279 lines
33 KiB
XML
279 lines
33 KiB
XML
<document id="8A55FA783B9ACA4D61C0B65A4A53A5C8" ID-CLB-Dataset="22249" ID-DOI="10.5852/ejt.2013.48" ID-GBIF-Dataset="02866e62-ab61-4cf4-9d98-0248e251084e" ID-ISSN="2118-9773" ID-Zenodo-Dep="3822836" ID-ZooBank="E7080722-E348-448D-96E5-D537F4865BB5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1589219740188" checkinUser="carolina" docAuthor="Thuy, Ben" docDate="2013" docId="9978976365D085F3D3622076FDD6984B" docLanguage="en" docName="ejt-2013-48.pdf.imf" docOrigin="European Journal of Taxonomy 48" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Alternacantha dilionessa Thuy 2013, sp. nov." docType="treatment" docUuid="FD230CAD-9C1E-4C38-B407-1EAB92E10F70" docUuidSource="ZooBank" docVersion="10" lastPageNumber="137" masterDocId="6541EF1B6555857BD1302529FFF89F78" masterDocTitle="Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)" masterLastPageNumber="242" masterPageNumber="1" pageNumber="134" updateTime="1698839159859" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="609FCB88770A8D3FE6950585359E3154">
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<mods:title id="89ACB4DB289AAAA5EB0999F10D65DAB5">Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)</mods:title>
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<mods:name id="2E67B458A39E7BF8822FD5D32E23E42F" type="personal">
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<mods:roleTerm id="2BFEABDE4136BC1A5CE4C247F9C7EBCD">Author</mods:roleTerm>
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<mods:namePart id="985181B2DDFAAB8F85C8C9052D1F3AD4">Thuy, Ben</mods:namePart>
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<mods:typeOfResource id="0B59CF50447A1DCDFA5F5513BA0F7767">text</mods:typeOfResource>
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<mods:title id="01939E1845A57ABF00F94261D7E02296">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="6EB0FD9F2DDDAFFEA8963B82862117D2">
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<mods:date id="3BECDBDF42581F6E1CC1B5787FDC91CF">2013</mods:date>
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<mods:number id="B991F18521514C70C5F4ED79A95855DD">2013-07-02</mods:number>
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<mods:detail id="CB83C54FE69894F7B986A72DACF16374" type="issue">
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<mods:number id="E15AF018CB3EDF6F316092370C5E934C">48</mods:number>
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<mods:end id="0651346BEABA86EBD405FE383875E2E3">242</mods:end>
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<mods:classification id="63FB0DCDA1DA5DE6A65CCF3A25B1B3AB">journal article</mods:classification>
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<mods:identifier id="152B8265E11CD363E30BDDC77EE5A49A" type="CLB-Dataset">22249</mods:identifier>
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<mods:identifier id="8E53C790BB7821D92715740B4010319C" type="DOI">10.5852/ejt.2013.48</mods:identifier>
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<mods:identifier id="9117341136395E389CC5210F6D4893CD" type="GBIF-Dataset">02866e62-ab61-4cf4-9d98-0248e251084e</mods:identifier>
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<mods:identifier id="9C76C5CA601B625893B950267294D005" type="ISSN">2118-9773</mods:identifier>
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<mods:identifier id="CD9EB098262C96F16AC2043C0BB72811" type="Zenodo-Dep">3822836</mods:identifier>
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<mods:identifier id="C0FC76DDD08F0D027AC16EAAD6AF3AB0" type="ZooBank">E7080722-E348-448D-96E5-D537F4865BB5</mods:identifier>
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</mods:mods>
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<treatment id="9978976365D085F3D3622076FDD6984B" ID-DOI="http://doi.org/10.5281/zenodo.3844167" ID-GBIF-Taxon="164044758" ID-Zenodo-Dep="3844167" LSID="urn:lsid:zoobank.org:act:FD230CAD-9C1E-4C38-B407-1EAB92E10F70" httpUri="http://treatment.plazi.org/id/9978976365D085F3D3622076FDD6984B" lastPageId="136" lastPageNumber="137" pageId="133" pageNumber="134">
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<subSubSection id="59CB75FE65D085FED3622076FC1A9A02" box="[594,994,1375,1402]" pageId="133" pageNumber="134" type="nomenclature">
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<paragraph id="116E267565D085FED3622076FC1A9A02" blockId="133.[361,1227,1375,1474]" box="[594,994,1375,1402]" pageId="133" pageNumber="134">
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<heading id="4A26911965D085FED3622076FC1A9A02" box="[594,994,1375,1402]" centered="true" fontSize="11" level="2" pageId="133" pageNumber="134" reason="2">
