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<document id="0C80069ED461A89562B398623F6375BF" ID-CLB-Dataset="301609" ID-DOI="10.11646/zootaxa.5491.1.1" ID-GBIF-Dataset="34c72ca9-f976-4c79-a123-f93af60fa26c" ID-ISSN="1175-5334" ID-Zenodo-Dep="13211849" ID-ZooBank="0612E62C-A53D-42BE-8578-68EC77D34627" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1722790779731" checkinUser="plazi" docAuthor="Ahrens, Dirk, Zhao, Ming-Zhi, Pham, Phu Van &amp; Liu, Wan-Gang" docDate="2024" docId="9A5F87E8FFDB7C6AFF4DAC39EE45FD0E" docLanguage="en" docName="zootaxa.5491.1.1.pdf" docOrigin="Zootaxa 5491 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5491.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Serica (Serica) gwangjuensis Ahrens, Zhao, Pham &amp; Liu 2024, new species" docType="treatment" docVersion="7" lastPageNumber="8" masterDocId="6666FF90FFDD7C6DFFDAAD56ED2BFFFF" masterDocTitle="Taxonomic updates on Pachyserica Brenske, 1898 and Serica MacLeay, 1819 reveal 38 new species and new challenges of Sericini systematics regarding DNA barcodes and genus-level diagnostic key characters (Coleoptera: Scarabaeidae: Sericinae)" masterLastPageNumber="89" masterPageNumber="1" pageNumber="7" updateTime="1732490179605" updateUser="ExternalLinkService">
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<mods:title id="CC27F946922C173C74A819CE96EB7138">Taxonomic updates on Pachyserica Brenske, 1898 and Serica MacLeay, 1819 reveal 38 new species and new challenges of Sericini systematics regarding DNA barcodes and genus-level diagnostic key characters (Coleoptera: Scarabaeidae: Sericinae)</mods:title>
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<mods:namePart id="5C9CABAE2B3922BFC3E85F401E6B2DAD">Ahrens, Dirk</mods:namePart>
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<mods:affiliation id="DA5DB88BA90F7CA02DC0F38659E31F99">Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 127, 53113 Bonn, Germany ahrens. dirk _ col @ gmx. de; https: // orcid. org / 0000 - 0003 - 3524 - 7153</mods:affiliation>
<mods:nameIdentifier id="CA152EA279F2055AB7996A111C694528" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
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<mods:namePart id="972DCC798FE7732342C2C6828402CCAF">Zhao, Ming-Zhi</mods:namePart>
<mods:nameIdentifier id="BCD8F601656F7C1A6E95A4DCDAF582C6" type="ORCID">0000-0002-3158-6177</mods:nameIdentifier>
<mods:affiliation id="C48D98FFE0CF0478E66CAF4C0AA01DE2">College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou 510642, P. R. China zhaomzhai @ 126. com; https: // orcid. org / 0000 - 0002 - 3158 - 6177</mods:affiliation>
<mods:nameIdentifier id="8FEF9E74B865A13B18BB8EB2B2E0CFBD" type="email">zhaomzhai@126.com</mods:nameIdentifier>
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<mods:namePart id="FDA910A96A1F76EB8E87F72CBA3AF292">Pham, Phu Van</mods:namePart>
<mods:nameIdentifier id="D3630001BC6113EEAFF5FD00B88C932C" type="ORCID">0000-0001-9369-6649</mods:nameIdentifier>
<mods:affiliation id="9898FA07D3187B51EB559404E94C0B2E">Institute of Ecology and Biological resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam phupham. iebr @ gmail. com; https: // orcid. org / 0000 - 0001 - 9369 - 6649</mods:affiliation>
<mods:nameIdentifier id="B16E59F8B6F939CF20EB8CCA67CB9977" type="email">phupham.iebr@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FD92BCE3363F5AC9F7400462B8CC3DEA">Liu, Wan-Gang</mods:namePart>
