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<document id="E8BDA896964AF5C84109A4437193FA26" ID-CLB-Dataset="36889" ID-DOI="10.11646/zootaxa.3774.1.1" ID-GBIF-Dataset="41d311ee-850e-4125-9a40-5a96a2aa9065" ID-ISSN="1175-5326" ID-Zenodo-Dep="285693" ID-ZooBank="B5A2EF46-2BF6-4ED3-A5F4-5F9951400545" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460391910470" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2014" docId="9A6687E64F023038FF34F8C7FC49F958" docLanguage="en" docName="zt03774p030.pdf" docOrigin="Zootaxa 3774 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microscydmus (Scydmomicrus) australiensis Franz" docType="treatment" docVersion="8" lastPageNumber="19" masterDocId="665FFF9E4F13302AFFA3FFEEFFFEFFCA" masterDocTitle="The smallest Cyrtoscydmini of Australia: revision of Microscydmus Saulcy &amp; Croissandeau and Penicillidmus gen. n. (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="18" updateTime="1698298320569" updateUser="ExternalLinkService">
<mods:mods id="772AC95611BA103BA91A8CBDD60645F7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="398E981D7EA31902E026567F9CD64DEA">The smallest Cyrtoscydmini of Australia: revision of Microscydmus Saulcy &amp; Croissandeau and Penicillidmus gen. n. (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="191C3E27C5E4C39812D6CC28C4A6E217">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="26EE207B99B72673E435CD17BF120113">Zootaxa</mods:title>
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<mods:date id="8115C5D286EB396549CB034228A9F7AB">2014</mods:date>
<mods:detail id="C5B44DF3598AD4B0C64290D558608A00" type="volume">
<mods:number id="33EED4ED64ED3AEE2C2902A2102C80D0">3774</mods:number>
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<mods:identifier id="6B02A6FDEADCFF07D04423E7393E8EB0" type="DOI">10.11646/zootaxa.3774.1.1</mods:identifier>
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<treatment id="9A6687E64F023038FF34F8C7FC49F958" ID-DOI="http://doi.org/10.5281/zenodo.5664968" ID-GBIF-Taxon="127667230" ID-Zenodo-Dep="5664968" LSID="urn:lsid:plazi:treatment:9A6687E64F023038FF34F8C7FC49F958" httpUri="http://treatment.plazi.org/id/9A6687E64F023038FF34F8C7FC49F958" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<subSubSection id="5AD5657B4F02303BFF34F8C7FDFEF8AF" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="127036F04F02303BFF34F8C7FCF8F88E" blockId="17.[151,774,1833,1893]" box="[151,774,1833,1860]" pageId="17" pageNumber="18">
<heading id="4938819C4F02303BFF34F8C7FCF8F88E" bold="true" box="[151,774,1833,1860]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<taxonomicName id="D5CF4D734F02303BFF34F8C7FCF8F88E" ID-CoL="6RHX6" authority="Franz" authorityName="Franz" box="[151,774,1833,1860]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="australiensis" subGenus="Scydmomicrus">
<emphasis id="20BBEAE24F02303BFF34F8C7FCF8F88E" bold="true" box="[151,774,1833,1860]" pageId="17" pageNumber="18">
<emphasis id="20BBEAE24F02303BFF34F8C7FEB9F889" bold="true" box="[151,327,1833,1859]" italics="true" pageId="17" pageNumber="18">Microscydmus</emphasis>
(
<emphasis id="20BBEAE24F02303BFEFBF8C7FDF7F889" bold="true" box="[344,521,1833,1859]" italics="true" pageId="17" pageNumber="18">Scydmomicrus</emphasis>
)
<emphasis id="20BBEAE24F02303BFDB9F8C7FD4DF889" bold="true" box="[538,691,1833,1859]" italics="true" pageId="17" pageNumber="18">australiensis</emphasis>
Franz
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="127036F04F02303BFF34F8A2FDFEF8AF" blockId="17.[151,774,1833,1893]" box="[151,512,1868,1893]" pageId="17" pageNumber="18">
(