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<taxonomicName id="D6D15DF665D085FED3622076FC799A02" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[594,897,1375,1402]" class="Ophiuroidea" family="Ophiacanthidae" genus="Alternacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="133" pageNumber="134" phylum="Echinodermata" rank="species" species="dilionessa" status="sp. nov.">
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<emphasis id="23A5FA6765D085FED3622076FC799A02" bold="true" box="[594,897,1375,1402]" italics="true" pageId="133" pageNumber="134">Alternacantha dilionessa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3896471C65D085FED2B72049FC1A9A02" box="[903,994,1376,1402]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="59CB75FE65D085FED05920AAFC6B9AB9" pageId="133" pageNumber="134" type="description">
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<paragraph id="116E267565D085FED05920AAFB339AE5" blockId="133.[361,1227,1375,1474]" box="[361,1227,1411,1437]" pageId="133" pageNumber="134">
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<uri id="65402A7765D085FED05920AAFB339AE5" box="[361,1227,1411,1437]" pageId="133" pageNumber="134">
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urn:lsid:zoobank.org:act:
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<uuid id="65771CA065D085FED3BB20AAFB339AE5" box="[651,1227,1411,1437]" pageId="133" pageNumber="134">FD230CAD-9C1E-4C38-B407-1EAB92E10F70</uuid>
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</uri>
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</paragraph>
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<paragraph id="116E267565D085FED391208EFC6B9AB9" blockId="133.[361,1227,1375,1474]" box="[673,915,1447,1474]" pageId="133" pageNumber="134">
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<figureCitation id="89EA3AF065D085FED391208EFD059AB9" box="[673,765,1447,1473]" captionStart="Fig" captionStartId="132.[189,232,1789,1815]" captionTargetBox="[245,1342,265,1750]" captionTargetId="figure@132.[245,1342,265,1750]" captionTargetPageId="132" captionText="Fig. 24. Fossil lateral arm plates in external (a) and internal (b) views and articulated arm fragment of ophiacanthid brittle stars. 1-2. Alternacantha arges sp. nov. from the late Oxfordian (Late Jurassic) of Raedersdorf, France; M 610 in dorsal (1a) and ventral (1b) views and with details of dorsal (2a) and ventral (2b) arm plating. 3-6. Alternacantha dilionessa sp. nov. from the early Kimmeridgian of the Pointe du Chay, France. 3. GZG.INV.78662 (holotype), proximal LAP. 4. GZG.INV.78663 (paratype), proximal LAP. 5. GZG.INV.78664 (paratype), median LAP. 6. GZG.INV.78665 (paratype), distal LAP. One common scale bar for 1, for 2 and for 3-6, respectively." figureDoi="http://doi.org/10.5281/zenodo.3822889" httpUri="https://zenodo.org/record/3822889/files/figure.png" pageId="133" pageNumber="134">Figs 24</figureCitation>
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: 3-6; 25: 1-3
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</paragraph>
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</subSubSection>
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||
<subSubSection id="59CB75FE65D085FED18D20C0FD5F99E5" pageId="133" pageNumber="134" type="diagnosis">
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<paragraph id="116E267565D085FED18D20C0FECE997B" blockId="133.[189,310,1513,1539]" box="[189,310,1513,1539]" pageId="133" pageNumber="134">
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<heading id="4A26911965D085FED18D20C0FECE997B" bold="true" box="[189,310,1513,1539]" fontSize="11" level="3" pageId="133" pageNumber="134" reason="3">
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<emphasis id="23A5FA6765D085FED18D20C0FECE997B" bold="true" box="[189,310,1513,1539]" pageId="133" pageNumber="134">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="116E267565D085FED18D2331FD5F99E5" blockId="133.[189,1398,1560,1694]" pageId="133" pageNumber="134">
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Species of
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<taxonomicName id="D6D15DF665D085FED0702331FE10994A" authorityName="Thuy & Meyer" authorityYear="2013" box="[320,488,1560,1586]" class="Ophiuroidea" family="Ophiacanthidae" genus="Alternacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="133" pageNumber="134" phylum="Echinodermata" rank="genus">
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<emphasis id="23A5FA6765D085FED0702331FE10994A" box="[320,488,1560,1586]" italics="true" pageId="133" pageNumber="134">Alternacantha</emphasis>
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</taxonomicName>
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with very large LAPs displaying a moderately well-defined vertical striation on the outer surface; up to six large, well-defined spurs on the outer proximal edge, paralleled by large, relatively short, oval spurs on the inner distal edge; distal edge of ventral arm plates evenly convex; distal edge of dorsal arm plates parabolic.
