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<document ID-CLB-Dataset="25292" ID-DOI="http://dx.doi.org/10.3897/phytokeys.133.37773" ID-GBIF-Dataset="a6caa96b-fb3a-4877-9e16-b2f519188af6" ID-PMC="PMC6791900" ID-Pensoft-Pub="1314-2003-133-1" ID-Pensoft-UUID="99F75C9803D65C4880141BE7B6589FB5" ID-PubMed="31632179" ModsDocID="1314-2003-133-1" checkinTime="1570497209971" checkinUser="pensoft" docAuthor="Shalimov, Aleksandr Petrovich, Wu, Yu-Dong &amp; Zhang, Xian-Chun" docDate="2019" docId="99EF9ADD39395278A429FF3862668D09" docLanguage="en" docName="PhytoKeys 133: 1-76" docOrigin="PhytoKeys 133" docSource="http://dx.doi.org/10.3897/phytokeys.133.37773" docTitle="Selaginella pulvinata Maxim." docType="treatment" docVersion="8" id="99F75C9803D65C4880141BE7B6589FB5" lastPageNumber="1" masterDocId="99F75C9803D65C4880141BE7B6589FB5" masterDocTitle="A taxonomic revision of the genus Selaginella (Selaginellaceae) from Nepal" masterLastPageNumber="76" masterPageNumber="1" pageNumber="1" updateTime="1732680503569" updateUser="ExternalLinkService">
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<mods:title id="E29129C80070D1F96258C851F1F78A08">A taxonomic revision of the genus Selaginella (Selaginellaceae) from Nepal</mods:title>
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<mods:namePart id="9F4BEB0BDC111F74E9B0C1222B6B90AD">Shalimov, Aleksandr Petrovich</mods:namePart>
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<treatment id="99EF9ADD39395278A429FF3862668D09" ID-GBIF-Taxon="159654787" LSID="urn:lsid:plazi:treatment:99EF9ADD39395278A429FF3862668D09" httpUri="http://treatment.plazi.org/id/99EF9ADD39395278A429FF3862668D09" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Selaginellaceae">
<subSubSection id="8F1DEAAAFBE8A64ACB2E9816AC02CB9E" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="B14E540052689F6B361639A6854AC0FF" pageId="0" pageNumber="1">
<taxonomicName id="C2411B8DDD498D2713731A2318C6A7B0" LSID="99EF9ADD-3939-5278-A429-FF3862668D09" authority="(Hook. &amp; Grev.) Maxim." class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Selaginella pulvinata" order="Selaginellales" pageId="0" pageNumber="1" phylum="Pteridophyta" rank="species" species="pulvinata">Selaginella pulvinata (Hook. &amp; Grev.) Maxim.</taxonomicName>
<figureCitation id="63EE3803717782F00680329FC51B845C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Morphological diversity of the leaves of Nepalese Sealginella species: 1 A-C S. indica (Nakaike 1325, PE) 2 A-C S. pulvinata (Tabata et al. 3520, PE) 3 A-C S. bryopteris (Tabata et al. 11989, PE). A - Axillary leaves, B - Dorsal leaves, C - Ventral leaves. Scale bars: 0.2 mm." figureDoi="10.3897/phytokeys.133.37773.figure1" httpUri="https://binary.pensoft.net/fig/344897" pageId="0" pageNumber="1" tableDoi="10.3897/phytokeys.133.37773.figure1">Figs 1</figureCitation>
<figureCitation id="9077F6B6C3FE9EF41745FE347FE9B2E2" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Selaginella pulvinata (Hook. &amp; Grev.) Maxim. A Habit B fragment of the upper surface of the lateral branch showing dorsal leaves imbricate at branch (A Maire AMA 894, E; B Tabata et al. 3520, PE). Link: (http: // data. rbge. org. uk / herb / E 00670565)." figureDoi="10.3897/phytokeys.133.37773.figure13" httpUri="https://binary.pensoft.net/fig/344909" pageId="0" pageNumber="1" tableDoi="10.3897/phytokeys.133.37773.figure13">
(2
<normalizedToken id="2081BA3891F0E2AFD97365DDAC7BA31D" originalValue="AC">A-C</normalizedToken>
), 13
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="13627ABDD89F98E5ADFD4DB222A6BB63" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="59D22F529A9875BF3173AFAB47B86547" pageId="0" pageNumber="1">