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<mods:affiliation id="70907466D67BD569F2F51336010057B6">Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 127, 53113 Bonn, Germany ahrens. dirk _ col @ gmx. de; https: // orcid. org / 0000 - 0003 - 3524 - 7153 &amp; SKLLQG, Institute of Earth and Environment, Chinese Academy of Sciences, Yanxiang Road 97, Yanta District, Xi’an 710061 P. R. China liuwangang @ ieecas. cn; https: // orcid. org / 0000 - 0003 - 4788 - 7967 &amp; Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 127, 53113 Bonn, Germany ahrens. dirk _ col @ gmx. de; https: // orcid. org / 0000 - 0003 - 3524 - 7153</mods:affiliation>
<mods:nameIdentifier id="C9E258F581EDD7D23AB6E966E197D6BC" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
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<mods:date id="5C756578075CFA0087A427864D36EA8F">2024</mods:date>
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<treatment id="9A5F87E8FFDB7C6AFF4DAC39EE45FD0E" ID-DOI="http://doi.org/10.5281/zenodo.13212355" ID-GBIF-Taxon="238836485" ID-Zenodo-Dep="13212355" LSID="urn:lsid:plazi:treatment:9A5F87E8FFDB7C6AFF4DAC39EE45FD0E" httpUri="http://treatment.plazi.org/id/9A5F87E8FFDB7C6AFF4DAC39EE45FD0E" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" scope_family="Scarabaeidae" scope_order="Coleoptera" scope_tribe="Sericini">
<subSubSection id="5AEC6575FFDB7C6BFF4DAC39EED8FE76" box="[151,1011,367,393]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="124936FEFFDB7C6BFF4DAC39EED8FE76" blockId="6.[151,1011,367,429]" box="[151,1011,367,393]" pageId="6" pageNumber="7">
<heading id="49018192FFDB7C6BFF4DAC39EED8FE76" bold="true" box="[151,1011,367,393]" fontSize="11" level="2" pageId="6" pageNumber="7" reason="4">
<emphasis id="2082EAECFFDB7C6BFF4DAC39EED8FE76" bold="true" box="[151,1011,367,393]" pageId="6" pageNumber="7">
<taxonomicName id="D5F64D7DFFDB7C6BFF4DAC39EE7DFE76" ID-CoL="CHNKH" authority="Ahrens, Zhao, Pham &amp; Liu" authorityName="Ahrens, Zhao, Pham &amp; Liu" authorityYear="2024" box="[151,854,367,393]" class="Insecta" family="Scarabaeidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="gwangjuensis" status="sp. nov." subGenus="Serica">
<emphasis id="2082EAECFFDB7C6BFF4DAC39EDCFFE76" bold="true" box="[151,228,367,393]" italics="true" pageId="6" pageNumber="7">Serica</emphasis>
(
<emphasis id="2082EAECFFDB7C6BFF2FAC39EC69FE76" bold="true" box="[245,322,367,393]" italics="true" pageId="6" pageNumber="7">Serica</emphasis>
)
<emphasis id="2082EAECFFDB7C6BFE89AC39ECD3FE76" bold="true" box="[339,504,367,393]" italics="true" pageId="6" pageNumber="7">gwangjuensis</emphasis>
Ahrens, Zhao, Pham &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="3BB15797FFDB7C6BFCB9AC39EED8FE76" box="[867,1011,367,393]" pageId="6" pageNumber="7" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5AEC6575FFDB7C6BFF4DACC5EC27FE52" box="[151,268,403,429]" pageId="6" pageNumber="7" type="description">
<paragraph id="124936FEFFDB7C6BFF4DACC5EC27FE52" blockId="6.[151,1011,367,429]" box="[151,268,403,429]" pageId="6" pageNumber="7">
<figureCitation id="8ACD2A7BFFDB7C6BFF4DACC5EC27FE52" box="[151,268,403,429]" captionStart="FIGURE 1" captionStartId="66.[152,255,1943,1967]" captionTargetBox="[212,1376,192,1910]" captionTargetId="figure-21@66.[202,1386,181,1920]" captionTargetPageId="66" captionText="FIGURE 1. A–E: Serica gwangjuensis new species (holotype); F–J: S. assingi new species (holotype); A, H: aedeagus, left side lateral view; C, J: aedeagus, right side lateral view; B, I: parameres, dorsal view; D, F: habitus, dorsal view; E, G: habitus, lateral view. Scale bar: 0.5 mm. Habitus images not to scale." figureDoi="http://doi.org/10.5281/zenodo.13211853" httpUri="https://zenodo.org/record/13211853/files/figure.png" pageId="6" pageNumber="7">Fig. 1A–E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="5AEC6575FFDB7C6BFF4DAC8DECBEFD7A" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="124936FEFFDB7C6BFF4DAC8DECBEFD7A" blockId="6.[151,1437,475,2014]" pageId="6" pageNumber="7">