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</paragraph>
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</subSubSection>
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<subSubSection id="59CB75FE65D085FED18D23EFFDD8982E" pageId="133" pageNumber="134" type="etymology">
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<paragraph id="116E267565D085FED18D23EFFEBC9998" blockId="133.[189,324,1734,1760]" box="[189,324,1734,1760]" pageId="133" pageNumber="134">
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<heading id="4A26911965D085FED18D23EFFEBC9998" bold="true" box="[189,324,1734,1760]" fontSize="11" level="3" pageId="133" pageNumber="134" reason="3">
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<emphasis id="23A5FA6765D085FED18D23EFFEBC9998" bold="true" box="[189,324,1734,1760]" pageId="133" pageNumber="134">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="116E267565D085FED18D23DCFDD8982E" blockId="133.[189,1399,1780,1878]" pageId="133" pageNumber="134">
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Name composed of
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<emphasis id="23A5FA6765D085FED09F23DDFDF19876" box="[431,521,1780,1806]" italics="true" pageId="133" pageNumber="134">diligere</emphasis>
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, Latin for “to cherish”, and “lioness”, translating into
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<emphasis id="23A5FA6765D085FED5AC23DDFB389876" box="[1180,1216,1780,1806]" italics="true" pageId="133" pageNumber="134">lea</emphasis>
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in Latin, as an affectionate declaration of esteem to my beloved wife Lea, without whose support the present study would not have been possible.
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</paragraph>
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</subSubSection>
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<subSubSection id="59CB75FE65D085FDD18D2257FEBE9E15" lastPageId="134" lastPageNumber="135" pageId="133" pageNumber="134" type="materials_examined">
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<paragraph id="116E267565D085FED18D2257FE9498E0" blockId="133.[189,364,1918,1944]" box="[189,364,1918,1944]" pageId="133" pageNumber="134">
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<heading id="4A26911965D085FED18D2257FE9498E0" bold="true" box="[189,364,1918,1944]" fontSize="11" level="3" pageId="133" pageNumber="134" reason="3">
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<emphasis id="23A5FA6765D085FED18D2257FE9498E0" bold="true" box="[189,364,1918,1944]" pageId="133" pageNumber="134">Type material</emphasis>
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</heading>
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</paragraph>
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<paragraph id="116E267565D085FED1DA2284FE719893" blockId="133.[189,393,1965,2027]" pageId="133" pageNumber="134">
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<materialsCitation id="A1B92C2865D085FED1DA2284FE719893" ID-GBIF-Occurrence="2612041329" collectionCode="GZG" pageId="133" pageNumber="134" specimenCode="GZG.INV.78662" specimenCount="1" typeStatus="holotype">
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<emphasis id="23A5FA6765D085FED1DA2284FEA498BF" bold="true" box="[234,348,1965,1991]" pageId="133" pageNumber="134">
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<typeStatus id="CE6A98D765D085FED1DA2284FEA498BF" box="[234,348,1965,1991]" pageId="133" pageNumber="134" type="holotype">Holotype</typeStatus>
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</emphasis>
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<specimenCode id="41778E0E65D085FED18D22F8FE7E9893" box="[189,390,2001,2027]" pageId="133" pageNumber="134">GZG.INV.78662</specimenCode>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="116E267565D385FDD1DA2422FE9F9E5D" blockId="134.[189,1399,267,365]" box="[234,359,267,293]" pageId="134" pageNumber="135">
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<emphasis id="23A5FA6765D385FDD1DA2422FE9F9E5D" bold="true" box="[234,359,267,293]" pageId="134" pageNumber="135">
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<typeStatus id="CE6A98D765D385FDD1DA2422FE9F9E5D" box="[234,359,267,293]" pageId="134" pageNumber="135" type="paratype">Paratypes</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="116E267565D385FDD18D2406FEBE9E15" blockId="134.[189,1399,267,365]" pageId="134" pageNumber="135">GZG.INV.78663, GZG.INV.78664, GZG.INV.78665, GZG.INV.78666, GZG.INV.78667 and GZG. INV.78668.</paragraph>
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</subSubSection>
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<subSubSection id="59CB75FE65D385FDD18D24BCFEDA9D7A" pageId="134" pageNumber="135" type="distribution">
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<paragraph id="116E267565D385FDD18D24BCFE039ED7" blockId="134.[189,507,405,431]" box="[189,507,405,431]" pageId="134" pageNumber="135">
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<heading id="4A26911965D385FDD18D24BCFE039ED7" bold="true" box="[189,507,405,431]" fontSize="11" level="3" pageId="134" pageNumber="135" reason="3">
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<emphasis id="23A5FA6765D385FDD18D24BCFE039ED7" bold="true" box="[189,507,405,431]" pageId="134" pageNumber="135">Type locality and horizon</emphasis>
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</heading>
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</paragraph>
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<paragraph id="116E267565D385FDD18D24EDFEDA9D7A" blockId="134.[189,1398,452,514]" pageId="134" pageNumber="135">
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Pointe du Chay near La Rochelle,
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<collectingCountry id="69C666E565D385FDD37924ECFD659EA7" box="[585,669,453,479]" name="France" pageId="134" pageNumber="135">France</collectingCountry>
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;
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<emphasis id="23A5FA6765D385FDD39724EDFCFE9EA6" box="[679,774,452,478]" italics="true" pageId="134" pageNumber="135">Achilles</emphasis>
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Subzone,
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<taxonomicName id="D6D15DF665D385FDD24E24EDFBC69EA7" authority="Zone" authorityName="Zone" box="[894,1086,452,479]" class="Malacostraca" family="Sphaeromatidae" genus="Cymodoce" kingdom="Animalia" order="Isopoda" pageId="134" pageNumber="135" phylum="Arthropoda" rank="genus">
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<emphasis id="23A5FA6765D385FDD24E24EDFC029EA6" box="[894,1018,452,478]" italics="true" pageId="134" pageNumber="135">Cymodoce</emphasis>
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Zone
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</taxonomicName>
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, early Kimmeridgian, Late Jurassic.
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</paragraph>
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</subSubSection>
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<subSubSection id="59CB75FE65D385FDD18D2702FADE9D0D" pageId="134" pageNumber="135" type="materials_examined">
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<paragraph id="116E267565D385FDD18D2702FE4C9D3D" blockId="134.[189,436,555,581]" box="[189,436,555,581]" pageId="134" pageNumber="135">
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<heading id="4A26911965D385FDD18D2702FE4C9D3D" bold="true" box="[189,436,555,581]" fontSize="11" level="3" pageId="134" pageNumber="135" reason="3">
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<emphasis id="23A5FA6765D385FDD18D2702FE4C9D3D" bold="true" box="[189,436,555,581]" pageId="134" pageNumber="135">Additional material</emphasis>
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</heading>
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</paragraph>
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<paragraph id="116E267565D385FDD18D2773FADE9D0D" blockId="134.[189,1318,602,629]" box="[189,1318,602,629]" pageId="134" pageNumber="135">GZG.INV.78669 (512 dissociated LAPs) and GZG.INV.78670 (distal articulated arm fragment).</paragraph>
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</subSubSection>
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<subSubSection id="59CB75FE65D385F3D18D27B5FB0E9A67" lastPageId="136" lastPageNumber="137" pageId="134" pageNumber="135" type="description">
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<paragraph id="116E267565D385FDD18D27B5FEB69DCE" blockId="134.[189,334,668,694]" box="[189,334,668,694]" pageId="134" pageNumber="135">
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<heading id="4A26911965D385FDD18D27B5FEB69DCE" bold="true" box="[189,334,668,694]" fontSize="11" level="3" pageId="134" pageNumber="135" reason="3">
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<emphasis id="23A5FA6765D385FDD18D27B5FEB69DCE" bold="true" box="[189,334,668,694]" pageId="134" pageNumber="135">Description</emphasis>
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</heading>
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</paragraph>
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<paragraph id="116E267565D385FDD1DA27E5FEA49D9E" blockId="134.[189,1399,716,1570]" box="[234,348,716,742]" pageId="134" pageNumber="135">
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<emphasis id="23A5FA6765D385FDD1DA27E5FEA49D9E" bold="true" box="[234,348,716,742]" pageId="134" pageNumber="135">
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<typeStatus id="CE6A98D765D385FDD1DA27E5FEA49D9E" box="[234,348,716,742]" pageId="134" pageNumber="135" type="holotype">Holotype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="116E267565D385FDD18D27D9FAF79A7A" blockId="134.[189,1399,716,1570]" pageId="134" pageNumber="135">GZG.INV.78662 is a dissociated, very large, proximal LAP; approximately twice wider than high; dorsal edge oblique, nearly straight, with pointed dorso-proximal tip; distal edge slightly convex; proximal edge nearly straight, with six small to moderately large, well-defined, prominent, horizontally elongate, proximally pointed and slightly protruding spurs; second and third ventralmost spurs largest; ventral fifth of LAP strongly protruding ventro-proximalwards; ventro-distal edge of LAP round, weakly protruding ventralwards; outer surface with moderately well-developed, rather coarse, slightly undulose, vertical striation restricted to narrow band near spine articulations and composed of slightly overlapping lamellae of variable width; greater part of outer surface with finely meshed stereom. Five very large, ear-shaped spine articulations in shallow notches of elevated distal edge; ventral and dorsal lobes of spine articulations merged into continuous volute; spine articulations proximally sharply bordered by edge of notches rather than well-defined ridge or single distalmost lamella; notches incising outer surface; gap between spine articulations and distal edge of LAP slightly narrower than one spine articulation; dorsalward increase in size of spine articulations and of gaps separating them; dorsalmost spine articulation separated from the remaining four by very large gap with slightly more coarsely meshed stereom. Ventral edge of LAP with relatively small but deeply concave tentacle notch.</paragraph>
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<paragraph id="116E267565D385FDD18D2025FE6A995A" blockId="134.[189,1399,716,1570]" pageId="134" pageNumber="135">Inner side of LAP with very large, conspicuous, sharply defined, prominent, very narrow ridge; ventral part of ridge bent, pointing ventro-proximalwards; central part of ridge straight, oblique, and dorsal part straight and very slightly oblique, not reaching dorsal edge of LAP; all parts connected by rounded kinks; no thickened parts; inner side of distal edge of LAP with three large, moderately well-defined, prominent, relatively short and high oval spurs in the centre and two smaller, poorly defined spurs ventrally and dorsally bordering the three larger ones; inner side of tentacle notch relatively small, well defined laterally. Shallow, poorly defined furrow dorsally bordering tentacle notch, dorsally obscured by shell fragment.</paragraph>
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<paragraph id="116E267565D385FDD1DA2362FD48991C" blockId="134.[189,1399,1610,1889]" box="[234,688,1610,1637]" pageId="134" pageNumber="135">
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<emphasis id="23A5FA6765D385FDD1DA2362FD48991C" bold="true" box="[234,688,1610,1637]" pageId="134" pageNumber="135">
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<typeStatus id="CE6A98D765D385FDD1DA2362FEA4991D" box="[234,348,1611,1637]" pageId="134" pageNumber="135" type="paratype">Paratype</typeStatus>
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supplements and variation
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="116E267565D385FDD18D2347FE639844" blockId="134.[189,1399,1610,1889]" pageId="134" pageNumber="135">
|
||
GZG.INV.78663 is a dissociated, very large, proximal LAP; approximately twice wider than high; agreeing closely with
|
||
<typeStatus id="CE6A98D765D385FDD0FB23BBFDCD99D4" box="[459,565,1682,1708]" pageId="134" pageNumber="135" type="holotype">holotype</typeStatus>
|
||
; proximal edge with one large, central, well-defined, prominent and protruding spur and two smaller, dorsal and ventral, non-protruding spurs; striation on outer surface limited to a very narrow band near the three ventralmost spine articulations. Four spine articulations similar to those observed on
|
||
<typeStatus id="CE6A98D765D385FDD31223D7FD749860" box="[546,652,1790,1816]" pageId="134" pageNumber="135" type="holotype">holotype</typeStatus>
|
||
; dorsalmost one even more widely separated from remaining spine articulations.
|
||
</paragraph>
|
||
<paragraph id="116E267565D385FDD18D226EFD899818" blockId="134.[189,1399,1610,1889]" box="[189,625,1862,1889]" pageId="134" pageNumber="135">
|
||
Inner side similar to that of
|
||
<typeStatus id="CE6A98D765D385FDD333226FFD969818" box="[515,622,1862,1888]" pageId="134" pageNumber="135" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="116E267565D385FDD18D22A0FE399893" blockId="134.[189,1399,1929,2027]" pageId="134" pageNumber="135">
|
||
GZG.INV.78664 is a dissociated median LAP; slightly higher than wide; proximal edge with two large, well-defined, prominent, and strongly protruding spurs. Four spine articulations similar to those observed on
|
||
<typeStatus id="CE6A98D765D385FDD06322F8FE469893" box="[339,446,2001,2027]" pageId="134" pageNumber="135" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<caption id="45AE76FD65D285FCD18D23F3FA899893" ID-DOI="http://doi.org/10.5281/zenodo.3822891" ID-Zenodo-Dep="3822891" httpUri="https://zenodo.org/record/3822891/files/figure.png" pageId="135" pageNumber="136" startId="135.[189,231,1754,1780]" targetBox="[258,1329,264,1713]" targetPageId="135">
|
||
<paragraph id="116E267565D285FCD18D23F3FA899893" blockId="135.[189,1399,1754,2027]" pageId="135" pageNumber="136">
|
||
<emphasis id="23A5FA6765D285FCD18D23F3FEE3998C" bold="true" box="[189,283,1754,1780]" pageId="135" pageNumber="136">Fig. 25.</emphasis>
|
||
Fossil skeletal plates and articulated arm fragments of ophiacanthid brittle stars; lateral arm plates (LAPs) in external (a) and internal (b) views.
|
||
<emphasis id="23A5FA6765D285FCD20E23D4FC93986F" bold="true" box="[830,875,1789,1815]" pageId="135" pageNumber="136">1-3.</emphasis>
|
||
<taxonomicName id="D6D15DF665D285FCD24723D4FB63986F" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[887,1179,1789,1815]" class="Ophiuroidea" family="Ophiacanthidae" genus="Alternacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="135" pageNumber="136" phylum="Echinodermata" rank="species" species="dilionessa" status="sp. nov.">
|
||
<emphasis id="23A5FA6765D285FCD24723D4FB63986F" box="[887,1179,1789,1815]" italics="true" pageId="135" pageNumber="136">Alternacantha dilionessa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3896471C65D285FCD59723D7FAFC9860" box="[1191,1284,1790,1816]" pageId="135" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from the early Kimmeridgian of the Pointe du Chay, France.
|
||
<emphasis id="23A5FA6765D285FCD2122208FCC09843" bold="true" box="[802,824,1825,1851]" pageId="135" pageNumber="136">1.</emphasis>
|
||
GZG.INV.78666 (paratype), arm spine.
|
||
<emphasis id="23A5FA6765D285FCD4252208FAD29843" bold="true" box="[1301,1322,1825,1851]" pageId="135" pageNumber="136">2.</emphasis>
|
||
GZG. INV.78667 (paratype), median arm fragment in ventral view.
|
||
<emphasis id="23A5FA6765D285FCD246226DFC739826" bold="true" box="[886,907,1860,1886]" pageId="135" pageNumber="136">3.</emphasis>
|
||
GZG.INV.78668 (paratype), proximal arm fragment in lateral (a) and dorsal (b) views.
|
||
<emphasis id="23A5FA6765D285FCD39A224EFD2F98F9" bold="true" box="[682,727,1895,1921]" pageId="135" pageNumber="136">4-5.</emphasis>
|
||
<taxonomicName id="D6D15DF665D285FCD3ED2241FEF898DC" authority="(Kristan-Tollmann & Gramann, 1992)" authorityName="Thuy" authorityYear="2013" baseAuthorityName="Kristan-Tollmann & Gramann" baseAuthorityYear="1992" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="135" pageNumber="136" phylum="Echinodermata" rank="species" species="subtilirugosa" status="comb. nov.">
|
||
<emphasis id="23A5FA6765D285FCD3ED2241FBF098F9" box="[733,1032,1895,1921]" italics="true" pageId="135" pageNumber="136">Dermocoma subtilirugosa</emphasis>
|
||
(
|
||
<bibRefCitation id="75405B8465D285FCD525224EFF0098DC" author="Kristan-Tollmann E. & Gramann F." pageId="135" pageNumber="136" pagination="463 - 474" refId="ref124199" refString="Kristan-Tollmann E. & Gramann F. 1992. Paleontological evidence for the Triassic age of rocks dredged from the northern Exmouth Plateau (Tethyan foraminifers, echinoderms and ostracodes). In: Rad U. von, Haq B. U. et al. (eds) Proceedings of the Ocean Drilling Program, Scientific Results 122: 463 - 474. College Station, Texas. http: // dx. doi. org / 10.2973 / odp. proc. sr. 122.186.1992" type="journal article" year="1992">Kristan-Tollmann & Gramann, 1992</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3896471C65D285FCD03622A3FE7C98DD" box="[262,388,1930,1957]" pageId="135" pageNumber="136" rank="species">comb. nov.</taxonomicNameLabel>
|
||
from the Rhaetian (Late Triassic) of Fischerwiese, Austria.
|
||
<emphasis id="23A5FA6765D285FCD51622A3FBC398DC" bold="true" box="[1062,1083,1930,1956]" pageId="135" pageNumber="136">4.</emphasis>
|
||
GZG.INV.78671, proximal LAP.
|
||
<emphasis id="23A5FA6765D285FCD1CE2287FEEB98B0" bold="true" box="[254,275,1966,1992]" pageId="135" pageNumber="136">5.</emphasis>
|
||
GZG.INV.78672, median LAP.
|
||
<emphasis id="23A5FA6765D285FCD3B32284FD6198BF" bold="true" box="[643,665,1965,1991]" pageId="135" pageNumber="136">6.</emphasis>
|
||
<taxonomicName id="D6D15DF665D285FCD3AF2287FCD598BF" authorityName="Hess" authorityYear="1964" box="[671,813,1966,1991]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="135" pageNumber="136" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="23A5FA6765D285FCD3AF2287FCD598BF" box="[671,813,1966,1991]" italics="true" pageId="135" pageNumber="136">Dermocoma</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. innom. 1 from the Rhaetian (Late Triassic) of Fischerwiese, Austria; GZG.INV.78674, median LAP. One common scale bar per species except for 1.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="116E267565DD85F3D18D2425FB359E08" blockId="136.[189,1398,267,369]" pageId="136" pageNumber="137">
|
||
Inner side of LAP with ridge similar to that observed on
|
||
<typeStatus id="CE6A98D765DD85F3D2622422FC449E5D" box="[850,956,267,293]" pageId="136" pageNumber="137" type="holotype">holotype</typeStatus>
|
||
; inner side of distal edge of LAP with two large, moderately well-defined, weakly prominent spurs. Shallow, moderately well-defined, slightly oblique furrow with two very small spine articulations dorsally bordering tentacle notch.
|
||
</paragraph>
|
||
<paragraph id="116E267565DD85F3D18D24B0FB3B9D31" blockId="136.[189,1399,409,735]" pageId="136" pageNumber="137">
|
||
GZG.INV.78665 is a dissociated distal LAP; slightly wider than high; dorsal edge concave as a result of a weakly developed constriction; proximal edge with single, moderately well-defined, prominent and slightly protruding spur; vertical striation well developed on distal two-fifths of the outer surface. Three spine articulations similar to those observed on
|
||
<typeStatus id="CE6A98D765DD85F3D3D62723FCA89D5C" box="[742,848,522,548]" pageId="136" pageNumber="137" type="holotype">holotype</typeStatus>
|
||
; dorsal gap between spine articulations slightly wider than ventral one. Ventral edge of LAP with small, weakly concave tentacle notch.
|
||
</paragraph>
|
||
<paragraph id="116E267565DD85F3D18D277CFDA59DA7" blockId="136.[189,1399,409,735]" pageId="136" pageNumber="137">Inner side of LAP with sharply defined, prominent ridge composed of strongly widened ventral part and ventro-proximally pointing orsal part; inner side of distal edge with single well-defined, slightly prominent spur; inner side of tentacle notch small, sharply defined laterally, pointing ventralwards. No perforations or furrows discernible.</paragraph>
|
||
<paragraph id="116E267565DD85F3D18D2621FCD89C16" blockId="136.[189,1399,776,878]" pageId="136" pageNumber="137">GZG.INV.78666 is a dissociated, large arm spine; cylindrical; slowly tapering; straight; with finely meshed, slightly longitudinally elongate stereom beset with scattered, minute granules; tip broken, exposing lumen filled with loosely meshed stereom.</paragraph>
|
||
<paragraph id="116E267565DD85F3D18D26BFFAD69BCE" blockId="136.[189,1399,918,1207]" pageId="136" pageNumber="137">
|
||
GZG.INV.78667 is an articulated arm fragment counting five proximal segments; LAPs well in agreement with
|
||
<typeStatus id="CE6A98D765DD85F3D1C82695FE9A9CAE" box="[248,354,956,982]" pageId="136" pageNumber="137" type="holotype">holotype</typeStatus>
|
||
; ventral arm plates very large, approximately bell shaped, slightly wider than high, widest distally, with strongly and evenly convex distal edge, concave lateral and latero-distal edges and pointed proximal angle; ventral arm plates overlapping, covering ventral portions of LAPs, the latter meeting underneath ventral arm plates; tentacle pores small, no tentacles scales preserved; dorsal arm plates large, nearly as wide as long, with nearly straight, short proximal edge; straight, distalwards diverging lateral edges and convex, parabolic distal edge; short, well-defined and prominent longitudinal ridge in proximal third of dorsal arm plate outer surface; dorsal arm plates overlapping, separating LAPs.
|
||
</paragraph>
|
||
<paragraph id="116E267565DD85F3D18D21C9FB0E9A67" blockId="136.[189,1399,1248,1312]" pageId="136" pageNumber="137">
|
||
GZG.INV.78668 is an articulated arm fragment counting six proximal to median segments; well in agreement with other arm fragment
|
||
<typeStatus id="CE6A98D765DD85F3D354202FFD359A58" box="[612,717,1286,1312]" pageId="136" pageNumber="137" type="paratype">paratype</typeStatus>
|
||
; tentacle pores with two small, leaf-like scales.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59CB75FE65DD85F3D18D2060FB4B99C6" pageId="136" pageNumber="137" type="discussion">
|
||
<paragraph id="116E267565DD85F3D18D2060FED79A1B" blockId="136.[189,303,1353,1379]" box="[189,303,1353,1379]" pageId="136" pageNumber="137">
|
||
<heading id="4A26911965DD85F3D18D2060FED79A1B" bold="true" box="[189,303,1353,1379]" fontSize="11" level="3" pageId="136" pageNumber="137" reason="3">
|
||
<emphasis id="23A5FA6765DD85F3D18D2060FED79A1B" bold="true" box="[189,303,1353,1379]" pageId="136" pageNumber="137">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="116E267565DD85F3D18D2050FB4B99C6" blockId="136.[189,1399,1401,1727]" pageId="136" pageNumber="137">
|
||
D’Orbigny (1850) already mentioned an articulated arm fragment from the Kimmeridgian of Chay peninsula,
|
||
<collectingCountry id="69C666E565DD85F3D00D20B6FE689AC1" box="[317,400,1439,1465]" name="France" pageId="136" pageNumber="137">France</collectingCountry>
|
||
, listing it as “
|
||
<taxonomicName id="D6D15DF665DD85F3D30620B6FC069AC2" authority="d'Orbigny, 1850" authorityName="d'Orbigny" authorityYear="1850" box="[566,1022,1439,1466]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Amphilepidida" pageId="136" pageNumber="137" phylum="Echinodermata" rank="species" species="bispinosa">
|
||
<emphasis id="23A5FA6765DD85F3D30620B6FCD69AC1" box="[566,814,1439,1465]" italics="true" pageId="136" pageNumber="137">Ophiurella bispinosa</emphasis>
|
||
d’Orbigny, 1850
|
||
</taxonomicName>
|
||
”, and it is more than likely that he was referring to the same species as the one described here. However, since
|
||
<taxonomicName id="D6D15DF665DD85F3D54D20EDFA8E9AA6" authorityName="d'Orbigny" authorityYear="1850" box="[1149,1398,1476,1502]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Amphilepidida" pageId="136" pageNumber="137" phylum="Echinodermata" rank="species" species="bispinosa">
|
||
<emphasis id="23A5FA6765DD85F3D54D20EDFA8E9AA6" box="[1149,1398,1476,1502]" italics="true" pageId="136" pageNumber="137">Ophiurella bispinosa</emphasis>
|
||
</taxonomicName>
|
||
has never been described, it must be considered as
|
||
<emphasis id="23A5FA6765DD85F3D22120C3FC4E997C" box="[785,950,1514,1540]" italics="true" pageId="136" pageNumber="137">nomen nudum</emphasis>
|
||
. The present specimens share greatest similarities with
|
||
<taxonomicName id="D6D15DF665DD85F3D0B02326FD7D9951" authorityName="Thuy & Meyer" authorityYear="2013" box="[384,645,1551,1577]" class="Ophiuroidea" family="Ophiacanthidae" genus="Alternacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="136" pageNumber="137" phylum="Echinodermata" rank="species" species="occulta">
|
||
<emphasis id="23A5FA6765DD85F3D0B02326FD7D9951" box="[384,645,1551,1577]" italics="true" pageId="136" pageNumber="137">Alternacantha occulta</emphasis>
|
||
</taxonomicName>
|
||
which, however, differs in having smaller proximal LAPs with a pointed to tongue-shaped distal edge, as well as fewer and less well-defined spurs on the outer proximal and inner distal edges. In
|
||
<taxonomicName id="D6D15DF665DD85F3D0C22373FD1E990C" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[498,742,1626,1652]" class="Ophiuroidea" family="Ophiacanthidae" genus="Alternacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="136" pageNumber="137" phylum="Echinodermata" rank="species" species="schwermannorum" status="sp. nov.">
|
||
<emphasis id="23A5FA6765DD85F3D0C22373FD1E990C" box="[498,742,1626,1652]" italics="true" pageId="136" pageNumber="137">A. schwermannorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3896471C65DD85F3D3DF2373FCB3990C" box="[751,843,1626,1652]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(see above), the distal edge of the ventral arm plates is slightly truncated rather than evenly convex, and the spurs on the inner side of the distal edge of the LAPs are very thin, horizontally elongate and pointed rather than high and oval.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59CB75FE65DD85F3D18D23C1FDD6984B" box="[189,558,1768,1843]" pageId="136" pageNumber="137" type="distribution">
|
||
<paragraph id="116E267565DD85F3D18D23C1FEB7987A" blockId="136.[189,335,1768,1794]" box="[189,335,1768,1794]" pageId="136" pageNumber="137">
|
||
<heading id="4A26911965DD85F3D18D23C1FEB7987A" bold="true" box="[189,335,1768,1794]" fontSize="11" level="3" pageId="136" pageNumber="137" reason="3">
|
||
<emphasis id="23A5FA6765DD85F3D18D23C1FEB7987A" bold="true" box="[189,335,1768,1794]" pageId="136" pageNumber="137">Occurrence</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="116E267565DD85F3D18D2230FDD6984B" blockId="136.[189,558,1817,1843]" box="[189,558,1817,1843]" pageId="136" pageNumber="137">
|
||
Early Kimmeridgian of
|
||
<collectingCountry id="69C666E565DD85F3D0E62230FDD2984B" box="[470,554,1817,1843]" name="France" pageId="136" pageNumber="137">France</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |