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<mods:title id="0AD479D3231E74659235241CE12C532E">Revision of the genus Lamprima Latreille, 1804 (Coleoptera: Lucanidae)</mods:title>
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<taxonomicName id="D5A54D5DFFD2FF9BFF4BFD7EFDF2FD50" ID-CoL="3S3Y8" authority="(Gestro 1875)" authorityName=": Nagel" authorityYear="1922" baseAuthorityName="Gestro" baseAuthorityYear="1875" box="[151,597,722,748]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="adolphinae">
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<emphasis id="20D1EACCFFD2FF9BFF4BFD7EFE03FD50" bold="true" box="[151,420,722,748]" italics="true" pageId="10" pageNumber="161">Lamprima adolphinae</emphasis>
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<figureCitation id="8A9E2A5BFFD2FF9BFF43FD59FF42FCB2" box="[159,229,757,782]" captionStart="FIGURES 14" captionStartId="13.[151,264,1945,1968]" captionTargetBox="[354,1219,217,1877]" captionTargetId="figure@13.[292,1290,195,1923]" captionTargetPageId="13" captionText="FIGURES 14. Male Lamprima species, dorsal view: 1, L. adolphinae (Gestro, 1875); 2, L. aenea (Fabricius, 1792); 3, L. imberbis Carter, 1926 (holotype); 4, L. insularis W.J. Macleay, 1885. Figure 3 courtesy Peter Hudson, South Australian Museum. All figures to same scale." httpUri="https://zenodo.org/record/1454257/files/figure.png" pageId="10" pageNumber="161">Figs 1</figureCitation>
,
<figureCitation id="8A9E2A5BFFD2FF9BFF2FFD59FEB6FCB2" box="[243,273,757,782]" captionStart="FIGURES 1520" captionStartId="15.[151,264,1787,1810]" captionTargetBox="[182,1405,249,1699]" captionTargetId="figure@15.[151,1436,217,1732]" captionTargetPageId="15" captionText="FIGURES 1520. Female Lamprima species, dorsal view: 15, L. adolphinae (Gestro, 1875); 16, L. aenea (Fabricius, 1792); 17, L. aurata Latreille, 1804, Sydney, New South Wales; 18, L. aurata, Sydney, New South Wales, blue form; 19, L. aurata, Western Australia; 20, L. insularis W.J. Macleay, 1885." httpUri="https://zenodo.org/record/1454261/files/figure.png" pageId="10" pageNumber="161">15</figureCitation>
,
<figureCitation id="8A9E2A5BFFD2FF9BFEC1FD5AFEC1FCB2" box="[285,358,758,782]" captionStart="FIGURES 2124" captionStartId="16.[151,264,1916,1939]" captionTargetBox="[264,1330,215,1867]" captionTargetId="figure@16.[151,1436,193,1895]" captionTargetPageId="16" captionText="FIGURES 2124. Male Lamprima species, dorsal of head and pronotum: 21, L. adolphinae (Gestro, 1875), elongate mandible form; 22, L. adolphinae, short mandible form; 23, L. aenea (Fabricius, 1792); 24, L. insularis W.J. Macleay, 1885." httpUri="https://zenodo.org/record/1454263/files/figure.png" pageId="10" pageNumber="161">2122</figureCitation>
,
<figureCitation id="8A9E2A5BFFD2FF9BFEADFD5AFE28FCB2" box="[369,399,758,782]" captionStart="FIGURES 3335" captionStartId="18.[151,264,1628,1651]" captionTargetBox="[175,1404,217,1578]" captionTargetId="figure@18.[151,1436,193,1607]" captionTargetPageId="18" captionText="FIGURES 3335. Female Lamprima species, dorsal of head and pronotum: 33, L. adolphinae (Gestro, 1875); 34, L. aenea (Fabricius, 1792); 35, L. insularis W.J. Macleay, 1885." httpUri="https://zenodo.org/record/1454267/files/figure.png" pageId="10" pageNumber="161">33</figureCitation>
,
<figureCitation id="8A9E2A5BFFD2FF9BFE47FD59FE43FCB2" box="[411,484,757,782]" captionStart="FIGURES 4553" captionStartId="27.[151,264,1889,1912]" captionTargetBox="[248,1353,211,1852]" captionTargetId="figure@27.[223,1364,193,1868]" captionTargetPageId="27" captionText="FIGURES 4553. Male Lamprima species, mandibles, lateral view: 45, L. adolphinae (Gestro, 1875), elongate mandible form; 46, L. adolphinae, short mandible form; 47, L. aenea (Fabricius, 1792); 48, L. aurata Latreille, 1804, elongate mandible form, northern Queensland; 49, L. aurata, New South Wales, large male; 50, L. aurata, New South Wales, small male; 51, L. aurata, Western Australia; 52, L. imberbis Carter, 1926, holotype; 53, L. insularis W.J. Macleay, 1885. Figure 52 courtesy Peter Hudson, South Australian Museum." httpUri="https://zenodo.org/record/1454271/files/figure.png" pageId="10" pageNumber="161">4546</figureCitation>
,
<figureCitation id="8A9E2A5BFFD2FF9BFE33FD59FD9FFCB2" box="[495,568,757,782]" captionStart="FIGURES 5457" captionStartId="28.[151,264,1626,1649]" captionTargetBox="[182,1381,217,1560]" captionTargetId="figure@28.[151,1436,193,1605]" captionTargetPageId="28" captionText="FIGURES 5457. Male Lamprima species, elytral disc microsculpture, at low and high magnification: 5455, L. adolphinae (Gestro, 1875); 5657, L. aurata Latreille, 1804. Large punctures approximately 25 microns in diameter." httpUri="https://zenodo.org/record/1454273/files/figure.png" pageId="10" pageNumber="161">5455</figureCitation>
,
<figureCitation id="8A9E2A5BFFD2FF9BFD9FFD59FDC5FCB2" box="[579,610,757,782]" captionStart="FIGURES 6265" captionStartId="32.[151,264,1930,1953]" captionTargetBox="[171,1410,193,1886]" captionTargetId="figure@32.[151,1436,193,1909]" captionTargetPageId="32" captionText="FIGURES 6265. Male Lamprima species, protibiae: 62, L. adolphinae (Gestro, 1875); 63, L. aenea (Fabricius, 1792); 64, L. insularis W.J. Macleay, 1885; 65, L. imberbis Carter, 1926. Figure 64 courtesy Peter Hudson, South Australian Museum." httpUri="https://zenodo.org/record/1454279/files/figure.png" pageId="10" pageNumber="161">62</figureCitation>
,
<figureCitation id="8A9E2A5BFFD2FF9BFDB2FD59FD2BFCB2" box="[622,652,757,782]" captionStart="FIGURES 7683" captionStartId="34.[151,264,1918,1941]" captionTargetBox="[251,1365,218,1854]" captionTargetId="figure@34.[219,1366,193,1894]" captionTargetPageId="34" captionText="FIGURES 7683. Lamprima species, aedeagi (ventral, lateral and dorsal): 76, L. adolphinae (Gestro, 1875); 77, L. aenea (Fabricius, 1792); 78, L. aurata Latreille, 1804, northern Queensland; 79, L. aurata, Snowy River, New South Wales; 80, L. aurata, Kingston, Tasmania; 81, L. aurata, South Australia; 82, L. aurata, Cue, Western Australia; 83, L. insularis W.J. Macleay, 1885." httpUri="https://zenodo.org/record/1454283/files/figure.png" pageId="10" pageNumber="161">76</figureCitation>
,
<figureCitation id="8A9E2A5BFFD2FF9BFD4BFD5AFD12FCB2" box="[663,693,758,782]" captionStart="FIGURES 8994" captionStartId="36.[151,264,1908,1931]" captionTargetBox="[179,1401,208,1855]" captionTargetId="figure@36.[151,1436,193,1888]" captionTargetPageId="36" captionText="FIGURES 8994. Lamprima species, ovipositor and spermatheca: 89, L. adolphinae (Gestro, 1875); 90, L. aenea (Fabricius, 1792); 91, L. aurata Latreille, 1804 (Bridport, Tasmania); Lamprima species, spermathecae: 92, L. aurata (Ourimbah, central New South Wales); 93, L. aurata (Nadgee, south New South Wales); 94, L. insularis W.J. Macleay, 1885." httpUri="https://zenodo.org/record/1454287/files/figure.png" pageId="10" pageNumber="161">89</figureCitation>
,
<figureCitation id="8A9E2A5BFFD2FF9BFD1DFD59FD79FCB2" box="[705,734,757,782]" captionStart="FIGURES 9596. 95" captionStartId="37.[151,264,1938,1961]" captionTargetBox="[279,1309,203,1889]" captionTargetId="figure@37.[250,1338,193,1921]" captionTargetPageId="37" captionText="FIGURES 9596. 95, Map of Australia and New Guinea to show Lamprima species: O= L. adolphinae (Gestro, 1875), Ɨ= L. aurata Latreille, 1804. 96, Map of Lord Howe to show distribution of Lamprima insularis W.J. Macleay, 1885 (shaded area = 500 + m)" httpUri="https://zenodo.org/record/1454289/files/figure.png" pageId="10" pageNumber="161">95</figureCitation>
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<taxonomicName id="D5A54D5DFFD2FF9BFF4BFC91FDF0FCEF" ID-CoL="46K26" authority="Gestro, 1875: 997" authorityName="Gestro" authorityPageNumber="997" authorityYear="1875" box="[151,599,829,851]" class="Insecta" family="Lucanidae" genus="Neolamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="adolphinae">
<emphasis id="20D1EACCFFD2FF9BFF4BFC91FE3DFCEF" box="[151,410,829,851]" italics="true" pageId="10" pageNumber="161">Neolamprima adolphinae</emphasis>
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locality: Hatam, Arfak Mountains).
