treatments-xml/data/9A/66/87/9A6687E64F06303CFF34F9FCFD9EFBB7.xml

<document id="FA5FA9ADDE9692157075A24BF554A3A6" ID-CLB-Dataset="36889" ID-DOI="10.11646/zootaxa.3774.1.1" ID-GBIF-Dataset="41d311ee-850e-4125-9a40-5a96a2aa9065" ID-ISSN="1175-5326" ID-Zenodo-Dep="285693" ID-ZooBank="B5A2EF46-2BF6-4ED3-A5F4-5F9951400545" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460391910470" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2014" docId="9A6687E64F06303CFF34F9FCFD9EFBB7" docLanguage="en" docName="zt03774p030.pdf" docOrigin="Zootaxa 3774 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microscydmus (Scydmomicrus) styxianus Jałoszyński, 2014, sp. n." docType="treatment" docVersion="8" lastPageNumber="23" masterDocId="665FFF9E4F13302AFFA3FFEEFFFEFFCA" masterDocTitle="The smallest Cyrtoscydmini of Australia: revision of Microscydmus Saulcy &amp; Croissandeau and Penicillidmus gen. n. (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="22" updateTime="1698298320569" updateUser="ExternalLinkService">
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<mods:title id="19125229EF715CE7CA3A82813F673B94">The smallest Cyrtoscydmini of Australia: revision of Microscydmus Saulcy &amp; Croissandeau and Penicillidmus gen. n. (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<paragraph id="127036F04F06303FFF34F9FCFD28F9E7" blockId="21.[151,726,1554,1614]" box="[151,726,1554,1581]" pageId="21" pageNumber="22">
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<emphasis id="20BBEAE24F06303FFF34F9FCFD28F9E7" bold="true" box="[151,726,1554,1581]" pageId="21" pageNumber="22">
<taxonomicName id="D5CF4D734F06303FFF34F9FCFD72F9E6" ID-CoL="435TK" box="[151,652,1554,1581]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="styxianus" status="sp. nov." subGenus="Scydmomicrus">
<emphasis id="20BBEAE24F06303FFF34F9FCFEB9F9E6" bold="true" box="[151,327,1554,1580]" italics="true" pageId="21" pageNumber="22">Microscydmus</emphasis>
(
<emphasis id="20BBEAE24F06303FFEFBF9FCFDF7F9E6" bold="true" box="[344,521,1554,1580]" italics="true" pageId="21" pageNumber="22">Scydmomicrus</emphasis>
) 
<emphasis id="20BBEAE24F06303FFDB9F9FCFD72F9E6" bold="true" box="[538,652,1554,1580]" italics="true" pageId="21" pageNumber="22">styxianus</emphasis>
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<taxonomicNameLabel id="3B8857994F06303FFD37F9FDFD28F9E7" box="[660,726,1555,1581]" pageId="21" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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<paragraph id="127036F04F06303FFF34F9DBFE7CF984" blockId="21.[151,726,1554,1614]" box="[151,386,1589,1614]" pageId="21" pageNumber="22">
(
<figureCitation id="8AF42A754F06303FFF3CF9DBFF02F984" box="[159,252,1589,1614]" captionStart="FIGURES 20 – 28" captionStartId="9.[151,264,1586,1609]" captionTargetBox="[159,1435,210,1555]" captionTargetId="figure@9.[151,1436,193,1565]" captionTargetPageId="9" captionText="FIGURES 20 – 28. Dorsal habitus of holotypes. Microscydmus (Scydmomicrus) australiensis Franz, female (20); M. (Sc.) nasicornis Franz, female (21); M. (Sc.) capitiseboraci sp. n., male (22); M. (Sc.) queenslandicus sp. n., male (23); M. (Sc.) styxianus sp. n., male (24); M. (Sc.) edithensis sp. n., male (25); M. (Sc.) tooloomensis sp. n., male (26); Penicillidmus masseyensis sp. n., male (27); P. unicolor sp. n., male (28)." httpUri="https://zenodo.org/record/285699/files/figure.png" pageId="21" pageNumber="22">Figs. 24</figureCitation>