<taxonomicName id="AB74549B3141238FE6AB4D254928C958" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Selaginella pulvinata" order="Selaginellales" pageId="0" pageNumber="1" phylum="Pteridophyta" rank="species" species="pulvinata">Selaginella pulvinata</taxonomicName>
(Hook. &amp; Grev.)Maxim.,
<normalizedToken id="159439B664CB52B2275DDA2D3DD432FE" originalValue="Mém">Mem</normalizedToken>
. Acad. Imp. Sci. Saint
<normalizedToken id="5E3D32963E74D38E55F8664A6E287AF8" originalValue="Pétersb">Petersb</normalizedToken>
. (
<normalizedToken id="8F7E9E8BAEA60DE3348A2E24A52BFD29" originalValue="Sér">Ser</normalizedToken>
. 7) 9: 335. 1859;
<bibRefCitation id="FE42D842C1BEC25FDD7AD1DD8C4CD382" author="Dixit, RD" journalOrPublisher="Dehra Dun, Bishen Singh Mahendra Pal Singh" pageId="0" pageNumber="1" refId="B4" refString="Dixit, RD, 1992. Selaginellaceae of India. Dehra Dun, Bishen Singh Mahendra Pal Singh" title="Selaginellaceae of India." year="1992">Dixit 1992</bibRefCitation>
;
<bibRefCitation id="0F4639B997A29C238E391E31576725C5" author="Thapa, N" journalOrPublisher="Department Plant Resources" pageId="0" pageNumber="1" pagination="1 - 175" refId="B27" refString="Thapa, N, 2002. Pteridophytes of Nepal. Bull. Department Plant Resources 19: 1 - 175" title="Pteridophytes of Nepal. Bull." volume="19" year="2002">Thapa 2002</bibRefCitation>
;
<bibRefCitation id="18FDD92F8E905D126923C4B618B286EF" author="Zhang, XC" editor="Zhang, XC" journalOrPublisher="Science Press, Beijing, Vol. 6" pageId="0" pageNumber="1" pagination="86 - 219" refId="B33" refString="Zhang, XC, 2004. Selaginellaceae. In: Zhang, XC, Ed., Flora Republica Popularis Sinicae. Science Press, Beijing, Vol. 6: 86 - 219" title="Selaginellaceae." volumeTitle="Flora Republica Popularis Sinicae." year="2004">Zhang 2004</bibRefCitation>
;
<bibRefCitation id="5771607DA24EEAD7D7F6030160977305" author="Zhang, XC" editor="Wu, ZY" journalOrPublisher="Science Press, Beijing and Missouri Botanical Garden Press, St. Louis, Vols 2 - 3" pageId="0" pageNumber="1" pagination="37 - 66" refId="B35" refString="Zhang, XC, Nooteboom, HP, Kato, M, 2013. Selaginellaceae. In: Wu, ZY, Raven, PH, Hong, DY, Eds., Flora of China. Science Press, Beijing and Missouri Botanical Garden Press, St. Louis, Vols 2-3: 37 - 66" title="Selaginellaceae." volumeTitle="Flora of China." year="2013">Zhang et al. 2013</bibRefCitation>
;
<bibRefCitation id="29F8263EE89B0874A8CB704002C78AB1" author="Fraser-Jenkins, CR" journalOrPublisher="National Herbarium and Plant Laboratories, Department of Plant Resources, Ministry of Forests and Soil Conservation, Kathmandu" pageId="0" pageNumber="1" refId="B7" refString="Fraser-Jenkins, CR, Kandel, DR, Pariyar, S, 2015. Ferns and fern-allies of Nepal. Vol. 1. National Herbarium and Plant Laboratories, Department of Plant Resources, Ministry of Forests and Soil Conservation, Kathmandu" title="Ferns and fern-allies of Nepal. Vol. 1." year="2015">Fraser-Jenkins et al. 2015</bibRefCitation>
;
<bibRefCitation id="A903936F5B07C51DAB6CAED51313F3D4" author="Fraser-Jenkins, CR" journalOrPublisher="National Herbarium and Plant Laboratories, Department of Plant Resources, Ministry of Forests and Soil Conservation, Kathmandu" pageId="0" pageNumber="1" refId="B8" refString="Fraser-Jenkins, CR, Gandhi, KN, Kholia, BS, Benniamin, A, 2017. An annotated checklist of Indian Pteridophytes, Part - I (Lycopodiaceae to Thelypteridace). Bishen Singh Mahendra Pal Singh Dehra Dun, 562 pp." title="An annotated checklist of Indian Pteridophytes, Part - I (Lycopodiaceae to Thelypteridace). Bishen Singh Mahendra Pal Singh Dehra Dun, 562 pp." year="2017">Fraser-Jenkins et al. 2017</bibRefCitation>
.
</paragraph>
<paragraph id="C322AD45E475C533502E500BCE8A393E" pageId="0" pageNumber="1">
<taxonomicName id="8270B84E6A76F11E7BB9D69A57FC05C1" authorityName="Hook &amp; Grev" authorityYear="1831" class="Lycopodiopsida" family="Selaginellaceae" genus="Lycopodium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycopodium pulvinatum" order="Selaginellales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pulvinatum">Lycopodium pulvinatum</taxonomicName>
Hook. &amp; Grev.,
<normalizedToken id="C19CF7C5521DAC66D37BFAC5D75E6674" originalValue="Hookers">Hooker's</normalizedToken>
J. Bot. Kew Gard. Misc. 2: 381. 1831.
</paragraph>
<paragraph id="76B9F2980B86355DA5EF1DA573A56244" pageId="0" pageNumber="1">
<taxonomicName id="6D52C41C97C2C4EE8A84F3068C737F28" authorityName="Alston" authorityYear="1934" baseAuthorityName="Hook. &amp; Grev." class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="treatment-meta" infraspecific-rank="var." kingdom="Plantae" lsidName="Selaginella tamariscina var. pulvinata" order="Selaginellales" pageId="0" pageNumber="1" phylum="Pteridophyta" rank="variety" species="tamariscina" variety="pulvinata">Selaginella tamariscina var. pulvinata</taxonomicName>
(Hook. &amp; Grev.) Alston, Bull. Fan Mem. Inst. Biol., Bot. 5(6): 271. 1934.
</paragraph>
<paragraph id="75CB3290DBE5D1A9335DC1672F5B99F2" pageId="0" pageNumber="1">
(
<taxonomicName id="E4150DA3BAACFF9DC7BFA23B447DCDE5" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Selaginella pulvinata" order="Selaginellales" pageId="0" pageNumber="1" phylum="Pteridophyta" rank="species" species="pulvinata">Selaginella pulvinata</taxonomicName>
(Hook. &amp; Grev.) Hand.-Mazz., Symb. Sin. 6: 5. 1929, later isonym).
</paragraph>
<paragraph id="2FDB1017F8DA87934A0E5830C5F15BA4" pageId="0" pageNumber="1">
<taxonomicName id="5544B7651A529B80EA1D1ADF39E65CEE" authorityName="H. S. Kung" authorityYear="1988" baseAuthorityName="Hook. &amp; Grev." class="Lycopodiopsida" family="Selaginellaceae" genus="Lycopodioides" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lycopodioides pulvinata" order="Selaginellales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pulvinata">Lycopodioides pulvinata</taxonomicName>
(Hook. &amp; Grev.) H.S. Kung, Fl. Sichuanica 6: 64, t. 18, f. 1-3. 1988.
<emphasis id="432C43396AF4A84CD910A411A961631A" bold="true" pageId="0" pageNumber="1">Type.</emphasis>
INDIA. E. India, Kamoon.
<emphasis id="F4B494BE4FC2F488B19A864E57BD5B4E" italics="true" pageId="0" pageNumber="1">Dr. Wallich</emphasis>
s.n. (holotype: K).
</paragraph>
</subSubSection>
<subSubSection id="01F709C9F2ED9A2C2CD0163737170AFF" pageId="0" pageNumber="1" type="description">
<paragraph id="8AD1515F412447288EF28E7F1D0D302D" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="7A7FDDCF7EF8907C59EF2559F1D38A23" pageId="0" pageNumber="1">
Stems 2
<normalizedToken id="44F33D8F53707AE8E7C1BE062134F1E0" originalValue="15(">-15(-</normalizedToken>
20) cm, many forming rosette at top of thick rootstock, branched from base. Main stems branched near and above base, primary branches pinnately branched, second branches 2-3 forked, stramineous or brown, main stem c. 1 mm in diam. at lower part. Axillary leaves ovate to triangular, c. 2.5
<normalizedToken id="B5960031B26563DA5954F4D907DB3D83" originalValue="×">x</normalizedToken>
1 mm, base exauriculate, margin lacerate-ciliolate, apex acute. Ventral leaves ovate, 2.9-3.2
<normalizedToken id="2A401352400A941CDE175FFAC8597D48" originalValue="×">x</normalizedToken>
1.4-1.5 mm, rotundate-cordate at base, margin lacerate, apex acuminate. Dorsal leaves ovate, 2.8-3.1
<normalizedToken id="2D07768EA2AA2B1D96F7B5B1A78C10B4" originalValue="×">x</normalizedToken>
0.9-1.2 mm, base truncate, entire to obscurely denticulate, posterior side thickened, apex aristate. Strobili solitary, terminal, compact, tetragonal, 7
<normalizedToken id="F8A011CC0259DB47BC1B91035A4D46D1" originalValue="15(">-15(-</normalizedToken>
20)
<normalizedToken id="EEAD2F54AEB50C9199C4A0478D5EF308" originalValue="×">x</normalizedToken>
1.5-2 mm. Sporophylls monomorphic, ovate, at base cordate, margin slight denticulate, apex acuminate. Megaspores white-yellow, surface verrucate; microspores yellow, surface irregularly papillate.