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<treatmentCitationGroup id="32B511F0FFD2FF9BFF4BFCF1FD88FCCE" box="[151,559,860,882]" pageId="10" pageNumber="161">
<taxonomicName id="D5A54D5DFFD2FF9BFF4BFCF1FD8CFCCE" ID-CoL="3S3Y8" authority=": Nagel, 1922: 16" authorityName=": Nagel" authorityPageNumber="16" authorityYear="1922" baseAuthorityName="Gestro" baseAuthorityYear="1875" box="[151,555,860,882]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="adolphinae">
<emphasis id="20D1EACCFFD2FF9BFF4BFCF1FEDEFCCE" box="[151,377,860,882]" italics="true" pageId="10" pageNumber="161">Lamprima adolphinae</emphasis>
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<treatmentCitation id="930410CFFFD2FF9BFE5AFCF0FD8CFCCE" author="" box="[390,555,860,882]" page="16" pageId="10" pageNumber="161">
<bibRefCitation id="76344B2FFFD2FF9BFE5AFCF0FDA2FCCE" box="[390,517,860,882]" pageId="10" pageNumber="161" refId="ref33998" refString="Nagel, P. (1922) Neolamprima Gestro = Lamprima Latr. Neolamprima mandibularis M'Leay = Lamprima latreillei M'Leay (Col. Lucan.). Entomologische Mitteilungen, 11, 16 - 17." type="journal article">Nagel, 1922</bibRefCitation>
: 16
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.
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<taxonomicName id="D5A54D5DFFD2FF9BFF4BFCD7FD3EFC2C" ID-CoL="7GZ37" authority="Nagel, 1930: 88" authorityName="Nagel" authorityPageNumber="88" authorityYear="1930" box="[151,665,890,912]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="subSpecies" species="adolphinae" subSpecies="olivacea">
<emphasis id="20D1EACCFFD2FF9BFF4BFCD7FE47FC2C" box="[151,480,890,912]" italics="true" pageId="10" pageNumber="161">Lamprima adolphinae olivacea</emphasis>
<treatmentCitation id="930410CFFFD2FF9BFE30FCD6FD3EFC2C" author="" box="[492,665,890,912]" page="88" pageId="10" pageNumber="161">
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: 88
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(
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locality: Komba, Finisterre Mountains);
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<bibRefCitation id="76344B2FFFD2FF9BFB48FCD6FA84FC2C" box="[1172,1315,890,912]" pageId="10" pageNumber="161" refId="ref33119" refString="Krajcik, M. (2001) Lucanidae of the world. Catalogue - part 1. Checklist of the stag beetles of the world (Coleoptera: Lucanidae). Krajcik, Most, vi + 108 pp." type="book">Krajcik 2001</bibRefCitation>
: 25
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(junior synonym of
<taxonomicName id="D5A54D5DFFD2FF9BFE99FC36FE75FC13" ID-CoL="3S3Y8" authorityName=": Nagel" authorityYear="1922" baseAuthorityName="Gestro" baseAuthorityYear="1875" box="[325,466,921,943]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="adolphinae">
<emphasis id="20D1EACCFFD2FF9BFE99FC36FE75FC13" box="[325,466,921,943]" italics="true" pageId="10" pageNumber="161">L. adolphinae</emphasis>
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).
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<paragraph id="121A36DEFFD2FF9BFF4BFC15FEE0FC50" blockId="10.[151,1437,829,1096]" pageId="10" pageNumber="161">
<treatmentCitationGroup id="32B511F0FFD2FF9BFF4BFC15FEE0FC50" pageId="10" pageNumber="161">
<taxonomicName id="D5A54D5DFFD2FF9BFF4BFC15FD3FFC72" ID-CoL="7GZ2V" authority="Kriesche, 1940: 39" authorityName="Kriesche" authorityPageNumber="39" authorityYear="1940" box="[151,664,952,974]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="subSpecies" species="adolphinae" subSpecies="lulua">
<emphasis id="20D1EACCFFD2FF9BFF4BFC15FE18FC72" box="[151,447,952,974]" italics="true" pageId="10" pageNumber="161">Lamprima adolphinae lulua</emphasis>
<treatmentCitation id="930410CFFFD2FF9BFE10FC14FD3FFC72" author="" box="[460,664,952,974]" page="39" pageId="10" pageNumber="161">
<bibRefCitation id="76344B2FFFD2FF9BFE10FC14FDC8FC72" box="[460,623,952,974]" pageId="10" pageNumber="161" refId="ref33159" refString="Kriesche, R. (1940) Vier neue Lucaniden. Entomologische Blaetter, Krefeld, 36, 39 - 40." type="journal article">Kriesche, 1940</bibRefCitation>
: 39
</treatmentCitation>
</taxonomicName>
(type locality: New Guinea);
<treatmentCitation id="930410CFFFD2FF9BFC3AFC14FB39FC72" author="" box="[998,1182,952,974]" page="25" pageId="10" pageNumber="161">
<bibRefCitation id="76344B2FFFD2FF9BFC3AFC14FBD2FC72" box="[998,1141,952,974]" pageId="10" pageNumber="161" refId="ref33119" refString="Krajcik, M. (2001) Lucanidae of the world. Catalogue - part 1. Checklist of the stag beetles of the world (Coleoptera: Lucanidae). Krajcik, Most, vi + 108 pp." type="book">Krajcik 2001</bibRefCitation>
: 25
</treatmentCitation>
(junior synonym of
<taxonomicName id="D5A54D5DFFD2FF9BFA5BFC15FE9EFC50" ID-CoL="3S3Y8" authorityName=": Nagel" authorityYear="1922" baseAuthorityName="Gestro" baseAuthorityYear="1875" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="adolphinae">
<emphasis id="20D1EACCFFD2FF9BFA5BFC15FE9EFC50" italics="true" pageId="10" pageNumber="161">L. adolphinae</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="121A36DEFFD2FF9BFF4BFC59FE47FB96" blockId="10.[151,1437,829,1096]" pageId="10" pageNumber="161">
<treatmentCitationGroup id="32B511F0FFD2FF9BFF4BFC59FE47FB96" pageId="10" pageNumber="161">
<taxonomicName id="D5A54D5DFFD2FF9BFF4BFC59FCEEFBB7" authority="Didier &amp; Seguy, 1952: 222" authorityName="Didier &amp; Seguy" authorityPageNumber="222" authorityYear="1952" box="[151,841,1013,1035]" class="Insecta" family="Lucanidae" genus="Neolamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="subSpecies" species="adolphinae" subSpecies="chalciditis">
<emphasis id="20D1EACCFFD2FF9BFF4BFC59FDB1FBB7" box="[151,534,1013,1035]" italics="true" pageId="10" pageNumber="161">Neolamprima adolphinae chalciditis</emphasis>
<treatmentCitation id="930410CFFFD2FF9BFDFDFC59FCEEFBB7" author="" box="[545,841,1013,1035]" page="222" pageId="10" pageNumber="161">
<bibRefCitation id="76344B2FFFD2FF9BFDFDFC59FCB4FBB7" box="[545,787,1013,1035]" pageId="10" pageNumber="161" refId="ref32304" refString="Didier, R. &amp; Seguy, E. (1952) Notes sur quelques especes de lucanides et descriptions de formes nouvelles. Revue Francaise d'Entomologie, 19, 220 - 233." type="journal article">Didier &amp; Séguy, 1952</bibRefCitation>
: 222
</treatmentCitation>
</taxonomicName>
(
<typeStatus id="CD1E887CFFD2FF9BFC80FC59FC2EFBB7" box="[860,905,1013,1035]" pageId="10" pageNumber="161">type</typeStatus>
locality: New
<collectingCountry id="6AB2764EFFD2FF9BFBF3FC59FBDAFBB7" box="[1071,1149,1013,1035]" name="Guinea-Bissau" pageId="10" pageNumber="161">Guinea</collectingCountry>
);
<treatmentCitation id="930410CFFFD2FF9BFB48FC59FAEDFBB7" author="" box="[1172,1354,1013,1035]" page="25" pageId="10" pageNumber="161">
<bibRefCitation id="76344B2FFFD2FF9BFB48FC59FA85FBB7" box="[1172,1314,1013,1035]" pageId="10" pageNumber="161" refId="ref33119" refString="Krajcik, M. (2001) Lucanidae of the world. Catalogue - part 1. Checklist of the stag beetles of the world (Coleoptera: Lucanidae). Krajcik, Most, vi + 108 pp." type="book">Krajcik 2001</bibRefCitation>
: 25
</treatmentCitation>
(junior synonym of
<taxonomicName id="D5A54D5DFFD2FF9BFE99FBB9FE75FB96" ID-CoL="46K26" authorityName="Gestro" authorityYear="1875" box="[325,466,1044,1066]" class="Insecta" family="Lucanidae" genus="Neolamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="adolphinae">
<emphasis id="20D1EACCFFD2FF9BFE99FBB9FE75FB96" box="[325,466,1044,1066]" italics="true" pageId="10" pageNumber="161">L. adolphinae</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="121A36DEFFD2FF9BFF4BFB9EFB0CFBF4" blockId="10.[151,1437,829,1096]" box="[151,1195,1074,1096]" pageId="10" pageNumber="161">
<treatmentCitationGroup id="32B511F0FFD2FF9BFF4BFB9EFB0CFBF4" box="[151,1195,1074,1096]" pageId="10" pageNumber="161">
<taxonomicName id="D5A54D5DFFD2FF9BFF4BFB9EFD48FBF4" ID-CoL="7HJHN" authority="Darge &amp; Seguy, 1953: 252" authorityName="Darge &amp; Seguy" authorityPageNumber="252" authorityYear="1953" box="[151,751,1074,1096]" class="Insecta" family="Lucanidae" genus="Neolamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="subSpecies" species="adolphinae" subSpecies="bohni">
<emphasis id="20D1EACCFFD2FF9BFF4BFB9EFE7CFBF4" box="[151,475,1074,1096]" italics="true" pageId="10" pageNumber="161">Neolamprima adolphinae bohni</emphasis>
<treatmentCitation id="930410CFFFD2FF9BFE3DFB9FFD48FBF4" author="" box="[481,751,1074,1096]" page="252" pageId="10" pageNumber="161">
<bibRefCitation id="76344B2FFFD2FF9BFE3DFB9FFD18FBF4" box="[481,703,1074,1096]" pageId="10" pageNumber="161" refId="ref32155" refString="Darge, R. &amp; Seguy, E. (1953) Une variete nouvelle du Neolamprima adolphinae Gestro (Coleopt. Lucanidae). Revue Francaise d'Entomologie, 20, 252 - 253." type="journal article">Darge &amp; Séguy, 1953</bibRefCitation>
: 252
</treatmentCitation>
</taxonomicName>
(type locality: New Guinea); new synonym
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="5ABF6555FFD2FF9BFF4BFBD9FD56F9A6" pageId="10" pageNumber="161" type="materials_examined">
<paragraph id="121A36DEFFD2FF9BFF4BFBD9FD56F9A6" blockId="10.[151,1437,1141,2030]" pageId="10" pageNumber="161">
<materialsCitation id="A2CD3C83FFD2FF9BFF4BFBD9FEEBFB0E" ID-GBIF-Occurrence="1915495899" collectionCode="AMS, CMNC" country="Indonesia" latitude="-1.15" location="Arfak" longLatPrecision="1308" longitude="133.96666" municipality="Material" pageId="10" pageNumber="161" specimenCount="1">
<emphasis id="20D1EACCFFD2FF9BFF4BFBD9FEDDFB32" bold="true" box="[151,378,1141,1166]" pageId="10" pageNumber="161">
<collectingMunicipality id="F27EACA4FFD2FF9BFF4BFBD9FF58FB32" box="[151,255,1141,1166]" pageId="10" pageNumber="161">Material</collectingMunicipality>
examined
</emphasis>
(about 350; locality data only; * = specimen dissected).