, 
<figureCitation id="8AF42A754F06303FFEABF9D8FED8F984" box="[264,294,1590,1614]" captionStart="FIGURES 29 – 37" captionStartId="10.[151,264,1512,1535]" captionTargetBox="[157,1418,212,1473]" captionTargetId="figure@10.[151,1436,193,1492]" captionTargetPageId="10" captionText="FIGURES 29 – 37. Body silhouette of holotypes. Microscydmus (Scydmomicrus) australiensis Franz, female (29); M. (Sc.) nasicornis Franz, female (30); M. (Sc.) capitiseboraci sp. n., male (31); M. (Sc.) queenslandicus sp. n., male (32); M. (Sc.) styxianus sp. n., male (33); M. (Sc.) edithensis sp. n., male (34); M. (Sc.) tooloomensis sp. n., male (35); Penicillidmus masseyensis sp. n., male (36); P. unicolor sp. n., male (37)." httpUri="https://zenodo.org/record/285700/files/figure.png" pageId="21" pageNumber="22">33</figureCitation>
, 
<figureCitation id="8AF42A754F06303FFE91F9DBFEAFF984" box="[306,337,1589,1614]" captionStart="FIGURES 54 – 61" captionStartId="16.[151,264,1561,1584]" captionTargetBox="[152,1427,213,1530]" captionTargetId="figure@16.[151,1436,193,1540]" captionTargetPageId="16" captionText="FIGURES 54 – 61. Aedeagi in ventral views. M. (Sc.) capitiseboraci sp. n. (54); M. (Sc.) queenslandicus sp. n. (55); M. (Sc.) styxianus sp. n. (56); M. (Sc.) edithensis sp. n. (57); M. (Sc.) tooloomensis sp. n. (58 – 59); Penicillidmus masseyensis sp. n. (60); P. unicolor sp. n. (61)." httpUri="https://zenodo.org/record/285705/files/figure.png" pageId="21" pageNumber="22">56</figureCitation>
, 
<figureCitation id="8AF42A754F06303FFEFEF9DBFE84F984" box="[349,378,1589,1614]" captionStart="FIGURES 62 – 63" captionStartId="17.[151,264,1608,1631]" captionTargetBox="[151,1434,1055,1586]" captionTargetId="figure@17.[151,1436,1055,1586]" captionTargetPageId="17" captionText="FIGURES 62 – 63. Distribution of Microscydmus (Scydmomicrus subgen. n.) (62) and Penicillidmus gen. n. (63). Microscydmus (Scydmomicrus) australiensis Franz (62 a); M. (Sc.) nasicornis Franz (62 b); M. (Sc.) capitiseboraci sp. n. (2 c); M. (Sc.) queenslandicus sp. n. (62 d); M. (Sc.) styxianus sp. n. (62 e); M. (Sc.) edithensis sp. n. (62 f); M. (Sc.) tooloomensis sp. n. (62 g); Microscydmus (Sc.) sp. (62 h); Penicillidmus masseyensis sp. n. (63 a); P. unicolor sp. n. (63 b); Penicillidmus sp. (63 c)." httpUri="https://zenodo.org/record/285706/files/figure.png" pageId="21" pageNumber="22">62</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5AD5657B4F06303FFF34F993FCD8F8C8" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="127036F04F06303FFF34F993FCD8F8C8" blockId="21.[151,1436,1661,2010]" pageId="21" pageNumber="22">
<emphasis id="20BBEAE24F06303FFF34F993FEA3F95C" bold="true" box="[151,349,1661,1686]" pageId="21" pageNumber="22">Material studied</emphasis>
. 
<typeStatus id="CD7488524F06303FFEC8F993FE26F95C" box="[363,472,1661,1686]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
: ♂: three labels: &quot; 
<geoCoordinate id="77FB50374F06303FFD3FF990FD10F95C" box="[668,750,1662,1686]" direction="south" orientation="latitude" pageId="21" pageNumber="22" precision="555" value="-30.44">30.44S</geoCoordinate>
<geoCoordinate id="77FB50374F06303FFD57F993FCACF95C" box="[756,850,1661,1686]" direction="east" orientation="longitude" pageId="21" pageNumber="22" precision="555" value="152.05">152.05E</geoCoordinate>
<collectionCode id="74DEAE354F06303FFCFAF990FC69F95C" LSID="urn:lsid:biocol.org:col:15627" box="[857,919,1662,1686]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15627" name="Royal Botanic Gardens, National Herbarium of New South Wales" pageId="21" pageNumber="22">NSW</collectionCode>
/ Styx River SF., / Cedar Pit Flor. Res. / 
<quantity id="D5379B154F06303FFAFFF993FA62F95C" box="[1372,1436,1661,1686]" metricMagnitude="4" metricUnit="m" metricValue="4.2" pageId="21" pageNumber="22" unit="km" value="42.0">42km</quantity>
SE Wollomombi / 
<quantity id="D5379B154F06303FFECBF94FFE54F970" box="[360,426,1697,1722]" metricMagnitude="2" metricUnit="m" metricValue="9.35" pageId="21" pageNumber="22" unit="m" value="935.0">935m</quantity>
, 
<date id="667110304F06303FFE15F94CFDB2F970" box="[438,588,1698,1722]" pageId="21" pageNumber="22" value="1993-06-10">10 June 1993</date>
/ D.S. Chandler&quot; [white, printed], &quot;Berlesate 
<collectionCode id="74DEAE354F06303FFBE8F94CFB71F970" LSID="urn:lsid:biocol.org:col:32981" box="[1099,1167,1698,1722]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="21" pageNumber="22">ANIC</collectionCode>
1638 / old subtr. closed forest / mixed litter&quot; [white, printed], &quot; 
<taxonomicName id="D5CF4D734F06303FFD23F928FCA8F917" box="[640,854,1734,1757]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="20BBEAE24F06303FFD23F928FCA8F917" box="[640,854,1734,1757]" italics="true" pageId="21" pageNumber="22">MICROSCYDMUS</emphasis>
</taxonomicName>
/ (
<emphasis id="20BBEAE24F06303FFC2AF928FB9EF917" box="[905,1120,1734,1757]" italics="true" pageId="21" pageNumber="22">SCYDMOMICRUS</emphasis>
) / 
<taxonomicName id="D5CF4D734F06303FFB32F928FB07F917" box="[1169,1273,1734,1757]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="styxianus">
<emphasis id="20BBEAE24F06303FFB32F928FB07F917" box="[1169,1273,1734,1757]" italics="true" pageId="21" pageNumber="22">styxianus</emphasis>
</taxonomicName>
m. / det. P. Jałoszyński, 2013 / 
<typeStatus id="CD7488524F06303FFED7F904FDE0F8C8" box="[372,542,1770,1794]" pageId="21" pageNumber="22" type="holotype">HOLOTYPUS</typeStatus>
&quot; [red, printed] (
<collectionCode id="74DEAE354F06303FFD6CF904FCE4F8C8" LSID="urn:lsid:biocol.org:col:32981" box="[719,794,1770,1794]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="21" pageNumber="22">ANIC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="5AD5657B4F06303FFF64F8E3FCBCF810" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="127036F04F06303FFF64F8E3FCBCF810" blockId="21.[151,1436,1661,2010]" pageId="21" pageNumber="22">
<emphasis id="20BBEAE24F06303FFF64F8E3FEBFF8EC" bold="true" box="[199,321,1805,1830]" pageId="21" pageNumber="22">Diagnosis.</emphasis>
Male: body uniformly light brown; frons subtriangular with tiny antero-median projection; punctures on head fine but distinct, clearly more distinct than those on pronotum; BL &lt;
<quantity id="D5379B154F06303FFB34F8DFFAFEF883" box="[1175,1280,1841,1866]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="21" pageNumber="22" unit="mm" value="0.6">0.60 mm</quantity>
; pronotum as broad as head, with ante-basal transverse groove; EI &lt;1.50; aedeagus with distinct internal armature containing a pair of lightly sclerotized strongly elongate sub-median sclerites surrounded at the level of basal foramen by a pair of darkly sclerotized loops and with subapical lateral groups of densely assembled microscopic needle-like structures. Females and their diagnostic characters unknown.
