262 lines
30 KiB
XML
262 lines
30 KiB
XML
<document id="7D386A65CCF5AC0FFEED3359A5A78FDE" ID-CLB-Dataset="22249" ID-DOI="10.5852/ejt.2013.48" ID-GBIF-Dataset="02866e62-ab61-4cf4-9d98-0248e251084e" ID-ISSN="2118-9773" ID-Zenodo-Dep="3822836" ID-ZooBank="E7080722-E348-448D-96E5-D537F4865BB5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1589219740188" checkinUser="carolina" docAuthor="Thuy, Ben" docDate="2013" docId="9978976365E085CCD37524D6FE319EC1" docLanguage="en" docName="ejt-2013-48.pdf.imf" docOrigin="European Journal of Taxonomy 48" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ophiomalleus beneficarum Thuy 2013, sp. nov." docType="treatment" docUuid="7B911036-1E01-4EDA-AF2C-E56F0027B8F4" docUuidSource="ZooBank" docVersion="10" lastPageNumber="184" masterDocId="6541EF1B6555857BD1302529FFF89F78" masterDocTitle="Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)" masterLastPageNumber="242" masterPageNumber="1" pageNumber="182" updateTime="1698839159859" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0F53BCC9CB9D1F28201BAA471857001A">Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)</mods:title>
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<mods:roleTerm id="629B79EDDB14BA09B201C4881A7EBECF">Author</mods:roleTerm>
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<mods:namePart id="AFCCF0FA30F19A6BAE99A81C02690AFC">Thuy, Ben</mods:namePart>
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<mods:typeOfResource id="1DC89F04D7693119302B34AA7BC91F56">text</mods:typeOfResource>
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<mods:title id="43AAC7AF62715EACD1DE184D2601C60E">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="A7771A53D0B64C2AD8B47C3162183841">
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<mods:date id="C78CD3EA4228CD2843D984ACB362DD0A">2013</mods:date>
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<mods:number id="EDDD99E16B23BAEA03F9D51CFF1915FF">2013-07-02</mods:number>
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<mods:detail id="D54A9A58D6092C3EB72BFF9C4C3629B3" type="issue">
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<mods:number id="AF33E47C5169F14C855833F967A2650E">48</mods:number>
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<mods:identifier id="31FA11D8DD9916D0AB7F4799B885B80B" type="DOI">10.5852/ejt.2013.48</mods:identifier>
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<mods:identifier id="33511CAA8558A932AD07C2718CA58135" type="Zenodo-Dep">3822836</mods:identifier>
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<mods:identifier id="AA09C6F207C9661321C249577485D2E7" type="ZooBank">E7080722-E348-448D-96E5-D537F4865BB5</mods:identifier>
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<treatment id="9978976365E085CCD37524D6FE319EC1" ID-DOI="http://doi.org/10.5281/zenodo.3844115" ID-GBIF-Taxon="164044747" ID-Zenodo-Dep="3844115" LSID="urn:lsid:zoobank.org:act:7B911036-1E01-4EDA-AF2C-E56F0027B8F4" httpUri="http://treatment.plazi.org/id/9978976365E085CCD37524D6FE319EC1" lastPageId="183" lastPageNumber="184" pageId="181" pageNumber="182">
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<subSubSection id="59CB75FE65E085CED37524D6FC179D62" box="[581,1007,511,538]" pageId="181" pageNumber="182" type="nomenclature">
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<paragraph id="116E267565E085CED37524D6FC179D62" blockId="181.[369,1219,511,609]" box="[581,1007,511,538]" pageId="181" pageNumber="182">
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<heading id="4A26911965E085CED37524D6FC179D62" box="[581,1007,511,538]" centered="true" fontSize="11" level="2" pageId="181" pageNumber="182" reason="2">
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<taxonomicName id="D6D15DF665E085CED37524D6FC769D62" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[581,910,511,538]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiomalleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="181" pageNumber="182" phylum="Echinodermata" rank="species" species="beneficarum" status="sp. nov.">