<emphasis id="20D1EACCFFD2FF9BFBD8FBD9FB25FB32" bold="true" box="[1028,1154,1141,1166]" pageId="10" pageNumber="161">
<collectingCountry id="6AB2764EFFD2FF9BFBD8FBD9FBDAFB32" box="[1028,1149,1141,1166]" name="Indonesia" pageId="10" pageNumber="161">Indonesia</collectingCountry>
:
</emphasis>
<location id="177A6005FFD2FF9BFB57FBD9FB6BFB32" LSID="urn:lsid:plazi:treatment:9A0C87C8FFD2FF9DFF4BFD7EFB21FA7C:177A6005FFD2FF9BFB57FBD9FB6BFB32" box="[1163,1228,1141,1166]" country="Indonesia" latitude="-1.15" longLatPrecision="1308" longitude="133.96666" municipality="Material" name="Arfak" pageId="10" pageNumber="161">Arfak</location>
[
<geoCoordinate id="77915019FFD2FF9BFB00FBDAFA82FB32" box="[1244,1317,1142,1166]" direction="south" orientation="latitude" pageId="10" pageNumber="161" precision="925" value="-1.15">1°09'S</geoCoordinate>
<geoCoordinate id="77915019FFD2FF9BFAF2FBD9FA32FB32" box="[1326,1429,1141,1166]" direction="east" orientation="longitude" pageId="10" pageNumber="161" precision="925" value="133.96666">133°58'E</geoCoordinate>
] (
<collectionCode id="74B4AE1BFFD2FF9BFF43FB36FF47FB0E" box="[159,224,1178,1202]" pageId="10" pageNumber="161">AMS</collectionCode>
,
<collectionCode id="74B4AE1BFFD2FF9BFF30FB36FEE3FB0E" box="[236,324,1178,1202]" pageId="10" pageNumber="161">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="A2CD3C83FFD2FF9BFE84FB35FD6FFB0E" ID-GBIF-Occurrence="1915495867" box="[344,712,1177,1202]" collectingDate="2003-12" collectionCode="AMS" country="Indonesia" location="Irian Jaya" pageId="10" pageNumber="161" specimenCount="6" specimenCount-male="6" stateProvince="Irian Jaya">
<specimenCount id="04A3FD57FFD2FF9BFE84FB35FE22FB0D" box="[344,389,1177,1202]" pageId="10" pageNumber="161" type="male">6 ♂</specimenCount>
,
<collectingRegion id="D061F83CFFD2FF9BFE4FFB35FDA3FB0E" box="[403,516,1177,1202]" country="Indonesia" name="Papua Barat" pageId="10" pageNumber="161">Irian Jaya</collectingRegion>
,
<date id="661B101EFFD2FF9BFDCDFB35FDCAFB0E" box="[529,621,1177,1202]" pageId="10" pageNumber="161" value="2003-12">
<collectingDate id="765FE9F6FFD2FF9BFDCDFB35FDCAFB0E" box="[529,621,1177,1202]" pageId="10" pageNumber="161" value="2003-12">xii.2003</collectingDate>
</date>
(
<collectionCode id="74B4AE1BFFD2FF9BFDA2FB36FD67FB0E" box="[638,704,1178,1202]" pageId="10" pageNumber="161">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="A2CD3C83FFD2FF9BFD09FB35FC43FB0E" ID-GBIF-Occurrence="1915495857" box="[725,996,1177,1202]" collectionCode="CMNC" country="Indonesia" location="Wissell Lakes" pageId="10" pageNumber="161" specimenCount="1" stateProvince="Irian Jaya">
<location id="177A6005FFD2FF9BFD09FB35FCD4FB0E" LSID="urn:lsid:plazi:treatment:9A0C87C8FFD2FF9DFF4BFD7EFB21FA7C:177A6005FFD2FF9BFD09FB35FCD4FB0E" box="[725,883,1177,1202]" country="Indonesia" name="Wissell Lakes" pageId="10" pageNumber="161" stateProvince="Irian Jaya">Wissell Lakes</location>
(
<collectionCode id="74B4AE1BFFD2FF9BFC5FFB36FC7BFB0E" box="[899,988,1178,1202]" pageId="10" pageNumber="161">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="A2CD3C83FFD2FF9BFC2CFB35FD37FB6A" ID-GBIF-Occurrence="1915495853" collectionCode="ANIC, NAIC" country="Papua New Guinea" latitude="-6.3333335" location="Aiyura" longLatPrecision="1304" longitude="145.9" pageId="10" pageNumber="161" specimenCount="1" stateProvince="Eastern Highlands">
<emphasis id="20D1EACCFFD2FF9BFC2CFB35FB44FB0E" bold="true" box="[1008,1251,1177,1202]" pageId="10" pageNumber="161">
<collectingCountry id="6AB2764EFFD2FF9BFC2CFB35FB7AFB0E" box="[1008,1245,1177,1202]" name="Papua New Guinea" pageId="10" pageNumber="161">Papua New Guinea</collectingCountry>
:
</emphasis>
<location id="177A6005FFD2FF9BFB37FB35FA9BFB0E" LSID="urn:lsid:plazi:treatment:9A0C87C8FFD2FF9DFF4BFD7EFB21FA7C:177A6005FFD2FF9BFB37FB35FA9BFB0E" box="[1259,1340,1177,1202]" country="Papua New Guinea" latitude="-6.3333335" longLatPrecision="1304" longitude="145.9" name="Aiyura" pageId="10" pageNumber="161" stateProvince="Eastern Highlands">Aiyura</location>
,
<collectingRegion id="D061F83CFFD2FF9BFA94FB36FEADFB6A" country="Papua New Guinea" name="Eastern Highlands" pageId="10" pageNumber="161">
<collectingRegion id="D061F83CFFD2FF9BFA94FB36FA3BFB0D" box="[1352,1436,1178,1201]" country="American Samoa" name="Eastern" pageId="10" pageNumber="161">Eastern</collectingRegion>
Highlands
</collectingRegion>
[
<geoCoordinate id="77915019FFD2FF9BFEC5FB11FEC4FB6A" box="[281,355,1213,1238]" direction="south" orientation="latitude" pageId="10" pageNumber="161" precision="925" value="-6.3333335">6°20'S</geoCoordinate>
<geoCoordinate id="77915019FFD2FF9BFEB7FB11FE74FB6A" box="[363,467,1213,1238]" direction="east" orientation="longitude" pageId="10" pageNumber="161" precision="925" value="145.9">145°54'E</geoCoordinate>
] (
<collectionCode id="74B4AE1BFFD2FF9BFE35FB12FD95FB6A" box="[489,562,1214,1238]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="10" pageNumber="161">ANIC</collectionCode>
,
<collectionCode id="74B4AE1BFFD2FF9BFDE2FB12FD20FB6A" box="[574,647,1214,1238]" country="Papua New Guinea" httpUri="http://grbio.org/cool/7mi7-dyvm" name="National Agricultural Insect Collection" pageId="10" pageNumber="161">NAIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="A2CD3C83FFD2FF9BFD40FB11FC0CFB6A" ID-GBIF-Occurrence="1915495872" box="[668,939,1213,1238]" collectionCode="AMS" country="Papua New Guinea" location="Bulolo" pageId="10" pageNumber="161" specimenCount="1" stateProvince="Morobe">
<location id="177A6005FFD2FF9BFD40FB11FD4CFB6A" LSID="urn:lsid:plazi:treatment:9A0C87C8FFD2FF9DFF4BFD7EFB21FA7C:177A6005FFD2FF9BFD40FB11FD4CFB6A" box="[668,747,1213,1238]" country="Papua New Guinea" name="Bulolo" pageId="10" pageNumber="161" stateProvince="Morobe">Bulolo</location>
,
<collectingRegion id="D061F83CFFD2FF9BFD2BFB11FCF7FB6A" box="[759,848,1213,1238]" country="Papua New Guinea" name="Morobe" pageId="10" pageNumber="161">Morobe</collectingRegion>
(
<collectionCode id="74B4AE1BFFD2FF9BFCBDFB12FC04FB6A" box="[865,931,1214,1238]" pageId="10" pageNumber="161">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="A2CD3C83FFD2FF9BFC64FB11FA3FFB6A" ID-GBIF-Occurrence="1915495866" box="[952,1432,1213,1238]" collectionCode="NAIC" country="Papua New Guinea" latitude="-6.1666665" location="Chuave" longLatPrecision="1305" longitude="145.75" pageId="10" pageNumber="161" specimenCount="1" stateProvince="Simbu">
<location id="177A6005FFD2FF9BFC64FB11FBB7FB6A" LSID="urn:lsid:plazi:treatment:9A0C87C8FFD2FF9DFF4BFD7EFB21FA7C:177A6005FFD2FF9BFC64FB11FBB7FB6A" box="[952,1040,1213,1238]" country="Papua New Guinea" latitude="-6.1666665" longLatPrecision="1305" longitude="145.75" name="Chuave" pageId="10" pageNumber="161" stateProvince="Simbu">Chuave</location>
,
<collectingRegion id="D061F83CFFD2FF9BFBC1FB11FBC1FB6A" box="[1053,1126,1213,1238]" country="Papua New Guinea" name="Chimbu" pageId="10" pageNumber="161">Simbu</collectingRegion>
[
<geoCoordinate id="77915019FFD2FF9BFBA9FB11FB18FB6A" box="[1141,1215,1213,1238]" direction="south" orientation="latitude" pageId="10" pageNumber="161" precision="925" value="-6.1666665">6°10'S</geoCoordinate>
<geoCoordinate id="77915019FFD2FF9BFB1BFB11FA89FB6A" box="[1223,1326,1213,1238]" direction="east" orientation="longitude" pageId="10" pageNumber="161" precision="925" value="145.75">145°45'E</geoCoordinate>
] (
<collectionCode id="74B4AE1BFFD2FF9BFA99FB12FA28FB6A" box="[1349,1423,1214,1238]" country="Papua New Guinea" httpUri="http://grbio.org/cool/7mi7-dyvm" name="National Agricultural Insect Collection" pageId="10" pageNumber="161">NAIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="A2CD3C83FFD2FF9BFF4BFB4DFD43FB46" ID-GBIF-Occurrence="1915495875" box="[151,740,1249,1274]" collectionCode="NAIC" country="Papua New Guinea" latitude="-5.9833336" location="Daulo" longLatPrecision="1305" longitude="145.31667" pageId="10" pageNumber="161" specimenCount="1" stateProvince="Eastern Highlands">
<location id="177A6005FFD2FF9BFF4BFB4DFF79FB46" LSID="urn:lsid:plazi:treatment:9A0C87C8FFD2FF9DFF4BFD7EFB21FA7C:177A6005FFD2FF9BFF4BFB4DFF79FB46" box="[151,222,1249,1274]" country="Papua New Guinea" latitude="-5.9833336" longLatPrecision="1305" longitude="145.31667" name="Daulo" pageId="10" pageNumber="161" stateProvince="Eastern Highlands">Daulo</location>
,
<collectingRegion id="D061F83CFFD2FF9BFF35FB4EFE10FB46" box="[233,439,1249,1274]" country="Papua New Guinea" name="Eastern Highlands" pageId="10" pageNumber="161">
<collectingRegion id="D061F83CFFD2FF9BFF35FB4EFE9AFB45" box="[233,317,1250,1273]" country="American Samoa" name="Eastern" pageId="10" pageNumber="161">Eastern</collectingRegion>
Highlands
</collectingRegion>
[
<geoCoordinate id="77915019FFD2FF9BFE19FB4DFDA9FB46" box="[453,526,1249,1274]" direction="south" orientation="latitude" pageId="10" pageNumber="161" precision="925" value="-5.9833336">5°59'S</geoCoordinate>
<geoCoordinate id="77915019FFD2FF9BFDC9FB4DFDDBFB46" box="[533,636,1249,1274]" direction="east" orientation="longitude" pageId="10" pageNumber="161" precision="925" value="145.31667">145°19'E</geoCoordinate>
] (
<collectionCode id="74B4AE1BFFD2FF9BFD4EFB4EFD7BFB46" box="[658,732,1250,1274]" country="Papua New Guinea" httpUri="http://grbio.org/cool/7mi7-dyvm" name="National Agricultural Insect Collection" pageId="10" pageNumber="161">NAIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="A2CD3C83FFD2FF9BFD33FB4DFC79FB46" ID-GBIF-Occurrence="1915495869" box="[751,990,1249,1274]" collectionCode="ANIC" country="Papua New Guinea" location="Finschhaven" pageId="10" pageNumber="161" specimenCount="1" stateProvince="Eastern Highlands">
<location id="177A6005FFD2FF9BFD33FB4DFCDAFB46" LSID="urn:lsid:plazi:treatment:9A0C87C8FFD2FF9DFF4BFD7EFB21FA7C:177A6005FFD2FF9BFD33FB4DFCDAFB46" box="[751,893,1249,1274]" country="Papua New Guinea" name="Finschhaven" pageId="10" pageNumber="161" stateProvince="Eastern Highlands">Finschhaven</location>
(
<collectionCode id="74B4AE1BFFD2FF9BFC50FB4EFC71FB46" box="[908,982,1250,1274]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="10" pageNumber="161">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="A2CD3C83FFD2FF9BFC35FB4DFA9AFB46" ID-GBIF-Occurrence="1915495868" box="[1001,1341,1249,1274]" collectionCode="ANIC" country="Papua New Guinea" location="Habu River" municipality="Frigano" pageId="10" pageNumber="161" specimenCount="1" stateProvince="Eastern Highlands">
<collectingMunicipality id="F27EACA4FFD2FF9BFC35FB4DFBE7FB46" box="[1001,1088,1249,1274]" pageId="10" pageNumber="161">Frigano</collectingMunicipality>
[
<location id="177A6005FFD2FF9BFB92FB4DFB72FB46" LSID="urn:lsid:plazi:treatment:9A0C87C8FFD2FF9DFF4BFD7EFB21FA7C:177A6005FFD2FF9BFB92FB4DFB72FB46" box="[1102,1237,1249,1274]" country="Papua New Guinea" municipality="Frigano" name="Habu River" pageId="10" pageNumber="161" stateProvince="Eastern Highlands">Habu River</location>
] (
<collectionCode id="74B4AE1BFFD2FF9BFB36FB4EFA92FB46" box="[1258,1333,1250,1274]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="10" pageNumber="161">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="A2CD3C83FFD2FF9BFA94FB4DFF58FAA2" ID-GBIF-Occurrence="1915495864" collectionCode="CMNC" country="Papua New Guinea" location="Goroka" pageId="10" pageNumber="161" specimenCount="1" stateProvince="Eastern Highlands">
<location id="177A6005FFD2FF9BFA94FB4DFA3BFB46" LSID="urn:lsid:plazi:treatment:9A0C87C8FFD2FF9DFF4BFD7EFB21FA7C:177A6005FFD2FF9BFA94FB4DFA3BFB46" box="[1352,1436,1249,1274]" country="Papua New Guinea" name="Goroka" pageId="10" pageNumber="161" stateProvince="Eastern Highlands">Goroka</location>
(
<collectionCode id="74B4AE1BFFD2FF9BFF43FAAAFF5FFAA2" box="[159,248,1286,1310]" pageId="10" pageNumber="161">CMNC</collectionCode>
)
</materialsCitation>
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[
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.
</paragraph>
</subSubSection>
<subSubSection id="5ABF6555FFD2FF9DFF1BF989FB24FF44" lastPageId="12" lastPageNumber="163" pageId="10" pageNumber="161" type="description">
<paragraph id="121A36DEFFD2FF9BFF1BF989FE53F8E2" blockId="10.[151,1437,1141,2030]" pageId="10" pageNumber="161">
<emphasis id="20D1EACCFFD2FF9BFF1BF989FEFEF982" bold="true" box="[199,345,1573,1598]" pageId="10" pageNumber="161">Description.</emphasis>
Male: length 24¯
<quantity id="D55D9B3BFFD2FF9BFDF1F989FD20F981" box="[557,647,1573,1598]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="161" unit="mm" value="60.0">60 mm</quantity>
; cylindrical, pronotum slightly broader than elytra; colour: usually pronotum and elytra entirely dull metallic bronze or green, head bronze-black, antennae and tarsomeres darker, almost black, tibiae slightly lighter bronze coloured than femora, and all except base of outer margins of mandibles purplish black; less often pronotum and elytra yellowish green or bluish green; head purplish red, tibiae coppery red or purplish red; rarely pronotum and elytra blue. Pronotum minutely, densely, rugosely microreticulate; reticulations obscured by dense micropunctures and dull (with a metallic sheen but not shiny); elytra slightly shinier, microreticulate; reticulations flat and distinct; anterior half of head shallowly microreticulate, shiny, basal half often duller; upper surfaces of pronotum and elytra apparently glabrous, but each puncture with minute simple seta arising from anterior edge.
</paragraph>
<paragraph id="121A36DEFFD2FF9AFF1BF8C5FF44FD70" blockId="10.[151,1437,1141,2030]" lastBlockId="11.[151,1437,151,2013]" lastPageId="11" lastPageNumber="162" pageId="10" pageNumber="161">
Head: sides and apex with sparse, inconspicuous setae; head length slightly less than half width; sides convergent, from small but angularly projecting temples, along feebly convex eyes, to usually slightly convex genae and right-angled (small males) to acute anteriorly projecting anterior angles (large males); projecting temples broadly grooved in lateral view. Anterior margin truncate to concave, vertically declivous to labrum; dorsum with two smooth ridges from anterior angles to midline of base of head where they meet at about 100°; area subtended by these ridges strongly punctate, punctures varying greatly in density but not confluent; area between ridges and sides of head also strongly punctate. Antennomere 2 transverse to quadrate; 34 elongate (lengths 1.52.0x widths); 5 usually elongate, rarely quadrate; 6 transverse to quadrate, rarely ridged on anterior edge; 7 cupuliform with lateral extension about as wide as antennomere, Mandible length 2137% of overall length; inner faces of mandibles almost entirely densely setose; mandibles straight sided in large males, slightly convex in small males, almost symmetrical, approximately equal in length; two
<typeStatus id="CD1E887CFFD3FF9AFCE5FEDCFCD3FE34" box="[825,884,368,392]" pageId="11" pageNumber="162">types</typeStatus>
of mandible: small males (
<figureCitation id="8A9E2A5BFFD3FF9AFB1FFEC3FAB8FE34" box="[1219,1311,367,392]" captionStart="FIGURES 2124" captionStartId="16.[151,264,1916,1939]" captionTargetBox="[264,1330,215,1867]" captionTargetId="figure@16.[151,1436,193,1895]" captionTargetPageId="16" captionText="FIGURES 2124. Male Lamprima species, dorsal of head and pronotum: 21, L. adolphinae (Gestro, 1875), elongate mandible form; 22, L. adolphinae, short mandible form; 23, L. aenea (Fabricius, 1792); 24, L. insularis W.J. Macleay, 1885." httpUri="https://zenodo.org/record/1454263/files/figure.png" pageId="11" pageNumber="162">Figs 22</figureCitation>
,
<figureCitation id="8A9E2A5BFFD3FF9AFAEDFEC3FAF7FE34" box="[1329,1360,367,392]" captionStart="FIGURES 4553" captionStartId="27.[151,264,1889,1912]" captionTargetBox="[248,1353,211,1852]" captionTargetId="figure@27.[223,1364,193,1868]" captionTargetPageId="27" captionText="FIGURES 4553. Male Lamprima species, mandibles, lateral view: 45, L. adolphinae (Gestro, 1875), elongate mandible form; 46, L. adolphinae, short mandible form; 47, L. aenea (Fabricius, 1792); 48, L. aurata Latreille, 1804, elongate mandible form, northern Queensland; 49, L. aurata, New South Wales, large male; 50, L. aurata, New South Wales, small male; 51, L. aurata, Western Australia; 52, L. imberbis Carter, 1926, holotype; 53, L. insularis W.J. Macleay, 1885. Figure 52 courtesy Peter Hudson, South Australian Museum." httpUri="https://zenodo.org/record/1454271/files/figure.png" pageId="11" pageNumber="162">46</figureCitation>
), with mandible mostly straight in lateral view, apical 1/3 turned upwards, upper edge sharply keeled, preapical dorsal acute tooth about ¼ from tip, apex bifid with two acute teeth, inner (lower) margin with broad acutely tipped lobe 1/3 from apex and irregular short projections between lobe and apex; large males (
<figureCitation id="8A9E2A5BFFD3FF9AFB8DFE70FB0DFE48" box="[1105,1194,476,501]" captionStart="FIGURES 2124" captionStartId="16.[151,264,1916,1939]" captionTargetBox="[264,1330,215,1867]" captionTargetId="figure@16.[151,1436,193,1895]" captionTargetPageId="16" captionText="FIGURES 2124. Male Lamprima species, dorsal of head and pronotum: 21, L. adolphinae (Gestro, 1875), elongate mandible form; 22, L. adolphinae, short mandible form; 23, L. aenea (Fabricius, 1792); 24, L. insularis W.J. Macleay, 1885." httpUri="https://zenodo.org/record/1454263/files/figure.png" pageId="11" pageNumber="162">Figs 21</figureCitation>
,
<figureCitation id="8A9E2A5BFFD3FF9AFB64FE70FB70FE49" box="[1208,1239,476,501]" captionStart="FIGURES 4553" captionStartId="27.[151,264,1889,1912]" captionTargetBox="[248,1353,211,1852]" captionTargetId="figure@27.[223,1364,193,1868]" captionTargetPageId="27" captionText="FIGURES 4553. Male Lamprima species, mandibles, lateral view: 45, L. adolphinae (Gestro, 1875), elongate mandible form; 46, L. adolphinae, short mandible form; 47, L. aenea (Fabricius, 1792); 48, L. aurata Latreille, 1804, elongate mandible form, northern Queensland; 49, L. aurata, New South Wales, large male; 50, L. aurata, New South Wales, small male; 51, L. aurata, Western Australia; 52, L. imberbis Carter, 1926, holotype; 53, L. insularis W.J. Macleay, 1885. Figure 52 courtesy Peter Hudson, South Australian Museum." httpUri="https://zenodo.org/record/1454271/files/figure.png" pageId="11" pageNumber="162">45</figureCitation>
), with mandibles strongly curved from base to apex in lateral view, upper edge rounded in basal half, preapical dorsal tooth migrated to extreme apex of mandible, which therefore has 3 acute teeth, apical third of upper surface either smooth or irregularly toothed before apex, inner (lower) margin with small acute or blunt tipped lobe ½1/3 from base, sometimes finely serrate edged just before lobe, and with 515 irregular, often partly fused, peg-like projections beyond this lobe; mandibles laterally smooth, finely punctate and setose at base; mentum flat, closely punctate and setose.
</paragraph>
<paragraph id="121A36DEFFD3FF9AFF1BFD7BFB91FAC4" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="162">
Thorax: pronotum strongly convex, faintly dimpled near lateral angles, shape typical for
<taxonomicName id="D5A54D5DFFD3FF9AFB10FD74FAE7FD53" authorityName="Latreille" authorityYear="1804" box="[1228,1344,728,751]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="20D1EACCFFD3FF9AFB10FD74FAE7FD53" box="[1228,1344,728,751]" italics="true" pageId="11" pageNumber="162">Lamprima</emphasis>
</taxonomicName>
, almost hexagonal but sides slightly concave anterior to greatest width at just behind middle; anterior margin truncate with slightly protruding anterior angles, basal margin strongly sinuate, anterior and posterior angles obtuse, anterior margin narrower than basal margin, margination complete, laterally without crenulation, Punctures of disc indistinct, much smaller than on head, more-or-less obliterated by surface sculpture, separated by about 36 diameters, punctures only distinct at margins. Hypomeron finely and closely punctate in posterior half, shallowly wrinkled in anterior half, with mostly recumbent setae; prosternum strongly and densely punctate, with mostly recumbent setae; scutellum semicircular to heart-shaped, with sparse, small punctures. Elytra with or without small tubercle at base of epipleural upper margin, slightly expanded posterior to humeri, then contracted to rounded apices, sides narrowly explanate in posterior 2/3; elytral base often distinctly bevelled to accommodate base of pronotum; elytral disc minutely and sparsely punctate, punctures similar to or smaller than on pronotal disc and separated by 510 diameters, and shallowly, irregularly strigose, the deeper grooves mostly longitudinal; mesometaventral process with scattered, recumbent setae, and shiny, apex blunt, approximately 80° in lateral view, sides usually slightly concave. Protibia with pair of curved, elongate apical teeth, outer wider than inner, and external margin with 35 well-spaced triangular approximately right-angled teeth, confined to apical half of tibia in large specimens; inner lobe of protibia large and rounded, with basal dense tuft of convergent red setae and anterior greatly expanded spur (width 6580% of length); upper surface of protibia with scattered punctures on inner half and an irregular line of punctures on outer half, short recumbent setae arising from punctures, plus tuft of elongate setae at tarsal insertion; mesotibiae and metatibiae with 05 minute, external teeth.
</paragraph>
<paragraph id="121A36DEFFD3FF9AFF1BFA28FEB5F9C8" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="162">Abdomen: sides of ventrites IV similar to pterothoracic venter, with dense, small punctures (partly coalescent, interspaces less than diameters) and setae; middle third of ventrites more sparsely punctate (insterspaces = several diameters), surface microreticulate, dull; apex ventrite V truncate to shallowly concave. Genitalia: apical half of phallobase dorsally with irregular short oblique ridges or tubercles either side of shallow median groove, apical margin with V-shaped notch, deepened at base; venter of phallobase smooth, apex more deeply notched; parameres setose dorsally, tips triangular but usually incurved; penis with oblique basal ridges, apex of penis beyond apices of parameres.</paragraph>
<paragraph id="121A36DEFFD3FF9DFF1BF9D3FB24FF44" blockId="11.[151,1437,151,2013]" lastBlockId="12.[151,1437,151,1472]" lastPageId="12" lastPageNumber="163" pageId="11" pageNumber="162">
Female. As male, except: length 18¯
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; pronotum and elytra brilliantly shiny, without evident microsculpture, bronze, dark brown, green, coppery or blue, head generally bronze-black or similar in colour to pronotum; pronotum more strongly and densely punctate, punctures almost as large as on head, interspaces 13 puncture diameters on pronotal disc; pronotum narrower than elytra; head with 7590° anterior angles, apices of smooth ridges rounded not projecting. Antennomere 3 elongate, 46 transverse, 6 with sharp outer ridge; dorsally visible part of mandibles shorter than head; mandibles in dorsal view with elongate-rectangular (rarely triangular) dorsal tubercle from base to almost half mandible length, remainder of dorsal surface excavate with sharp outer edge; pronotum margins distinctly crenulate on apical half but basal half complete or with &lt;5 shallow notches. Outer edge protibia with 68 triangular teeth, generally increasing in size from base to apex, inner edge without internal lobe, spur elongate triangular; outer edges mesotibiae and metatibiae with 57 prominent spines; venter shiny, otherwise similar to male; apex ventrite V rounded; apex tergite IX transparent, attenuated and sharp or narrowly rounded; gonocoxite transverse, with both inner and outer edges expanded from base; spermatheca tapered from blunt apex to base, slightly bent, spermathecal duct long and densely coiled.
</paragraph>
</subSubSection>
<subSubSection id="5ABF6555FFD4FF9DFF1BFEA8FEEAFBB3" pageId="12" pageNumber="163" type="materials_examined">
<paragraph id="121A36DEFFD4FF9DFF1BFEA8FE76FD70" blockId="12.[151,1437,151,1472]" pageId="12" pageNumber="163">
<emphasis id="20D1EACCFFD4FF9DFF1BFEA8FEEDFEA1" bold="true" box="[199,330,260,285]" pageId="12" pageNumber="163">Taxonomy.</emphasis>
<taxonomicName id="D5A54D5DFFD4FF9DFE85FEA9FDFBFEA0" authorityName=": Nagel" authorityYear="1922" baseAuthorityName="Gestro" baseAuthorityYear="1875" box="[345,604,260,284]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="adolphinae">
<emphasis id="20D1EACCFFD4FF9DFE85FEA9FDFBFEA0" box="[345,604,260,284]" italics="true" pageId="12" pageNumber="163">Lamprima adolphinae</emphasis>
</taxonomicName>
was first described in 1875. However, 40 years earlier,
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<emphasis id="20D1EACCFFD4FF9DFAFAFEA9FA3BFEA0" box="[1318,1436,260,284]" italics="true" pageId="12" pageNumber="163">L. fulgida</emphasis>
<bibRefCitation id="76344B2FFFD4FF9DFF4BFE8BFEF6FEFC" box="[151,337,295,320]" pageId="12" pageNumber="163" refId="ref31694" refString="Boisduval, [J. A.] (1835) Faune entomologique de l'Oceanie, comprenant les Coleopteres, les Hemipteres, les Nevropteres, les Hymenopteres et les Dipteres. Librairie Encyclopedique du Roret, Paris, 705 pp." type="book">Boisduval, 1835</bibRefCitation>
</taxonomicName>
, was described from the island of Waigeo, west of New
<collectingCountry id="6AB2764EFFD4FF9DFC18FE8BFBBFFEFC" box="[964,1048,295,320]" name="Guinea-Bissau" pageId="12" pageNumber="163">Guinea</collectingCountry>
. Since all records of
<taxonomicName id="D5A54D5DFFD4FF9DFADDFE85FA3BFEFC" authorityName=": Nagel" authorityYear="1922" baseAuthorityName="Gestro" baseAuthorityYear="1875" box="[1281,1436,296,320]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="adolphinae">
<emphasis id="20D1EACCFFD4FF9DFADDFE85FA3BFEFC" box="[1281,1436,296,320]" italics="true" pageId="12" pageNumber="163">L. adolphinae</emphasis>
</taxonomicName>
show that it is endemic to New
<collectingCountry id="6AB2764EFFD4FF9DFE2BFEE0FDEFFED9" box="[503,584,332,357]" name="Guinea-Bissau" pageId="12" pageNumber="163">Guinea</collectingCountry>
and other species of
<taxonomicName id="D5A54D5DFFD4FF9DFCEFFEE1FC0FFED8" authorityName="Latreille" authorityYear="1804" box="[819,936,333,356]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="20D1EACCFFD4FF9DFCEFFEE1FC0FFED8" box="[819,936,333,356]" italics="true" pageId="12" pageNumber="163">Lamprima</emphasis>
</taxonomicName>
are unknown there, it might be assumed that
<taxonomicName id="D5A54D5DFFD4FF9DFF4BFEDDFEAFFE34" authorityName="Boisduval" authorityYear="1835" box="[151,264,368,392]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="fulgida">
<emphasis id="20D1EACCFFD4FF9DFF4BFEDDFEAFFE34" box="[151,264,368,392]" italics="true" pageId="12" pageNumber="163">L. fulgida</emphasis>
</taxonomicName>
is the oldest name for this insect.