</paragraph>
</subSubSection>
<subSubSection id="1EF6E127E96DAA758F15BAE1BE69062E" pageId="0" pageNumber="1" type="ecology">
<paragraph id="E294A15720F773F91079B40EE8BF8359" pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph id="FF7A17023CEE8EA9EB7ACCEE0AF1977F" pageId="0" pageNumber="1">Terrestrial or epilithic, xerophytic. Alt. 1800-4400 m.</paragraph>
</subSubSection>
<subSubSection id="EA03537855D408A55FFF8A128718FF33" pageId="0" pageNumber="1" type="distribution">
<paragraph id="9E1E9B520CF710318A543679F551495C" pageId="0" pageNumber="1">Distribution in Nepal.</paragraph>
<paragraph id="F1927CA21841449AAFC2086CCE02DE72" pageId="0" pageNumber="1">W.</paragraph>
<paragraph id="8B6C68C521547B0F504E1CFD3E78BAC2" pageId="0" pageNumber="1">
Nepalese threatened status: EN (
<bibRefCitation id="27128338573039494A397730EA611231" author="Fraser-Jenkins, CR" journalOrPublisher="National Herbarium and Plant Laboratories, Department of Plant Resources, Ministry of Forests and Soil Conservation, Kathmandu" pageId="0" pageNumber="1" refId="B7" refString="Fraser-Jenkins, CR, Kandel, DR, Pariyar, S, 2015. Ferns and fern-allies of Nepal. Vol. 1. National Herbarium and Plant Laboratories, Department of Plant Resources, Ministry of Forests and Soil Conservation, Kathmandu" title="Ferns and fern-allies of Nepal. Vol. 1." year="2015">Fraser-Jenkins et al. 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="57C767FFEA0BBB3C8E24150278D09B4F" pageId="0" pageNumber="1" type="distribution">
<paragraph id="179DA587491914331CFF79C62FB36EC2" pageId="0" pageNumber="1">General distribution.</paragraph>
<paragraph id="0B1687136264DEFA4B1FC2978CFF3F58" pageId="0" pageNumber="1">CHINA (Chongqing, S Gansu, Guangxi, Guizhou, Hebei, Henan, Liaoning, Shaanxi, Shanxi, Sichuan, Xizang), INDIA (Uttarakhand), KOREA, MONGOLIA, RUSSIA (Siberia), THAILAND, VIETNAM.</paragraph>
</subSubSection>
<subSubSection id="88F8960C5500FC158E2376D4C0F12E3E" pageId="0" pageNumber="1" type="chromosome number">
<paragraph id="2416E9D9FE2F10156D7ECBE32926A636" pageId="0" pageNumber="1">Chromosome number.</paragraph>
<paragraph id="AB12F70D8A40CF8787DE00B4707177C7" pageId="0" pageNumber="1">not available data.</paragraph>
<paragraph id="5051100FBA4FA92001B7CAC2C3A6E7BF" pageId="0" pageNumber="1">Selected specimens examined:</paragraph>
<paragraph id="F2B2328AE0180A0F1BAC8B819FCD8083" pageId="0" pageNumber="1">
<emphasis id="4DB80E52A9904BA221A72A7F1A6B3DFF" bold="true" pageId="0" pageNumber="1">W Nepal: BAJHANG</emphasis>
: &quot;Bauligad, on open rock, rooting on crevices, alt. 1830 m, 6 Jul 1980,
<emphasis id="2E3E6B7EB5FF54A85DCAFDB228D31B46" italics="true" pageId="0" pageNumber="1">P.R. Shakya</emphasis>
,
<emphasis id="A6E136ED96B71C353C9D1E9C57B92019" italics="true" pageId="0" pageNumber="1">L.R. Sharma</emphasis>
,
<emphasis id="A5B085774DBB86BD5A14E856986ECBF5" italics="true" pageId="0" pageNumber="1">K.R. Amatya 6328</emphasis>
&quot; (KATH).
</paragraph>
<paragraph id="3AE8CF7894DD899EC1521FAF457F2363" pageId="0" pageNumber="1">
<emphasis id="56A67F8038391C99434F73D544A0B399" bold="true" pageId="0" pageNumber="1">DOLPA</emphasis>
: &quot;between Besagad and Shahartara, 14 Sep 1976,
<emphasis id="C1F6B04D970843D36ABB9B54382997B6" italics="true" pageId="0" pageNumber="1">H. Tabata</emphasis>
,
<emphasis id="9D9B9F61EE2E684C3DCB4E4D0675EA25" italics="true" pageId="0" pageNumber="1">K.R. Rajbhandari</emphasis>
,
<emphasis id="BF6EEAD2681047FEB0C5C43599FD13ED" italics="true" pageId="0" pageNumber="1">Y. Shimizu 3520</emphasis>
&quot; (PE).
</paragraph>
</subSubSection>
</treatment>
</document>