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<document id="1FF27BB4A9F209C219070485020CB85F" ID-CLB-Dataset="22249" ID-DOI="10.5852/ejt.2013.48" ID-GBIF-Dataset="02866e62-ab61-4cf4-9d98-0248e251084e" ID-ISSN="2118-9773" ID-Zenodo-Dep="3822836" ID-ZooBank="E7080722-E348-448D-96E5-D537F4865BB5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1589219740188" checkinUser="carolina" docAuthor="Thuy, Ben" docDate="2013" docId="9978976365CA85DAD3552055FCB29B45" docLanguage="en" docName="ejt-2013-48.pdf.imf" docOrigin="European Journal of Taxonomy 48" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dermacantha Thuy 2013, gen. nov." docType="treatment" docUuid="D73D5436-6A0A-43C1-8298-C5C4BE5D5CA3" docUuidSource="ZooBank" docVersion="10" lastPageNumber="162" masterDocId="6541EF1B6555857BD1302529FFF89F78" masterDocTitle="Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)" masterLastPageNumber="242" masterPageNumber="1" pageNumber="160" updateTime="1698839159859" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2454FB3882703FF0467319A20DD418AF">Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)</mods:title>
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<mods:role id="84F078D4C726783EA07DDD0C57A6F63B">
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<mods:namePart id="D8350B2D678447A33671843F36459359">Thuy, Ben</mods:namePart>
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<mods:title id="288C34472033A1EC29350EF75B4F13CD">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="CACD6F19740620D4FA850734049D227C">
<mods:date id="AA19CEBAF82FB08547F82873E9B33BE7">2013</mods:date>
<mods:detail id="87AA72741B23FB01109A73E08A26C940" type="pubDate">
<mods:number id="264C4226A65A4DE8288D2EF19491202B">2013-07-02</mods:number>
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<mods:detail id="F4C8F1554D2CE4F8A198A487C9A0A333" type="issue">
<mods:number id="BB59047A9934FF24619DE9D9C68E54BE">48</mods:number>
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<mods:classification id="96D5D11B1BCA1C7632650E2BD241D4C4">journal article</mods:classification>
<mods:identifier id="74A5A9BCD53682562C963018CD3F3A80" type="CLB-Dataset">22249</mods:identifier>
<mods:identifier id="32046D5D465CF13C572CD4291A37474A" type="DOI">10.5852/ejt.2013.48</mods:identifier>
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<mods:identifier id="445BEFEE0742E6C9E6A017963777E1A5" type="Zenodo-Dep">3822836</mods:identifier>
<mods:identifier id="3BE9B15D9549634961B9C6B5A53AA77E" type="ZooBank">E7080722-E348-448D-96E5-D537F4865BB5</mods:identifier>
</mods:mods>
<treatment id="9978976365CA85DAD3552055FCB29B45" ID-DOI="http://doi.org/10.5281/zenodo.3844245" ID-GBIF-Taxon="164044763" ID-Zenodo-Dep="3844245" LSID="urn:lsid:zoobank.org:act:D73D5436-6A0A-43C1-8298-C5C4BE5D5CA3" httpUri="http://treatment.plazi.org/id/9978976365CA85DAD3552055FCB29B45" lastPageId="161" lastPageNumber="162" pageId="159" pageNumber="160">
<subSubSection id="59CB75FE65CA85E4D3552055FC379AEE" box="[613,975,1403,1430]" pageId="159" pageNumber="160" type="nomenclature">
<paragraph id="116E267565CA85E4D3552055FC379AEE" blockId="159.[359,1228,1403,1467]" box="[613,975,1403,1430]" pageId="159" pageNumber="160">
<heading id="4A26911965CA85E4D3552055FC379AEE" box="[613,975,1403,1430]" centered="true" fontSize="11" level="2" pageId="159" pageNumber="160" reason="2">
Genus
<taxonomicName id="D6D15DF665CA85E4D3872052FCA59AEE" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[695,861,1403,1430]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="159" pageNumber="160" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="23A5FA6765CA85E4D3872052FCA59AEE" bold="true" box="[695,861,1403,1430]" italics="true" pageId="159" pageNumber="160">Dermacantha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C65CA85E4D2542055FC379AEE" box="[868,975,1404,1430]" pageId="159" pageNumber="160" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE65CA85E4D0572089FB349AC3" box="[359,1228,1440,1467]" pageId="159" pageNumber="160" type="description">
<paragraph id="116E267565CA85E4D0572089FB349AC3" blockId="159.[359,1228,1403,1467]" box="[359,1228,1440,1467]" pageId="159" pageNumber="160">
<uri id="65402A7765CA85E4D0572089FB349AC3" box="[359,1228,1440,1467]" pageId="159" pageNumber="160">
urn:lsid:zoobank.org:act:
<uuid id="65771CA065CA85E4D3BB2089FB349AC3" box="[651,1228,1440,1467]" pageId="159" pageNumber="160">D73D5436-6A0A-43C1-8298-C5C4BE5D5CA3</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE65CA85E4D18D20CAFCB69954" pageId="159" pageNumber="160" type="type_taxon">
<paragraph id="116E267565CA85E4D18D20CAFEA09A84" blockId="159.[189,344,1506,1533]" box="[189,344,1506,1533]" pageId="159" pageNumber="160">
<heading id="4A26911965CA85E4D18D20CAFEA09A84" bold="true" box="[189,344,1506,1533]" fontSize="11" level="3" pageId="159" pageNumber="160" reason="3">
<emphasis id="23A5FA6765CA85E4D18D20CAFEA09A84" bold="true" box="[189,344,1506,1533]" pageId="159" pageNumber="160">
<typeStatus id="CE6A98D765CA85E4D18D20CAFF029A85" box="[189,250,1507,1533]" pageId="159" pageNumber="160">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="116E267565CA85E4D18D233BFCB69954" blockId="159.[189,846,1554,1581]" box="[189,846,1554,1581]" pageId="159" pageNumber="160">
<taxonomicName id="D6D15DF665CA85E4D18D233BFE369954" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[189,462,1554,1580]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="159" pageNumber="160" phylum="Echinodermata" rank="species" species="leonorae" status="sp. nov.">
<emphasis id="23A5FA6765CA85E4D18D233BFE369954" box="[189,462,1554,1580]" italics="true" pageId="159" pageNumber="160">Dermacantha leonorae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C65CA85E4D0E5233AFDCA9955" box="[469,562,1555,1581]" pageId="159" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE65CA85E4D18D237CFEE099BB" pageId="159" pageNumber="160" type="discussion">
<paragraph id="116E267565CA85E4D18D237CFE209917" blockId="159.[189,472,1621,1647]" box="[189,472,1621,1647]" pageId="159" pageNumber="160">
<heading id="4A26911965CA85E4D18D237CFE209917" bold="true" box="[189,472,1621,1647]" fontSize="11" level="3" pageId="159" pageNumber="160" reason="3">
<emphasis id="23A5FA6765CA85E4D18D237CFE209917" bold="true" box="[189,472,1621,1647]" pageId="159" pageNumber="160">Other species included</emphasis>
</heading>
</paragraph>
<paragraph id="116E267565CA85E4D18D23ADFEE099BB" blockId="159.[189,1398,1668,1731]" pageId="159" pageNumber="160">
<taxonomicName id="D6D15DF665CA85E4D18D23ADFE3799E6" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[189,463,1668,1694]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="159" pageNumber="160" phylum="Echinodermata" rank="species" species="pattyana" status="sp. nov.">
<emphasis id="23A5FA6765CA85E4D18D23ADFE3799E6" box="[189,463,1668,1694]" italics="true" pageId="159" pageNumber="160">Dermacantha pattyana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C65CA85E4D0E823ACFDC099E7" box="[472,568,1669,1695]" pageId="159" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="D6D15DF665CA85E4D37623ADFBAF99E7" authority="Kutscher &amp; Hary, 1991" authorityName="Lutken" authorityYear="1855" box="[582,1111,1668,1695]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiocten" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="159" pageNumber="160" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA6765CA85E4D37623ADFD3899E6" box="[582,704,1668,1694]" italics="true" pageId="159" pageNumber="160">Ophiocten</emphasis>
?
<emphasis id="23A5FA6765CA85E4D3E723ACFCCC99E6" box="[727,820,1669,1694]" italics="true" pageId="159" pageNumber="160">seeweni</emphasis>
<bibRefCitation id="75405B8465CA85E4D20E23ADFBAF99E7" author="Kutscher M. &amp; Hary A." box="[830,1111,1668,1695]" pageId="159" pageNumber="160" pagination="37 - 72" refId="ref124680" refString="Kutscher M. &amp; Hary A. 1991. Echinodermen im Unteren Lias (Bucklandi- und Semicostatum-Zone) zwischen Ellange und Elvange (SE-Luxemburg). Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 182: 37 - 72." type="journal article" year="1991">Kutscher &amp; Hary, 1991</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="D6D15DF665CA85E4D5A423ADFA8E99E6" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[1172,1398,1668,1694]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="159" pageNumber="160" phylum="Echinodermata" rank="species" species="carli" status="sp. nov.">
<emphasis id="23A5FA6765CA85E4D5A423ADFA8E99E6" box="[1172,1398,1668,1694]" italics="true" pageId="159" pageNumber="160">Dermacantha carli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C65CA85E4D18D2380FEE099BB" box="[189,280,1705,1731]" pageId="159" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE65CA85E4D18D23C2FCC79893" pageId="159" pageNumber="160" type="diagnosis">
<paragraph id="116E267565CA85E4D18D23C2FECE987D" blockId="159.[189,310,1771,1797]" box="[189,310,1771,1797]" pageId="159" pageNumber="160">
<heading id="4A26911965CA85E4D18D23C2FECE987D" bold="true" box="[189,310,1771,1797]" fontSize="11" level="3" pageId="159" pageNumber="160" reason="3">
<emphasis id="23A5FA6765CA85E4D18D23C2FECE987D" bold="true" box="[189,310,1771,1797]" pageId="159" pageNumber="160">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="116E267565CA85E4D18D2232FCC79893" blockId="159.[189,1399,1819,2027]" pageId="159" pageNumber="160">Ophiacanthid with small to moderately large LAPs; small height/width ratio, even in proximal LAPs; relatively few (up to five) small spine articulations in notches of slightly elevated distal portion of LAP; ventral and dorsal lobes of spine articulations proximally separated by small knob; ventral portion of LAP generally strongly protruding ventro-proximalwards; up to two poorly to well-defined spurs on outer proximal and inner distal edges; inner side of LAPs with simple, slender ridge devoid of sharp kinks or strongly thickened parts; tentacle notch small.</paragraph>
</subSubSection>
<caption id="45AE76FD65F585DBD18D2358FD1D9893" ID-DOI="http://doi.org/10.5281/zenodo.3822899" ID-Zenodo-Dep="3822899" httpUri="https://zenodo.org/record/3822899/files/figure.png" pageId="160" pageNumber="161" startId="160.[189,232,1649,1675]" targetBox="[297,1290,265,1608]" targetPageId="160">
<paragraph id="116E267565F585DBD18D2358FD1D9893" blockId="160.[189,1399,1649,2027]" pageId="160" pageNumber="161">
<emphasis id="23A5FA6765F585DBD18D2358FEE599F3" bold="true" box="[189,285,1649,1675]" pageId="160" pageNumber="161">Fig. 29.</emphasis>
Fossil lateral arm plates (LAPs) of ophiacanthid brittle stars in external (a) and internal (b) views.
<emphasis id="23A5FA6765F585DBD02023BDFEDE99D6" bold="true" box="[272,294,1684,1710]" pageId="160" pageNumber="161">1.</emphasis>
<taxonomicName id="D6D15DF665F585DBD01D23BCFE4699D6" authorityName="Hess" authorityYear="1964" box="[301,446,1685,1710]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="160" pageNumber="161" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA6765F585DBD01D23BCFE4699D6" box="[301,446,1685,1710]" italics="true" pageId="160" pageNumber="161">Dermocoma</emphasis>
</taxonomicName>
sp. nov. innom. 4 from the early Albian (Early Cretaceous) of Leighton Buzzard, Great Britain; NHM EE 14844, proximal LAP.
<emphasis id="23A5FA6765F585DBD3CB239EFCD199A9" bold="true" box="[763,809,1719,1745]" pageId="160" pageNumber="161">2-5.</emphasis>
<taxonomicName id="D6D15DF665F585DBD203239EFBBD99A9" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[819,1093,1719,1745]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="160" pageNumber="161" phylum="Echinodermata" rank="species" species="pattyana" status="gen. et sp. nov.">
<emphasis id="23A5FA6765F585DBD203239EFBBD99A9" box="[819,1093,1719,1745]" italics="true" pageId="160" pageNumber="161">Dermacantha pattyana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C65F585DBD57F2391FAFE99AA" box="[1103,1286,1720,1746]" pageId="160" pageNumber="161" rank="species">gen. et sp. nov.</taxonomicNameLabel>
from the Hettangian (Early Jurassic) of Fontenoille, Belgium.
<emphasis id="23A5FA6765F585DBD27E23F3FC9C998C" bold="true" box="[846,868,1754,1780]" pageId="160" pageNumber="161">2.</emphasis>
MnhnL HE414 (holotype), proximal LAP.
<emphasis id="23A5FA6765F585DBD18D23D4FF2B986F" bold="true" box="[189,211,1789,1815]" pageId="160" pageNumber="161">3.</emphasis>
MnhnL HE415 (paratype), proximal LAP.
<emphasis id="23A5FA6765F585DBD3D723D4FD05986F" bold="true" box="[743,765,1789,1815]" pageId="160" pageNumber="161">4.</emphasis>
MnhnL HE416 (paratype), median LAP.
<emphasis id="23A5FA6765F585DBD5CE23D7FAEC9860" bold="true" box="[1278,1300,1790,1816]" pageId="160" pageNumber="161">5.</emphasis>
MnhnL HE417 (paratype), distal LAP.
<emphasis id="23A5FA6765F585DBD3142208FDAB9843" bold="true" box="[548,595,1825,1851]" pageId="160" pageNumber="161">6-8.</emphasis>
<taxonomicName id="D6D15DF665F585DBD3692208FB7F9843" authority="(Kutscher &amp; Hary, 1991)" authorityName="Thuy" authorityYear="2013" baseAuthorityName="Kutscher &amp; Hary" baseAuthorityYear="1991" box="[601,1159,1825,1851]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="160" pageNumber="161" phylum="Echinodermata" rank="species" species="seeweni" status="comb. nov.">
<emphasis id="23A5FA6765F585DBD3692208FCA69843" box="[601,862,1825,1851]" italics="true" pageId="160" pageNumber="161">Dermacantha seeweni</emphasis>
(
<bibRefCitation id="75405B8465F585DBD25C2208FB869843" author="Kutscher M. &amp; Hary A." box="[876,1150,1825,1851]" pageId="160" pageNumber="161" pagination="37 - 72" refId="ref124680" refString="Kutscher M. &amp; Hary A. 1991. Echinodermen im Unteren Lias (Bucklandi- und Semicostatum-Zone) zwischen Ellange und Elvange (SE-Luxemburg). Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 182: 37 - 72." type="journal article" year="1991">Kutscher &amp; Hary, 1991</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="3896471C65F585DBD5BD2208FAF59843" box="[1165,1293,1825,1851]" pageId="160" pageNumber="161" rank="species">comb. nov.</taxonomicNameLabel>
from the late Sinemurian (Early Jurassic) of Bishop’s Cleeve, Great Britain.
<emphasis id="23A5FA6765F585DBD2E7226DFC159826" bold="true" box="[983,1005,1860,1886]" pageId="160" pageNumber="161">6.</emphasis>
GZG.INV.78706, proximal LAP.
<emphasis id="23A5FA6765F585DBD18D224EFF2B98F9" bold="true" box="[189,211,1895,1921]" pageId="160" pageNumber="161">7.</emphasis>
GZG.INV.78707, median LAP.
<emphasis id="23A5FA6765F585DBD360224EFD9E98F9" bold="true" box="[592,614,1895,1921]" pageId="160" pageNumber="161">8.</emphasis>
GZG.INV.78708, distal LAP.
<emphasis id="23A5FA6765F585DBD2FC224EFBFF98F9" bold="true" box="[972,1031,1895,1921]" pageId="160" pageNumber="161">9-11.</emphasis>
<taxonomicName id="D6D15DF665F585DBD53F224EFAD898F9" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[1039,1312,1895,1921]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="160" pageNumber="161" phylum="Echinodermata" rank="species" species="leonorae" status="gen. et sp. nov.">
<emphasis id="23A5FA6765F585DBD53F224EFAD898F9" box="[1039,1312,1895,1921]" italics="true" pageId="160" pageNumber="161">Dermacantha leonorae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C65F585DBD4182241FEE298DD" pageId="160" pageNumber="161" rank="species">gen. et sp. nov.</taxonomicNameLabel>
from the early Pliensbachian (Early Jurassic) of Blockley, Great Britain.
<emphasis id="23A5FA6765F585DBD5A222A3FB5098DC" bold="true" box="[1170,1192,1930,1956]" pageId="160" pageNumber="161">9.</emphasis>
GZG.INV.78710 (holotype), proximal LAP.
<emphasis id="23A5FA6765F585DBD0C32284FDE098BF" bold="true" box="[499,536,1965,1991]" pageId="160" pageNumber="161">10.</emphasis>
GZG.INV.78711 (paratype), median LAP.
<emphasis id="23A5FA6765F585DBD5352284FBDE98BF" bold="true" box="[1029,1062,1965,1991]" pageId="160" pageNumber="161">11.</emphasis>
GZG.INV.78712 (paratype), distal LAP. One common scale bar per species.
</paragraph>
</caption>
<subSubSection id="59CB75FE65F485DAD18D2422FEAD9EE3" pageId="161" pageNumber="162" type="etymology">
<paragraph id="116E267565F485DAD18D2422FEBC9E5D" blockId="161.[189,324,267,293]" box="[189,324,267,293]" pageId="161" pageNumber="162">
<heading id="4A26911965F485DAD18D2422FEBC9E5D" bold="true" box="[189,324,267,293]" fontSize="11" level="3" pageId="161" pageNumber="162" reason="3">
<emphasis id="23A5FA6765F485DAD18D2422FEBC9E5D" bold="true" box="[189,324,267,293]" pageId="161" pageNumber="162">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="116E267565F485DAD18D2413FEAD9EE3" blockId="161.[189,1399,314,411]" pageId="161" pageNumber="162">
Name composed of
<emphasis id="23A5FA6765F485DAD09B2413FE0E9E2C" box="[427,502,314,340]" italics="true" pageId="161" pageNumber="162">derma</emphasis>
, Greek for “skin”, and
<taxonomicName id="D6D15DF665F485DAD23D2413FC979E2C" authorityName="Laporte &amp; Gory" authorityYear="1838" box="[781,879,314,340]" class="Insecta" family="Buprestidae" genus="Acantha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="161" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="23A5FA6765F485DAD23D2413FC979E2C" box="[781,879,314,340]" italics="true" pageId="161" pageNumber="162">Acantha</emphasis>
</taxonomicName>
, a nymph in Greek mythology whose name literally translates as “thorny”, in reference to the striking similarities in LAP morphology shared with
<taxonomicName id="D6D15DF665F485DAD18D24A8FEB69EE2" authorityName="Hess" authorityYear="1964" box="[189,334,385,410]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="161" pageNumber="162" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA6765F485DAD18D24A8FEB69EE2" box="[189,334,385,410]" italics="true" pageId="161" pageNumber="162">Dermocoma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE65F485DAD18D24EBFCB29B45" pageId="161" pageNumber="162" type="discussion">
<paragraph id="116E267565F485DAD18D24EBFED79EA4" blockId="161.[189,303,450,476]" box="[189,303,450,476]" pageId="161" pageNumber="162">
<heading id="4A26911965F485DAD18D24EBFED79EA4" bold="true" box="[189,303,450,476]" fontSize="11" level="3" pageId="161" pageNumber="162" reason="3">
<emphasis id="23A5FA6765F485DAD18D24EBFED79EA4" bold="true" box="[189,303,450,476]" pageId="161" pageNumber="162">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="116E267565F485DAD18D24D8FACF9CE9" blockId="161.[189,1399,497,913]" pageId="161" pageNumber="162">
The ophiacanthid fossil record includes a number of dissociated LAP
<typeStatus id="CE6A98D765F485DAD2CE24DBFBC49D74" box="[1022,1084,498,524]" pageId="161" pageNumber="162">types</typeStatus>
which, at first sight, seem compatible with the LAP morphological diagnosis of
<taxonomicName id="D6D15DF665F485DAD260273CFC199D56" authorityName="Hess" authorityYear="1964" box="[848,993,533,558]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="161" pageNumber="162" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA6765F485DAD260273CFC199D56" box="[848,993,533,558]" italics="true" pageId="161" pageNumber="162">Dermocoma</emphasis>
</taxonomicName>
, in particular with respect to the strongly protruding ventral portion of the LAPs, the position of the spine articulations, the presence of spurs on the outer proximal and inner distal edges and the shape of the ridge on the inner side. The spine articulation morphology, however, differs markedly. In fact, while the LAPs of
<taxonomicName id="D6D15DF665F485DAD57827A9FB219DE1" authorityName="Hess" authorityYear="1964" box="[1096,1241,640,665]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="161" pageNumber="162" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA6765F485DAD57827A9FB219DE1" box="[1096,1241,640,665]" italics="true" pageId="161" pageNumber="162">Dermocoma</emphasis>
</taxonomicName>
display spine articulations with a continuous volute, those of the LAP
<typeStatus id="CE6A98D765F485DAD272278AFC8D9DC5" box="[834,885,675,701]" pageId="161" pageNumber="162">type</typeStatus>
in question have the ventral and dorsal lobes proximally separated by one or several small knobs. This significant difference is generally combined with a smaller height/width ratio, even in proximal LAPs, and with smaller spine articulations than in
<taxonomicName id="D6D15DF665F485DAD18D2624FEB69C5E" authorityName="Hess" authorityYear="1964" box="[189,334,781,806]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="161" pageNumber="162" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA6765F485DAD18D2624FEB69C5E" box="[189,334,781,806]" italics="true" pageId="161" pageNumber="162">Dermocoma</emphasis>
</taxonomicName>
. Since these differences co-occur consistently,
<taxonomicName id="D6D15DF665F485DAD2422624FBEB9C5F" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[882,1043,781,807]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="161" pageNumber="162" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="23A5FA6765F485DAD2422624FBEB9C5F" box="[882,1043,781,807]" italics="true" pageId="161" pageNumber="162">Dermacantha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C65F485DAD52A2624FB7C9C5F" box="[1050,1156,781,807]" pageId="161" pageNumber="162" rank="genus">gen. nov.</taxonomicNameLabel>
is introduced here to accommodate the
<taxonomicName id="D6D15DF665F485DAD0A12618FDDA9C32" authorityName="Hess" authorityYear="1964" box="[401,546,817,842]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="161" pageNumber="162" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA6765F485DAD0A12618FDDA9C32" box="[401,546,817,842]" italics="true" pageId="161" pageNumber="162">Dermocoma</emphasis>
</taxonomicName>
-like LAP
<typeStatus id="CE6A98D765F485DAD3AC2618FD229C33" box="[668,730,817,843]" pageId="161" pageNumber="162">types</typeStatus>
with small spine articulations displaying a discontinuous volute. The striking similarities in LAP morphology, however, strongly suggest that
<taxonomicName id="D6D15DF665F485DAD581267DFABA9C15" authorityName="Hess" authorityYear="1964" box="[1201,1346,852,877]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermocoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="161" pageNumber="162" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA6765F485DAD581267DFABA9C15" box="[1201,1346,852,877]" italics="true" pageId="161" pageNumber="162">Dermocoma</emphasis>
</taxonomicName>
and
<taxonomicName id="D6D15DF665F485DAD18D265EFEA69CE9" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[189,350,887,913]" class="Ophiuroidea" family="Ophiacanthidae" genus="Dermacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="161" pageNumber="162" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="23A5FA6765F485DAD18D265EFEA69CE9" box="[189,350,887,913]" italics="true" pageId="161" pageNumber="162">Dermacantha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C65F485DAD056265EFE299CE9" box="[358,465,887,913]" pageId="161" pageNumber="162" rank="genus">gen. nov.</taxonomicNameLabel>
share close phylogenetic ties, as suggested by the name of the new genus.
</paragraph>
<paragraph id="116E267565F485DAD18D2691FCB29B45" blockId="161.[189,1399,952,1085]" pageId="161" pageNumber="162">
Superficial similarities are shared with the LAPs of
<taxonomicName id="D6D15DF665F485DAD2052690FC4C9CAB" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[821,948,953,979]" class="Ophiuroidea" family="Ophiacanthidae" genus="Lapidaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="161" pageNumber="162" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="23A5FA6765F485DAD2052690FC4C9CAB" box="[821,948,953,979]" italics="true" pageId="161" pageNumber="162">Lapidaster</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C65F485DAD28F2690FBD49CAB" box="[959,1068,953,979]" pageId="161" pageNumber="162" rank="genus">gen. nov.</taxonomicNameLabel>
with respect to the weakly elevated distal portion of the LAPs and the spurs on the outer proximal and inner distal edges. In
<taxonomicName id="D6D15DF665F485DAD18D26D6FEC49B61" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[189,316,1023,1049]" class="Ophiuroidea" family="Ophiacanthidae" genus="Lapidaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="161" pageNumber="162" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="23A5FA6765F485DAD18D26D6FEC49B61" box="[189,316,1023,1049]" italics="true" pageId="161" pageNumber="162">Lapidaster</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C65F485DAD0752129FE4B9B62" box="[325,435,1024,1050]" pageId="161" pageNumber="162" rank="genus">gen. nov.</taxonomicNameLabel>
, however, the spine articulations are never sunken into notches of the distal LAP portion, and the tentacle notch generally is much larger.
</paragraph>
</subSubSection>
</treatment>
</document>
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