<bibRefCitation id="76344B2FFFD4FF9DFD7FFEC3FCE1FE34" box="[675,838,367,392]" pageId="12" pageNumber="163" refId="ref34381" refString="Reiche, L. J. (1841) Note sur le genre Lamprima, et description d'une nouvelle espece. Revue Zoologique, 4, 50 - 51." type="journal article">Reiche (1841)</bibRefCitation>
placed all hitherto described species of
<emphasis id="20D1EACCFFD4FF9DFAFCFEDCFA3BFE3B" box="[1312,1436,368,391]" italics="true" pageId="12" pageNumber="163">
<taxonomicName id="D5A54D5DFFD4FF9DFAFCFEDCFA30FE3B" authorityName="Latreille" authorityYear="1804" box="[1312,1431,368,391]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="genus">Lamprima</taxonomicName>
,
</emphasis>
including
<emphasis id="20D1EACCFFD4FF9DFED4FE39FEDAFE10" box="[264,381,404,428]" italics="true" pageId="12" pageNumber="163">
<taxonomicName id="D5A54D5DFFD4FF9DFED4FE39FEDEFE10" authorityName="Boisduval" authorityYear="1835" box="[264,377,404,428]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="fulgida">L. fulgida</taxonomicName>
,
</emphasis>
under the name
<taxonomicName id="D5A54D5DFFD4FF9DFDE7FE39FD3BFE10" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[571,668,405,428]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="aenea">
<emphasis id="20D1EACCFFD4FF9DFDE7FE39FD3BFE10" box="[571,668,405,428]" italics="true" pageId="12" pageNumber="163">L. aenea</emphasis>
</taxonomicName>
.
<bibRefCitation id="76344B2FFFD4FF9DFD77FE38FCDEFE10" box="[683,889,404,429]" pageId="12" pageNumber="163" refId="ref31899" refString="Burmeister, H. C. C. (1847) Handbuch der Entomologie. Fol. 5. Coleoptera Lamellicornia Xylophila et Pectinicornia. Enslin, Berlin, viii + 584 pp." type="book">Burmeister (1847)</bibRefCitation>
and
<bibRefCitation id="76344B2FFFD4FF9DFC6DFE38FBCAFE11" box="[945,1133,404,429]" pageId="12" pageNumber="163" refId="ref34685" refString="Thomson, J. (1862) Catalogue des lucanides de la collection de M. James Thomson suivi d'un appendix renfermant la description des coupes generiques et specifiques nouvelles. Annales de la Societe Entomologique de France, Series 4, 2, 389 - 436." type="journal article">Thomson (1862)</bibRefCitation>
accepted the validity of
<taxonomicName id="D5A54D5DFFD4FF9DFA59FE39FF41FE6C" authorityName="Boisduval" authorityYear="1835" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="fulgida">
<emphasis id="20D1EACCFFD4FF9DFA59FE39FF41FE6C" italics="true" pageId="12" pageNumber="163">L. fulgida</emphasis>
</taxonomicName>
, but the latter did not follow the rule of priority.
<bibRefCitation id="76344B2FFFD4FF9DFCD3FE14FC39FE6C" box="[783,926,439,464]" pageId="12" pageNumber="163" refId="ref34084" refString="Parry, F. J. S. (1864) A catalogue of lucanoid Coleoptera; with illustrations and descriptions of various new and interesting species. Transactions of the Entomological Society of London, Series 3, 2, 1 - 113, 12 pls." type="book chapter">Parry (1864)</bibRefCitation>
queried the status of
<taxonomicName id="D5A54D5DFFD4FF9DFB4DFE15FB59FE6C" authorityName="Boisduval" authorityYear="1835" box="[1169,1278,440,464]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="fulgida">
<emphasis id="20D1EACCFFD4FF9DFB4DFE15FB59FE6C" box="[1169,1278,440,464]" italics="true" pageId="12" pageNumber="163">L. fulgida</emphasis>
</taxonomicName>
, suggesting it was a junior synonym of
<taxonomicName id="D5A54D5DFFD4FF9DFE6CFE71FDBFFE48" authorityName="Latreille" authorityYear="1817" box="[432,536,477,500]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="aurata">
<emphasis id="20D1EACCFFD4FF9DFE6CFE71FDBFFE48" box="[432,536,477,500]" italics="true" pageId="12" pageNumber="163">L. aurata</emphasis>
</taxonomicName>
. Following
<bibRefCitation id="76344B2FFFD4FF9DFD43FE71FC88FE49" box="[671,815,476,501]" pageId="12" pageNumber="163" refId="ref34084" refString="Parry, F. J. S. (1864) A catalogue of lucanoid Coleoptera; with illustrations and descriptions of various new and interesting species. Transactions of the Entomological Society of London, Series 3, 2, 1 - 113, 12 pls." type="book chapter">Parry (1864)</bibRefCitation>
,
<taxonomicName id="D5A54D5DFFD4FF9DFCE6FE71FC01FE48" authorityName="Boisduval" authorityYear="1835" box="[826,934,476,500]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="fulgida">
<emphasis id="20D1EACCFFD4FF9DFCE6FE71FC01FE48" box="[826,934,476,500]" italics="true" pageId="12" pageNumber="163">L. fulgida</emphasis>
</taxonomicName>
has always been treated as a junior synonym of
<taxonomicName id="D5A54D5DFFD4FF9DFF6AFDADFEB8FDA4" authorityName="Latreille" authorityYear="1817" box="[182,287,513,536]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="aurata">
<emphasis id="20D1EACCFFD4FF9DFF6AFDADFEB8FDA4" box="[182,287,513,536]" italics="true" pageId="12" pageNumber="163">L. aurata</emphasis>
</taxonomicName>
(
<bibRefCitation id="76344B2FFFD4FF9DFEF2FE53FE66FDA4" box="[302,449,511,536]" pageId="12" pageNumber="163" refId="ref32874" refString="Harold, B. von. (1868) Catalogus coleoptorum hucusque descriptorum synonymicus et systematicus. Tom. III. Familia XXXII. Lucanidae. E. H. Gummi, Munich, 38 pp. [pp. 941 - 978]" type="book chapter">Harold 1868</bibRefCitation>
;
<bibRefCitation id="76344B2FFFD4FF9DFE11FE53FDDAFDA4" box="[461,637,511,536]" pageId="12" pageNumber="163" refId="ref33643" refString="Macleay, W. [J.] (1885 a) Revision of the genus Lamprima of Latreille, with descriptions of new species. Proceedings of the Linnean Society of New South Wales, 10, 129 - 140." type="journal article">Macleay 1885a</bibRefCitation>
;
<bibRefCitation id="76344B2FFFD4FF9DFD56FE53FC83FDA4" box="[650,804,511,536]" pageId="12" pageNumber="163" refId="ref31658" refString="Boileau, H. (1913) Note sur lucanides conserves dans les collections de l'Universite d'Oxford et du British Museum. Transactions of the Entomological Society of London, 1913, 213 - 272." type="journal article">Boileau 1913</bibRefCitation>
;
<bibRefCitation id="76344B2FFFD4FF9DFCECFE53FC61FDA4" box="[816,966,511,536]" pageId="12" pageNumber="163" refId="ref31636" refString="Benesh, B. (1960) Lucanidea [sic]. Coleopterorum Catalogus Supplementa, 8, 1 - 178." type="journal article">Benesh 1960</bibRefCitation>
;
<bibRefCitation id="76344B2FFFD4FF9DFC0EFDACFB6AFDA4" box="[978,1229,511,536]" pageId="12" pageNumber="163" refId="ref33922" refString="Moore, B. P. &amp; Cassis, G. (1992) Lucanidae. In: Cassis, G., Houston, W. W. K., Weir, T. A. &amp; Moore, B. P. (Eds.), Zoological Catalogue of Australia. Fol. 9. Coleoptera: Scarabaeoidea. Australian Government Publishing Service, Canberra, pp. 4 - 19." type="book chapter">Moore &amp; Cassis 1992</bibRefCitation>
).
<typeStatus id="CD1E887CFFD4FF9DFB3EFDACFABDFDA4" box="[1250,1306,512,536]" pageId="12" pageNumber="163">Type</typeStatus>
material of
<taxonomicName id="D5A54D5DFFD4FF9DFF4BFD89FEA2FD80" authorityName="Boisduval" authorityYear="1835" box="[151,261,548,572]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="fulgida">
<emphasis id="20D1EACCFFD4FF9DFF4BFD89FEA2FD80" box="[151,261,548,572]" italics="true" pageId="12" pageNumber="163">L. fulgida</emphasis>
</taxonomicName>
seems to be missing, except for a possible female
<typeStatus id="CD1E887CFFD4FF9DFC98FD89FC3CFD81" box="[836,923,549,573]" pageId="12" pageNumber="163" type="syntype">syntype</typeStatus>
in the Hope collection, Oxford, examined by Boileau who identified it as
<taxonomicName id="D5A54D5DFFD4FF9DFE00FDE5FC9FFDDC" authority="(Boileau 1913: 216)" authorityPageNumber="216" baseAuthorityName="Boileau" baseAuthorityYear="1913" box="[476,824,583,608]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="aurata">
<emphasis id="20D1EACCFFD4FF9DFE00FDE5FDE0FDDC" box="[476,583,585,608]" italics="true" pageId="12" pageNumber="163">L. aurata</emphasis>
(
<bibRefCitation id="76344B2FFFD4FF9DFD8BFDEBFC97FDDC" box="[599,816,583,608]" pageId="12" pageNumber="163" refId="ref31658" refString="Boileau, H. (1913) Note sur lucanides conserves dans les collections de l'Universite d'Oxford et du British Museum. Transactions of the Entomological Society of London, 1913, 213 - 272." type="journal article">Boileau 1913: 216</bibRefCitation>
)
</taxonomicName>
. Boisduval's description of the male is in comparison with
<taxonomicName id="D5A54D5DFFD4FF9DFF0CFDC1FE97FD38" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[208,304,621,644]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="aenea">
<emphasis id="20D1EACCFFD4FF9DFF0CFDC1FE97FD38" box="[208,304,621,644]" italics="true" pageId="12" pageNumber="163">L. aenea</emphasis>
</taxonomicName>
. The male of
<taxonomicName id="D5A54D5DFFD4FF9DFE11FDC1FD9DFD38" authorityName="Boisduval" authorityYear="1835" box="[461,570,620,644]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="fulgida">
<emphasis id="20D1EACCFFD4FF9DFE11FDC1FD9DFD38" box="[461,570,620,644]" italics="true" pageId="12" pageNumber="163">L. fulgida</emphasis>
</taxonomicName>
was golden-green with a coppery-red head, larger and had shorter mandibles (
<bibRefCitation id="76344B2FFFD4FF9DFF42FD23FE2BFD14" box="[158,396,655,680]" pageId="12" pageNumber="163" refId="ref31694" refString="Boisduval, [J. A.] (1835) Faune entomologique de l'Oceanie, comprenant les Coleopteres, les Hemipteres, les Nevropteres, les Hymenopteres et les Dipteres. Librairie Encyclopedique du Roret, Paris, 705 pp." type="book">Boisduval 1835: 231</bibRefCitation>
). This is much more like
<taxonomicName id="D5A54D5DFFD4FF9DFD78FD3DFCAAFD14" baseAuthorityName="Boileau" baseAuthorityYear="1913" box="[676,781,657,680]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="aurata">
<emphasis id="20D1EACCFFD4FF9DFD78FD3DFCAAFD14" box="[676,781,657,680]" italics="true" pageId="12" pageNumber="163">L. aurata</emphasis>
</taxonomicName>
than
<taxonomicName id="D5A54D5DFFD4FF9DFC95FD3DFC43FD14" authorityName=": Nagel" authorityYear="1922" baseAuthorityName="Gestro" baseAuthorityYear="1875" box="[841,996,656,680]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="adolphinae">
<emphasis id="20D1EACCFFD4FF9DFC95FD3DFC43FD14" box="[841,996,656,680]" italics="true" pageId="12" pageNumber="163">L. adolphinae</emphasis>
</taxonomicName>
, therefore we agree with the synonymy of
<taxonomicName id="D5A54D5DFFD4FF9DFF69FD19FE84FD70" authorityName="Boisduval" authorityYear="1835" box="[181,291,692,716]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="fulgida">
<emphasis id="20D1EACCFFD4FF9DFF69FD19FE84FD70" box="[181,291,692,716]" italics="true" pageId="12" pageNumber="163">L. fulgida</emphasis>
</taxonomicName>
with
<taxonomicName id="D5A54D5DFFD4FF9DFEBEFD19FE6DFD70" baseAuthorityName="Boileau" baseAuthorityYear="1913" box="[354,458,693,716]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="aurata">
<emphasis id="20D1EACCFFD4FF9DFEBEFD19FE6DFD70" box="[354,458,693,716]" italics="true" pageId="12" pageNumber="163">L. aurata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="121A36DEFFD4FF9DFF1BFD7BFCF7FCE1" blockId="12.[151,1437,151,1472]" pageId="12" pageNumber="163">
Like other species of
<taxonomicName id="D5A54D5DFFD4FF9DFE62FD74FD94FD53" authorityName="Latreille" authorityYear="1804" box="[446,563,728,751]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="20D1EACCFFD4FF9DFE62FD74FD94FD53" box="[446,563,728,751]" italics="true" pageId="12" pageNumber="163">Lamprima</emphasis>
</taxonomicName>
,
<taxonomicName id="D5A54D5DFFD4FF9DFD9FFD75FD47FD4C" authorityName=": Nagel" authorityYear="1922" baseAuthorityName="Gestro" baseAuthorityYear="1875" box="[579,736,728,752]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="adolphinae">
<emphasis id="20D1EACCFFD4FF9DFD9FFD75FD47FD4C" box="[579,736,728,752]" italics="true" pageId="12" pageNumber="163">L. adolphinae</emphasis>
</taxonomicName>
shows some variation in colour and these variants have been named. These colour forms have no taxonomic validity and are therefore all treated here as junior synonyms of
<taxonomicName id="D5A54D5DFFD4FF9DFA59FD51FEB2FC84" authorityName=": Nagel" authorityYear="1922" baseAuthorityName="Gestro" baseAuthorityYear="1875" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="adolphinae">
<emphasis id="20D1EACCFFD4FF9DFA59FD51FEB2FC84" italics="true" pageId="12" pageNumber="163">L. adolphinae</emphasis>
</taxonomicName>
.
<bibRefCitation id="76344B2FFFD4FF9DFEF8FCB3FE1AFC84" box="[292,445,799,824]" pageId="12" pageNumber="163" refId="ref33119" refString="Krajcik, M. (2001) Lucanidae of the world. Catalogue - part 1. Checklist of the stag beetles of the world (Coleoptera: Lucanidae). Krajcik, Most, vi + 108 pp." type="book">Krajik (2001)</bibRefCitation>
has already made the formal synonymy of three names.
<taxonomicName id="D5A54D5DFFD4FF9DFBE2FC8CFA9EFC84" authorityName=": Nagel" authorityYear="1922" baseAuthorityName="Gestro" baseAuthorityYear="1875" box="[1086,1337,800,824]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="adolphinae">
<emphasis id="20D1EACCFFD4FF9DFBE2FC8CFA9EFC84" box="[1086,1337,800,824]" italics="true" pageId="12" pageNumber="163">Lamprima adolphinae</emphasis>
</taxonomicName>
is also a senior synonym of
<taxonomicName id="D5A54D5DFFD4FF9DFEB2FCE9FCEBFCE1" authority="Darge &amp; Seguy, 1953" authorityName="Darge &amp; Seguy" authorityYear="1953" box="[366,844,836,861]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="subSpecies" species="adolphinae" subSpecies="bohni">
<emphasis id="20D1EACCFFD4FF9DFEB2FCE9FDF6FCE0" box="[366,593,836,860]" italics="true" pageId="12" pageNumber="163">L. adolphinae bohni</emphasis>
<bibRefCitation id="76344B2FFFD4FF9DFD8BFCE9FCEBFCE1" box="[599,844,836,861]" pageId="12" pageNumber="163" refId="ref32155" refString="Darge, R. &amp; Seguy, E. (1953) Une variete nouvelle du Neolamprima adolphinae Gestro (Coleopt. Lucanidae). Revue Francaise d'Entomologie, 20, 252 - 253." type="journal article">Darge &amp; Séguy, 1953</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="121A36DEFFD4FF9DFF1BFCC4FEEAFBB3" blockId="12.[151,1437,151,1472]" pageId="12" pageNumber="163">
<taxonomicName id="D5A54D5DFFD4FF9DFF1BFCC4FE60FC3C" authorityName=": Nagel" authorityYear="1922" baseAuthorityName="Gestro" baseAuthorityYear="1875" box="[199,455,872,896]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="adolphinae">
<emphasis id="20D1EACCFFD4FF9DFF1BFCC4FE60FC3C" box="[199,455,872,896]" italics="true" pageId="12" pageNumber="163">Lamprima adolphinae</emphasis>
</taxonomicName>
in New
<collectingCountry id="6AB2764EFFD4FF9DFDE6FCCBFD2CFC3C" box="[570,651,871,896]" name="Guinea-Bissau" pageId="12" pageNumber="163">Guinea</collectingCountry>
has two male mandible forms, like
<taxonomicName id="D5A54D5DFFD4FF9DFB95FCC4FB19FCC3" authorityName="Latreille" authorityYear="1804" box="[1097,1214,872,895]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="20D1EACCFFD4FF9DFB95FCC4FB19FCC3" box="[1097,1214,872,895]" italics="true" pageId="12" pageNumber="163">Lamprima</emphasis>
</taxonomicName>
males in northern
<collectingRegion id="D061F83CFFD4FF9DFF4BFC20FEBCFC19" box="[151,283,908,933]" country="Australia" name="Queensland" pageId="12" pageNumber="163">Queensland</collectingRegion>
(
<bibRefCitation id="76344B2FFFD4FF9DFEECFC20FE02FC18" box="[304,421,908,932]" pageId="12" pageNumber="163" refId="ref33532" refString="Lea, A. M. (1929) Notes on some miscellaneous Coleoptera, with descriptions on new species. Part VII. Transactions and Proceedings of the Royal Society of Southern Australia, 53, 203 - 244." type="journal article">Lea 1929</bibRefCitation>
). However both forms in New
<collectingCountry id="6AB2764EFFD4FF9DFCFAFC20FCD0FC19" box="[806,887,908,933]" name="Guinea-Bissau" pageId="12" pageNumber="163">Guinea</collectingCountry>
are distinguishable from their counterparts in northern
<collectingRegion id="D061F83CFFD4FF9DFF22FC03FE25FC74" box="[254,386,943,968]" country="Australia" name="Queensland" pageId="12" pageNumber="163">Queensland</collectingRegion>
by constant differences in the male mandibles and by the male genitalia. We have no doubt that
<taxonomicName id="D5A54D5DFFD4FF9DFF1BFC79FE66FC50" authorityName=": Nagel" authorityYear="1922" baseAuthorityName="Gestro" baseAuthorityYear="1875" box="[199,449,980,1004]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="adolphinae">
<emphasis id="20D1EACCFFD4FF9DFF1BFC79FE66FC50" box="[199,449,980,1004]" italics="true" pageId="12" pageNumber="163">Lamprima adolphinae</emphasis>
</taxonomicName>
is a valid species although the females are almost identical to those of
<taxonomicName id="D5A54D5DFFD4FF9DFB0AFC79FA3BFC50" baseAuthorityName="Boileau" baseAuthorityYear="1913" box="[1238,1436,981,1004]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="aurata">
<emphasis id="20D1EACCFFD4FF9DFB0AFC79FA3BFC50" box="[1238,1436,981,1004]" italics="true" pageId="12" pageNumber="163">Lamprima aurata</emphasis>
</taxonomicName>
and
<taxonomicName id="D5A54D5DFFD4FF9DFF1AFC55FEE1FBAC" authorityName="W. J. Macleay" authorityYear="1885" box="[198,326,1016,1040]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="20D1EACCFFD4FF9DFF1AFC55FEE1FBAC" box="[198,326,1016,1040]" italics="true" pageId="12" pageNumber="163">L. insularis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="5ABF6555FFD4FF9DFF1BFBB0FD6EFB1C" pageId="12" pageNumber="163" type="description">
<paragraph id="121A36DEFFD4FF9DFF1BFBB0FD6EFB1C" blockId="12.[151,1437,151,1472]" pageId="12" pageNumber="163">
<emphasis id="20D1EACCFFD4FF9DFF1BFBB0FDF1FB89" bold="true" box="[199,598,1052,1077]" pageId="12" pageNumber="163">Natural history and distribution.</emphasis>
The major male appears to be the dominant form in collections, but this may reflect collector bias.
<bibRefCitation id="76344B2FFFD4FF9DFE1AFBECFDE4FBE4" box="[454,579,1088,1112]" pageId="12" pageNumber="163" refId="ref33532" refString="Lea, A. M. (1929) Notes on some miscellaneous Coleoptera, with descriptions on new species. Part VII. Transactions and Proceedings of the Royal Society of Southern Australia, 53, 203 - 244." type="journal article">Lea (1929)</bibRefCitation>
noted considerable variation in male mandible development in this species and the small mandible form occurs throughout its range.
<bibRefCitation id="76344B2FFFD4FF9DFCEDFBC9FC63FBC1" box="[817,964,1124,1149]" pageId="12" pageNumber="163" refId="ref33573" refString="Levet, B. (2016) Observations sur Lamprima adolphinae (Gestro, 1875) (Coleoptera, Scarabaeoidea, Lucanidae). Le Coleopteriste, 19, 106 - 107." type="journal article">Levet (2016)</bibRefCitation>
provides notes on rearing
<taxonomicName id="D5A54D5DFFD4FF9DFB2BFBC9FA32FBC0" authorityName=": Nagel" authorityYear="1922" baseAuthorityName="Gestro" baseAuthorityYear="1875" box="[1271,1429,1124,1148]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="adolphinae">
<emphasis id="20D1EACCFFD4FF9DFB2BFBC9FA32FBC0" box="[1271,1429,1124,1148]" italics="true" pageId="12" pageNumber="163">L. adolphinae</emphasis>
</taxonomicName>
, recommending a temperature regime of 2028 °C.
</paragraph>
</subSubSection>
<subSubSection id="5ABF6555FFD4FF9DFF1BFB01FB21FA7C" pageId="12" pageNumber="163" type="reference_group">
<paragraph id="121A36DEFFD4FF9DFF1BFB01FE22FAC4" blockId="12.[151,1437,151,1472]" pageId="12" pageNumber="163">
<taxonomicName id="D5A54D5DFFD4FF9DFF1BFB01FE62FB78" ID-CoL="3S3Y8" authorityName=": Nagel" authorityYear="1922" baseAuthorityName="Gestro" baseAuthorityYear="1875" box="[199,453,1196,1220]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="adolphinae">
<emphasis id="20D1EACCFFD4FF9DFF1BFB01FE62FB78" box="[199,453,1196,1220]" italics="true" pageId="12" pageNumber="163">Lamprima adolphinae</emphasis>
</taxonomicName>
is largely montane, from
<quantity id="D55D9B3BFFD4FF9DFD22FB00FCEEFB78" box="[766,841,1196,1221]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="163" unit="m" value="500.0">500 m</quantity>
to high elevation, for example
<quantity id="D55D9B3BFFD4FF9DFB18FB00FAB9FB78" box="[1220,1310,1196,1220]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="12" pageNumber="163" unit="m" value="2800.0">2800 m</quantity>
on Mount Giluwe. Most sites are in the cloud forest zone,
<quantity id="D55D9B3BFFD4FF9DFD0DFB63FCD7FB54" box="[721,880,1231,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="12" pageNumber="163" unit="m" value="1750.0" valueMax="2500.0" valueMin="1000.0">10002500 m</quantity>
(
<bibRefCitation id="76344B2FFFD4FF9DFC5FFB7CFBABFB54" box="[899,1036,1231,1256]" pageId="12" pageNumber="163" refId="ref33573" refString="Levet, B. (2016) Observations sur Lamprima adolphinae (Gestro, 1875) (Coleoptera, Scarabaeoidea, Lucanidae). Le Coleopteriste, 19, 106 - 107." type="journal article">Levet 2016</bibRefCitation>
). The few lowland sites may only indicate the lowland ports from which specimens have been supplied to collectors (
<emphasis id="20D1EACCFFD4FF9DFBBCFB59FB2FFAB0" box="[1120,1160,1269,1292]" italics="true" pageId="12" pageNumber="163">e.g.</emphasis>
, Fak Fak:
<bibRefCitation id="76344B2FFFD4FF9DFAD6FB59FA37FAB1" box="[1290,1424,1268,1293]" pageId="12" pageNumber="163" refId="ref33573" refString="Levet, B. (2016) Observations sur Lamprima adolphinae (Gestro, 1875) (Coleoptera, Scarabaeoidea, Lucanidae). Le Coleopteriste, 19, 106 - 107." type="journal article">Levet 2016</bibRefCitation>
).
<taxonomicName id="D5A54D5DFFD4FF9DFF4BFAB4FE33FA8C" ID-CoL="3S3Y8" authorityName=": Nagel" authorityYear="1922" baseAuthorityName="Gestro" baseAuthorityYear="1875" box="[151,404,1304,1328]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="adolphinae">
<emphasis id="20D1EACCFFD4FF9DFF4BFAB4FE33FA8C" box="[151,404,1304,1328]" italics="true" pageId="12" pageNumber="163">Lamprima adolphinae</emphasis>
</taxonomicName>
is widespread on the island of New
<collectingCountry id="6AB2764EFFD4FF9DFC95FABBFC3DFA8C" box="[841,922,1303,1328]" name="Guinea-Bissau" pageId="12" pageNumber="163">Guinea</collectingCountry>
(
<figureCitation id="8A9E2A5BFFD4FF9DFC71FABBFBA3FA8C" box="[941,1028,1303,1328]" captionStart="FIGURES 9596. 95" captionStartId="37.[151,264,1938,1961]" captionTargetBox="[279,1309,203,1889]" captionTargetId="figure@37.[250,1338,193,1921]" captionTargetPageId="37" captionText="FIGURES 9596. 95, Map of Australia and New Guinea to show Lamprima species: O= L. adolphinae (Gestro, 1875), Ɨ= L. aurata Latreille, 1804. 96, Map of Lord Howe to show distribution of Lamprima insularis W.J. Macleay, 1885 (shaded area = 500 + m)" httpUri="https://zenodo.org/record/1454289/files/figure.png" pageId="12" pageNumber="163">Fig. 95</figureCitation>
). On the distribution map we have included records of
<taxonomicName id="D5A54D5DFFD4FF9DFE4DFA91FD91FAE8" ID-CoL="3S3Y8" authorityName=": Nagel" authorityYear="1922" baseAuthorityName="Gestro" baseAuthorityYear="1875" box="[401,566,1340,1364]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="adolphinae">
<emphasis id="20D1EACCFFD4FF9DFE4DFA91FD91FAE8" box="[401,566,1340,1364]" italics="true" pageId="12" pageNumber="163">L. adolphinae</emphasis>
</taxonomicName>
from Ononge, Admisibil and Walmak, all photographed on a website (
<bibRefCitation id="76344B2FFFD4FF9DFF42FACCFEDEFAC4" box="[158,377,1376,1400]" pageId="12" pageNumber="163" refId="ref31367" refString="Anonymous (2017 c) The insects of Papua Indonesia. Online. Available from: http: // www. papua-insects. nl (accessed 18 March 2018)" type="url">Anonymous 2017c</bibRefCitation>
).
</paragraph>
<paragraph id="121A36DEFFD4FF9DFF1BFA28FB21FA7C" blockId="12.[151,1437,151,1472]" pageId="12" pageNumber="163">
Conservation status.
<taxonomicName id="D5A54D5DFFD4FF9DFE6BFA29FD13FA20" ID-CoL="3S3Y8" authorityName=": Nagel" authorityYear="1922" baseAuthorityName="Gestro" baseAuthorityYear="1875" box="[439,692,1412,1436]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="adolphinae">
<emphasis id="20D1EACCFFD4FF9DFE6BFA29FD13FA20" box="[439,692,1412,1436]" italics="true" pageId="12" pageNumber="163">Lamprima adolphinae</emphasis>
</taxonomicName>
is widely distributed in New
<collectingCountry id="6AB2764EFFD4FF9DFBCFFA28FBC3FA21" box="[1043,1124,1412,1437]" name="Guinea-Bissau" pageId="12" pageNumber="163">Guinea</collectingCountry>
and does not appear to be under immediate threat, although it dominates the commercial trade in
<taxonomicName id="D5A54D5DFFD4FF9DFC6EFA04FB80FA03" ID-CoL="62XGM" authorityName="Latreille" authorityYear="1804" box="[946,1063,1448,1471]" class="Insecta" family="Lucanidae" genus="Lamprima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="genus">
<emphasis id="20D1EACCFFD4FF9DFC6EFA04FB80FA03" box="[946,1063,1448,1471]" italics="true" pageId="12" pageNumber="163">Lamprima</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
</treatment>
</document>