph>
<paragraph id="13450801FFB0FFAFFF62CE99FB8D5D35" blockId="1.[151,1437,834,1977]" pageId="1" pageNumber="232">
Pronotum (
<figureCitation id="8BC11484FFB0FFAFFEE1CE99FE735EA1" box="[324,422,1411,1437]" captionStart="FIGURES 18" captionStartId="2.[151,264,1848,1872]" captionTargetBox="[163,1430,193,1812]" captionTargetId="figure-19@2.[151,1436,179,1823]" captionTargetPageId="2" captionText="FIGURES 18. Paraneseuthia libanica sp. n., male. Dorsal habitus (1); head in ventral view (2); prothorax in ventral view (3); pterothorax in ventral view (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.4411801" httpUri="https://zenodo.org/record/4411801/files/figure.png" pageId="1" pageNumber="232">Figs 1, 3</figureCitation>
) subquadrate with all margins rounded, broadest near anterior third; PL
<quantity id="D402A5E4FFB0FFAFFB6ECE99FAE55EA1" box="[1227,1328,1411,1437]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="232" unit="mm" value="0.2">0.20 mm</quantity>
, PW
<quantity id="D402A5E4FFB0FFAFFACECE99FF135EFD" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="232" unit="mm" value="0.2">0.20 mm</quantity>
; posterior and anterior corners obtuse-angled and blunt; pronotal base with small but distinct median pit and two pairs of similarly small and distinct lateral pits, of which the lateralmost pair is developed as elongate impressions. Punctures on pronotal disc fine and inconspicuous; setae short, sparse and recumbent.
</paragraph>
<paragraph id="13450801FFB0FFAFFF62CD09FEBA5DA5" blockId="1.[151,1437,834,1977]" pageId="1" pageNumber="232">
Elytra (
<figureCitation id="8BC11484FFB0FFAFFEBBCD09FEB75D11" box="[286,354,1555,1581]" captionStart="FIGURES 18" captionStartId="2.[151,264,1848,1872]" captionTargetBox="[163,1430,193,1812]" captionTargetId="figure-19@2.[151,1436,179,1823]" captionTargetPageId="2" captionText="FIGURES 18. Paraneseuthia libanica sp. n., male. Dorsal habitus (1); head in ventral view (2); prothorax in ventral view (3); pterothorax in ventral view (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.4411801" httpUri="https://zenodo.org/record/4411801/files/figure.png" pageId="1" pageNumber="232">Fig. 1</figureCitation>
) together oval, broadest between middle and anterior third and relatively strongly narrowing posterad, evenly convex; EL
<quantity id="D402A5E4FFB0FFAFFE45CD2CFD925D6D" box="[480,583,1590,1617]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="232" unit="mm" value="0.45">0.45 mm</quantity>
, EW
<quantity id="D402A5E4FFB0FFAFFD20CD2DFD395D6D" box="[645,748,1591,1617]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="232" unit="mm" value="0.3">0.30 mm</quantity>
, EI 1.50; humeral calli and basal impressions lacking; apices separately rounded. Punctures on elytra inconspicuous, dense but small and diffuse; setae sparse and recumbent. Hind wings absent.
</paragraph>
<paragraph id="13450801FFB0FFAFFF62CDB9FC245D81" blockId="1.[151,1437,834,1977]" box="[199,1009,1699,1725]" pageId="1" pageNumber="232">Legs moderately long and slender; unmodified, all tibiae nearly straight.</paragraph>
<paragraph id="13450801FFB0FFAFFF62CDDDFDCC5C71" blockId="1.[151,1437,834,1977]" pageId="1" pageNumber="232">
Aedeagus (
<figureCitation id="8BC11484FFB0FFAFFEE3CDDDFE7E5DDD" box="[326,427,1734,1761]" captionStart="FIGURES 18" captionStartId="2.[151,264,1848,1872]" captionTargetBox="[163,1430,193,1812]" captionTargetId="figure-19@2.[151,1436,179,1823]" captionTargetPageId="2" captionText="FIGURES 18. Paraneseuthia libanica sp. n., male. Dorsal habitus (1); head in ventral view (2); prothorax in ventral view (3); pterothorax in ventral view (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.4411801" httpUri="https://zenodo.org/record/4411801/files/figure.png" pageId="1" pageNumber="232">Figs 58</figureCitation>
) moderately slender, AeL
<quantity id="D402A5E4FFB0FFAFFD7CCDDDFC975DDD" box="[729,834,1735,1761]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="232" unit="mm" value="0.24">0.24 mm</quantity>
; median lobe in ventral view with symmetrical basal half and asymmetrical distal half; apex with a pair of elongate projections directed distodorsad; diaphragm oblique in relation to the long axis of aedeagus, situated in subapical region; parameres slender, each with slightly broadened apical region, lacking apical setae.
</paragraph>
<paragraph id="13450801FFB0FFAFFF62CC4DFB0D5CA9" blockId="1.[151,1437,834,1977]" pageId="1" pageNumber="232">
Female. Externally indistinguishable from male. BL
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; HL
<quantity id="D402A5E4FFB0FFAFFBADCC4DFB7A5C4D" box="[1032,1199,1879,1905]" metricMagnitude="-5" metricUnit="m" metricValue="8.499999999999998" metricValueMax="9.0" metricValueMin="8.0" pageId="1" pageNumber="232" unit="mm" value="0.08499999999999999" valueMax="0.09" valueMin="0.08">0.080.09 mm</quantity>
, HW
<quantity id="D402A5E4FFB0FFAFFB50CC4CFA885C4D" box="[1269,1373,1878,1905]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="232" unit="mm" value="0.15">0.15 mm</quantity>
, AnL
<quantity id="D402A5E4FFB0FFAFFF32CC60FF2B5CA9" box="[151,254,1914,1941]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="232" unit="mm" value="0.25">0.25 mm</quantity>
; PL
<quantity id="D402A5E4FFB0FFAFFE95CC61FE435CA9" box="[304,406,1915,1941]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="232" unit="mm" value="0.2">0.20 mm</quantity>
, PW
<quantity id="D402A5E4FFB0FFAFFE77CC61FDAD5CA9" box="[466,632,1915,1941]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="1" pageNumber="232" unit="mm" value="0.22" valueMax="0.23" valueMin="0.21">0.210.23 mm</quantity>
; EL
<quantity id="D402A5E4FFB0FFAFFD09CC61FC845CA9" box="[684,849,1915,1941]" metricMagnitude="-4" metricUnit="m" metricValue="4.05" metricValueMax="4.3" metricValueMin="3.8" pageId="1" pageNumber="232" unit="mm" value="0.405" valueMax="0.43" valueMin="0.38">0.380.43 mm</quantity>
, EW
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, EI 1.361.55.
</paragraph>
</subSubSection>
<subSubSection id="5BE05B8AFFB0FFAFFF62CC84FDB05C85" box="[199,613,1950,1977]" pageId="1" pageNumber="232" type="distribution">
<paragraph id="13450801FFB0FFAFFF62CC84FDB05C85" blockId="1.[151,1437,834,1977]" box="[199,613,1950,1977]" pageId="1" pageNumber="232">
<emphasis id="218ED413FFB0FFAFFF62CC84FEB45C84" bold="true" box="[199,353,1950,1976]" pageId="1" pageNumber="232">Distribution.</emphasis>
West-central
<collectingCountry id="6BED4891FFB0FFAFFE5ECC85FDB45C85" box="[507,609,1951,1977]" name="Lebanon" pageId="1" pageNumber="232">Lebanon</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="47855889FFB3FFACFF32CC22FC065C48" ID-DOI="http://doi.org/10.5281/zenodo.4411801" ID-Zenodo-Dep="4411801" httpUri="https://zenodo.org/record/4411801/files/figure.png" pageId="2" pageNumber="233" startId="2.[151,264,1848,1872]" targetBox="[163,1430,193,1812]" targetPageId="2">
<paragraph id="13450801FFB3FFACFF32CC22FC065C48" blockId="2.[151,1436,1848,1909]" pageId="2" pageNumber="233">
<emphasis id="218ED413FFB3FFACFF32CC22FEEF5C6C" bold="true" box="[151,314,1848,1872]" pageId="2" pageNumber="233">FIGURES 18.</emphasis>
<taxonomicName id="D4FA7382FFB3FFACFEE5CC22FDF95C6C" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[320,556,1848,1872]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="species" species="libanica" status="sp. nov.">
<emphasis id="218ED413FFB3FFACFEE5CC22FDF95C6C" box="[320,556,1848,1872]" italics="true" pageId="2" pageNumber="233">Paraneseuthia libanica</emphasis>
</taxonomicName>
<emphasis id="218ED413FFB3FFACFD97CC23FDBF5C6D" bold="true" box="[562,618,1849,1873]" pageId="2" pageNumber="233">
<taxonomicNameLabel id="3ABD6968FFB3FFACFD97CC23FDBF5C6D" box="[562,618,1849,1873]" pageId="2" pageNumber="233" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male. Dorsal habitus (1); head in ventral view (2); prothorax in ventral view (3); pterothorax in ventral view (4); aedeagus in ventral (5, 7) and lateral (6, 8) views.
</paragraph>
</caption>
<subSubSection id="5BE05B8AFFB3FFACFF62CCB9FD055C81" box="[199,720,1955,1981]" pageId="2" pageNumber="233" type="etymology">
<paragraph id="13450801FFB3FFACFF62CCB9FD055C81" blockId="2.[199,1436,1955,2017]" box="[199,720,1955,1981]" pageId="2" pageNumber="233">
<emphasis id="218ED413FFB3FFACFF62CCB9FE9B5C81" bold="true" box="[199,334,1955,1981]" pageId="2" pageNumber="233">Etymology.</emphasis>
After the Latinized country name.
</paragraph>
</subSubSection>
<subSubSection id="5BE05B8AFFB3FFADFF62CCDDFBEC5E69" lastPageId="3" lastPageNumber="234" pageId="2" pageNumber="233" type="discussion">
<paragraph id="13450801FFB3FFADFF62CCDDFD3F5AC9" blockId="2.[199,1436,1955,2017]" lastBlockId="3.[151,1437,151,1365]" lastPageId="3" lastPageNumber="234" pageId="2" pageNumber="233">
<emphasis id="218ED413FFB3FFACFF62CCDDFEEE5CDD" bold="true" box="[199,315,1991,2017]" pageId="2" pageNumber="233">Remarks.</emphasis>
<taxonomicName id="D4FA7382FFB3FFACFEE7CCDDFD9D5CDD" authorityName="Jałoszyński" authorityYear="2020" box="[322,584,1991,2017]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="233" phylum="Arthropoda" rank="species" species="libanica">
<emphasis id="218ED413FFB3FFACFEE7CCDDFD9D5CDD" box="[322,584,1991,2017]" italics="true" pageId="2" pageNumber="233">Paraneseuthia libanica</emphasis>
</taxonomicName>
is the second species of this genus known to occur in the Mediterranean basin, far away from all remaining congeners that inhabit sub-Himalayan
<collectingCountry id="6BED4891FFB2FFADFC10CB8DFC245B8D" box="[949,1009,151,177]" name="India" pageId="3" pageNumber="234">India</collectingCountry>
, SE Asia, Far East, New
<collectingCountry id="6BED4891FFB2FFADFAACCB8DFA885B8D" box="[1289,1373,151,177]" name="Guinea-Bissau" pageId="3" pageNumber="234">Guinea</collectingCountry>
, Australia and
<collectingCountry id="6BED4891FFB2FFADFEA2CBA1FEE75BE9" box="[263,306,187,213]" name="Fiji" pageId="3" pageNumber="234">Fiji</collectingCountry>
. Both western Palaearctic species, i.e., the Turkish
<taxonomicName id="D4FA7382FFB2FFADFCCACBA1FB1A5BE9" authority="Jaloszynski, 2015" authorityName="Jaloszynski" authorityYear="2015" box="[879,1231,186,213]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="meybohmi">
<emphasis id="218ED413FFB2FFADFCCACBA1FC285BE9" box="[879,1021,187,213]" italics="true" pageId="3" pageNumber="234">P. meybohmi</emphasis>
<bibRefCitation id="776B75F0FFB2FFADFBA0CBA1FB1A5BE9" author="Jaloszynski, P." box="[1029,1231,186,213]" pageId="3" pageNumber="234" pagination="287 - 295" refId="ref5118" refString="Jaloszynski, P. (2015) Paraneseuthia Franz in the Indian Himalayan Region and Western Palaearctic (Coleoptera, Staphylini- dae, Scydmaeninae). Zootaxa, 3948 (2), 287 - 295. https: // doi. org / 10.11646 / zootaxa. 3948.2.9" type="journal article" year="2015">Jałoszyński, 2015</bibRefCitation>
</taxonomicName>
, and the Lebanese
<taxonomicName id="D4FA7382FFB2FFADFF32CBC5FED85BC5" authorityName="Jałoszyński" authorityYear="2020" box="[151,269,223,249]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="libanica">
<emphasis id="218ED413FFB2FFADFF32CBC5FED85BC5" box="[151,269,223,249]" italics="true" pageId="3" pageNumber="234">P. libanica</emphasis>
</taxonomicName>
, have small (
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), relatively slender and convex adults, and share the same form of the aedeagus. The latter is of a unique shape, not known in any other group of species within this genus. The diaphragm, normally situated in the sub-basal or submedian region of the median lobe and parallel to its long axis, in
<taxonomicName id="D4FA7382FFB2FFADFB61CA3DFA875A7D" authorityName="Jaloszynski" authorityYear="2015" box="[1220,1362,295,321]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="meybohmi">
<emphasis id="218ED413FFB2FFADFB61CA3DFA875A7D" box="[1220,1362,295,321]" italics="true" pageId="3" pageNumber="234">P. meybohmi</emphasis>
</taxonomicName>
and
<taxonomicName id="D4FA7382FFB2FFADFA2DCA3DFF265A59" authorityName="Jałoszyński" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="libanica">
<emphasis id="218ED413FFB2FFADFA2DCA3DFF265A59" italics="true" pageId="3" pageNumber="234">P. libanica</emphasis>
</taxonomicName>
is shifted toward the subapical region and situated on a step-wise platform oblique or almost perpendicular in relation to the long axis of the median lobe. Aedeagi of both species also have asymmetrical apices, are curved in lateral view, and have parameres lacking apical setae.
<taxonomicName id="D4FA7382FFB2FFADFCAFCA89FBC55A91" authorityName="Jałoszyński" authorityYear="2020" box="[778,1040,403,429]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="libanica">
<emphasis id="218ED413FFB2FFADFCAFCA89FBC55A91" box="[778,1040,403,429]" italics="true" pageId="3" pageNumber="234">Paraneseuthia libanica</emphasis>
</taxonomicName>
has smaller eyes than
<taxonomicName id="D4FA7382FFB2FFADFAADCA89FA405A91" authorityName="Jaloszynski" authorityYear="2015" box="[1288,1429,403,429]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="meybohmi">
<emphasis id="218ED413FFB2FFADFAADCA89FA405A91" box="[1288,1429,403,429]" italics="true" pageId="3" pageNumber="234">P. meybohmi</emphasis>
</taxonomicName>
, a different shape of the pronotum, and a clearly different shape of the apical region of the aedeagus (with a pair of elongate lateral projections, lacking in
<taxonomicName id="D4FA7382FFB2FFADFDEECAC1FD0C5AC9" authorityName="Jaloszynski" authorityYear="2015" box="[587,729,475,501]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="meybohmi">
<emphasis id="218ED413FFB2FFADFDEECAC1FD0C5AC9" box="[587,729,475,501]" italics="true" pageId="3" pageNumber="234">P. meybohmi</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="13450801FFB2FFADFF62CAE5FBEC5E69" blockId="3.[151,1437,151,1365]" pageId="3" pageNumber="234">
Morphological structures of the head and thorax were compared in detail for Far Eastern (Japanese and Russian) and SE Asian groups of
<taxonomicName id="D4FA7382FFB2FFADFE7AC939FD575901" authorityName="Franz" authorityYear="1986" box="[479,642,547,573]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="genus">
<emphasis id="218ED413FFB2FFADFE7AC939FD575901" box="[479,642,547,573]" italics="true" pageId="3" pageNumber="234">Paraneseuthia</emphasis>
</taxonomicName>
that markedly differ in body forms and genital structures. These groups were found to include congeneric species, as their cephalic and ventral thoracic character states were demonstrated to be the same (
<bibRefCitation id="776B75F0FFB2FFADFEEEC971FDC659B9" author="Jaloszynski, P." box="[331,531,619,645]" pageId="3" pageNumber="234" pagination="181 - 193" refId="ref4865" refString="Jaloszynski, P. (2010) Paraneseuthia Franz, 1986 of Australo-Oriental Region, with notes on Palearctic species (Coleoptera: Staphylinidae: Scydmaeninae). Annales Zoologici, 60 (2), 181 - 193. https: // doi. org / 10.3161 / 000345410 x 516830" type="journal article" year="2010">Jałoszyński 2010</bibRefCitation>
). A similar morphological study was carried out for the sub-Himalayan
<taxonomicName id="D4FA7382FFB2FFADFAE5C971FEB45995" authority="Jaloszynski, 2015" authorityName="Jaloszynski" authorityYear="2015" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="loebli">
<emphasis id="218ED413FFB2FFADFAE5C971FA4959B9" box="[1344,1436,619,645]" italics="true" pageId="3" pageNumber="234">P. loebli</emphasis>
<bibRefCitation id="776B75F0FFB2FFADFF32C995FEB45995" author="Jaloszynski, P." box="[151,353,654,681]" pageId="3" pageNumber="234" pagination="287 - 295" refId="ref5118" refString="Jaloszynski, P. (2015) Paraneseuthia Franz in the Indian Himalayan Region and Western Palaearctic (Coleoptera, Staphylini- dae, Scydmaeninae). Zootaxa, 3948 (2), 287 - 295. https: // doi. org / 10.11646 / zootaxa. 3948.2.9" type="journal article" year="2015">Jałoszyński, 2015</bibRefCitation>
</taxonomicName>
, with similar conclusions (
<bibRefCitation id="776B75F0FFB2FFADFD28C995FC865995" author="Jaloszynski, P." box="[653,851,654,681]" pageId="3" pageNumber="234" pagination="287 - 295" refId="ref5118" refString="Jaloszynski, P. (2015) Paraneseuthia Franz in the Indian Himalayan Region and Western Palaearctic (Coleoptera, Staphylini- dae, Scydmaeninae). Zootaxa, 3948 (2), 287 - 295. https: // doi. org / 10.11646 / zootaxa. 3948.2.9" type="journal article" year="2015">Jałoszyński 2015</bibRefCitation>
).
<emphasis id="218ED413FFB2FFADFCC3C995FB5F5995" box="[870,1162,655,681]" italics="true" pageId="3" pageNumber="234">
<taxonomicName id="D4FA7382FFB2FFADFCC3C995FB535995" authorityName="Jaloszynski" authorityYear="2015" box="[870,1158,655,681]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="meybohmi">Paraneseuthia meybohmi</taxonomicName>
,
</emphasis>
the first known Western Palaearctic species, was not available in a sufficient number of specimens to allow for disarticulations. Among specimens of
<taxonomicName id="D4FA7382FFB2FFADFE94C9CDFE7D59CD" authorityName="Jałoszyński" authorityYear="2020" box="[305,424,727,753]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="libanica">
<emphasis id="218ED413FFB2FFADFE94C9CDFE7D59CD" box="[305,424,727,753]" italics="true" pageId="3" pageNumber="234">P. libanica</emphasis>
</taxonomicName>
, two males deposited at MHNG were found already disarticulated and mounted in euparal by the collector of the
<typeStatus id="CC41B6A3FFB2FFADFE3DC9E1FE1D5829" box="[408,456,763,789]" pageId="3" pageNumber="234">type</typeStatus>
series, Claude Besuchet. Ventral structures of the head, pro- and pterothorax of one of these specimens are illustrated in
<figureCitation id="8BC11484FFB2FFADFDB7C805FDAD5805" box="[530,632,799,825]" captionStart="FIGURES 18" captionStartId="2.[151,264,1848,1872]" captionTargetBox="[163,1430,193,1812]" captionTargetId="figure-19@2.[151,1436,179,1823]" captionTargetPageId="2" captionText="FIGURES 18. Paraneseuthia libanica sp. n., male. Dorsal habitus (1); head in ventral view (2); prothorax in ventral view (3); pterothorax in ventral view (4); aedeagus in ventral (5, 7) and lateral (6, 8) views." figureDoi="http://doi.org/10.5281/zenodo.4411801" httpUri="https://zenodo.org/record/4411801/files/figure.png" pageId="3" pageNumber="234">Figs 24</figureCitation>
. Apart from the unusually slender body of
<taxonomicName id="D4FA7382FFB2FFADFBFEC805FB065805" authorityName="Jałoszyński" authorityYear="2020" box="[1115,1235,799,825]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="libanica">
<emphasis id="218ED413FFB2FFADFBFEC805FB065805" box="[1115,1235,799,825]" italics="true" pageId="3" pageNumber="234">P. libanica</emphasis>
</taxonomicName>
, this species does not show any important differences in relation to its previously studied relatives. The only major structure that was found variable within the genus is the pair of procoxal rests on the anterior region of the mesoventrite. In Far Eastern (Japanese) species, the rests are not demarcated posteriorly. In SE Asian species each procoxal rest has a sharply marked posterior carina, which originates at the mesoventral intercoxal process, near its median region, runs laterad, and is arcuate, connecting to the mesal corner of the mesanepisternum. In the sub-Himalayan
<taxonomicName id="D4FA7382FFB2FFADFAB5C8C9FAB958D1" authorityName="Jaloszynski" authorityYear="2015" box="[1296,1388,979,1005]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="loebli">
<emphasis id="218ED413FFB2FFADFAB5C8C9FAB958D1" box="[1296,1388,979,1005]" italics="true" pageId="3" pageNumber="234">P. loebli</emphasis>
</taxonomicName>
, the procoxal rests are demarcated posteriorly by a transverse carina parallel to the anterior margin of mesoventrite, the carina reaches only to middle between the mesoventral intercoxal process and the mesanepisternum; and the lateral portion of mesoventrite, between the procoxal rest and mesanepisternum + mesepimeron is occupied by a large pit filled with setae. In the Mediterranean
<taxonomicName id="D4FA7382FFB2FFADFDE0CF79FD6E5F41" authorityName="Jałoszyński" authorityYear="2020" box="[581,699,1123,1149]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="libanica">
<emphasis id="218ED413FFB2FFADFDE0CF79FD6E5F41" box="[581,699,1123,1149]" italics="true" pageId="3" pageNumber="234">P. libanica</emphasis>
</taxonomicName>
, the procoxal rests on the mesoventrite are posteriorly demarcated by a sharp carina which extends laterad nearly to mesanapisterna (it is obliterated shortly before reaching them), the carina is parallel in relation to the anterior mesoventral margin, and the mesoventrite lacks lateral pits. These differences in mesothoracic structures are not sufficient to divide the genus into subgenera, especially that
<taxonomicName id="D4FA7382FFB2FFADFB5CCFD5FA495FD5" authorityName="Franz" authorityYear="1986" box="[1273,1436,1231,1257]" class="Insecta" family="Staphylinidae" genus="Paraneseuthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="234" phylum="Arthropoda" rank="genus">
<emphasis id="218ED413FFB2FFADFB5CCFD5FA495FD5" box="[1273,1436,1231,1257]" italics="true" pageId="3" pageNumber="234">Paraneseuthia</emphasis>
</taxonomicName>
is the most diverse genus of Eutheiini in terms of body shapes, male secondary dimorphic structures, and especially the aedeagus. Certainly the Mediterranean species form a distinct group, differing from all other congeners in the structure of the aedeagus, but not in any other, taxonomically important characters.
</paragraph>
</subSubSection>
</treatment>
</document>