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<emphasis id="23A5FA6765E085CED37524D6FC769D62" bold="true" box="[581,910,511,538]" italics="true" pageId="181" pageNumber="182">Ophiomalleus beneficarum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3896471C65E085CED2A42729FC179D62" box="[916,1007,512,538]" pageId="181" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="59CB75FE65E085CED041270AFCA79D19" pageId="181" pageNumber="182" type="description">
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<paragraph id="116E267565E085CED041270AFB3B9D46" blockId="181.[369,1219,511,609]" box="[369,1219,547,574]" pageId="181" pageNumber="182">
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<uri id="65402A7765E085CED041270AFB3B9D46" box="[369,1219,547,574]" pageId="181" pageNumber="182">
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urn:lsid:zoobank.org:act:
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<uuid id="65771CA065E085CED3BF270AFB3B9D46" box="[655,1219,547,574]" pageId="181" pageNumber="182">7B911036-1E01-4EDA-AF2C-E56F0027B8F4</uuid>
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</uri>
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</paragraph>
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<paragraph id="116E267565E085CED3E5276EFCA79D19" blockId="181.[369,1219,511,609]" box="[725,863,583,609]" pageId="181" pageNumber="182">
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<figureCitation id="89EA3AF065E085CED3E5276EFCD59D19" box="[725,813,583,609]" captionStart="Fig" captionStartId="179.[189,232,1684,1710]" captionTargetBox="[284,1303,265,1644]" captionTargetId="figure@179.[284,1303,265,1644]" captionTargetPageId="179" captionText="Fig. 32. Fossil skeletal plates of ophiacanthid brittle stars; lateral arm plates (LAPs) in external (a) and internal (b) views. 1. Ophiomitrella? sp. 2 from the Bathonian (Middle Jurassic) of Jumara, India; GZG. INV.78740, median to distal LAP. 2. Ophiomitrella sp. nov. from the late Maastrichtian (Late Cretaceous) of Maastricht, the Netherlands; NHMM JJ 5104, proximal LAP. 3-6. Ophiomalleus beneficarum gen. et sp. nov. from the Callovian of Bauer-Wehrland (3), Althüttendorf (4-5) and Hohensaaten (6), Germany. 3. GZG.INV.78741 (holotype), proximal to median LAP. 4. GZG.INV.78742 (paratype), proximal LAP. 5. GZG.INV.78743 (paratype), distal LAP. 6. GZG.INV.78744 (paratype), arm spine. 7-9. Ophiomalleus stevenwilsoni gen. et sp. nov. from the latest Hauterivian (Early Cretaceous) of Sarstedt, Germany. 7. GZG.INV.78748 (holotype), proximal LAP. 8. GZG.INV.78749 (paratype), median LAP. 9. GZG. INV.78750 (paratype), distal LAP. One common scale bar per species except for 6." figureDoi="http://doi.org/10.5281/zenodo.3822905" httpUri="https://zenodo.org/record/3822905/files/figure.png" pageId="181" pageNumber="182">Fig. 32</figureCitation>
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: 3-6
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</paragraph>
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</subSubSection>
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<subSubSection id="59CB75FE65E085CED18D27A0FD839DB0" pageId="181" pageNumber="182" type="reference_group">
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<paragraph id="116E267565E085CED18D27A0FD839DB0" blockId="181.[189,1399,649,712]" pageId="181" pageNumber="182">
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<treatmentCitationGroup id="31C1015B65E085CED18D27A0FD839DB0" pageId="181" pageNumber="182">
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<taxonomicName id="D6D15DF665E085CED18D27A0FE109DDB" authorityName="Muller & Troschel" authorityYear="1842" box="[189,488,649,675]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="181" pageNumber="182" phylum="Echinodermata" rank="genus">
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<emphasis id="23A5FA6765E085CED18D27A0FEAE9DDB" box="[189,342,649,675]" italics="true" pageId="181" pageNumber="182">Ophiacantha</emphasis>
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?