<emphasis id="2082EAECFFDB7C6BFF4DAC8DECE6FE0A" bold="true" box="[151,461,475,501]" pageId="6" pageNumber="7">Type material examined.</emphasis>
<materialsCitation id="A29E3CA3FFDB7C6BFE01AC8DEF31FDE6" ID-GBIF-Occurrence="4923345321" collectingDate="2010-06-07" collectionCode="ZFMK" collectorName="T. Kolkebeck" country="South Korea" location="Gwangju" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Gwangju" typeStatus="holotype">
<typeStatus id="CD4D885CFFDB7C6BFE01AC8DEF68FE0A" box="[475,579,475,501]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
<specimenCount id="04F0FD77FFDB7C6BFD88AC8AEF4CFE09" box="[594,615,476,502]" count="1" pageId="6" pageNumber="7" type="male">♂</specimenCount>
“
<date id="6648103EFFDB7C6BFDA4AC8DEFD4FE0A" box="[638,767,475,501]" pageId="6" pageNumber="7" value="2010-06-07">
<collectingDate id="760CE9D6FFDB7C6BFDA4AC8DEFD4FE0A" box="[638,767,475,501]" pageId="6" pageNumber="7" value="2010-06-07">06.07.2010</collectingDate>
</date>
Mudeungsan,
<collectingRegion id="D032F81CFFDB7C6BFC69AC8DE931FE0A" box="[947,1050,475,501]" country="South Korea" name="Gwangju" pageId="6" pageNumber="7">Gwangju</collectingRegion>
(
<collectingCountry id="6AE1766EFFDB7C6BFBF5AC8DE9B7FE0A" box="[1071,1180,475,501]" name="South Korea" pageId="6" pageNumber="7">Südkorea</collectingCountry>
) leg.
<collectorName id="BF035328FFDB7C6BFB33AC8AE8ADFE0A" box="[1257,1414,475,502]" pageId="6" pageNumber="7">T. Kölkebeck</collectorName>
/ Asia
<taxonomicName id="D5F64D7DFFDB7C6BFF0EACA9EC00FDE6" authorityName="Kirby" authorityYear="1837" box="[212,299,511,537]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="66" phylum="Arthropoda" rank="tribe" tribe="Sericini">Sericini</taxonomicName>
spec. 862” (
<collectionCode id="74E7AE3BFFDB7C6BFE67ACA9EF3AFDE6" box="[445,529,511,537]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="6" pageNumber="7" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="A29E3CA3FFDB7C6BFDFCAF56ECA1FDC2" ID-GBIF-Occurrence="4923345418" collectingDate="2010-06-07" collectionCode="ZFMK" collectorName="T. Kolkebeck" country="South Korea" location="Mudeungsan" pageId="6" pageNumber="7" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Gwangju" typeStatus="paratype">
<typeStatus id="CD4D885CFFDB7C6BFDFCAF56EFBCFDE5" box="[550,663,512,538]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
:
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,
<specimenCount id="04F0FD77FFDB7C6BFD3AACA9EE0AFDE5" box="[736,801,511,538]" count="3" pageId="6" pageNumber="7" type="female">3 ♀♀</specimenCount>
“
<date id="6648103EFFDB7C6BFCEEACA9EE9EFDE6" box="[820,949,511,537]" pageId="6" pageNumber="7" value="2010-06-07">
<collectingDate id="760CE9D6FFDB7C6BFCEEACA9EE9EFDE6" box="[820,949,511,537]" pageId="6" pageNumber="7" value="2010-06-07">06.07.2010</collectingDate>
</date>
<location id="17296025FFDB7C6BFC64ACA9E979FDE6" LSID="urn:lsid:plazi:treatment:9A5F87E8FFDB7C6AFF4DAC39EE45FD0E:17296025FFDB7C6BFC64ACA9E979FDE6" box="[958,1106,511,537]" country="South Korea" name="Mudeungsan" pageId="6" pageNumber="7" stateProvince="Gwangju">Mudeungsan</location>
,
<collectingRegion id="D032F81CFFDB7C6BFB85ACA9E9EDFDE6" box="[1119,1222,511,537]" country="South Korea" name="Gwangju" pageId="6" pageNumber="7">Gwangju</collectingRegion>
(
<collectingCountry id="6AE1766EFFDB7C6BFB0CACA9E868FDE6" box="[1238,1347,511,537]" name="South Korea" pageId="6" pageNumber="7">Südkorea</collectingCountry>
) leg.
<collectorName id="BF035328FFDB7C6BFA5CAF56EC38FDC2" pageId="6" pageNumber="7">T. Kölkebeck</collectorName>
” (
<collectionCode id="74E7AE3BFFDB7C6BFEF5AF75ECA8FDC2" box="[303,387,547,573]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="6" pageNumber="7" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="A29E3CA3FFDB7C6BFE42AF75E84AFDC2" ID-GBIF-Occurrence="4923345370" box="[408,1377,547,574]" collectingDate="2010-06-27" collectionCode="ZFMK" collectorName="T. Kolkebeck" country="South Korea" location="Beomeosa" pageId="6" pageNumber="7" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Busan" typeStatus="paratype">
<specimenCount id="04F0FD77FFDB7C6BFE42AF75ECF7FDC1" box="[408,476,547,574]" count="4" pageId="6" pageNumber="7" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="04F0FD77FFDB7C6BFE31AF75EF33FDC1" box="[491,536,547,574]" count="1" pageId="6" pageNumber="7" type="female">1 ♀</specimenCount>
“
<date id="6648103EFFDB7C6BFDF6AF75EF9DFDC2" box="[556,694,547,573]" pageId="6" pageNumber="7" value="2010-06-27">
<collectingDate id="760CE9D6FFDB7C6BFDF6AF75EF9DFDC2" box="[556,694,547,573]" pageId="6" pageNumber="7" value="2010-06-27">27.06. 2010</collectingDate>
</date>
<location id="17296025FFDB7C6BFD1AAF72EE13FDC1" LSID="urn:lsid:plazi:treatment:9A5F87E8FFDB7C6AFF4DAC39EE45FD0E:17296025FFDB7C6BFD1AAF72EE13FDC1" box="[704,824,548,574]" country="South Korea" name="Beomeosa" pageId="6" pageNumber="7" stateProvince="Busan">Beomeosa</location>
,
<collectingRegion id="D032F81CFFDB7C6BFC9CAF72EEA7FDC1" box="[838,908,548,574]" country="South Korea" name="Busan" pageId="6" pageNumber="7">Busan</collectingRegion>
(
<collectingCountry id="6AE1766EFFDB7C6BFC47AF75E921FDC2" box="[925,1034,547,573]" name="South Korea" pageId="6" pageNumber="7">Südkorea</collectingCountry>
) leg.
<collectorName id="BF035328FFDB7C6BFB94AF72E9C1FDC2" box="[1102,1258,547,574]" pageId="6" pageNumber="7">T. Kölkebeck</collectorName>
” (
<collectionCode id="74E7AE3BFFDB7C6BFADCAF75E871FDC2" box="[1286,1370,547,573]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="6" pageNumber="7" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="A29E3CA3FFDB7C6BFAB5AF75ECA4FD7A" ID-GBIF-Occurrence="4923345302" collectingDate="2018-06-15" collectionCode="ZFMK" country="South Korea" elevation="581" location="Walter Grosser" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Gyeongnam" typeStatus="paratype">
<specimenCount id="04F0FD77FFDB7C6BFAB5AF75E8B7FDC1" box="[1391,1436,547,574]" count="1" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
“
<collectingCountry id="6AE1766EFFDB7C6BFF7BAF11EC56FD9D" box="[161,381,583,610]" name="South Korea" pageId="6" pageNumber="7">Republic of Korea</collectingCountry>
<collectingRegion id="D032F81CFFDB7C6BFE50AF11EF47FD9E" box="[394,620,583,609]" country="South Korea" name="Gyeongnam" pageId="6" pageNumber="7">Gyeongsangnam-do</collectingRegion>
Jirisan N.P. N35°22.241′, E127°46.853′ 12.–
<date id="6648103EFFDB7C6BFBA5AF11E9DEFD9D" box="[1151,1269,583,610]" pageId="6" pageNumber="7" value="2018-06-15">
<collectingDate id="760CE9D6FFDB7C6BFBA5AF11E9DEFD9D" box="[1151,1269,583,610]" pageId="6" pageNumber="7" value="2018-06-15">15.6.2018</collectingDate>
</date>
,
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<elevation id="99DBD1CDFFDB7C6BFADCAF11E86DFD9D" box="[1286,1350,583,610]" metricMagnitude="2" metricUnit="m" metricValue="5.81" pageId="6" pageNumber="7" unit="m" value="581.0">581m</elevation>
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<location id="17296025FFDB7C6BFA88AF11EDC5FD7A" LSID="urn:lsid:plazi:treatment:9A5F87E8FFDB7C6AFF4DAC39EE45FD0E:17296025FFDB7C6BFA88AF11EDC5FD7A" country="South Korea" name="Walter Grosser" pageId="6" pageNumber="7" stateProvince="Gyeongnam">Walter Grosser</location>
lgt.” (
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)
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.
</paragraph>
</subSubSection>
<subSubSection id="5AEC6575FFDB7C6AFF1DAFD9EF81FE9A" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="description">
<paragraph id="124936FEFFDB7C6BFF1DAFD9EC13FD0D" blockId="6.[151,1437,475,2014]" pageId="6" pageNumber="7">
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Description of
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.
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Length:
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, length of elytra:
<quantity id="D50E9B1BFFDB7C6BFCAEAFD9EEE6FD55" box="[884,973,655,682]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="6" pageNumber="7" unit="mm" value="5.5">5.5 mm</quantity>
, width:
<quantity id="D50E9B1BFFDB7C6BFBF0AFD9E9AFFD55" box="[1066,1156,655,682]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="6" pageNumber="7" unit="mm" value="4.1">4.1 mm</quantity>
. Body oblong, including antenna and legs yellowish brown, eyes and frons blackish, elytra without dark spots, dorsal surface shiny and finely shortly setose.
</paragraph>
<paragraph id="124936FEFFDB7C6BFF1DAFADEEDFFB5E" blockId="6.[151,1437,475,2014]" pageId="6" pageNumber="7">
Labroclypeus little wider than long, widest at base, lateral margins convex and strongly convergent, anterior angles almost blunt, anterior margin deeply but narrowly emarginate medially, anterior and lateral margins weakly reflexed; surface flat and shiny, finely and densely punctate, distance between punctures smaller than their diameter, partly producing some very superficial transverse wrinkles; surface throughout with dense, long, erect setae; frontoclypeal suture indistinctly incised, slightly elevated and weakly curved; smooth area anterior to eye moderately large and weakly convex, approximately 1.2 times as wide as long; ocular canthus moderately long and slender (1/4 of ocular diameter), smooth, with one very short terminal seta. Frons flat and shiny; surface with fine and dense punctures and fine, dense, erect pilosity as in labroclypeus and the rest of the body surface. Eyes moderately large, ratio diameter/interocular width: 0.65. Antenna with nine antennomeres; antennomeres three and four slightly longer than wide, antennomeres five and six transverse and short; club with three antennomeres, 2.3 times as long as remaining antennomeres combined and reflexed.
<collectingRegion id="D032F81CFFDB7C6BFC54A932EEC6FB81" box="[910,1005,1124,1150]" country="Myanmar" name="Chin" pageId="6" pageNumber="7">Mentum</collectingRegion>
weakly elevated, anteriorly flattened. Labrum transverse, short, moderately produced, deeply emarginate medially.
</paragraph>
<paragraph id="124936FEFFDB7C6BFF1DA9FAEF40FA85" blockId="6.[151,1437,475,2014]" pageId="6" pageNumber="7">Pronotum subtrapezoidal, wider than long, widest at base, lateral margins moderately evenly convex and strongly convergent anteriorly, anterior angles weakly produced, blunt, posterior angles blunt, weakly rounded at tip; anterior margin medially with a complete but fine marginal line and strongly convexly produced; surface densely and finely punctate, with fine, dense, posteriorly bent setae; anterior and lateral margins with moderately long and sparse setae; hypomeron carinate at base. Scutellum slender and long, triangular, apex well rounded, finely and evenly, densely punctate, with short setae as in pronotum.</paragraph>
<paragraph id="124936FEFFDB7C6BFF1DA8D5EF5EF9D2" blockId="6.[151,1437,475,2014]" pageId="6" pageNumber="7">Elytra oblong, widest at apical third, striae strongly impressed, finely and densely punctate, intervals convex, with fine, dense punctures concentrated along striae, uniformly yellow, with the sutural and epipleura slightly darker; punctures on intervals with fine, short setae which are bent posteriorly; epipleural edge fine, ending at moderately curved external apical angle of elytra, epipleura densely setose, apical border chitinous, without membranous rim of fine microtrichomes (magnification 100x).</paragraph>
<paragraph id="124936FEFFDB7C6BFF1DAB61ECF3F8FA" blockId="6.[151,1437,475,2014]" pageId="6" pageNumber="7">Ventral surface moderately dull, partly weakly shiny, finely and densely punctate, moderately densely setose; metacoxa glabrous, with a few long setae only laterally; abdominal sternites finely and densely punctate, with a transverse row of coarse punctures, each bearing a short seta. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.54. Pygidium weakly convex and weakly shiny, finely and densely punctate, smooth midline lacking, with partly dense, fine and short setae (similar to those in the rest of dorsal surface).</paragraph>
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Legs slender; femora with two longitudinal rows of setae, between rows finely and not densely punctate; metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior margin, ventral posterior margin serrated in apical half and not widened, dorsal posterior margin completely serrated, in basal half with a few long setae which are more than half as long as width of metafemur. Metatibia slender and long, widest at apex, ratio of width/length: 1/4.6; dorsal margin sharply carinate, with two groups of spines, basal group at first third, apical group at two thirds of metatibial length, basally with a few single, robust setae; lateral face longitudinally concave, superficially punctate, entirely covered with irregular wrinkles; ventral margin serrated, with two widely separated robust setae; medial face with a shallow longitudinal groove medially, sparsely punctate and glabrous; apex interiorly near tarsal articulation distinctly but bluntly truncate. Tarsomeres ventrally with sparse, very minute setae, meso- and metatarsomeres missing in the
<typeStatus id="CD4D885CFFDA7C6AFD75AC55EE3EFEE2" box="[687,789,259,285]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
; protibia moderately long, bidentate, external margin with a few small teeth in base half; anterior claws asymmetrical, basal tooth of inner claw small and lobiform.
</paragraph>
<paragraph id="124936FEFFDA7C6AFF1DAC1DEF81FE9A" blockId="7.[151,1437,151,753]" box="[199,682,331,357]" pageId="7" pageNumber="8">
Aedeagus:
<figureCitation id="8ACD2A7BFFDA7C6AFE9EAC1DEC96FE9A" box="[324,445,331,357]" captionStart="FIGURE 1" captionStartId="66.[152,255,1943,1967]" captionTargetBox="[212,1376,192,1910]" captionTargetId="figure-21@66.[202,1386,181,1920]" captionTargetPageId="66" captionText="FIGURE 1. A–E: Serica gwangjuensis new species (holotype); F–J: S. assingi new species (holotype); A, H: aedeagus, left side lateral view; C, J: aedeagus, right side lateral view; B, I: parameres, dorsal view; D, F: habitus, dorsal view; E, G: habitus, lateral view. Scale bar: 0.5 mm. Habitus images not to scale." figureDoi="http://doi.org/10.5281/zenodo.13211853" httpUri="https://zenodo.org/record/13211853/files/figure.png" pageId="7" pageNumber="8">Fig. 1A–C</figureCitation>
. Habitus:
<figureCitation id="8ACD2A7BFFDA7C6AFDF4AC1DEF8CFE9A" box="[558,679,331,357]" captionStart="FIGURE 1" captionStartId="66.[152,255,1943,1967]" captionTargetBox="[212,1376,192,1910]" captionTargetId="figure-21@66.[202,1386,181,1920]" captionTargetPageId="66" captionText="FIGURE 1. A–E: Serica gwangjuensis new species (holotype); F–J: S. assingi new species (holotype); A, H: aedeagus, left side lateral view; C, J: aedeagus, right side lateral view; B, I: parameres, dorsal view; D, F: habitus, dorsal view; E, G: habitus, lateral view. Scale bar: 0.5 mm. Habitus images not to scale." figureDoi="http://doi.org/10.5281/zenodo.13211853" httpUri="https://zenodo.org/record/13211853/files/figure.png" pageId="7" pageNumber="8">Fig. 1D–E</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5AEC6575FFDA7C6AFF1DAC38EC0BFE0A" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="124936FEFFDA7C6AFF1DAC38EC0BFE0A" blockId="7.[151,1437,151,753]" pageId="7" pageNumber="8">
<emphasis id="2082EAECFFDA7C6AFF1DAC38EC69FE77" bold="true" box="[199,322,366,392]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
<taxonomicName id="D5F64D7DFFDA7C6AFE90AC39EF19FE77" authorityName="Ahrens &amp; Zhao &amp; Pham &amp; Liu" authorityYear="2024" box="[330,562,367,393]" class="Insecta" family="Scarabaeidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gwangjuensis" status="sp. nov.">
<emphasis id="2082EAECFFDA7C6AFE90AC39EF19FE77" box="[330,562,367,393]" italics="true" pageId="7" pageNumber="8">Serica gwangjuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3BB15797FFDA7C6AFDE1AC39EFEFFE76" box="[571,708,367,393]" pageId="7" pageNumber="8" rank="species">new species</taxonomicNameLabel>
is in shape of genitalia similar to
<taxonomicName id="D5F64D7DFFDA7C6AFB97AC39E8B2FE76" authority="Kim &amp; Kim, 2003" authorityName="Kim &amp; Kim" authorityYear="2003" box="[1101,1433,367,393]" class="Insecta" family="Scarabaeidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="2082EAECFFDA7C6AFB97AC39E991FE76" box="[1101,1210,367,393]" italics="true" pageId="7" pageNumber="8">S. hirsuta</emphasis>
Kim &amp; Kim, 2003
</taxonomicName>
.
<taxonomicName id="D5F64D7DFFDA7C6AFF4DACC5ECABFE53" authorityName="Ahrens &amp; Zhao &amp; Pham &amp; Liu" authorityYear="2024" box="[151,384,403,429]" class="Insecta" family="Scarabaeidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gwangjuensis" status="sp. nov.">
<emphasis id="2082EAECFFDA7C6AFF4DACC5ECABFE53" box="[151,384,403,429]" italics="true" pageId="7" pageNumber="8">Serica gwangjuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3BB15797FFDA7C6AFE53ACC5EF3AFE52" box="[393,529,403,429]" pageId="7" pageNumber="8" rank="species">new species</taxonomicNameLabel>
differs from the latter by the narrower right paramere which is sharply pointed at apex and the almost reduced, small, sickle-shaped left paramere, which is barely visibly under the phallobasal membranes.
</paragraph>
</subSubSection>
<subSubSection id="5AEC6575FFDA7C6AFF1DACA9EC4DFDC2" pageId="7" pageNumber="8" type="etymology">
<paragraph id="124936FEFFDA7C6AFF1DACA9EC4DFDC2" blockId="7.[151,1437,151,753]" pageId="7" pageNumber="8">
<emphasis id="2082EAECFFDA7C6AFF1DACA9EC65FDE6" bold="true" box="[199,334,511,537]" pageId="7" pageNumber="8">Etymology.</emphasis>
The name of this new species (adjective in singular nominative case) is derived from its
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locality,
<collectingRegion id="D032F81CFFDA7C6AFF2DAF75EC4AFDC2" box="[247,353,547,573]" country="South Korea" name="Gwangju" pageId="7" pageNumber="8">Gwangju</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="5AEC6575FFDA7C6AFF1DAF10EE45FD0E" pageId="7" pageNumber="8" type="description">
<paragraph id="124936FEFFDA7C6AFF1DAF10EE45FD0E" blockId="7.[151,1437,151,753]" pageId="7" pageNumber="8">
<emphasis id="2082EAECFFDA7C6AFF1DAF10EC14FD9F" bold="true" box="[199,319,582,608]" pageId="7" pageNumber="8">Variation.</emphasis>
Length:
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, length of elytra:
<quantity id="D50E9B1BFFDA7C6AFD2FAF10EE55FD9E" box="[757,894,582,609]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="5.6" metricValueMin="5.4" pageId="7" pageNumber="8" unit="mm" value="5.5" valueMax="5.6" valueMin="5.4">5.4–5.6 mm</quantity>
, width: 4.0–
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. Meso- and metatarsomeres dorsally with longitudinal wrinkles; metatarsomeres laterally and dorsally carinate, with a strongly serrated ridge ventrally, ventrally glabrous; first metatarsomere little shorter than following two tarsomeres combined and almost twice as long as dorsal tibial spur. Female: Antennal club distinctly shorter than remaining antennomeres combined, eyes smaller than in male: ratio diameter/interocular width: 0.46.
</paragraph>
</subSubSection>
</treatment>
</document>
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