<figureCitation id="8AF42A754F02303BFF3CF8A2FF02F8AF" box="[159,252,1868,1893]" captionStart="FIGURES 20 – 28" captionStartId="9.[151,264,1586,1609]" captionTargetBox="[159,1435,210,1555]" captionTargetId="figure@9.[151,1436,193,1565]" captionTargetPageId="9" captionText="FIGURES 20 – 28. Dorsal habitus of holotypes. Microscydmus (Scydmomicrus) australiensis Franz, female (20); M. (Sc.) nasicornis Franz, female (21); M. (Sc.) capitiseboraci sp. n., male (22); M. (Sc.) queenslandicus sp. n., male (23); M. (Sc.) styxianus sp. n., male (24); M. (Sc.) edithensis sp. n., male (25); M. (Sc.) tooloomensis sp. n., male (26); Penicillidmus masseyensis sp. n., male (27); P. unicolor sp. n., male (28)." httpUri="https://zenodo.org/record/285699/files/figure.png" pageId="17" pageNumber="18">Figs. 20</figureCitation>
,
<figureCitation id="8AF42A754F02303BFEABF8A3FED8F8AF" box="[264,294,1869,1893]" captionStart="FIGURES 29 – 37" captionStartId="10.[151,264,1512,1535]" captionTargetBox="[157,1418,212,1473]" captionTargetId="figure@10.[151,1436,193,1492]" captionTargetPageId="10" captionText="FIGURES 29 – 37. Body silhouette of holotypes. Microscydmus (Scydmomicrus) australiensis Franz, female (29); M. (Sc.) nasicornis Franz, female (30); M. (Sc.) capitiseboraci sp. n., male (31); M. (Sc.) queenslandicus sp. n., male (32); M. (Sc.) styxianus sp. n., male (33); M. (Sc.) edithensis sp. n., male (34); M. (Sc.) tooloomensis sp. n., male (35); Penicillidmus masseyensis sp. n., male (36); P. unicolor sp. n., male (37)." httpUri="https://zenodo.org/record/285700/files/figure.png" pageId="17" pageNumber="18">29</figureCitation>
,
<figureCitation id="8AF42A754F02303BFE91F8A3FEAFF8AF" box="[306,337,1869,1893]" captionStart="FIGURES 38 – 41" captionStartId="11.[151,264,1708,1731]" captionTargetBox="[166,1418,221,1674]" captionTargetId="figure@11.[151,1436,193,1687]" captionTargetPageId="11" captionText="FIGURES 38 – 41. Head capsule in dorsal (38, 40), dorso-posterior (39) and lateral (41) views. Microscydmus (Scydmomicrus) australiensis Franz, female (38); M. (Sc.) capitiseboraci sp. n., male (39); Penicillidmus sp., female (40 – 41). Abbreviations: cl, clypeus; fr, frons; gen, gena; lb, labrum; md, mandible; mn, mentum; mxp, maxillar palpus; nr, neck region; oc, occipital constriction; occ, occipital constriction; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/285701/files/figure.png" pageId="17" pageNumber="18">38</figureCitation>
,
<figureCitation id="8AF42A754F02303BFEFEF8A3FE85F8AF" box="[349,379,1869,1893]" captionStart="FIGURES 42 – 45" captionStartId="12.[151,264,1820,1843]" captionTargetBox="[182,1408,310,1798]" captionTargetId="figure@12.[171,1409,301,1808]" captionTargetPageId="12" captionText="FIGURES 42 – 45. Head capsule in ventral (42, 45), and antero-ventral (43 – 44) views. Microscydmus (Scydmomicrus) australiensis Franz, female (42); M. (Sc.) capitiseboraci sp. n., male (43); M. (Sc.) tooloomensis sp. n., male (44); Penicillidmus sp., female (45). Abbreviations: alc, anterior arm of longitudinal carina; bst, basistipes; cd, cardo; fo, foramen occipitale; fr, frons; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lb, labrum; lg, ligula; lp, labial palpus; md, mandible; mn, mentum; mst, mesostipes; mxp 1 – 4, maxillary palpomere I – IV; plc, posterior arm of longitudinal carina; ppf, palpifer; ptp, posterior tentorial pit; smn, submental carina; smn, submentum; tc, transverse carina." httpUri="https://zenodo.org/record/285702/files/figure.png" pageId="17" pageNumber="18">42</figureCitation>
,
<figureCitation id="8AF42A754F02303BFE25F8A2FE5AF8AF" box="[390,420,1868,1893]" captionStart="FIGURES 46 – 49" captionStartId="13.[151,264,1431,1454]" captionTargetBox="[154,1421,208,1405]" captionTargetId="figure@13.[151,1436,193,1411]" captionTargetPageId="13" captionText="FIGURES 46 – 49. Prothorax in ventral view. Microscydmus (Scydmomicrus) australiensis Franz, female (46); M. (Sc.) capitiseboraci sp. n., male (47); M. (Sc.) tooloomensis sp. n., male (48); Penicillidmus sp., female (49). Abbreviations: hy, hypomeron; hyr, hypomeral ridge; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pf, profurcal fovea; plp, postero-lateral brush; psp, prosternal intercoxal process." httpUri="https://zenodo.org/record/285703/files/figure.png" pageId="17" pageNumber="18">46</figureCitation>
,
<figureCitation id="8AF42A754F02303BFE13F8A2FE30F8AF" box="[432,462,1868,1893]" captionStart="FIGURES 50 – 53" captionStartId="15.[151,264,1823,1846]" captionTargetBox="[221,1330,199,1771]" captionTargetId="figure@15.[210,1365,193,1795]" captionTargetPageId="15" captionText="FIGURES 50 – 53. Pterothorax in ventral view. Microscydmus (Scydmomicrus) australiensis Franz, female (50); M. (Sc.) capitiseboraci sp. n., male (51); M. (Sc.) tooloomensis sp. n., male (52); Penicillidmus sp., female (53). Abbreviations: aest 2, mesanepisternum; ai, asetose impression of mesoventrite; ar, anterior ridge of mesoventrite; cl, coxal line; lmfa; lateral metafurcal arm; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pl, posterior lobe of mesocoxal projection; pre, prepectus; sp, spermatheca; v 3, metaventrite." httpUri="https://zenodo.org/record/285704/files/figure.png" pageId="17" pageNumber="18">50</figureCitation>
,
<figureCitation id="8AF42A754F02303BFE79F8A2FE09F8AF" box="[474,503,1868,1893]" captionStart="FIGURES 62 – 63" captionStartId="17.[151,264,1608,1631]" captionTargetBox="[151,1434,1055,1586]" captionTargetId="figure@17.[151,1436,1055,1586]" captionTargetPageId="17" captionText="FIGURES 62 – 63. Distribution of Microscydmus (Scydmomicrus subgen. n.) (62) and Penicillidmus gen. n. (63). Microscydmus (Scydmomicrus) australiensis Franz (62 a); M. (Sc.) nasicornis Franz (62 b); M. (Sc.) capitiseboraci sp. n. (2 c); M. (Sc.) queenslandicus sp. n. (62 d); M. (Sc.) styxianus sp. n. (62 e); M. (Sc.) edithensis sp. n. (62 f); M. (Sc.) tooloomensis sp. n. (62 g); Microscydmus (Sc.) sp. (62 h); Penicillidmus masseyensis sp. n. (63 a); P. unicolor sp. n. (63 b); Penicillidmus sp. (63 c)." httpUri="https://zenodo.org/record/285706/files/figure.png" pageId="17" pageNumber="18">62</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5AD5657B4F02303BFF34F87AFD95F860" box="[151,619,1940,1962]" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="127036F04F02303BFF34F87AFD95F860" blockId="17.[151,619,1940,1962]" box="[151,619,1940,1962]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="32DF11DE4F02303BFF34F87AFD95F860" box="[151,619,1940,1962]" pageId="17" pageNumber="18">
<taxonomicName id="D5CF4D734F02303BFF34F87AFD99F860" ID-CoL="6RHX6" authority="Franz, 1975: 271" authorityName="Franz" authorityPageNumber="271" authorityYear="1975" box="[151,615,1940,1962]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="20BBEAE24F02303BFF34F87AFE4CF860" box="[151,434,1940,1962]" italics="true" pageId="17" pageNumber="18">Microscydmus australiensis</emphasis>
<treatmentCitation id="936E10E14F02303BFE14F87BFD99F860" author="Franz" box="[439,615,1940,1962]" page="271" pageId="17" pageNumber="18" year="1975">
<bibRefCitation id="765E4B014F02303BFE14F87BFDCAF860" author="Franz" box="[439,564,1940,1962]" pageId="17" pageNumber="18" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312." type="journal article" year="1975">Franz, 1975</bibRefCitation>
: 271
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="5AD5657B4F023038FF34F839FB26FE8A" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="127036F04F02303BFF34F839FA63F825" blockId="17.[151,1437,2007,2032]" box="[151,1437,2007,2032]" pageId="17" pageNumber="18">
<emphasis id="20BBEAE24F02303BFF34F839FEA0F83A" bold="true" box="[151,350,2007,2032]" pageId="17" pageNumber="18">Material studied</emphasis>
.
<typeStatus id="CD7488524F02303BFECFF839FE27F83A" box="[364,473,2007,2032]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
: ♀: four labels: &quot;Trapped by sticky seeds / of
<taxonomicName id="D5CF4D734F02303BFC7CF839FB49F83A" box="[991,1207,2007,2032]" class="Magnoliopsida" family="Nyctaginaceae" genus="Pisonia" kingdom="Plantae" order="Caryophyllales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="brunoniana">Pisonia brunoniana</taxonomicName>
/ Cairns dist.,: F. P.
</paragraph>
<paragraph id="127036F04F013038FF34FF79FC3DFF1E" blockId="18.[151,1436,151,1184]" pageId="18" pageNumber="19">
Dodd&quot; [white, printed], &quot;
<taxonomicName id="D5CF4D734F013038FE1AFF76FDA5FF7A" box="[441,603,152,176]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="20BBEAE24F013038FE1AFF76FDA5FF7A" box="[441,603,152,176]" italics="true" pageId="18" pageNumber="19">Microscydmus</emphasis>
</taxonomicName>
/
<emphasis id="20BBEAE24F013038FDDBFF76FD14FF7A" box="[632,746,152,176]" italics="true" pageId="18" pageNumber="19">australiae</emphasis>
[sic!] m. / det. H. Franz&quot; [white, handwritten and printed], &quot;
<typeStatus id="CD7488524F013038FF03FF52FF17FF1E" box="[160,233,188,212]" pageId="18" pageNumber="19">Typus</typeStatus>
&quot; [red, handwritten], &quot;
<collectionCode id="74DEAE354F013038FE7AFF52FDD3FF1E" LSID="urn:lsid:biocol.org:col:34982" box="[473,557,188,212]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="18" pageNumber="19">SAMA</collectionCode>
Database / No. 25-037045&quot; (
<collectionCode id="74DEAE354F013038FCD6FF52FC49FF1E" box="[885,951,188,212]" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="18" pageNumber="19">SAM</collectionCode>
).
</paragraph>
<paragraph id="127036F04F013038FF64FF31FB26FE8A" blockId="18.[151,1436,151,1184]" pageId="18" pageNumber="19">
<emphasis id="20BBEAE24F013038FF64FF31FE54FF32" bold="true" box="[199,426,223,248]" pageId="18" pageNumber="19">Revised diagnosis.</emphasis>
Female: body uniformly yellowish-brown; frons subtriangular; punctures on head inconspicuous, only slightly more distinct than those on pronotum; BL &lt;
<quantity id="D5379B154F013038FC45FEEAFBB1FED6" box="[998,1103,260,285]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="19" unit="mm" value="0.6">0.60 mm</quantity>
; pronotum 1.2x broader than head, with ante-basal transverse groove; EI 1.50. Males and their diagnostic characters unknown.
</paragraph>
</subSubSection>
<subSubSection id="5AD5657B4F013038FF64FEA2FE76FC02" pageId="18" pageNumber="19" type="description">
<paragraph id="127036F04F013038FF64FEA2FE8DFE67" blockId="18.[151,1436,151,1184]" pageId="18" pageNumber="19">
<emphasis id="20BBEAE24F013038FF64FEA2FE8AFEAF" bold="true" box="[199,372,332,357]" pageId="18" pageNumber="19">Redescription.</emphasis>
Body of female (
<figureCitation id="8AF42A754F013038FDE8FEA2FD5FFEAE" box="[587,673,332,357]" captionStart="FIGURES 20 – 28" captionStartId="9.[151,264,1586,1609]" captionTargetBox="[159,1435,210,1555]" captionTargetId="figure@9.[151,1436,193,1565]" captionTargetPageId="9" captionText="FIGURES 20 – 28. Dorsal habitus of holotypes. Microscydmus (Scydmomicrus) australiensis Franz, female (20); M. (Sc.) nasicornis Franz, female (21); M. (Sc.) capitiseboraci sp. n., male (22); M. (Sc.) queenslandicus sp. n., male (23); M. (Sc.) styxianus sp. n., male (24); M. (Sc.) edithensis sp. n., male (25); M. (Sc.) tooloomensis sp. n., male (26); Penicillidmus masseyensis sp. n., male (27); P. unicolor sp. n., male (28)." httpUri="https://zenodo.org/record/285699/files/figure.png" pageId="18" pageNumber="19">Fig. 20</figureCitation>
) moderately convex, elongate and slender, with moderately long appendages, BL
<quantity id="D5379B154F013038FEFFFE81FE28FE4D" box="[348,470,367,392]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" pageId="18" pageNumber="19" unit="mm" value="0.525">0.525 mm</quantity>
; cuticle glossy, body uniformly yellowish-brown with slightly lighter appendages, vestiture yellowish.
</paragraph>
<paragraph id="127036F04F013038FF64FE59FC01FDAA" blockId="18.[151,1436,151,1184]" pageId="18" pageNumber="19">
Head (
<figureCitation id="8AF42A754F013038FEB7FE59FE96FE1A" box="[276,360,439,464]" captionStart="FIGURES 38 – 41" captionStartId="11.[151,264,1708,1731]" captionTargetBox="[166,1418,221,1674]" captionTargetId="figure@11.[151,1436,193,1687]" captionTargetPageId="11" captionText="FIGURES 38 – 41. Head capsule in dorsal (38, 40), dorso-posterior (39) and lateral (41) views. Microscydmus (Scydmomicrus) australiensis Franz, female (38); M. (Sc.) capitiseboraci sp. n., male (39); Penicillidmus sp., female (40 – 41). Abbreviations: cl, clypeus; fr, frons; gen, gena; lb, labrum; md, mandible; mn, mentum; mxp, maxillar palpus; nr, neck region; oc, occipital constriction; occ, occipital constriction; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/285701/files/figure.png" pageId="18" pageNumber="19">Fig. 38</figureCitation>
) subtrapezoidal, broadest at eyes, HL
<quantity id="D5379B154F013038FC8BFE59FC61FE05" box="[808,927,439,464]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" pageId="18" pageNumber="19" unit="mm" value="0.075">0.075 mm</quantity>
, HW
<quantity id="D5379B154F013038FC46FE59FBA3FE05" box="[997,1117,439,464]" metricMagnitude="-4" metricUnit="m" metricValue="1.25" pageId="18" pageNumber="19" unit="mm" value="0.125">0.125 mm</quantity>
; tempora shorter than eyes, relatively weakly convergent caudad; vertex weakly convex; frons subtriangular, with distinct antero-median projection. Eyes moderately large and moderately projecting laterally from the head silhouette, finely faceted. Punctures on head dorsum fine and sparse, inconspicuous; setae short, sparse and suberect. Antennae slender, AnL 0.200 mm; antennomeres I–II elongate, III–X transverse; XI as long as broad.
</paragraph>
<paragraph id="127036F04F013038FF64FD83FE5AFD3A" blockId="18.[151,1436,151,1184]" pageId="18" pageNumber="19">
Pronotum in dorsal view oval, broadest near anterior third, PL
<quantity id="D5379B154F013038FC33FD82FBFBFD4E" box="[912,1029,620,645]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="19" unit="mm" value="0.15">0.150 mm</quantity>
, PW
<quantity id="D5379B154F013038FBE7FD82FB44FD4E" box="[1092,1210,620,645]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="19" unit="mm" value="0.15">0.150 mm</quantity>
; anterior and lateral margins rounded; posterior margin shallowly bisinuate; base of pronotum with two pairs of small and shallow pits, internal pair connected by transverse impression. Punctures on pronotal disc fine and inconspicuous; setae sparse, thin, short and suberect.
</paragraph>
<paragraph id="127036F04F013038FF64FD12FDA3FC4A" blockId="18.[151,1436,151,1184]" pageId="18" pageNumber="19">
Elytra oval, slightly more convex than pronotum, broadest slightly anterior to middle, EL 0.300 mm, EW 0.200 mm, EI 1.500; basal impressions short; elytral apices separately rounded. Punctures on elytral disc fine but slightly more distinct and denser than those on pronotum; setae short, sparse and suberect.
<collectingCountry id="6AD876604F013038FB7BFCAAFAEEFC97" box="[1240,1296,836,861]" name="India" pageId="18" pageNumber="19">Hind</collectingCountry>
wings well developed, about twice as long as elytra.
</paragraph>
<paragraph id="127036F04F013038FF64FC63FCB0FC6F" blockId="18.[151,1436,151,1184]" box="[199,846,908,933]" pageId="18" pageNumber="19">Legs moderately long and slender, without modifications.</paragraph>
<paragraph id="127036F04F013038FF64FC41FE76FC02" blockId="18.[151,1436,151,1184]" box="[199,392,943,968]" pageId="18" pageNumber="19">
<emphasis id="20BBEAE24F013038FF64FC41FEFAFC02" bold="true" box="[199,260,943,968]" pageId="18" pageNumber="19">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="5AD5657B4F013038FF64FC3AFCCEFC27" box="[199,816,980,1005]" pageId="18" pageNumber="19" type="distribution">
<paragraph id="127036F04F013038FF64FC3AFCCEFC27" blockId="18.[151,1436,151,1184]" box="[199,816,980,1005]" pageId="18" pageNumber="19">
<emphasis id="20BBEAE24F013038FF64FC3AFEA7FC27" bold="true" box="[199,345,980,1005]" pageId="18" pageNumber="19">Distribution</emphasis>
(
<figureCitation id="8AF42A754F013038FECBFC3AFE42FC27" box="[360,444,980,1005]" captionStart="FIGURES 62 – 63" captionStartId="17.[151,264,1608,1631]" captionTargetBox="[151,1434,1055,1586]" captionTargetId="figure@17.[151,1436,1055,1586]" captionTargetPageId="17" captionText="FIGURES 62 – 63. Distribution of Microscydmus (Scydmomicrus subgen. n.) (62) and Penicillidmus gen. n. (63). Microscydmus (Scydmomicrus) australiensis Franz (62 a); M. (Sc.) nasicornis Franz (62 b); M. (Sc.) capitiseboraci sp. n. (2 c); M. (Sc.) queenslandicus sp. n. (62 d); M. (Sc.) styxianus sp. n. (62 e); M. (Sc.) edithensis sp. n. (62 f); M. (Sc.) tooloomensis sp. n. (62 g); Microscydmus (Sc.) sp. (62 h); Penicillidmus masseyensis sp. n. (63 a); P. unicolor sp. n. (63 b); Penicillidmus sp. (63 c)." httpUri="https://zenodo.org/record/285706/files/figure.png" pageId="18" pageNumber="19">Fig. 62</figureCitation>
a). NE
<collectingCountry id="6AD876604F013038FDAAFC3AFD8EFC27" box="[521,624,980,1005]" name="Australia" pageId="18" pageNumber="19">Australia</collectingCountry>
(N Queensland).
</paragraph>
</subSubSection>
<subSubSection id="5AD5657B4F013038FF64FC19FC49F958" pageId="18" pageNumber="19" type="discussion">
<paragraph id="127036F04F013038FF64FC19FAA9FB6A" blockId="18.[151,1436,151,1184]" pageId="18" pageNumber="19">
<emphasis id="20BBEAE24F013038FF64FC19FEC5FBDA" bold="true" box="[199,315,1015,1040]" pageId="18" pageNumber="19">Remarks.</emphasis>
<taxonomicName id="D5CF4D734F013038FEE2FC16FD87FBDA" box="[321,633,1016,1040]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="20BBEAE24F013038FEE2FC16FD87FBDA" box="[321,633,1016,1040]" italics="true" pageId="18" pageNumber="19">Microscydmus australiensis</emphasis>
</taxonomicName>
is known from a single female only, which is most similar to the female of
<taxonomicName id="D5CF4D734F013038FF1FFBF3FEA3FBFE" box="[188,349,1053,1076]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="nasicornis">
<emphasis id="20BBEAE24F013038FF1FFBF3FEA3FBFE" box="[188,349,1053,1076]" italics="true" pageId="18" pageNumber="19">M. nasicornis</emphasis>
</taxonomicName>
. Both species have lightly pigmented bodies, similar proportions of body parts, distinctly subtriangular frons and unremarkable, fine punctures on the head dorsum.
<taxonomicName id="D5CF4D734F013038FC79FBAEFAEDFB92" box="[986,1299,1088,1112]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="20BBEAE24F013038FC79FBAEFAEDFB92" box="[986,1299,1088,1112]" italics="true" pageId="18" pageNumber="19">Microscydmus australiensis</emphasis>
</taxonomicName>
is distinctly smaller than
<taxonomicName id="D5CF4D734F013038FE89FB8BFE3BFBB6" box="[298,453,1125,1148]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="nasicornis">
<emphasis id="20BBEAE24F013038FE89FB8BFE3BFBB6" box="[298,453,1125,1148]" italics="true" pageId="18" pageNumber="19">M. nasicornis</emphasis>
</taxonomicName>
(BL
<quantity id="D5379B154F013038FDA0FB8AFD88FBB6" box="[515,630,1124,1149]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" pageId="18" pageNumber="19" unit="mm" value="0.525">0.525 mm</quantity>
vs.
<quantity id="D5379B154F013038FD05FB8AFCE4FBB6" box="[678,794,1124,1149]" metricMagnitude="-4" metricUnit="m" metricValue="6.25" pageId="18" pageNumber="19" unit="mm" value="0.625">0.625 mm</quantity>
) and has broader head in relation to pronotum (PW/HW 1.2 vs. 1.4).
<taxonomicName id="D5CF4D734F013038FE81FB66FDA4FB6A" box="[290,602,1160,1184]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="20BBEAE24F013038FE81FB66FDA4FB6A" box="[290,602,1160,1184]" italics="true" pageId="18" pageNumber="19">Microscydmus australiensis</emphasis>
</taxonomicName>
differs from other Australian congeners in the following characters:
</paragraph>
<paragraph id="127036F04F013038FF34FB24FC45FA70" blockId="18.[151,1436,1225,1682]" pageId="18" pageNumber="19">
- from
<taxonomicName id="D5CF4D734F013038FEABFB24FE2EFB2B" box="[264,464,1225,1249]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="capitiseboraci" status="sp. nov.">
<emphasis id="20BBEAE24F013038FEABFB24FE2EFB2B" box="[264,464,1225,1249]" italics="true" pageId="18" pageNumber="19">M. capitiseboraci</emphasis>
</taxonomicName>
<emphasis id="20BBEAE24F013038FE78FB27FDE0FB28" bold="true" box="[475,542,1225,1250]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="3B8857994F013038FE78FB27FDE0FB28" box="[475,542,1225,1250]" pageId="18" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="D5CF4D734F013038FD8CFB24FCFAFB2B" box="[559,772,1225,1249]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="queenslandicus" status="sp. nov.">
<emphasis id="20BBEAE24F013038FD8CFB24FCFAFB2B" box="[559,772,1225,1249]" italics="true" pageId="18" pageNumber="19">M. queenslandicus</emphasis>
</taxonomicName>
<emphasis id="20BBEAE24F013038FCADFB27FCAFFB28" bold="true" box="[782,849,1225,1250]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="3B8857994F013038FCADFB27FCAFFB28" box="[782,849,1225,1250]" pageId="18" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="D5CF4D734F013038FC2DFB24FBE1FB2B" box="[910,1055,1226,1249]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="styxianus" status="sp. nov.">
<emphasis id="20BBEAE24F013038FC2DFB24FBE1FB2B" box="[910,1055,1226,1249]" italics="true" pageId="18" pageNumber="19">M. styxianus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3B8857994F013038FB89FB27FB92FB28" box="[1066,1132,1225,1250]" pageId="18" pageNumber="19" rank="species">
<emphasis id="20BBEAE24F013038FB89FB27FB92FB28" bold="true" box="[1066,1132,1225,1250]" pageId="18" pageNumber="19">sp. n.</emphasis>
</taxonomicNameLabel>
in the frons covered with inconspicuous punctures and more distinctly narrowing anteriorly; in these three species punctures on the head dorsum are distinct and the frons is shorter and less narrowed (
<taxonomicName id="D5CF4D734F013038FC16FAFCFB7AFAE3" box="[949,1156,1297,1321]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="capitiseboraci">
<emphasis id="20BBEAE24F013038FC16FAFCFB7AFAE3" box="[949,1156,1297,1321]" italics="true" pageId="18" pageNumber="19">M. capitiseboraci</emphasis>
</taxonomicName>
), broadly rounded (
<taxonomicName id="D5CF4D734F013038FADEFAFCFEBDFA87" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="20BBEAE24F013038FADEFAFCFEBDFA87" italics="true" pageId="18" pageNumber="19">M. queenslandicus</emphasis>
</taxonomicName>
) or more subtrapezoidal than subtriangular (
<taxonomicName id="D5CF4D734F013038FCF7FAD8FC1BFA87" box="[852,997,1334,1357]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="styxianus">
<emphasis id="20BBEAE24F013038FCF7FAD8FC1BFA87" box="[852,997,1334,1357]" italics="true" pageId="18" pageNumber="19">M. styxianus</emphasis>
</taxonomicName>
); additionally in
<taxonomicName id="D5CF4D734F013038FB16FAD8FA90FA87" box="[1205,1390,1333,1357]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="20BBEAE24F013038FB16FAD8FA90FA87" box="[1205,1390,1333,1357]" italics="true" pageId="18" pageNumber="19">M. australiensis</emphasis>
</taxonomicName>
the pronotum is as long as broad (slightly broader than long in
<taxonomicName id="D5CF4D734F013038FC89FAB4FC11FABB" box="[810,1007,1369,1393]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="capitiseboraci">
<emphasis id="20BBEAE24F013038FC89FAB4FC11FABB" box="[810,1007,1369,1393]" italics="true" pageId="18" pageNumber="19">M. capitiseboraci</emphasis>
</taxonomicName>
); the elytral index is low (EI 1.500 vs.
<quantity id="D5379B154F013038FF34FA90FF0DFA5C" box="[151,243,1405,1430]" metricMagnitude="-2" metricUnit="m" metricValue="4.3434" pageId="18" pageNumber="19" unit="in" value="1.71">1.710 in</quantity>
<taxonomicName id="D5CF4D734F013038FF58FA90FE3FFA5F" box="[251,449,1405,1429]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="capitiseboraci">
<emphasis id="20BBEAE24F013038FF58FA90FE3FFA5F" box="[251,449,1405,1429]" italics="true" pageId="18" pageNumber="19">M. capitiseboraci</emphasis>
</taxonomicName>
); and the ratio EL/PL is also remarkably low (2.000 vs.
<quantity id="D5379B154F013038FBE2FA90FB60FA5C" box="[1089,1182,1405,1430]" metricMagnitude="-2" metricUnit="m" metricValue="6.096" pageId="18" pageNumber="19" unit="in" value="2.4">2.400 in</quantity>
<taxonomicName id="D5CF4D734F013038FB06FA90FA92FA5F" box="[1189,1388,1405,1429]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="capitiseboraci">
<emphasis id="20BBEAE24F013038FB06FA90FA92FA5F" box="[1189,1388,1405,1429]" italics="true" pageId="18" pageNumber="19">M. capitiseboraci</emphasis>
</taxonomicName>
and in
<taxonomicName id="D5CF4D734F013038FF17FA4CFE7BFA73" box="[180,389,1441,1465]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="20BBEAE24F013038FF17FA4CFE7BFA73" box="[180,389,1441,1465]" italics="true" pageId="18" pageNumber="19">M. queenslandicus</emphasis>
</taxonomicName>
, while in
<taxonomicName id="D5CF4D734F013038FE56FA4CFD7DFA73" box="[501,643,1442,1465]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="styxianus">
<emphasis id="20BBEAE24F013038FE56FA4CFD7DFA73" box="[501,643,1442,1465]" italics="true" pageId="18" pageNumber="19">M. styxianus</emphasis>
</taxonomicName>
EL/PL is as high as 2.540);
</paragraph>
<paragraph id="127036F04F013038FF34FA28FE8DF980" blockId="18.[151,1436,1225,1682]" pageId="18" pageNumber="19">
- from
<taxonomicName id="D5CF4D734F013038FEA4FA28FE61FA17" box="[263,415,1477,1501]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="edithensis" status="sp. nov.">
<emphasis id="20BBEAE24F013038FEA4FA28FE61FA17" box="[263,415,1477,1501]" italics="true" pageId="18" pageNumber="19">M. edithensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3B8857994F013038FE0AFA2BFE15FA14" box="[425,491,1477,1502]" pageId="18" pageNumber="19" rank="species">
<emphasis id="20BBEAE24F013038FE0AFA2BFE15FA14" bold="true" box="[425,491,1477,1502]" pageId="18" pageNumber="19">sp. n.</emphasis>
</taxonomicNameLabel>
in the uniformly yellowish-brown body pigmentation (
<taxonomicName id="D5CF4D734F013038FBC9FA28FAFCFA17" box="[1130,1282,1477,1501]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="edithensis">
<emphasis id="20BBEAE24F013038FBC9FA28FAFCFA17" box="[1130,1282,1477,1501]" italics="true" pageId="18" pageNumber="19">M. edithensis</emphasis>
</taxonomicName>
has the head distinctly darker than the pronotum and elytra); elytral sculpture (fine, inconspicuous punctures in
<taxonomicName id="D5CF4D734F013038FB44FA04FA62F9CB" box="[1255,1436,1513,1537]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="australiensis">
<emphasis id="20BBEAE24F013038FB44FA04FA62F9CB" box="[1255,1436,1513,1537]" italics="true" pageId="18" pageNumber="19">M. australiensis</emphasis>
</taxonomicName>
vs. dense and fine punctures forming fine transverse wrinkles in
<taxonomicName id="D5CF4D734F013038FCCEF9E0FBFDF9EF" box="[877,1027,1549,1573]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="edithensis">
<emphasis id="20BBEAE24F013038FCCEF9E0FBFDF9EF" box="[877,1027,1549,1573]" italics="true" pageId="18" pageNumber="19">M. edithensis</emphasis>
</taxonomicName>
); and distinctly smaller body (
<quantity id="D5379B154F013038FAF8F9E3FF3DF983" metricMagnitude="-4" metricUnit="m" metricValue="5.25" pageId="18" pageNumber="19" unit="mm" value="0.525">0.525 mm</quantity>
vs.
<quantity id="D5379B154F013038FF53F9DFFE98F980" box="[240,358,1585,1610]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="18" pageNumber="19" unit="mm" value="0.65">0.650 mm</quantity>
);
</paragraph>
<paragraph id="127036F04F013038FF34F9B8FC49F958" blockId="18.[151,1436,1225,1682]" pageId="18" pageNumber="19">
- from
<taxonomicName id="D5CF4D734F013038FEA5F9B8FE3FF9A7" box="[262,449,1621,1645]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="tooloomensis">
<emphasis id="20BBEAE24F013038FEA5F9B8FE3FF9A7" box="[262,449,1621,1645]" italics="true" pageId="18" pageNumber="19">M. tooloomensis</emphasis>
</taxonomicName>
in the presence of the transverse ante-basal impression on the pronotum (absent in
<taxonomicName id="D5CF4D734F013038FADEF9B8FED5F95B" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="tooloomensis">
<emphasis id="20BBEAE24F013038FADEF9B8FED5F95B" italics="true" pageId="18" pageNumber="19">M. tooloomensis</emphasis>
</taxonomicName>
); and slightly shorter body (BL
<quantity id="D5379B154F013038FD36F997FCF9F95B" box="[661,775,1657,1682]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" pageId="18" pageNumber="19" unit="mm" value="0.525">0.525 mm</quantity>
vs. 0.600 mm).
</paragraph>
</subSubSection>
</treatment>
</document>
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