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<emphasis id="23A5FA6765E085CED04327A3FE109DDB" box="[371,488,650,675]" italics="true" pageId="181" pageNumber="182">constricta</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="9070006465E085CED32727A0FCE89DDB" author="Kutscher M." box="[535,784,649,675]" page="61" pageId="181" pageNumber="182" year="1987">
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<bibRefCitation id="75405B8465E085CED32727A0FCE89DDB" author="Kutscher M." box="[535,784,649,675]" pageId="181" pageNumber="182" pagination="50 - 104" refId="ref124575" refString="Kutscher M. 1987 a. Die Echinodermen der Callovien-Geschiebe. Der Geschiebesammler 21: 50 - 104." type="journal article" year="1987">Kutscher 1987a: 61</bibRefCitation>
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</treatmentCitation>
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, pl. 3 figs 1-2 (material incorrectly assigned to
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<taxonomicName id="D6D15DF665E085CED18D2784FD969DB0" authority="Hess, 1966" authorityName="Hess" authorityYear="1966" box="[189,622,685,712]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="181" pageNumber="182" phylum="Echinodermata" rank="species" species="constricta">
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<emphasis id="23A5FA6765E085CED18D2784FEAE9DBF" box="[189,342,685,711]" italics="true" pageId="181" pageNumber="182">Ophiacantha</emphasis>
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?
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<emphasis id="23A5FA6765E085CED05A2787FE279DBF" box="[362,479,686,711]" italics="true" pageId="181" pageNumber="182">constricta</emphasis>
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<bibRefCitation id="75405B8465E085CED0D72787FD969DB0" author="Hess H." box="[487,622,685,712]" pageId="181" pageNumber="182" pagination="1025 - 1063" refId="ref122420" refString="Hess H. 1966. Mikropalaontologische Untersuchungen an Ophiuren V: Die Ophiuren aus dem Argovien (unteres Ober-Oxford) vom Guldental (Kt. Solothurn) und von Savigna (Dept. Jura). Eclogae geologicae Helvetiae 59: 1025 - 1063." type="journal article" year="1966">Hess, 1966</bibRefCitation>
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</taxonomicName>
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).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="59CB75FE65E085CED18D27C6FDEB9CBF" pageId="181" pageNumber="182" type="diagnosis">
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<paragraph id="116E267565E085CED18D27C6FECE9C71" blockId="181.[189,310,751,777]" box="[189,310,751,777]" pageId="181" pageNumber="182">
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<heading id="4A26911965E085CED18D27C6FECE9C71" bold="true" box="[189,310,751,777]" fontSize="11" level="3" pageId="181" pageNumber="182" reason="3">
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<emphasis id="23A5FA6765E085CED18D27C6FECE9C71" bold="true" box="[189,310,751,777]" pageId="181" pageNumber="182">Diagnosis</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="116E267565E085CED18D2637FDEB9CBF" blockId="181.[189,1399,798,967]" pageId="181" pageNumber="182">
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Species of
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<taxonomicName id="D6D15DF665E085CED00B2637FE189C40" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[315,480,798,824]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiomalleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="181" pageNumber="182" phylum="Echinodermata" rank="genus" status="gen. nov.">
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<emphasis id="23A5FA6765E085CED00B2637FE189C40" box="[315,480,798,824]" italics="true" pageId="181" pageNumber="182">Ophiomalleus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3896471C65E085CED0D62636FDA89C41" box="[486,592,799,825]" pageId="181" pageNumber="182" rank="genus">gen. nov.</taxonomicNameLabel>
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with very large LAPs displaying a poorly developed vertical striation on narrow band near spine articulations; at least six poorly defined spurs on proximal outer edge in proximal LAPs; up to five spine articulations with finely corrugated proximal edge and strong dorsalward increase in size except for often very small dorsalmost one; single, moderately broad, strongly prominent, S-shaped ridge on inner side.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="59CB75FE65E085CED18D26C6FB709B24" pageId="181" pageNumber="182" type="etymology">
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<paragraph id="116E267565E085CED18D26C6FEBC9B71" blockId="181.[189,324,1007,1033]" box="[189,324,1007,1033]" pageId="181" pageNumber="182">
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<heading id="4A26911965E085CED18D26C6FEBC9B71" bold="true" box="[189,324,1007,1033]" fontSize="11" level="3" pageId="181" pageNumber="182" reason="3">
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<emphasis id="23A5FA6765E085CED18D26C6FEBC9B71" bold="true" box="[189,324,1007,1033]" pageId="181" pageNumber="182">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="116E267565E085CED18D2136FB709B24" blockId="181.[189,1398,1054,1116]" pageId="181" pageNumber="182">
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Name formed after
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<emphasis id="23A5FA6765E085CED0912137FDE09B40" box="[417,536,1054,1080]" italics="true" pageId="181" pageNumber="182">beneficum</emphasis>
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, Latin for “charitable”, in reference to the infamous
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<emphasis id="23A5FA6765E085CED54B2137FA979B40" box="[1147,1391,1054,1080]" italics="true" pageId="181" pageNumber="182">malleus maleficarum</emphasis>
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, to recall that witch hunts, regrettably, did not go extinct for good in the Dark Ages.
