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<document id="6EF8F7E327DD5F8FF56C5232BBFE2DB6" ID-CLB-Dataset="29524" ID-DOI="10.11646/zootaxa.4446.1.4" ID-GBIF-Dataset="d35cedde-4442-4a14-be2d-d5213a12ab29" ID-ISSN="1175-5326" ID-Zenodo-Dep="1437384" ID-ZooBank="E5ABBD4C-C643-4091-9AD4-D9D4028A218A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538121156330" checkinUser="plazi" docAuthor="Ban, Teruaki" docDate="2018" docId="9B474B53FFE83D66FF01B7A8FB7B50C4" docLanguage="en" docName="zootaxa.4446.1.4.pdf" docOrigin="Zootaxa 4446 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sadoletus melasmus Ban 2018, sp. nov." docType="treatment" docVersion="8" lastPageNumber="60" masterDocId="677E332BFFEE3D6FFF96B264FFD05011" masterDocTitle="The lygaeoid bug family Heterogastridae (Hemiptera: Heteroptera: Lygaeoidea) from Laos and Thailand, with descriptions of three new species" masterLastPageNumber="67" masterPageNumber="51" pageNumber="57" updateTime="1698555679930" updateUser="ExternalLinkService">
<mods:mods id="05D2C42445185723E04A4978D2E429F9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6E815CD05729D908D34E5143EC25C287">
<mods:title id="6CCA36C2FF0631B2AE9C9D5CD7587241">The lygaeoid bug family Heterogastridae (Hemiptera: Heteroptera: Lygaeoidea) from Laos and Thailand, with descriptions of three new species</mods:title>
</mods:titleInfo>
<mods:name id="3EE0499134C5F1A0B83F3C2CE3D32EEB" type="personal">
<mods:role id="D7AAF0F65064EDDDB5363C82C26D54BB">
<mods:roleTerm id="D0CA940284C055DE83D2099DB7730B10">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D496C580D53AC7C8AD1B06E4E5986FC2">Ban, Teruaki</mods:namePart>
</mods:name>
<mods:typeOfResource id="41DDAA1532FFA585A3B12009F4804090">text</mods:typeOfResource>
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<mods:titleInfo id="2AD376B3244F1D43A8CA04391478CF96">
<mods:title id="C49C727962F74336A65E7729A8A08228">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="60B34B08224C738EEDE6AC9B71283023">
<mods:date id="0557D3A95CFB6F7D85C51721FA31954B">2018</mods:date>
<mods:detail id="9770FE3BC7D3EC8D42E58D3B2ED7D6A1" type="pubDate">
<mods:number id="49E2F86058E4A913CE538AB0C2C7B7DF">2018-07-16</mods:number>
</mods:detail>
<mods:detail id="2AFAA485BB126DFA31BACD5A393C34AC" type="volume">
<mods:number id="5F361F9D0A575EAE0269EA762F3FF758">4446</mods:number>
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<mods:number id="566F71A7BCC22BA8A390C9756595B8C5">1</mods:number>
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<mods:end id="28E92047E1C4BC4E8FF2EB132CED4F05">67</mods:end>
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<mods:classification id="B630DC2181617BDF08D582A92F5A7CBA">journal article</mods:classification>
<mods:identifier id="71E01CF297EAA5F1C476B5FD6C65430E" type="CLB-Dataset">29524</mods:identifier>
<mods:identifier id="EE2AB0709584B58C56C52A315158BAFD" type="DOI">10.11646/zootaxa.4446.1.4</mods:identifier>
<mods:identifier id="3291B1B51073B42676DCF9CFC4AC61FD" type="GBIF-Dataset">d35cedde-4442-4a14-be2d-d5213a12ab29</mods:identifier>
<mods:identifier id="C0E115A2A39DBBB4EBA1B175DDCBB333" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="94D54227D6C127082B473098F2E11AB1" type="Zenodo-Dep">1437384</mods:identifier>
<mods:identifier id="C4772C5EDB8B14124C59865A01B32FA2" type="ZooBank">E5ABBD4C-C643-4091-9AD4-D9D4028A218A</mods:identifier>
</mods:mods>
<treatment id="9B474B53FFE83D66FF01B7A8FB7B50C4" ID-DOI="http://doi.org/10.5281/zenodo.3507637" ID-GBIF-Taxon="148403380" ID-Zenodo-Dep="3507637" LSID="urn:lsid:plazi:treatment:9B474B53FFE83D66FF01B7A8FB7B50C4" httpUri="http://treatment.plazi.org/id/9B474B53FFE83D66FF01B7A8FB7B50C4" lastPageId="9" lastPageNumber="60" pageId="6" pageNumber="57">
<subSubSection id="5BF4A9CEFFE83D69FF01B7A8FE6A5616" pageId="6" pageNumber="57" type="nomenclature">
<paragraph id="1351FA45FFE83D69FF01B7A8FE2155F7" blockId="6.[151,497,1484,1543]" box="[151,497,1484,1510]" pageId="6" pageNumber="57">
<heading id="48194D29FFE83D69FF01B7A8FE2155F7" bold="true" box="[151,497,1484,1510]" fontSize="11" level="1" pageId="6" pageNumber="57" reason="1">
<emphasis id="219A2657FFE83D69FF01B7A8FE2155F7" bold="true" box="[151,497,1484,1510]" pageId="6" pageNumber="57">
<taxonomicName id="D4EE81C6FFE83D69FF01B7A8FE5C55F7" ID-CoL="8QP8R" authority="Ban, 2018" authorityName="Ban" authorityYear="2018" box="[151,396,1484,1510]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="melasmus" status="sp. nov.">
<emphasis id="219A2657FFE83D69FF01B7A8FE5C55F7" bold="true" box="[151,396,1484,1510]" italics="true" pageId="6" pageNumber="57">Sadoletus melasmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3AA99B2CFFE83D69FE05B7A8FE2155F7" box="[403,497,1484,1510]" pageId="6" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="1351FA45FFE83D69FF01B78AFE6A5616" blockId="6.[151,497,1484,1543]" box="[151,442,1518,1543]" pageId="6" pageNumber="57">
(
<figureCitation id="8BD5E6C0FFE83D69FF09B78AFEDA5616" box="[159,266,1518,1543]" captionStart="FIGURES 6–9" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 6–9. Habitus of Heterogastridae species from Laos and Thailand. 6–7, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 8–9, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="6" pageNumber="57">Figs. 8–9</figureCitation>
,
<figureCitation id="8BD5E6C0FFE83D69FE80B78AFEE45616" box="[278,308,1518,1543]" captionStart="FIGURES 14–16" captionStartId="11.[151,264,768,791]" captionTargetBox="[199,1371,188,745]" captionTargetId="figure@11.[181,1405,183,761]" captionTargetPageId="11" captionText="FIGURES 14–16. Females of each new Sadoletus species, ventral view. 14, Sadoletus laosensis sp. nov.; 15, S. melasmus sp. nov.; 16, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437392/files/figure.png" pageId="6" pageNumber="57">15</figureCitation>
,
<figureCitation id="8BD5E6C0FFE83D69FED6B78BFE8E5616" box="[320,350,1519,1543]" captionStart="FIGURES 17–19" captionStartId="11.[151,264,1221,1244]" captionTargetBox="[173,1370,892,1151]" captionTargetId="figure@11.[151,1436,861,1218]" captionTargetPageId="11" captionText="FIGURES 17–19. Abdominal sternite VI and ovipositer of each new Sadoletus species, ventral view. 17, Sadoletus laosensis sp. nov.; 18, S. melasmus sp. nov.; 19, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437394/files/figure.png" pageId="6" pageNumber="57">18</figureCitation>
,
<figureCitation id="8BD5E6C0FFE83D69FEFCB78AFE615616" box="[362,433,1518,1543]" captionStart="FIGURES 25–29" captionStartId="13.[151,264,1961,1984]" captionTargetBox="[152,1431,355,1907]" captionTargetId="figure@13.[151,1436,318,1925]" captionTargetPageId="13" captionText="FIGURES 25–29. Sadoletus melasmus sp. nov. 25, Pygophore, dorsal view (setae omitted from left half); 26, left paramere, dorsal view; 27–28, phallus (endosoma not everted), ventral (27) and lateral (28) views; 29, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437398/files/figure.png" pageId="6" pageNumber="57">25–29</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5BF4A9CEFFE83D69FF01B452FDD756AA" pageId="6" pageNumber="57" type="materials_examined">
<paragraph id="1351FA45FFE83D69FF01B452FDD756AA" blockId="6.[151,1437,1590,2011]" pageId="6" pageNumber="57">
<materialsCitation id="A386F018FFE83D69FF01B452FD7C5662" ID-GBIF-Occurrence="1913858566" collectingDate="2015-03-17" country="Thailand" elevation="502" location="Teruaki Ban" municipality="Rayong Prov." pageId="6" pageNumber="57" specimenCount="1" specimenCount-female="1" stateProvince="Rayong" typeStatus="holotype">
<emphasis id="219A2657FFE83D69FF01B452FE84565E" bold="true" box="[151,340,1590,1615]" pageId="6" pageNumber="57">
<typeStatus id="CC5544E7FFE83D69FF01B452FF02565E" box="[151,210,1590,1615]" pageId="6" pageNumber="57">Type</typeStatus>
Materials.
</emphasis>
<typeStatus id="CC5544E7FFE83D69FECAB452FE19565E" box="[348,457,1590,1615]" pageId="6" pageNumber="57" type="holotype">Holotype</typeStatus>
:
<specimenCount id="05E831CCFFE83D69FE43B453FE3A565F" box="[469,490,1591,1614]" pageId="6" pageNumber="57" type="female">♀</specimenCount>
,
<emphasis id="219A2657FFE83D69FE6CB452FD4A565E" bold="true" box="[506,666,1590,1615]" pageId="6" pageNumber="57">
<collectingCountry id="6BF9BAD5FFE83D69FE6CB452FD45565E" box="[506,661,1590,1615]" name="Thailand" pageId="6" pageNumber="57">THAILAND</collectingCountry>
:
</emphasis>
12°40'14“–30“N, 101°24'50“–52”E, alt.
<elevation id="98C31D76FFE83D69FBF9B452FB21565F" box="[1135,1265,1590,1615]" metricMagnitude="2" metricUnit="m" metricValue="5.0200000000000005" metricValueMax="5.5200000000000005" metricValueMin="4.5200000000000005" pageId="6" pageNumber="57" unit="m" value="502.0" valueMax="552.0" valueMin="452.0">
<quantity id="D41657A0FFE83D69FBF9B452FB21565F" box="[1135,1265,1590,1615]" metricMagnitude="2" metricUnit="m" metricValue="5.0200000000000005" metricValueMax="5.5200000000000005" metricValueMin="4.5200000000000005" pageId="6" pageNumber="57" unit="m" value="502.0" valueMax="552.0" valueMin="452.0">452–552 m</quantity>
</elevation>
, Near
<collectingRegion id="D12A34A7FFE83D69FAA8B453FA48565E" box="[1342,1432,1591,1615]" country="Thailand" name="Rayong" pageId="6" pageNumber="57">Rayong</collectingRegion>
,
<collectingMunicipality id="F335603FFFE83D69FF01B43FFEE35663" box="[151,307,1627,1651]" pageId="6" pageNumber="57">Rayong Prov.</collectingMunicipality>
,
<collectingDate id="7714256DFFE83D69FEA8B43EFE6E5662" box="[318,446,1626,1651]" pageId="6" pageNumber="57" value="2015-03-17">
<date id="6750DC85FFE83D69FEA8B43EFE6E5662" box="[318,446,1626,1651]" pageId="6" pageNumber="57" value="2015-03-17">17.III.2015</date>
</collectingDate>
,
<location id="1631AC9EFFE83D69FE5DB43EFD855663" LSID="urn:lsid:plazi:treatment:9B474B53FFE83D66FF01B7A8FB7B50C4:1631AC9EFFE83D69FE5DB43EFD855663" box="[459,597,1626,1651]" country="Thailand" municipality="Rayong Prov." name="Teruaki Ban" pageId="6" pageNumber="57" stateProvince="Rayong">Teruaki Ban</location>
(TUA)
</materialsCitation>
.
<materialsCitation id="A386F018FFE83D69FD21B43FFB545686" ID-GBIF-Occurrence="1913858575" collectingDate="2015-03-14" country="Thailand" elevation="66" latitude="12.820694" location="Teruaki Ban" longLatPrecision="1" longitude="102.127945" municipality="Krathing Country Resort" pageId="6" pageNumber="57" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Chanthaburi" typeStatus="paratype">
<typeStatus id="CC5544E7FFE83D69FD21B43FFCF95662" box="[695,809,1627,1651]" pageId="6" pageNumber="57" type="paratype">Paratypes</typeStatus>
:
<emphasis id="219A2657FFE83D69FCA2B43EFC045662" bold="true" box="[820,980,1626,1651]" pageId="6" pageNumber="57">
<collectingCountry id="6BF9BAD5FFE83D69FCA2B43EFC1F5662" box="[820,975,1626,1651]" name="Thailand" pageId="6" pageNumber="57">THAILAND</collectingCountry>
:
</emphasis>
<specimenCount id="05E831CCFFE83D69FC4AB43FFBD75663" box="[988,1031,1627,1651]" pageId="6" pageNumber="57" type="male">1 ♂</specimenCount>
<specimenCount id="05E831CCFFE83D69FB98B43FFBE95663" box="[1038,1081,1627,1651]" pageId="6" pageNumber="57" type="female">2 ♀</specimenCount>
,
<geoCoordinate id="76DA9C82FFE83D69FBDEB43EFB345662" box="[1096,1252,1626,1651]" direction="north" orientation="latitude" pageId="6" pageNumber="57" precision="1" value="12.820694">12°49'14.5“N</geoCoordinate>
,
<geoCoordinate id="76DA9C82FFE83D69FB66B43EFA475662" box="[1264,1431,1626,1651]" direction="east" orientation="longitude" pageId="6" pageNumber="57" precision="1" value="102.127945">102°07'40.6”E</geoCoordinate>
, alt.
<elevation id="98C31D76FFE83D69FF55B41AFF2F5687" box="[195,255,1662,1687]" metricMagnitude="1" metricUnit="m" metricValue="6.6" pageId="6" pageNumber="57" unit="m" value="66.0">
<quantity id="D41657A0FFE83D69FF55B41AFF2F5687" box="[195,255,1662,1687]" metricMagnitude="1" metricUnit="m" metricValue="6.6" pageId="6" pageNumber="57" unit="m" value="66.0">66 m</quantity>
</elevation>
,
<collectingMunicipality id="F335603FFFE83D69FE9BB41AFDF95687" box="[269,553,1662,1687]" pageId="6" pageNumber="57">Krathing Country Resort</collectingMunicipality>
,
<collectingRegion id="D12A34A7FFE83D69FDA0B41AFD105686" box="[566,704,1662,1687]" country="Thailand" name="Chanthaburi" pageId="6" pageNumber="57">Chanthaburi</collectingRegion>
Prov.,
<collectingDate id="7714256DFFE83D69FC85B41AFC435686" box="[787,915,1662,1687]" pageId="6" pageNumber="57" value="2015-03-14">
<date id="6750DC85FFE83D69FC85B41AFC435686" box="[787,915,1662,1687]" pageId="6" pageNumber="57" value="2015-03-14">14.III.2015</date>
</collectingDate>
,
<location id="1631AC9EFFE83D69FC37B41AFBFC5687" LSID="urn:lsid:plazi:treatment:9B474B53FFE83D66FF01B7A8FB7B50C4:1631AC9EFFE83D69FC37B41AFBFC5687" box="[929,1068,1662,1687]" country="Thailand" latitude="12.820694" longLatPrecision="1" longitude="102.127945" municipality="Krathing Country Resort" name="Teruaki Ban" pageId="6" pageNumber="57" stateProvince="Chanthaburi">Teruaki Ban</location>
(TUA)
</materialsCitation>
;
<materialsCitation id="A386F018FFE83D69FB07B41BFDD756AA" ID-GBIF-Occurrence="1913858577" collectingDate="2015-03-15" country="Thailand" elevation="66" latitude="12.820694" location="Teruaki Ban" longLatPrecision="1" longitude="102.127945" municipality="Krathing Country Resort" pageId="6" pageNumber="57" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Chanthaburi" typeStatus="paratype">
<specimenCount id="05E831CCFFE83D69FB07B41BFB6D5687" box="[1169,1213,1663,1687]" pageId="6" pageNumber="57" type="male">1 ♂</specimenCount>
<specimenCount id="05E831CCFFE83D69FB53B41BFB215687" box="[1221,1265,1663,1687]" pageId="6" pageNumber="57" type="female">2 ♀</specimenCount>
, same locality,
<collectingDate id="7714256DFFE83D69FF01B4C6FEC856AA" box="[151,280,1698,1723]" pageId="6" pageNumber="57" value="2015-03-15">
<date id="6750DC85FFE83D69FF01B4C6FEC856AA" box="[151,280,1698,1723]" pageId="6" pageNumber="57" value="2015-03-15">15.III.2015</date>
</collectingDate>
, Teruaki Ban (TUA).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="5BF4A9CEFFE83D69FF51B4A2FCAD57CA" pageId="6" pageNumber="57" type="diagnosis">
<paragraph id="1351FA45FFE83D69FF51B4A2FCAD57CA" blockId="6.[151,1437,1590,2011]" pageId="6" pageNumber="57">
<emphasis id="219A2657FFE83D69FF51B4A2FE9156CE" bold="true" box="[199,321,1734,1759]" pageId="6" pageNumber="57">Diagnosis.</emphasis>
Recognized among congeners of
<taxonomicName id="D4EE81C6FFE83D69FD47B4A2FCEE56CF" authorityName="Distant" authorityYear="1903" box="[721,830,1734,1758]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="219A2657FFE83D69FD47B4A2FCEE56CF" box="[721,830,1734,1758]" italics="true" pageId="6" pageNumber="57">Sadoletus</emphasis>
</taxonomicName>
by the following characters: Head, anterior lobe of pronotum, basal half of scutellum and apical third of corium black (
<figureCitation id="8BD5E6C0FFE83D69FC07B48EFC2E5712" box="[913,1022,1770,1795]" captionStart="FIGURES 6–9" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 6–9. Habitus of Heterogastridae species from Laos and Thailand. 6–7, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 8–9, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="6" pageNumber="57">Figs. 8, 9</figureCitation>
); labium exceeding posterior margin of prothorax (
<figureCitation id="8BD5E6C0FFE83D69FEA6B56AFE535736" box="[304,387,1806,1831]" captionStart="FIGURES 14–16" captionStartId="11.[151,264,768,791]" captionTargetBox="[199,1371,188,745]" captionTargetId="figure@11.[181,1405,183,761]" captionTargetPageId="11" captionText="FIGURES 14–16. Females of each new Sadoletus species, ventral view. 14, Sadoletus laosensis sp. nov.; 15, S. melasmus sp. nov.; 16, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437392/files/figure.png" pageId="6" pageNumber="57">Fig. 15</figureCitation>
); posterior lobe of pronotum pale yellow, with submedian longitudinal band and 2 dark brown spots at lateral angle (
<figureCitation id="8BD5E6C0FFE83D69FE06B556FE2D575A" box="[400,509,1842,1867]" captionStart="FIGURES 6–9" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 6–9. Habitus of Heterogastridae species from Laos and Thailand. 6–7, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 8–9, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="6" pageNumber="57">Figs. 8, 9</figureCitation>
); corium white in basal two-thirds and blackened in apical third, with black spots around inner angle of corium and claval commissure (
<figureCitation id="8BD5E6C0FFE83D69FD67B532FC8B577E" box="[753,859,1878,1903]" captionStart="FIGURES 6–9" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 6–9. Habitus of Heterogastridae species from Laos and Thailand. 6–7, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 8–9, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="6" pageNumber="57">Figs. 8, 9</figureCitation>
); clavus white; femora and tibiae yellow and white, with brown or yellow annulations, metafemur and metatibia with brown annulations (
<figureCitation id="8BD5E6C0FFE83D69FB02B51EFADA5782" box="[1172,1290,1914,1939]" captionStart="FIGURES 6–9" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 6–9. Habitus of Heterogastridae species from Laos and Thailand. 6–7, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 8–9, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="6" pageNumber="57">Figs. 8, 9</figureCitation>
); abdominal sternum black; ovipositor strongly curved laterally (
<figureCitation id="8BD5E6C0FFE83D69FD7DB5FAFC9057A6" box="[747,832,1950,1975]" captionStart="FIGURES 17–19" captionStartId="11.[151,264,1221,1244]" captionTargetBox="[173,1370,892,1151]" captionTargetId="figure@11.[151,1436,861,1218]" captionTargetPageId="11" captionText="FIGURES 17–19. Abdominal sternite VI and ovipositer of each new Sadoletus species, ventral view. 17, Sadoletus laosensis sp. nov.; 18, S. melasmus sp. nov.; 19, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437394/files/figure.png" pageId="6" pageNumber="57">Fig. 18</figureCitation>
); posterior margin of pygophore and cuplike sclerite fused together (
<figureCitation id="8BD5E6C0FFE83D69FED3B5A6FE4957CA" box="[325,409,1986,2011]" captionStart="FIGURES 25–29" captionStartId="13.[151,264,1961,1984]" captionTargetBox="[152,1431,355,1907]" captionTargetId="figure@13.[151,1436,318,1925]" captionTargetPageId="13" captionText="FIGURES 25–29. Sadoletus melasmus sp. nov. 25, Pygophore, dorsal view (setae omitted from left half); 26, left paramere, dorsal view; 27–28, phallus (endosoma not everted), ventral (27) and lateral (28) views; 29, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437398/files/figure.png" pageId="6" pageNumber="57">Fig. 25</figureCitation>
); paramere sickle-shaped at apex (
<figureCitation id="8BD5E6C0FFE83D69FC8AB5A6FCA057CA" box="[796,880,1986,2011]" captionStart="FIGURES 25–29" captionStartId="13.[151,264,1961,1984]" captionTargetBox="[152,1431,355,1907]" captionTargetId="figure@13.[151,1436,318,1925]" captionTargetPageId="13" captionText="FIGURES 25–29. Sadoletus melasmus sp. nov. 25, Pygophore, dorsal view (setae omitted from left half); 26, left paramere, dorsal view; 27–28, phallus (endosoma not everted), ventral (27) and lateral (28) views; 29, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437398/files/figure.png" pageId="6" pageNumber="57">Fig. 26</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5BF4A9CEFFE93D68FF51B2F3FD8556AC" pageId="7" pageNumber="58" type="description">
<paragraph id="1351FA45FFE93D68FF51B2F3FC33522C" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="58">
<emphasis id="219A2657FFE93D68FF51B2F3FE8950A1" bold="true" box="[199,345,151,176]" pageId="7" pageNumber="58">Description.</emphasis>
Female (
<typeStatus id="CC5544E7FFE93D68FE50B2F3FDFF50A1" box="[454,559,151,176]" pageId="7" pageNumber="58" type="holotype">holotype</typeStatus>
): Venter almost black. Short decumbent setae on body silvery. Punctures on pronotum, scutellum, and corium dark brown. Head (
<figureCitation id="8BD5E6C0FFE93D68FC80B2D8FC8F50C5" box="[790,863,188,213]" captionStart="FIGURES 6–9" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 6–9. Habitus of Heterogastridae species from Laos and Thailand. 6–7, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 8–9, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="7" pageNumber="58">Fig. 9</figureCitation>
) black. Clypeus pale yellow. Antenna generally brownish yellow, apical three quarters of segment IV tinged with dark brown. Labium dark brown; segment III and basal three quarters of segment IV brownish yellow. Pronotum (
<figureCitation id="8BD5E6C0FFE93D68FCE0B360FC6C510D" box="[886,956,260,285]" captionStart="FIGURES 6–9" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 6–9. Habitus of Heterogastridae species from Laos and Thailand. 6–7, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 8–9, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="7" pageNumber="58">Fig. 9</figureCitation>
) black in anterior lobe and pale brownish yellow in posterior lobe, posterior lobe with longitudinal pale stripe along meson and 2 dark brown longitudinal band and 2 dark brown markings at around lateral angle. Venter of mesothorax and metathorax black; peritreme brown. Scutellum (
<figureCitation id="8BD5E6C0FFE93D68FEEFB30BFE135199" box="[377,451,367,392]" captionStart="FIGURES 6–9" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 6–9. Habitus of Heterogastridae species from Laos and Thailand. 6–7, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 8–9, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="7" pageNumber="58">Fig. 9</figureCitation>
) black in basal half and brown in apical half. Apical half of scutellum with pale longitudinal stripe along meson. Corium (
<figureCitation id="8BD5E6C0FFE93D68FD16B3F0FD1751BD" box="[640,711,404,429]" captionStart="FIGURES 6–9" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 6–9. Habitus of Heterogastridae species from Laos and Thailand. 6–7, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 8–9, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="7" pageNumber="58">Fig. 9</figureCitation>
) white in basal two-thirds and black in remaining part; black circular spot at inner angle; clavus white, with longitudinal black stripe along claval commissure; hemelytral membrane hyaline, with oval brownish spot at middle. Coxae dark brown; trochanters, femora and tibiae pale yellow; metafemur and metatibia with brown annulations; tarsomere I pale yellow, tarsomeres II and III pale brown. Abdomen (
<figureCitation id="8BD5E6C0FFE93D68FEFCB040FE6B522C" box="[362,443,548,573]" captionStart="FIGURES 14–16" captionStartId="11.[151,264,768,791]" captionTargetBox="[199,1371,188,745]" captionTargetId="figure@11.[181,1405,183,761]" captionTargetPageId="11" captionText="FIGURES 14–16. Females of each new Sadoletus species, ventral view. 14, Sadoletus laosensis sp. nov.; 15, S. melasmus sp. nov.; 16, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437392/files/figure.png" pageId="7" pageNumber="58">Fig. 15</figureCitation>
) black; each laterotergite entirely reddish brown.
</paragraph>
<paragraph id="1351FA45FFE93D68FF51B023FA4652E1" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="58">
Body (
<figureCitation id="8BD5E6C0FFE93D68FE82B023FE885271" box="[276,344,583,608]" captionStart="FIGURES 6–9" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 6–9. Habitus of Heterogastridae species from Laos and Thailand. 6–7, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 8–9, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="7" pageNumber="58">Fig. 9</figureCitation>
) 3.1 times as long as width across humeri. Head (
<figureCitation id="8BD5E6C0FFE93D68FC1BB023FC015271" box="[909,977,583,608]" captionStart="FIGURES 6–9" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 6–9. Habitus of Heterogastridae species from Laos and Thailand. 6–7, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 8–9, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="7" pageNumber="58">Fig. 9</figureCitation>
) 1.7 times as wide as its length, covered with both rough and fine punctures; tylus tumid upwards; clypeus weakly projecting anteriad; juga with decumbent setae. Eyes occupying nearly half area of head in lateral view, not reaching anterior margin of pronotum. Antennae covered with suberect setae; proportional lengths of segments I to IV 1.0: 1.7: 1.6: 2.4. Labium slightly exceeding posterior margin of prothorax, not reaching mesocoxae; proportional lengths of segments I to IV 1.4: 1.4: 1.1: 1.0.
</paragraph>
<paragraph id="1351FA45FFE93D68FF51B099FC5153FD" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="58">
Pronotum (
<figureCitation id="8BD5E6C0FFE93D68FED3B098FE595305" box="[325,393,764,789]" captionStart="FIGURES 6–9" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 6–9. Habitus of Heterogastridae species from Laos and Thailand. 6–7, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 8–9, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="7" pageNumber="58">Fig. 9</figureCitation>
) a little more than 1.4 times as wide as its length, nearly trapezoidal, lacking collar, covered with dense punctures, and with lateral margin of anterior lobe weakly serrate; posterior lobe 1.4 times as long as anterior lobe, slightly more convex than anterior lobe; posterior margin almost straight. Venter of prothorax, mesepisterna and metepisterna covered with decumbent setae. Scutellum (
<figureCitation id="8BD5E6C0FFE93D68FB81B103FBB25391" box="[1047,1122,871,896]" captionStart="FIGURES 6–9" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 6–9. Habitus of Heterogastridae species from Laos and Thailand. 6–7, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 8–9, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="7" pageNumber="58">Fig. 9</figureCitation>
) triangular, weakly tumid transversely in basal half, covered with long suberect setae. Hemelytra (
<figureCitation id="8BD5E6C0FFE93D68FC28B1E8FBD153B5" box="[958,1025,908,933]" captionStart="FIGURES 6–9" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 6–9. Habitus of Heterogastridae species from Laos and Thailand. 6–7, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 8–9, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="7" pageNumber="58">Fig. 9</figureCitation>
) exceeding apex of abdomen; apical margin of corium as long as claval suture. Profemur moderately expanded, armed with a spine ventrally at about apical third; tibiae and tarsi sparsely covered with long erect setae.
</paragraph>
<paragraph id="1351FA45FFE93D68FF51B193FC62546D" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="58">
Abdomen wider than hemelytral width; sternum covered with decumbent setae. Ovipositor (
<figureCitation id="8BD5E6C0FFE93D68FB4BB193FAE05401" box="[1245,1328,1015,1040]" captionStart="FIGURES 17–19" captionStartId="11.[151,264,1221,1244]" captionTargetBox="[173,1370,892,1151]" captionTargetId="figure@11.[151,1436,861,1218]" captionTargetPageId="11" captionText="FIGURES 17–19. Abdominal sternite VI and ovipositer of each new Sadoletus species, ventral view. 17, Sadoletus laosensis sp. nov.; 18, S. melasmus sp. nov.; 19, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437394/files/figure.png" pageId="7" pageNumber="58">Fig. 18</figureCitation>
) strongly curved laterally, reaching posterior margin of sternite VI, approximate oviposital length
<quantity id="D41657A0FFE93D68FB5AB678FAFD5425" box="[1228,1325,1052,1076]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="58" unit="mm" value="0.4">0.4 mm</quantity>
; the two hemiventrites of the medially completely bisected sternite VII are short, therefore posterior margins of sternites III–V straight, posterior margin of sternite VI weakly sinuate (
<figureCitation id="8BD5E6C0FFE93D68FCC7B600FC75546D" box="[849,933,1124,1149]" captionStart="FIGURES 17–19" captionStartId="11.[151,264,1221,1244]" captionTargetBox="[173,1370,892,1151]" captionTargetId="figure@11.[151,1436,861,1218]" captionTargetPageId="11" captionText="FIGURES 17–19. Abdominal sternite VI and ovipositer of each new Sadoletus species, ventral view. 17, Sadoletus laosensis sp. nov.; 18, S. melasmus sp. nov.; 19, S. subpellucidus sp. nov. Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437394/files/figure.png" pageId="7" pageNumber="58">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="1351FA45FFE93D68FF51B6E3FDA554B1" blockId="7.[151,1437,151,2013]" box="[199,629,1159,1184]" pageId="7" pageNumber="58">
Spermatheca (paratypes) as in
<figureCitation id="8BD5E6C0FFE93D68FD88B6E3FDA154B1" box="[542,625,1159,1184]" captionStart="FIGURES 25–29" captionStartId="13.[151,264,1961,1984]" captionTargetBox="[152,1431,355,1907]" captionTargetId="figure@13.[151,1436,318,1925]" captionTargetPageId="13" captionText="FIGURES 25–29. Sadoletus melasmus sp. nov. 25, Pygophore, dorsal view (setae omitted from left half); 26, left paramere, dorsal view; 27–28, phallus (endosoma not everted), ventral (27) and lateral (28) views; 29, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437398/files/figure.png" pageId="7" pageNumber="58">Fig. 29</figureCitation>
.
</paragraph>
<paragraph id="1351FA45FFE93D68FF51B6C8FAA05569" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="58">
Male (
<typeStatus id="CC5544E7FFE93D68FE86B6C8FE5354D5" box="[272,387,1196,1220]" pageId="7" pageNumber="58" type="paratype">paratypes</typeStatus>
): Male resembles female in general appearance. Body (
<figureCitation id="8BD5E6C0FFE93D68FC64B6C8FBE554D5" box="[1010,1077,1196,1221]" captionStart="FIGURES 6–9" captionStartId="8.[151,264,1896,1919]" captionTargetBox="[295,1268,427,1807]" captionTargetId="figure@8.[264,1323,373,1875]" captionTargetPageId="8" captionText="FIGURES 6–9. Habitus of Heterogastridae species from Laos and Thailand. 6–7, Sadoletus laosensis sp. nov., male (6), holotype, female (7); 8–9, S. melasmus sp. nov., male (8), holotype, female (9). Scales: 1.0 mm." httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="7" pageNumber="58">Fig. 8</figureCitation>
) evidently narrower; coloration of posterior lobe of pronotum and each leg paler than female; pygophore (
<figureCitation id="8BD5E6C0FFE93D68FC4DB6ABFBFD54F9" box="[987,1069,1231,1256]" captionStart="FIGURES 25–29" captionStartId="13.[151,264,1961,1984]" captionTargetBox="[152,1431,355,1907]" captionTargetId="figure@13.[151,1436,318,1925]" captionTargetPageId="13" captionText="FIGURES 25–29. Sadoletus melasmus sp. nov. 25, Pygophore, dorsal view (setae omitted from left half); 26, left paramere, dorsal view; 27–28, phallus (endosoma not everted), ventral (27) and lateral (28) views; 29, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437398/files/figure.png" pageId="7" pageNumber="58">Fig. 25</figureCitation>
) nearly spherical in dorsal view, covered with suberect setae; dorsal sinus of posterior aperture one quarter as wide at middle as pygophore, without collar along anterior margin; posterior margin of pygophore and cuplike sclerite fused together; paramere (
<figureCitation id="8BD5E6C0FFE93D68FAD4B773FA445521" box="[1346,1428,1303,1328]" captionStart="FIGURES 25–29" captionStartId="13.[151,264,1961,1984]" captionTargetBox="[152,1431,355,1907]" captionTargetId="figure@13.[151,1436,318,1925]" captionTargetPageId="13" captionText="FIGURES 25–29. Sadoletus melasmus sp. nov. 25, Pygophore, dorsal view (setae omitted from left half); 26, left paramere, dorsal view; 27–28, phallus (endosoma not everted), ventral (27) and lateral (28) views; 29, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437398/files/figure.png" pageId="7" pageNumber="58">Fig. 26</figureCitation>
) robust, twisted at basal third, sickle-shaped at apex, covered with several erect setae, sensory processes projecting laterally; phallosoma (
<figureCitation id="8BD5E6C0FFE93D68FE04B73BFDC85569" box="[402,536,1375,1400]" captionStart="FIGURES 25–29" captionStartId="13.[151,264,1961,1984]" captionTargetBox="[152,1431,355,1907]" captionTargetId="figure@13.[151,1436,318,1925]" captionTargetPageId="13" captionText="FIGURES 25–29. Sadoletus melasmus sp. nov. 25, Pygophore, dorsal view (setae omitted from left half); 26, left paramere, dorsal view; 27–28, phallus (endosoma not everted), ventral (27) and lateral (28) views; 29, spermatheca. Scales: 0.1 mm." httpUri="https://zenodo.org/record/1437398/files/figure.png" pageId="7" pageNumber="58">Figs. 27, 28</figureCitation>
) nearly twice as long as its width in ventral view, wrinkled at apical margin.
</paragraph>
<paragraph id="1351FA45FFE93D68FF51B7E0FD8556AC" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="58">
<emphasis id="219A2657FFE93D68FF51B7E0FEA6558C" bold="true" box="[199,374,1412,1437]" pageId="7" pageNumber="58">Measurements</emphasis>
[
<specimenCount id="05E831CCFFE93D68FE10B7E1FE4D558D" box="[390,413,1413,1436]" pageId="7" pageNumber="58" type="male">♂</specimenCount>
(n=2) /
<specimenCount id="05E831CCFFE93D68FE6AB7E1FDC1558D" box="[508,529,1413,1436]" pageId="7" pageNumber="58" type="female">♀</specimenCount>
(n=5), value for
<typeStatus id="CC5544E7FFE93D68FD40B7E0FCE8558C" box="[726,824,1412,1437]" pageId="7" pageNumber="58" type="holotype">holotype</typeStatus>
female in parentheses]. Body length 4.55–4.6 / 4.61– 5.2 (4.97); body height between coxae and dorsum 1.15–1.2 / 1.18–1.38 (1.3); head length 0.7–0.75 / 0.7–0.83 (0.75), head width across eyes 1.25–1.3 / 1.23–1.38 (1.31); length of antennal segments I–IV respectively 0.33– 0.34 / 0.31–0.35 (0.34), 0.59–0.63 / 0.52–0.65 (0.58), 0.58–0.59 / 0.5–0.6 (0.53), 0.85–0.86 / 0.76–0.88 (0.81); length of rostral segments I–IV respectively 0.41–0.43 / 0.41–0.43 (0.42), 0.43–0.44 / 0.44–0.45 (0.44), 0.33–0.35 / 0.31–0.35 (0.35), 0.3–0.31 / 0.29–0.33 (0.31); mesal pronotal length 1.08–1.13 / 1.06–1.31 (1.18); posterior pronotal width 1.38–1.4 / 1.41–1.73 (1.59); mesal scutellal length 0.74–0.76 / 0.76–0.96 (0.81); basal scutellal width 0.74–0.78 / 0.76–1.0 (0.88); hemelytral length 2.85–2.9 / 2.96–3.38 (3.25); maximum width across hemelytra 1.35–1.43 / 1.44–1.73 (1.53).
</paragraph>
</subSubSection>
<subSubSection id="5BF4A9CEFFE93D68FF51B4A3FC8D56F1" box="[199,861,1735,1761]" pageId="7" pageNumber="58" type="distribution">
<paragraph id="1351FA45FFE93D68FF51B4A3FC8D56F1" blockId="7.[151,1437,151,2013]" box="[199,861,1735,1761]" pageId="7" pageNumber="58">
<emphasis id="219A2657FFE93D68FF51B4A3FEB056F1" bold="true" box="[199,352,1735,1760]" pageId="7" pageNumber="58">Distribution.</emphasis>
<collectingCountry id="6BF9BAD5FFE93D68FEF1B4A3FE1B56F1" box="[359,459,1735,1760]" name="Thailand" pageId="7" pageNumber="58">Thailand</collectingCountry>
(
<collectingRegion id="D12A34A7FFE93D68FE4FB4A3FDB556F1" box="[473,613,1735,1760]" country="Thailand" name="Chanthaburi" pageId="7" pageNumber="58">Chanthaburi</collectingRegion>
Prov.,
<collectingRegion id="D12A34A7FFE93D68FD23B4ADFCDC56F0" box="[693,780,1737,1761]" country="Thailand" name="Rayong" pageId="7" pageNumber="58">Rayong</collectingRegion>
Prov.).
</paragraph>
</subSubSection>
<subSubSection id="5BF4A9CEFFE93D68FF51B488FCB55739" pageId="7" pageNumber="58" type="etymology">
<paragraph id="1351FA45FFE93D68FF51B488FCB55739" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="58">
<emphasis id="219A2657FFE93D68FF51B488FE9E5714" bold="true" box="[199,334,1772,1797]" pageId="7" pageNumber="58">Etymology.</emphasis>
The species epithet is the Greek adjective “
<taxonomicName id="D4EE81C6FFE93D68FCEBB488FC3B5715" authorityName="Ban" authorityYear="2018" box="[893,1003,1772,1796]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="melasmus">
<emphasis id="219A2657FFE93D68FCEBB488FC3B5715" box="[893,1003,1772,1796]" italics="true" pageId="7" pageNumber="58">melasmus</emphasis>
</taxonomicName>
” (meaning black spot) referring to characteristic black to dark brown spot at inner angle of corium.
</paragraph>
</subSubSection>
<subSubSection id="5BF4A9CEFFE93D66FF51B550FB7B50C4" lastPageId="9" lastPageNumber="60" pageId="7" pageNumber="58" type="discussion">
<paragraph id="1351FA45FFE93D67FF51B550FB685151" blockId="7.[151,1437,151,2013]" lastBlockId="8.[151,1437,151,320]" lastPageId="8" lastPageNumber="59" pageId="7" pageNumber="58">
<emphasis id="219A2657FFE93D68FF51B550FEEB575C" bold="true" box="[199,315,1844,1869]" pageId="7" pageNumber="58">Remarks.</emphasis>
This new species is most similar in general appearance to
<taxonomicName id="D4EE81C6FFE93D68FB90B550FB31575C" authority="Bergroth" authorityName="Bergroth" box="[1030,1249,1844,1869]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="bakeri">
<emphasis id="219A2657FFE93D68FB90B550FBBF575D" box="[1030,1135,1844,1868]" italics="true" pageId="7" pageNumber="58">S. bakeri</emphasis>
Bergroth
</taxonomicName>
,
<taxonomicName id="D4EE81C6FFE93D68FB64B550FF2A5761" authority="Bergroth" authorityName="Bergroth" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="montanellus">
<emphasis id="219A2657FFE93D68FB64B550FA4C575D" box="[1266,1436,1844,1868]" italics="true" pageId="7" pageNumber="58">S. montanellus</emphasis>
Bergroth
</taxonomicName>
and
<taxonomicName id="D4EE81C6FFE93D68FEA2B53CFDDC5761" authority="Ban" authorityName="Ban &amp; Ishikawa" authorityYear="2013" box="[308,524,1880,1904]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="ryukyuensis">
<emphasis id="219A2657FFE93D68FEA2B53CFE065761" box="[308,470,1880,1904]" italics="true" pageId="7" pageNumber="58">S. ryukyuensis</emphasis>
Ban
</taxonomicName>
et
<collectingRegion id="D12A34A7FFE93D68FDA4B533FD4A5761" box="[562,666,1879,1904]" country="Japan" name="Ishikawa" pageId="7" pageNumber="58">Ishikawa</collectingRegion>
, but it can be distinguished from the latter by a combination of the following characters: Body length more than
<quantity id="D41657A0FFE93D68FD09B518FD275785" box="[671,759,1916,1941]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="58" unit="mm" value="4.5">4.5 mm</quantity>
(in
<taxonomicName id="D4EE81C6FFE93D68FCB1B518FC1D5785" box="[807,973,1916,1940]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="montanellus">
<emphasis id="219A2657FFE93D68FCB1B518FC1D5785" box="[807,973,1916,1940]" italics="true" pageId="7" pageNumber="58">S. montanellus</emphasis>
</taxonomicName>
, 3.0 mm); antennal segment II less than twice as long as segment I (vs. antennal segment II more than twice as long as segment I); labium slightly exceeding posterior margin of prothorax, not reaching mesocoxae (vs. at least reaching mesocoxae); posterior lobe of pronotum pale brownish yellow (in
<taxonomicName id="D4EE81C6FFE63D67FDDCB2FCFD2050A1" box="[586,752,152,176]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="montanellus">
<emphasis id="219A2657FFE63D67FDDCB2FCFD2050A1" box="[586,752,152,176]" italics="true" pageId="8" pageNumber="59">S. montanellus</emphasis>
</taxonomicName>
, dark brown); coxae black (in
<taxonomicName id="D4EE81C6FFE63D67FBDDB2FCFB3C50A1" authorityName="Ban &amp; Ishikawa" authorityYear="2013" box="[1099,1260,152,176]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="ryukyuensis">
<emphasis id="219A2657FFE63D67FBDDB2FCFB3C50A1" box="[1099,1260,152,176]" italics="true" pageId="8" pageNumber="59">S. ryukyuensis</emphasis>
</taxonomicName>
, coxae brown); ground color of femora and tibiae pale yellow (vs. femora and tibiae yellowish brown); apical third of corium black in both sexes (in
<taxonomicName id="D4EE81C6FFE63D67FEC9B284FDD150E9" authorityName="Ban &amp; Ishikawa" authorityYear="2013" box="[351,513,224,248]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="ryukyuensis">
<emphasis id="219A2657FFE63D67FEC9B284FDD150E9" box="[351,513,224,248]" italics="true" pageId="8" pageNumber="59">S. ryukyuensis</emphasis>
</taxonomicName>
, apical third of corium white in female); hemelytra with dark markings at inner angle of corium and hemelytral membrane (vs. hemelytra lacking dark marking at inner angle of corium and hemelytral membrane); abdominal sterna black (in
<taxonomicName id="D4EE81C6FFE63D67FD42B34CFCE85151" box="[724,824,296,320]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="bakeri">
<emphasis id="219A2657FFE63D67FD42B34CFCE85151" box="[724,824,296,320]" italics="true" pageId="8" pageNumber="59">S. bakeri</emphasis>
</taxonomicName>
abdominal sterna reddish brown).
</paragraph>
<caption id="4791AACDFFE63D67FF01B50CFBB5578C" httpUri="https://zenodo.org/record/1437388/files/figure.png" pageId="8" pageNumber="59" startId="8.[151,264,1896,1919]" targetBox="[295,1268,427,1807]" targetPageId="8">
<paragraph id="1351FA45FFE63D67FF01B50CFBB5578C" blockId="8.[151,1436,1896,1949]" pageId="8" pageNumber="59">
<emphasis id="219A2657FFE63D67FF01B50CFEE7576C" bold="true" box="[151,311,1896,1919]" pageId="8" pageNumber="59">FIGURES 6–9</emphasis>
. Habitus of
<taxonomicName id="D4EE81C6FFE63D67FE28B50CFD8E576F" authorityName="Stal" authorityYear="1872" box="[446,606,1896,1918]" class="Insecta" family="Heterogastridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="family">Heterogastridae</taxonomicName>
species from Laos and Thailand. 6–7,
<taxonomicName id="D4EE81C6FFE63D67FC6BB50CFB16576F" authority="Ban, 2018" authorityName="Ban" authorityYear="2018" box="[1021,1222,1896,1918]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="laosensis" status="sp. nov.">
<emphasis id="219A2657FFE63D67FC6BB50CFB16576F" box="[1021,1222,1896,1918]" italics="true" pageId="8" pageNumber="59">Sadoletus laosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3AA99B2CFFE63D67FB46B50DFAF3576F" box="[1232,1315,1897,1918]" pageId="8" pageNumber="59" rank="species">
<emphasis id="219A2657FFE63D67FB46B50DFAF3576F" bold="true" box="[1232,1315,1897,1918]" pageId="8" pageNumber="59">sp. nov.</emphasis>
</taxonomicNameLabel>
, male (6), holotype, female (7); 8–9,
<taxonomicName id="D4EE81C6FFE63D67FE33B5ECFDF2578C" authority="Ban, 2018" authorityName="Ban" authorityYear="2018" box="[421,546,1927,1949]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="melasmus" status="sp. nov.">
<emphasis id="219A2657FFE63D67FE33B5ECFDF2578C" box="[421,546,1927,1949]" italics="true" pageId="8" pageNumber="59">S. melasmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3AA99B2CFFE63D67FDBEB5E3FDA8578D" box="[552,632,1927,1948]" pageId="8" pageNumber="59" rank="species">
<emphasis id="219A2657FFE63D67FDBEB5E3FDA8578D" bold="true" box="[552,632,1927,1948]" pageId="8" pageNumber="59">sp. nov.</emphasis>
</taxonomicNameLabel>
, male (8), holotype, female (9). Scales: 1.0 mm.
</paragraph>
</caption>
<paragraph id="1351FA45FFE73D66FF51B2F3FB7B50C4" blockId="9.[151,1436,151,213]" pageId="9" pageNumber="60">
<emphasis id="219A2657FFE73D66FF51B2F3FE5B50A1" bold="true" box="[199,395,151,176]" pageId="9" pageNumber="60">Biological notes.</emphasis>
The specimens were collected from trees of
<taxonomicName id="D4EE81C6FFE73D66FC19B2FCFC1D50BE" box="[911,973,152,175]" class="Magnoliopsida" family="Moraceae" genus="Ficus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Rosales" pageId="9" pageNumber="60" phylum="Tracheophyta" rank="genus">
<emphasis id="219A2657FFE73D66FC19B2FCFC1D50BE" box="[911,973,152,175]" italics="true" pageId="9" pageNumber="60">Ficus</emphasis>
</taxonomicName>
sp. in mountainous forests and the open forest adjacent to a resort hotel, with several other heterogastrid and rhyparochromid species.
</paragraph>
</subSubSection>
</treatment>
</document>
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