</paragraph>
</subSubSection>
<subSubSection id="5AD5657B4F05303CFF64FF79FE5CFC02" pageId="22" pageNumber="23" type="description">
<paragraph id="127036F04F05303CFF64FF79FEF5FF32" blockId="22.[151,1436,151,1149]" pageId="22" pageNumber="23">
<emphasis id="20BBEAE24F05303CFF64FF79FEA7FF7A" bold="true" box="[199,345,151,176]" pageId="22" pageNumber="23">Description.</emphasis>
Body of male (
<figureCitation id="8AF42A754F05303CFDA9FF79FDA2FF7A" box="[522,604,151,176]" captionStart="FIGURES 20 – 28" captionStartId="9.[151,264,1586,1609]" captionTargetBox="[159,1435,210,1555]" captionTargetId="figure@9.[151,1436,193,1565]" captionTargetPageId="9" captionText="FIGURES 20 – 28. Dorsal habitus of holotypes. Microscydmus (Scydmomicrus) australiensis Franz, female (20); M. (Sc.) nasicornis Franz, female (21); M. (Sc.) capitiseboraci sp. n., male (22); M. (Sc.) queenslandicus sp. n., male (23); M. (Sc.) styxianus sp. n., male (24); M. (Sc.) edithensis sp. n., male (25); M. (Sc.) tooloomensis sp. n., male (26); Penicillidmus masseyensis sp. n., male (27); P. unicolor sp. n., male (28)." httpUri="https://zenodo.org/record/285699/files/figure.png" pageId="22" pageNumber="23">Fig. 24</figureCitation>
) moderately convex, strongly elongate and slender, with moderately long appendages, BL 
<quantity id="D5379B154F05303CFEF7FF52FE34FF1E" box="[340,458,188,213]" metricMagnitude="-4" metricUnit="m" metricValue="5.869999999999999" pageId="22" pageNumber="23" unit="mm" value="0.587">0.587 mm</quantity>
; cuticle glossy; body uniformly light brown with slightly lighter appendages; vestiture yellowish.
</paragraph>
<paragraph id="127036F04F05303CFF64FEEAFC80FE67" blockId="22.[151,1436,151,1149]" pageId="22" pageNumber="23">
Head (
<figureCitation id="8AF42A754F05303CFEB7FEEAFE96FED6" box="[276,360,260,285]" captionStart="FIGURES 29 – 37" captionStartId="10.[151,264,1512,1535]" captionTargetBox="[157,1418,212,1473]" captionTargetId="figure@10.[151,1436,193,1492]" captionTargetPageId="10" captionText="FIGURES 29 – 37. Body silhouette of holotypes. Microscydmus (Scydmomicrus) australiensis Franz, female (29); M. (Sc.) nasicornis Franz, female (30); M. (Sc.) capitiseboraci sp. n., male (31); M. (Sc.) queenslandicus sp. n., male (32); M. (Sc.) styxianus sp. n., male (33); M. (Sc.) edithensis sp. n., male (34); M. (Sc.) tooloomensis sp. n., male (35); Penicillidmus masseyensis sp. n., male (36); P. unicolor sp. n., male (37)." httpUri="https://zenodo.org/record/285700/files/figure.png" pageId="22" pageNumber="23">Fig. 33</figureCitation>
) subtrapezoidal, broadest at eyes, HL 0.100 mm, HW 
<quantity id="D5379B154F05303CFC41FEEAFBA7FED6" box="[994,1113,260,285]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="23" unit="mm" value="0.15">0.150 mm</quantity>
; tempora much shorter than eyes, strongly convergent postero-mesally; vertex weakly convex; frons subtriangular with small antero-median projection. Eyes large but moderately projecting laterally from the head silhouette, coarsely faceted. Punctures on head dorsum sparse and small but distinct; setae short, sparse and suberect. Antennae slender, AnL 
<quantity id="D5379B154F05303CFABCFE81FA66FE4D" box="[1311,1432,367,392]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="22" pageNumber="23" unit="mm" value="0.25">0.250 mm</quantity>
; antennomeres I–II elongate, III–X transverse; XI as long as broad.
</paragraph>
<paragraph id="127036F04F05303CFF64FE56FDC9FDF7" blockId="22.[151,1436,151,1149]" pageId="22" pageNumber="23">
Pronotum (
<figureCitation id="8AF42A754F05303CFEE7FE59FE6BFE1A" box="[324,405,439,464]" captionStart="FIGURES 29 – 37" captionStartId="10.[151,264,1512,1535]" captionTargetBox="[157,1418,212,1473]" captionTargetId="figure@10.[151,1436,193,1492]" captionTargetPageId="10" captionText="FIGURES 29 – 37. Body silhouette of holotypes. Microscydmus (Scydmomicrus) australiensis Franz, female (29); M. (Sc.) nasicornis Franz, female (30); M. (Sc.) capitiseboraci sp. n., male (31); M. (Sc.) queenslandicus sp. n., male (32); M. (Sc.) styxianus sp. n., male (33); M. (Sc.) edithensis sp. n., male (34); M. (Sc.) tooloomensis sp. n., male (35); Penicillidmus masseyensis sp. n., male (36); P. unicolor sp. n., male (37)." httpUri="https://zenodo.org/record/285700/files/figure.png" pageId="22" pageNumber="23">Fig. 33</figureCitation>
) in dorsal view oval, broadest near anterior third, PL 
<quantity id="D5379B154F05303CFC4BFE56FBA5FE05" box="[1000,1115,440,464]" metricMagnitude="-4" metricUnit="m" metricValue="1.37" pageId="22" pageNumber="23" unit="mm" value="0.137">0.137 mm</quantity>
, PW 
<quantity id="D5379B154F05303CFB34FE59FAF5FE05" box="[1175,1291,439,464]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="23" unit="mm" value="0.15">0.150 mm</quantity>
; anterior and lateral margins rounded; posterior margin shallowly bisinuate; base of pronotum with two pairs of small and shallow pits, internal pair connected by transverse impression. Punctures on pronotal disc fine and inconspicuous; setae sparse, thin, short and suberect.
</paragraph>
<paragraph id="127036F04F05303CFF64FDA9FDA3FD07" blockId="22.[151,1436,151,1149]" pageId="22" pageNumber="23">
Elytra oval, slightly more convex than pronotum, broadest near middle, EL 
<quantity id="D5379B154F05303CFB9DFDA9FB48FD95" box="[1086,1206,583,608]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="22" pageNumber="23" unit="mm" value="0.35">0.350 mm</quantity>
, EW 
<quantity id="D5379B154F05303CFB58FDA6FA8CFD95" box="[1275,1394,584,608]" metricMagnitude="-4" metricUnit="m" metricValue="2.37" pageId="22" pageNumber="23" unit="mm" value="0.237">0.237 mm</quantity>
, EI 1.474; basal impressions short; elytral apices separately rounded. Punctures on elytral disc slightly more distinct than those on pronotum but still fine and unremarkable; setae short, sparse and suberect. 
<collectingCountry id="6AD876604F05303CFB75FD61FAF0FD62" box="[1238,1294,655,680]" name="India" pageId="22" pageNumber="23">Hind</collectingCountry>
wings well developed, about twice as long as elytra.
</paragraph>
<paragraph id="127036F04F05303CFF64FD36FCB0FD3A" blockId="22.[151,1436,151,1149]" box="[199,846,727,752]" pageId="22" pageNumber="23">Legs moderately long and slender, without modifications.</paragraph>
<paragraph id="127036F04F05303CFF64FD12FF34FC6E" blockId="22.[151,1436,151,1149]" pageId="22" pageNumber="23">
Aedeagus (
<figureCitation id="8AF42A754F05303CFEE7FD12FE68FCDF" box="[324,406,764,789]" captionStart="FIGURES 54 – 61" captionStartId="16.[151,264,1561,1584]" captionTargetBox="[152,1427,213,1530]" captionTargetId="figure@16.[151,1436,193,1540]" captionTargetPageId="16" captionText="FIGURES 54 – 61. Aedeagi in ventral views. M. (Sc.) capitiseboraci sp. n. (54); M. (Sc.) queenslandicus sp. n. (55); M. (Sc.) styxianus sp. n. (56); M. (Sc.) edithensis sp. n. (57); M. (Sc.) tooloomensis sp. n. (58 – 59); Penicillidmus masseyensis sp. n. (60); P. unicolor sp. n. (61)." httpUri="https://zenodo.org/record/285705/files/figure.png" pageId="22" pageNumber="23">Fig. 56</figureCitation>
) large, AeL 
<quantity id="D5379B154F05303CFD80FD12FD74FCDE" box="[547,650,764,788]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="22" pageNumber="23" unit="mm" value="0.13">0.13 mm</quantity>
; moderately slender, with distinct internal armature containing several well-defined components: a pair of lightly sclerotized strongly elongate sub-median sclerites extending from base to subapical region of median lobe, sub-basally located pair of dark lateral sclerites forming elongate loops, and subapical lateral groups of densely assembled microscopic needle-like structures; each paramere with long apical seta.
</paragraph>
<paragraph id="127036F04F05303CFF64FC41FE5CFC02" blockId="22.[151,1436,151,1149]" box="[199,418,943,968]" pageId="22" pageNumber="23">
<emphasis id="20BBEAE24F05303CFF64FC41FEE0FC02" bold="true" box="[199,286,943,968]" pageId="22" pageNumber="23">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="5AD5657B4F05303CFF64FC3AFC8FFC27" box="[199,881,980,1005]" pageId="22" pageNumber="23" type="distribution">
<paragraph id="127036F04F05303CFF64FC3AFC8FFC27" blockId="22.[151,1436,151,1149]" box="[199,881,980,1005]" pageId="22" pageNumber="23">
<emphasis id="20BBEAE24F05303CFF64FC3AFEA7FC27" bold="true" box="[199,345,980,1005]" pageId="22" pageNumber="23">Distribution</emphasis>
(
<figureCitation id="8AF42A754F05303CFECBFC3AFE42FC27" box="[360,444,980,1005]" captionStart="FIGURES 62 – 63" captionStartId="17.[151,264,1608,1631]" captionTargetBox="[151,1434,1055,1586]" captionTargetId="figure@17.[151,1436,1055,1586]" captionTargetPageId="17" captionText="FIGURES 62 – 63. Distribution of Microscydmus (Scydmomicrus subgen. n.) (62) and Penicillidmus gen. n. (63). Microscydmus (Scydmomicrus) australiensis Franz (62 a); M. (Sc.) nasicornis Franz (62 b); M. (Sc.) capitiseboraci sp. n. (2 c); M. (Sc.) queenslandicus sp. n. (62 d); M. (Sc.) styxianus sp. n. (62 e); M. (Sc.) edithensis sp. n. (62 f); M. (Sc.) tooloomensis sp. n. (62 g); Microscydmus (Sc.) sp. (62 h); Penicillidmus masseyensis sp. n. (63 a); P. unicolor sp. n. (63 b); Penicillidmus sp. (63 c)." httpUri="https://zenodo.org/record/285706/files/figure.png" pageId="22" pageNumber="23">Fig. 62</figureCitation>
e). E 
<collectingCountry id="6AD876604F05303CFE57FC3AFDA5FC27" box="[500,603,980,1005]" name="Australia" pageId="22" pageNumber="23">Australia</collectingCountry>
(NE New South 
<collectingCountry id="6AD876604F05303CFCBFFC3AFC9BFC27" box="[796,869,980,1005]" name="United Kingdom" pageId="22" pageNumber="23">Wales</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="5AD5657B4F05303CFF64FC19FB7BFBDA" box="[199,1157,1015,1040]" pageId="22" pageNumber="23" type="etymology">
<paragraph id="127036F04F05303CFF64FC19FB7BFBDA" blockId="22.[151,1436,151,1149]" box="[199,1157,1015,1040]" pageId="22" pageNumber="23">
<emphasis id="20BBEAE24F05303CFF64FC19FEB0FBDA" bold="true" box="[199,334,1015,1040]" pageId="22" pageNumber="23">Etymology.</emphasis>
Locotypical, after the Styx River, near which the 
<typeStatus id="CD7488524F05303CFCDFFC19FC20FBDA" box="[892,990,1015,1040]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
was collected.
</paragraph>
</subSubSection>
<subSubSection id="5AD5657B4F05303CFF64FBF2FD9EFBB7" pageId="22" pageNumber="23" type="discussion">
<paragraph id="127036F04F05303CFF64FBF2FD9EFBB7" blockId="22.[151,1436,151,1149]" pageId="22" pageNumber="23">
<emphasis id="20BBEAE24F05303CFF64FBF2FEC5FBFF" bold="true" box="[199,315,1052,1077]" pageId="22" pageNumber="23">Remarks.</emphasis>
<taxonomicName id="D5CF4D734F05303CFEE7FBF2FDA6FBFE" box="[324,600,1052,1076]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="styxianus">
<emphasis id="20BBEAE24F05303CFEE7FBF2FDA6FBFE" box="[324,600,1052,1076]" italics="true" pageId="22" pageNumber="23">Microscydmus styxianus</emphasis>
</taxonomicName>
is the most slender Australian species; it can be easily identified on the basis of the very short pronotum in relation to the elytra. The ratio EL/PL for 
<taxonomicName id="D5CF4D734F05303CFBB3FBAFFB61FB9D" box="[1040,1183,1088,1112]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="styxianus">
<emphasis id="20BBEAE24F05303CFBB3FBAFFB61FB9D" box="[1040,1183,1088,1112]" italics="true" pageId="22" pageNumber="23">M. styxianus</emphasis>
</taxonomicName>
is 2.545, while for all remaining species it equals 1.857–2.400.
</paragraph>
</subSubSection>
</treatment>
</document>