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</paragraph>
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</subSubSection>
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<subSubSection id="59CB75FE65E085CED18D21ADFC959A02" pageId="181" pageNumber="182" type="materials_examined">
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<paragraph id="116E267565E085CED18D21ADFE949BE5" blockId="181.[189,364,1155,1182]" box="[189,364,1155,1182]" pageId="181" pageNumber="182">
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<heading id="4A26911965E085CED18D21ADFE949BE5" bold="true" box="[189,364,1155,1182]" fontSize="11" level="3" pageId="181" pageNumber="182" reason="3">
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<emphasis id="23A5FA6765E085CED18D21ADFE949BE5" bold="true" box="[189,364,1155,1182]" pageId="181" pageNumber="182">Type material</emphasis>
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</heading>
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</paragraph>
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<paragraph id="116E267565E085CED1DA219AFE719B89" blockId="181.[189,393,1203,1265]" pageId="181" pageNumber="182">
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<materialsCitation id="A1B92C2865E085CED1DA219AFE719B89" ID-GBIF-Occurrence="2612041338" collectionCode="GZG" pageId="181" pageNumber="182" specimenCode="GZG.INV.78741" specimenCount="1" typeStatus="holotype">
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<emphasis id="23A5FA6765E085CED1DA219AFEA49BB5" bold="true" box="[234,348,1203,1229]" pageId="181" pageNumber="182">
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||
<typeStatus id="CE6A98D765E085CED1DA219AFEA49BB5" box="[234,348,1203,1229]" pageId="181" pageNumber="182" type="holotype">Holotype</typeStatus>
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</emphasis>
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||
<specimenCode id="41778E0E65E085CED18D21FEFE7D9B89" box="[189,389,1239,1265]" pageId="181" pageNumber="182">GZG.INV.78741</specimenCode>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="116E267565E085CED1DA2030FEA49A4B" blockId="181.[189,1398,1305,1402]" box="[234,348,1305,1331]" pageId="181" pageNumber="182">
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<emphasis id="23A5FA6765E085CED1DA2030FEA49A4B" bold="true" box="[234,348,1305,1331]" pageId="181" pageNumber="182">
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||
<typeStatus id="CE6A98D765E085CED1DA2030FEA49A4B" box="[234,348,1305,1331]" pageId="181" pageNumber="182" type="paratype">Paratype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="116E267565E085CED18D2015FC959A02" blockId="181.[189,1398,1305,1402]" pageId="181" pageNumber="182">
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GZG.INV.78742, GZG.INV.78743 Althütendorf,
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<collectingCountry id="69C666E565E085CED3C22015FC999A2E" box="[754,865,1340,1366]" name="Germany" pageId="181" pageNumber="182">Germany</collectingCountry>
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; and GZG.INV.78744 Hohensaaten,
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<collectingCountry id="69C666E565E085CED4332015FA8A9A2E" box="[1283,1394,1340,1366]" name="Germany" pageId="181" pageNumber="182">Germany</collectingCountry>
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; all from glacial erratic boulders, Callovian, Middle Jurassic.
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</paragraph>
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</subSubSection>
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<subSubSection id="59CB75FE65E085CED18D208BFB9E9A93" pageId="181" pageNumber="182" type="distribution">
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||
<paragraph id="116E267565E085CED18D208BFE039AC4" blockId="181.[189,507,1442,1468]" box="[189,507,1442,1468]" pageId="181" pageNumber="182">
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||
<heading id="4A26911965E085CED18D208BFE039AC4" bold="true" box="[189,507,1442,1468]" fontSize="11" level="3" pageId="181" pageNumber="182" reason="3">
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||
<emphasis id="23A5FA6765E085CED18D208BFE039AC4" bold="true" box="[189,507,1442,1468]" pageId="181" pageNumber="182">Type locality and horizon</emphasis>
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</heading>
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||
</paragraph>
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||
<paragraph id="116E267565E085CED18D20F8FB9E9A93" blockId="181.[189,1126,1489,1515]" box="[189,1126,1489,1515]" pageId="181" pageNumber="182">
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||
Bauer-Wehrland,
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||
<collectingCountry id="69C666E565E085CED0BC20F8FE059A93" box="[396,509,1489,1515]" name="Germany" pageId="181" pageNumber="182">Germany</collectingCountry>
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; glacial erratic boulders, Callovian, Middle Jurassic.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="59CB75FE65E085CED18D233AFC4199DB" pageId="181" pageNumber="182" type="materials_examined">
|
||
<paragraph id="116E267565E085CED18D233AFE4C9955" blockId="181.[189,436,1555,1581]" box="[189,436,1555,1581]" pageId="181" pageNumber="182">
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||
<heading id="4A26911965E085CED18D233AFE4C9955" bold="true" box="[189,436,1555,1581]" fontSize="11" level="3" pageId="181" pageNumber="182" reason="3">
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||
<emphasis id="23A5FA6765E085CED18D233AFE4C9955" bold="true" box="[189,436,1555,1581]" pageId="181" pageNumber="182">Additional material</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="116E267565E085CED18D236BFC4199DB" blockId="181.[189,1399,1602,1700]" pageId="181" pageNumber="182">
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||
GZG.INV.78745 (18 dissociated LAPs) from the Callovian of Bauer-Wehrland,
|
||
<collectingCountry id="69C666E565E085CED59E236BFAE79924" box="[1198,1311,1602,1628]" name="Germany" pageId="181" pageNumber="182">Germany</collectingCountry>
|
||
; GZG. INV.78746 (5 dissociated LAPs) from the Callovian of Althüttendorf,
|
||
<collectingCountry id="69C666E565E085CED534234FFB8D99F8" box="[1028,1141,1638,1664]" name="Germany" pageId="181" pageNumber="182">Germany</collectingCountry>
|
||
; GZG.INV.78747 (11 dissociated LAPs) from the Callovian of Hohensaaten,
|
||
<collectingCountry id="69C666E565E085CED27523A0FC4C99DB" box="[837,948,1673,1699]" name="Germany" pageId="181" pageNumber="182">Germany</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59CB75FE65E085CDD18D23E2FAB299DC" lastPageId="182" lastPageNumber="183" pageId="181" pageNumber="182" type="description">
|
||
<paragraph id="116E267565E085CED18D23E2FEB6999D" blockId="181.[189,334,1739,1765]" box="[189,334,1739,1765]" pageId="181" pageNumber="182">
|
||
<heading id="4A26911965E085CED18D23E2FEB6999D" bold="true" box="[189,334,1739,1765]" fontSize="11" level="3" pageId="181" pageNumber="182" reason="3">
|
||
<emphasis id="23A5FA6765E085CED18D23E2FEB6999D" bold="true" box="[189,334,1739,1765]" pageId="181" pageNumber="182">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="116E267565E085CED1DA23D2FEA4986D" blockId="181.[189,1399,1787,2027]" box="[234,348,1787,1813]" pageId="181" pageNumber="182">
|
||
<emphasis id="23A5FA6765E085CED1DA23D2FEA4986D" bold="true" box="[234,348,1787,1813]" pageId="181" pageNumber="182">
|
||
<typeStatus id="CE6A98D765E085CED1DA23D2FEA4986D" box="[234,348,1787,1813]" pageId="181" pageNumber="182" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="116E267565E085CDD18D2237FB0F9D1D" blockId="181.[189,1399,1787,2027]" lastBlockId="182.[189,1399,267,792]" lastPageId="182" lastPageNumber="183" pageId="181" pageNumber="182">GZG.INV.78741 is a dissociated, very large, proximal to median LAP; approximately 1.5 times higher than wide; main portion of LAP of rounded rectangular outline; dorsal and distal edges gently convex; proximal edge nearly straight, with four relatively small, poorly defined, almost indiscernible, horizontally elongate spurs; ventral portion of LAP very large, conspicuously protruding ventralwards; ventro-distal tip of LAP strongly protruding ventralwards, tongue shaped; outer surface of LAP with fine, rather irregular vertical striation composed of slender, non-overlapping lamellae, and restricted to distal third of outer surface bordering spine articulations; moderately finely meshed stereom on ventral portion of LAP and most of the median third of the outer surface; proximal third with very finely meshed and between spurs weakly horizontally striated stereom. Four very large, ear-shaped spine articulations freestanding on strongly elevated distal portion of LAP; very strong dorsalward increase in size of spine articulations and weaker dorsalward increase in size of gaps separating them; ventral and dorsal lobes of spine articulations merged into continuous volute; proximal edge of volute finely but irregularly corrugated; ventralmost spine articulation on protruding ventro-distal tip of LAP; dorsalmost spine articulation not at dorsal edge of LAP; row of spine articulations slightly oblique; gap between spine articulations and distal edge of LAP nearly as wide as half one spine articulation. Ventral edge of LAP nearly straight, with deeply incised, relatively small, more than semi-circular tentacle notch.</paragraph>
|
||
<paragraph id="116E267565E385CDD18D2759FC909C6F" blockId="182.[189,1399,267,792]" pageId="182" pageNumber="183">Inner side of LAP with large, moderately broad, strongly prominent, nearly S-shaped ridge, with ventral tip pointing ventro-proximalwards and confluent with ventral portion of LAP, and dorsal tip pointing dorsalwards and separated from dorsal edge of LAP; inner side of distal edge of LAP with two relatively large, well-defined, slightly prominent, horizontally elongate spurs. Three irregular perforations loosely arranged in vertical row dorsally bordering tentacle notch.</paragraph>
|
||
<paragraph id="116E267565E385CDD1DA2616FD489C21" blockId="182.[189,1399,831,1285]" box="[234,688,831,857]" pageId="182" pageNumber="183">
|
||
<emphasis id="23A5FA6765E385CDD1DA2616FD489C21" bold="true" box="[234,688,831,857]" pageId="182" pageNumber="183">
|
||
<typeStatus id="CE6A98D765E385CDD1DA2616FEA49C21" box="[234,348,831,857]" pageId="182" pageNumber="183" type="paratype">Paratype</typeStatus>
|
||
supplements and variation
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="116E267565E385CDD18D264AFDDB9B0E" blockId="182.[189,1399,831,1285]" pageId="182" pageNumber="183">
|
||
GZG.INV.78742 is a dissociated proximal LAP; ventral portion missing; LAP originally more than 1.5 times higher than wide; main portion of LAP nearly rectangular to trapezoid in outline; dorsal edge irregularly convex; distal edge nearly straight; proximal edge gently concave, with at least six small to moderately large, poorly defined, horizontally slightly elongate, weakly prominent, nonprotruding spurs; outer surface with moderately finely meshed stereom; trabeculae of stereom near spine articulations thickened and merged into very poorly developed, irregular vertical striation. Five spine articulations similar to those of
|
||
<typeStatus id="CE6A98D765E385CDD31C2111FD6E9B2A" box="[556,662,1080,1106]" pageId="182" pageNumber="183" type="holotype">holotype</typeStatus>
|
||
; strong dorsalward increase in size of spine articulations except for very small dorsalmost one.
|
||
</paragraph>
|
||
<paragraph id="116E267565E385CDD18D2156FBF59A7C" blockId="182.[189,1399,831,1285]" pageId="182" pageNumber="183">
|
||
Dorsal half of ridge on inner side preserved, uniformly broad, well defined, strongly prominent, composed of straight, near-vertical dorsal portion and oblique, slightly bent central part; both parts connected by round kink; at least three spurs similar to those observed on
|
||
<typeStatus id="CE6A98D765E385CDD2B121EEFC149B99" box="[897,1004,1223,1249]" pageId="182" pageNumber="183" type="holotype">holotype</typeStatus>
|
||
. Three very small, closely spaced perforations in very short vertical row dorsally bordering tentacle notch.
|
||
</paragraph>
|
||
<paragraph id="116E267565E385CDD18D2005FB369A80" blockId="182.[189,1399,1324,1599]" pageId="182" pageNumber="183">
|
||
GZG.INV.78743 is a dissociated distal LAP; nearly as wide as high; very well in agreement with
|
||
<typeStatus id="CE6A98D765E385CDD18D2079FEDF9A12" box="[189,295,1360,1386]" pageId="182" pageNumber="183" type="holotype">holotype</typeStatus>
|
||
; dorsal edge concave as a result of a constriction; proximal edge with three spurs similar to those of
|
||
<typeStatus id="CE6A98D765E385CDD019205AFE6B9AF5" box="[297,403,1395,1421]" pageId="182" pageNumber="183" type="holotype">holotype</typeStatus>
|
||
; outer surface with moderately finely meshed stereom, no vertical striation. Three spine articulations similar to those of
|
||
<typeStatus id="CE6A98D765E385CDD3B320BEFD159AC9" box="[643,749,1431,1457]" pageId="182" pageNumber="183" type="holotype">holotype</typeStatus>
|
||
, slightly smaller, in shallow notches of elevated distal edge of LAP and with better-developed fine corrugation on proximal edge of volute. Ventral edge of LAP slightly concave, with deeply incised slightly less than semi-circular tentacle notch.
|
||
</paragraph>
|
||
<paragraph id="116E267565E385CDD18D2328FEB49947" blockId="182.[189,1399,1324,1599]" pageId="182" pageNumber="183">
|
||
Ridge on inner side of LAP similar to that of
|
||
<typeStatus id="CE6A98D765E385CDD3E62328FCC59963" box="[726,829,1537,1563]" pageId="182" pageNumber="183" type="holotype">holotype</typeStatus>
|
||
except for a shorter, near-vertical dorsal portion of the ridge.
|
||
</paragraph>
|
||
<paragraph id="116E267565E385CDD18D234EFAB299DC" blockId="182.[189,1398,1639,1701]" pageId="182" pageNumber="183">GZG.INV.78744 is a very large, cylindrical, slightly bulgy arm spine; tip broken, exposing hollow lumen; spine with moderately coarse, irregularly longitudinal trabecular ridges beset with minute granules.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59CB75FE65E385CCD18D23E5FC739E31" lastPageId="183" lastPageNumber="184" pageId="182" pageNumber="183" type="discussion">
|
||
<paragraph id="116E267565E385CDD18D23E5FED7999E" blockId="182.[189,303,1740,1766]" box="[189,303,1740,1766]" pageId="182" pageNumber="183">
|
||
<heading id="4A26911965E385CDD18D23E5FED7999E" bold="true" box="[189,303,1740,1766]" fontSize="11" level="3" pageId="182" pageNumber="183" reason="3">
|
||
<emphasis id="23A5FA6765E385CDD18D23E5FED7999E" bold="true" box="[189,303,1740,1766]" pageId="182" pageNumber="183">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="116E267565E385CCD18D23D2FC739E31" blockId="182.[189,1399,1787,2027]" lastBlockId="183.[189,1398,267,329]" lastPageId="183" lastPageNumber="184" pageId="182" pageNumber="183">
|
||
The specimens at hand were initially described and figured as
|
||
<taxonomicName id="D6D15DF665E385CDD29123D2FBC2986D" authorityName="Muller & Troschel" authorityYear="1842" box="[929,1082,1787,1813]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="182" pageNumber="183" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="23A5FA6765E385CDD29123D2FBC2986D" box="[929,1082,1787,1813]" italics="true" pageId="182" pageNumber="183">Ophiacantha</emphasis>
|
||
</taxonomicName>
|
||
?
|
||
<emphasis id="23A5FA6765E385CDD56023D5FB3D986D" box="[1104,1221,1788,1813]" italics="true" pageId="182" pageNumber="183">constricta</emphasis>
|
||
Hess, 1965 by
|
||
<bibRefCitation id="75405B8465E385CDD18D2236FE729841" author="Kutscher M." box="[189,394,1823,1849]" pageId="182" pageNumber="183" pagination="50 - 104" refId="ref124575" refString="Kutscher M. 1987 a. Die Echinodermen der Callovien-Geschiebe. Der Geschiebesammler 21: 50 - 104." type="journal article" year="1987">Kutscher (1987a)</bibRefCitation>
|
||
as a result of the misleading inclusion of more than one type of LAPs in the concept of that species. The LAP type including the
|
||
<typeStatus id="CE6A98D765E385CDD3F0226BFCDF9824" box="[704,807,1858,1884]" pageId="182" pageNumber="183" type="holotype">holotype</typeStatus>
|
||
of
|
||
<emphasis id="23A5FA6765E385CDD27E226AFC929825" box="[846,874,1859,1885]" italics="true" pageId="182" pageNumber="183">O.</emphasis>
|
||
?
|
||
<emphasis id="23A5FA6765E385CDD24E226AFC0B9824" box="[894,1011,1859,1884]" italics="true" pageId="182" pageNumber="183">constricta</emphasis>
|
||
, here transferred to
|
||
<taxonomicName id="D6D15DF665E385CDD5D1226BFA8F9824" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[1249,1399,1858,1884]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiogaleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="182" pageNumber="183" phylum="Echinodermata" rank="genus" status="gen. nov.">
|
||
<emphasis id="23A5FA6765E385CDD5D1226BFA8F9824" box="[1249,1399,1858,1884]" italics="true" pageId="182" pageNumber="183">Ophiogaleus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3896471C65E385CDD18D224FFED298F8" box="[189,298,1894,1920]" pageId="182" pageNumber="183" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
(see above), however, fundamentally differs from the LAPs described above. The latter are here assigned to
|
||
<taxonomicName id="D6D15DF665E385CDD0B122A3FDDE98DC" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[385,550,1930,1956]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiomalleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="182" pageNumber="183" phylum="Echinodermata" rank="genus" status="gen. nov.">
|
||
<emphasis id="23A5FA6765E385CDD0B122A3FDDE98DC" box="[385,550,1930,1956]" italics="true" pageId="182" pageNumber="183">Ophiomalleus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3896471C65E385CDD31D22A3FD6298DC" box="[557,666,1930,1956]" pageId="182" pageNumber="183" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, as discussed above. Within this genus, these LAPs are unique in displaying up to five spine articulations, a poorly developed vertical striation in median and, to a lesser extent, proximal LAPs, up to six poorly defined spurs on the outer proximal edge, and a moderately broad, S-shaped ridge on the inner side. Since the LAPs described above display the highly distinctive features of the new genus best it is chosen as its type species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59CB75FE65E285CCD18D2459FE319EC1" box="[189,457,368,441]" pageId="183" pageNumber="184" type="distribution">
|
||
<paragraph id="116E267565E285CCD18D2459FEB79EF2" blockId="183.[189,335,368,394]" box="[189,335,368,394]" pageId="183" pageNumber="184">
|
||
<heading id="4A26911965E285CCD18D2459FEB79EF2" bold="true" box="[189,335,368,394]" fontSize="11" level="3" pageId="183" pageNumber="184" reason="3">
|
||
<emphasis id="23A5FA6765E285CCD18D2459FEB79EF2" bold="true" box="[189,335,368,394]" pageId="183" pageNumber="184">Occurrence</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="116E267565E285CCD18D24B6FE319EC1" blockId="183.[189,457,415,441]" box="[189,457,415,441]" pageId="183" pageNumber="184">
|
||
Callovian of
|
||
<collectingCountry id="69C666E565E285CCD06624B6FE3D9EC1" box="[342,453,415,441]" name="Germany" pageId="183" pageNumber="184">Germany</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |