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<document id="EE11008FEAD1781BD71A1E521A4B62BF" ID-CLB-Dataset="22249" ID-DOI="10.5852/ejt.2013.48" ID-GBIF-Dataset="02866e62-ab61-4cf4-9d98-0248e251084e" ID-ISSN="2118-9773" ID-Zenodo-Dep="3822836" ID-ZooBank="E7080722-E348-448D-96E5-D537F4865BB5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1589219740188" checkinUser="carolina" docAuthor="Thuy, Ben" docDate="2013" docId="99789763658985A6D089212FFBC49AFF" docLanguage="en" docName="ejt-2013-48.pdf.imf" docOrigin="European Journal of Taxonomy 48" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ophiojagtus alternatus Thuy 2013, comb. nov." docType="treatment" docVersion="9" lastPageNumber="222" masterDocId="6541EF1B6555857BD1302529FFF89F78" masterDocTitle="Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)" masterLastPageNumber="242" masterPageNumber="1" pageNumber="221" updateTime="1698839159859" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4D57FFDBF89EC8EEC1E88EF98A945C26">Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)</mods:title>
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<mods:namePart id="186D7DAA0A00F8BDC210FAC4A128EA32">Thuy, Ben</mods:namePart>
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<mods:title id="BD9B04C15E53E6EBF6A93F238166044C">European Journal of Taxonomy</mods:title>
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<mods:date id="0928081236DA272192C3C08E053ECF86">2013</mods:date>
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<mods:number id="E698CABAAF7DD6FD057CD20C7B308328">2013-07-02</mods:number>
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<mods:number id="82AA5709A166414C4D55603826ECC361">48</mods:number>
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<treatment id="99789763658985A6D089212FFBC49AFF" ID-DOI="http://doi.org/10.5281/zenodo.3844103" ID-GBIF-Taxon="164044802" ID-Zenodo-Dep="3844103" LSID="urn:lsid:plazi:treatment:99789763658985A6D089212FFBC49AFF" httpUri="http://treatment.plazi.org/id/99789763658985A6D089212FFBC49AFF" lastPageId="221" lastPageNumber="222" pageId="220" pageNumber="221">
<subSubSection id="59CB75FE658985A7D089212FFC9E9B3F" pageId="220" pageNumber="221" type="nomenclature">
<paragraph id="116E2675658985A7D089212FFC9E9B3F" blockId="220.[441,1147,1030,1095]" pageId="220" pageNumber="221">
<taxonomicName id="D6D15DF6658985A7D089212FFC0A9B58" authority="(Kutscher &amp; Jagt, 2000)" authorityName="Thuy" authorityYear="2013" baseAuthorityName="Kutscher &amp; Jagt" baseAuthorityYear="2000" box="[441,1010,1030,1057]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiojagtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="220" pageNumber="221" phylum="Echinodermata" rank="species" species="alternatus" status="comb. nov.">
<emphasis id="23A5FA67658985A7D089212FFD379B59" bold="true" box="[441,719,1030,1057]" italics="true" pageId="220" pageNumber="221">Ophiojagtus alternatus</emphasis>
(
<bibRefCitation id="75405B84658985A7D3EE212FFC119B58" author="Kutscher M. &amp; Jagt J. W. M." box="[734,1001,1030,1057]" pageId="220" pageNumber="221" pagination="45 - 107" refId="ref124720" refString="Kutscher M. &amp; Jagt J. W. M. 2000. Early Maastrichtian ophiuroids from Rugen (northeast Germany) and Mon (Denmark). In: Jagt J. W. M. 2000. Late Cretaceous-Early Palaeogene echinoderms and the K / T boundary in the southeast Netherlands and the northeast Belgium - Part 3: Ophiuroids. Scripta Geologica 121: 45 - 107." type="journal article" year="2000">Kutscher &amp; Jagt, 2000</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="3896471C658985A7D2C9212FFB839B59" box="[1017,1147,1030,1057]" pageId="220" pageNumber="221" rank="species">comb. nov.</taxonomicNameLabel>
<figureCitation id="89EA3AF0658985A7D3FE2104FCDC9B3F" box="[718,804,1069,1095]" captionStart="Fig" captionStartId="217.[189,232,1684,1710]" captionTargetBox="[284,1303,265,1644]" captionTargetId="figure@217.[284,1303,265,1644]" captionTargetPageId="217" captionText="Fig. 38. Fossil lateral arm plates (LAPs) of ophiacanthid brittle stars in external (a) and internal (b) views. 1-4. Ophiojagtus irimurai gen. et sp. nov. from the early Kimmeridgian (Late Jurassic) of the Pointe du Chay, France. 1. GZG.INV.78808 (holotype), proximal LAP. 2. GZG.INV.78809 (paratype), median LAP. 3. GZG.INV.78810 (paratype), median LAP. 4. GZG.INV.78811 (paratype), distal LAP. 5-7. Ophiojagtus acklesi gen. et sp. nov. from the late Aptian (Early Cretaceous) of Wizard Way, Texas. 5. GZG.INV.78814 (holotype), proximal LAP. 6. GZG.INV.78815 (paratype), median to distal LAP. 7. GZG.INV.78816 (paratype), proximal LAP. 8. Ophiojagtus sp. 1 from the early Cenomanian (Late Cretaceous) of Waco, Texas; proximal LAP. 9-10. Ophiojagtus alternatus (Kutscher &amp; Jagt, 2000) comb. nov. from the early late Maastrichtian (Late Cretaceous) of Haccourt, Belgium. 9. NHMM 2012 060, proximal to median LAP. 10. NHMM 2012 061, proximal LAP. One common scale bar per species." figureDoi="http://doi.org/10.5281/zenodo.3822917" httpUri="https://zenodo.org/record/3822917/files/figure.png" pageId="220" pageNumber="221">Fig. 38</figureCitation>
: 9-10
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE658985A7D18D2146FAE89A2D" pageId="220" pageNumber="221" type="reference_group">
<paragraph id="116E2675658985A7D18D2146FBB59BF1" blockId="220.[189,1224,1135,1198]" box="[189,1101,1135,1162]" pageId="220" pageNumber="221">
<treatmentCitationGroup id="31C1015B658985A7D18D2146FBB59BF1" box="[189,1101,1135,1162]" pageId="220" pageNumber="221">
<taxonomicName id="D6D15DF6658985A7D18D2146FC5D9BF2" authority="Kutscher &amp; Jagt, 2000 in Jagt 2000: 7" authorityName="Matsumoto" authorityYear="1915" box="[189,933,1135,1162]" class="Ophiuroidea" family="Ophiomyxidae" genus="Ophiosmilax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="220" pageNumber="221" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658985A7D18D2146FEAB9BF1" box="[189,339,1135,1161]" italics="true" pageId="220" pageNumber="221">Ophiosmilax</emphasis>
?
<treatmentCitation id="90700064658985A7D0572146FC5D9BF2" author="Kutscher M. &amp; Jagt J. W. M." box="[359,933,1135,1162]" page="7" pageId="220" pageNumber="221" year="2000">
<emphasis id="23A5FA67658985A7D0572146FE269BF1" box="[359,478,1135,1161]" italics="true" pageId="220" pageNumber="221">alternatus</emphasis>
<bibRefCitation id="75405B84658985A7D0D62146FD169BF1" author="Kutscher M. &amp; Jagt J. W. M." box="[486,750,1135,1162]" pageId="220" pageNumber="221" pagination="45 - 107" refId="ref124720" refString="Kutscher M. &amp; Jagt J. W. M. 2000. Early Maastrichtian ophiuroids from Rugen (northeast Germany) and Mon (Denmark). In: Jagt J. W. M. 2000. Late Cretaceous-Early Palaeogene echinoderms and the K / T boundary in the southeast Netherlands and the northeast Belgium - Part 3: Ophiuroids. Scripta Geologica 121: 45 - 107." type="journal article" year="2000">
Kutscher &amp; Jagt,
<quantity id="D6298B90658985A7D3832146FD169BF1" box="[691,750,1135,1161]" metricMagnitude="1" metricUnit="m" metricValue="5.08" pageId="220" pageNumber="221" unit="in" value="2000.0">2000</quantity>
</bibRefCitation>
<emphasis id="23A5FA67658985A7D3C52159FCF49BF1" box="[757,780,1136,1161]" italics="true" pageId="220" pageNumber="221">in</emphasis>
<bibRefCitation id="75405B84658985A7D2232159FC5D9BF2" author="Jagt J. W. M." box="[787,933,1135,1162]" pageId="220" pageNumber="221" pagination="1 - 179" refId="ref123589" refString="Jagt J. W. M. 2000. Late Cretaceous-Early Palaeogene echinoderms and the K / T boundary in the southeast Netherlands and nordeast Belgium - Part 3: Ophiuroids. With a chapter on Early Maastrichtian ophiuroids from Rugen (northeast Germany) and Mon (Denmark) by Manfred Kutscher &amp; John W. M. Jagt. Scripta Geologica 121: 1 - 179." type="journal article" year="2000">Jagt 2000: 7</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, pl. 1 figs 1-2.
</treatmentCitationGroup>
</paragraph>
<paragraph id="116E2675658985A7D18D21BAFB309BD5" blockId="220.[189,1224,1135,1198]" box="[189,1224,1171,1198]" pageId="220" pageNumber="221">
<treatmentCitationGroup id="31C1015B658985A7D18D21BAFB309BD5" box="[189,1224,1171,1198]" pageId="220" pageNumber="221">
<taxonomicName id="D6D15DF6658985A7D18D21BAFCBC9BD5" authority="Kutscher &amp; Jagt, 2000: 47 - 48" authorityName="Matsumoto" authorityPageNumber="47 - 48" authorityYear="1915" box="[189,836,1171,1198]" class="Ophiuroidea" family="Ophiomyxidae" genus="Ophiosmilax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="220" pageNumber="221" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658985A7D18D21BAFEAB9BD5" box="[189,339,1171,1197]" italics="true" pageId="220" pageNumber="221">Ophiosmilax</emphasis>
?
<treatmentCitation id="90700064658985A7D05721BAFCBC9BD5" author="Kutscher M. &amp; Jagt J. W. M." box="[359,836,1171,1198]" page="48" pageId="220" pageNumber="221" year="2000">
<emphasis id="23A5FA67658985A7D05721BAFE269BD5" box="[359,478,1171,1197]" italics="true" pageId="220" pageNumber="221">alternatus</emphasis>
<bibRefCitation id="75405B84658985A7D0D621BAFCBC9BD5" author="Kutscher M. &amp; Jagt J. W. M." box="[486,836,1171,1198]" pageId="220" pageNumber="221" pagination="45 - 107" refId="ref124720" refString="Kutscher M. &amp; Jagt J. W. M. 2000. Early Maastrichtian ophiuroids from Rugen (northeast Germany) and Mon (Denmark). In: Jagt J. W. M. 2000. Late Cretaceous-Early Palaeogene echinoderms and the K / T boundary in the southeast Netherlands and the northeast Belgium - Part 3: Ophiuroids. Scripta Geologica 121: 45 - 107." type="journal article" year="2000">Kutscher &amp; Jagt, 2000: 47-48</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, pl. 31 figs 8-12, pl. 33 figs 9-11.
</treatmentCitationGroup>
</paragraph>
<paragraph id="116E2675658985A7D18D21FCFCD09B97" blockId="220.[189,848,1237,1299]" box="[189,808,1237,1264]" pageId="220" pageNumber="221">
<treatmentCitationGroup id="31C1015B658985A7D18D21FCFCD09B97" box="[189,808,1237,1264]" pageId="220" pageNumber="221">
<taxonomicName id="D6D15DF6658985A7D18D21FCFEAB9B97" authorityName="Matsumoto" authorityYear="1915" box="[189,339,1237,1263]" class="Ophiuroidea" family="Ophiomyxidae" genus="Ophiosmilax" kingdom="Animalia" order="Ophiurida" pageId="220" pageNumber="221" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658985A7D18D21FCFEAB9B97" box="[189,339,1237,1263]" italics="true" pageId="220" pageNumber="221">Ophiosmilax</emphasis>
</taxonomicName>
? n. sp. –
<treatmentCitation id="90700064658985A7D0F321FFFD789B97" author="Jagt J. W. M." box="[451,640,1237,1264]" page="199" pageId="220" pageNumber="221" year="1999">
<bibRefCitation id="75405B84658985A7D0F321FFFD789B97" author="Jagt J. W. M." box="[451,640,1237,1264]" pageId="220" pageNumber="221" pagination="197 - 206" refId="ref123504" refString="Jagt J. W. M. 1999 a. Ophiuroid diversity in the type area of the Maastrichtian Stage. Geologie en Mijnbouw 78 (2): 197 - 206." type="journal article" year="1999">Jagt 1999a: 199</bibRefCitation>
</treatmentCitation>
, pl. 1 figs 1-2.
</treatmentCitationGroup>
</paragraph>
<paragraph id="116E2675658985A7D18D21D0FCA89A6B" blockId="220.[189,848,1237,1299]" box="[189,848,1273,1299]" pageId="220" pageNumber="221">
<taxonomicName id="D6D15DF6658985A7D18D21D0FE269A6B" authorityName="Matsumoto" authorityYear="1915" box="[189,478,1273,1299]" class="Ophiuroidea" family="Ophiomyxidae" genus="Ophiosmilax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="220" pageNumber="221" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658985A7D18D21D0FEAB9A6B" box="[189,339,1273,1299]" italics="true" pageId="220" pageNumber="221">Ophiosmilax</emphasis>
?
<emphasis id="23A5FA67658985A7D05721D0FE269A6B" box="[359,478,1273,1299]" italics="true" pageId="220" pageNumber="221">alternatus</emphasis>
</taxonomicName>
–
<treatmentCitation id="90700064658985A7D0CC21D0FD179A6B" box="[508,751,1273,1299]" pageId="220" pageNumber="221">
<bibRefCitation id="75405B84658985A7D0CC21D0FD179A6B" author="Jagt J. W. M. &amp; Riegraf W." box="[508,751,1273,1299]" pageId="220" pageNumber="221" pagination="61 - 63" refId="ref123756" refString="Jagt J. W. M. &amp; Riegraf W. 2003. Late cretaceous [sic] ophiuroids from the Munsterland Basin (Germany): preliminary observations. In: Feral J. - P. &amp; David B. (eds) Echinoderm Research 2001: 61 - 63. Swets &amp; Zeitlinger, Lisse." type="journal article" year="2003">Jagt &amp; Riegraf 2003</bibRefCitation>
</treatmentCitation>
: table 1.
</paragraph>
<paragraph id="116E2675658985A7D18D2015FAE89A2D" blockId="220.[189,1296,1337,1365]" box="[189,1296,1337,1365]" pageId="220" pageNumber="221">
<treatmentCitationGroup id="31C1015B658985A7D18D2015FAE89A2D" box="[189,1296,1337,1365]" pageId="220" pageNumber="221">
<emphasis id="23A5FA67658985A7D18D2015FF119A2D" box="[189,233,1340,1365]" italics="true" pageId="220" pageNumber="221">non</emphasis>
<taxonomicName id="D6D15DF6658985A7D1C02012FDEA9A2D" authorityName="Matsumoto" authorityYear="1915" box="[240,530,1339,1365]" class="Ophiuroidea" family="Ophiomyxidae" genus="Ophiosmilax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="220" pageNumber="221" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658985A7D1C02012FE7E9A2D" box="[240,390,1339,1365]" italics="true" pageId="220" pageNumber="221">Ophiosmilax</emphasis>
?
<emphasis id="23A5FA67658985A7D0AB2012FDEA9A2D" box="[411,530,1339,1365]" italics="true" pageId="220" pageNumber="221">alternatus</emphasis>
</taxonomicName>
–
<treatmentCitation id="90700064658985A7D31F2010FD159A2D" author="Storc R." box="[559,749,1337,1365]" page="397" pageId="220" pageNumber="221" year="2004">
<bibRefCitation id="75405B84658985A7D31F2010FD159A2D" author="Storc R." box="[559,749,1337,1365]" pageId="220" pageNumber="221" pagination="395 - 401" refId="ref128214" refString="Storc R. 2004. The Late Cretaceous brittle star Ophiosmilax? alternatus from Europe and northern Africa. Geobios 37 (3): 395 - 401. http: // dx. doi. org / 10.1016 / j. geobios. 2003.01.010" type="journal article" year="2004">Štorc 2004: 397</bibRefCitation>
</treatmentCitation>
, figs 6-8. —
<treatmentCitation id="90700064658985A7D2B52010FB6B9A2D" author="Storc R. &amp; Zitt J." box="[901,1171,1337,1365]" page="124" pageId="220" pageNumber="221" year="2008">
<bibRefCitation id="75405B84658985A7D2B52010FB6B9A2D" author="Storc R. &amp; Zitt J." box="[901,1171,1337,1365]" pageId="220" pageNumber="221" pagination="123 - 140" refId="ref128259" refString="Storc R. &amp; Zitt J. 2008. Late Turonian ophiuroids (Echinodermata) from the Bohemian Cretaceous Basin, Czech Republic. Bulletin of Geosciences 83 (2): 123 - 140. http: // dx. doi. org / 10.3140 / bull. geosci. 2008.02.123" type="journal article" year="2008">Štorc &amp; Žítt 2008: 124</bibRefCitation>
</treatmentCitation>
, fig. 4A-F.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE658985A7D18D2054FDEB9976" pageId="220" pageNumber="221" type="diagnosis">
<paragraph id="116E2675658985A7D18D2054FECE9AEF" blockId="220.[189,310,1405,1431]" box="[189,310,1405,1431]" pageId="220" pageNumber="221">
<heading id="4A269119658985A7D18D2054FECE9AEF" bold="true" box="[189,310,1405,1431]" fontSize="11" level="3" pageId="220" pageNumber="221" reason="3">
<emphasis id="23A5FA67658985A7D18D2054FECE9AEF" bold="true" box="[189,310,1405,1431]" pageId="220" pageNumber="221">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="116E2675658985A7D18D2085FDEB9976" blockId="220.[189,1399,1452,1550]" pageId="220" pageNumber="221">
Species of
<taxonomicName id="D6D15DF6658985A7D0752085FE2E9ABE" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[325,470,1452,1478]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiojagtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="220" pageNumber="221" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="23A5FA67658985A7D0752085FE2E9ABE" box="[325,470,1452,1478]" italics="true" pageId="220" pageNumber="221">Ophiojagtus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C658985A7D0D12084FDB79ABF" box="[481,591,1453,1479]" pageId="220" pageNumber="221" rank="genus">gen. nov.</taxonomicNameLabel>
with moderately large LAPs displaying an extremely long ventral portion, three to four spine articulations and an evenly bent, slender ridge on the inner side extending well onto the ventral portion.
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE658985A7D18D231FFD199876" pageId="220" pageNumber="221" type="materials_examined">
<paragraph id="116E2675658985A7D18D231FFE529928" blockId="220.[189,426,1590,1616]" box="[189,426,1590,1616]" pageId="220" pageNumber="221">
<heading id="4A269119658985A7D18D231FFE529928" bold="true" box="[189,426,1590,1616]" fontSize="11" level="3" pageId="220" pageNumber="221" reason="3">
<emphasis id="23A5FA67658985A7D18D231FFE529928" bold="true" box="[189,426,1590,1616]" pageId="220" pageNumber="221">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="116E2675658985A7D18D234CFD199876" blockId="220.[189,1399,1637,1806]" pageId="220" pageNumber="221">
34 dissociated LAPs in the Manfred Kutscher Collection in Sassnitz,
<collectingCountry id="69C666E5658985A7D50E234CFB549907" box="[1086,1196,1637,1663]" name="Germany" pageId="220" pageNumber="221">Germany</collectingCountry>
, from the early Maastrichtian of Rügen,
<collectingCountry id="69C666E5658985A7D0DB23A0FDA199DB" box="[491,601,1673,1699]" name="Germany" pageId="220" pageNumber="221">Germany</collectingCountry>
, the
<typeStatus id="CE6A98D7658985A7D3A723A0FD3299DB" box="[663,714,1673,1699]" pageId="220" pageNumber="221">type</typeStatus>
material of
<bibRefCitation id="75405B84658985A7D25323A0FB7D99DB" author="Kutscher M. &amp; Jagt J. W. M." box="[867,1157,1673,1699]" pageId="220" pageNumber="221" pagination="45 - 107" refId="ref124720" refString="Kutscher M. &amp; Jagt J. W. M. 2000. Early Maastrichtian ophiuroids from Rugen (northeast Germany) and Mon (Denmark). In: Jagt J. W. M. 2000. Late Cretaceous-Early Palaeogene echinoderms and the K / T boundary in the southeast Netherlands and the northeast Belgium - Part 3: Ophiuroids. Scripta Geologica 121: 45 - 107." type="journal article" year="2000">Kutscher &amp; Jagt (2000)</bibRefCitation>
;
<collectionCode id="77C0BEB0658985A7D5A423A0FB0B99DB" box="[1172,1267,1673,1699]" country="Netherlands" httpUri="http://grbio.org/cool/6t80-4rpg" name="Natuurhistorische Museum Maastricht" pageId="220" pageNumber="221">NHMM</collectionCode>
2012 060,
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2012 061 and
<collectionCode id="77C0BEB0658985A7D0E92384FDC099BF" box="[473,568,1709,1735]" country="Netherlands" httpUri="http://grbio.org/cool/6t80-4rpg" name="Natuurhistorische Museum Maastricht" pageId="220" pageNumber="221">NHMM</collectionCode>
JJ 3679 (13 dissociated LAPs) from the early late Maastrichtian of Haccourt,
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, the original material of
<bibRefCitation id="75405B84658985A7D38B23F8FCBB9992" author="Jagt J. W. M." box="[699,835,1744,1771]" pageId="220" pageNumber="221" pagination="1 - 179" refId="ref123589" refString="Jagt J. W. M. 2000. Late Cretaceous-Early Palaeogene echinoderms and the K / T boundary in the southeast Netherlands and nordeast Belgium - Part 3: Ophiuroids. With a chapter on Early Maastrichtian ophiuroids from Rugen (northeast Germany) and Mon (Denmark) by Manfred Kutscher &amp; John W. M. Jagt. Scripta Geologica 121: 1 - 179." type="journal article" year="2000">Jagt (2000)</bibRefCitation>
; GZG.INV.78819 (2 dissociated LAPs) from the late Campanian of Lägerdorf-Alsen,
<collectingCountry id="69C666E5658985A7D35E23DDFD259876" box="[622,733,1780,1806]" name="Germany" pageId="220" pageNumber="221">Germany</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE658985A6D18D221FFE829DE3" lastPageId="221" lastPageNumber="222" pageId="220" pageNumber="221" type="description">
<paragraph id="116E2675658985A7D18D221FFEB69828" blockId="220.[189,334,1846,1872]" box="[189,334,1846,1872]" pageId="220" pageNumber="221">
<heading id="4A269119658985A7D18D221FFEB69828" bold="true" box="[189,334,1846,1872]" fontSize="11" level="3" pageId="220" pageNumber="221" reason="3">
<emphasis id="23A5FA67658985A7D18D221FFEB69828" bold="true" box="[189,334,1846,1872]" pageId="220" pageNumber="221">Description</emphasis>
</heading>
</paragraph>
<paragraph id="116E2675658985A6D18D224CFEAF9E99" blockId="220.[189,1399,1893,2027]" lastBlockId="221.[189,1399,267,667]" lastPageId="221" lastPageNumber="222" pageId="220" pageNumber="221">Moderately large, dissociated LAPs of stout, massive and strongly curved aspect; proximal ones more than twice higher than wide, distal ones less than 1.5 times higher than wide; dorsal edge weakly concave as a result of the pointed dorso-proximal tip of the LAP slightly protruding towards arm midline; distal edge convex; dorsal part of proximal edge evenly convex, ventral part concave; ventral portion of LAP narrow, extremely long, accounting for more than half of the total LAP height in proximal LAPs to slightly less than half of the total LAP height in distal ones; outer surface with finely meshed stereom. Three to four large, ear-shaped spine articulations freestanding on strongly bulging distal portion of LAP and composed of thick, continuous volute; dorsalward increase in size of spine articulations and of gaps separating them; gap between spine articulations and distal edge of LAP narrow.</paragraph>
<paragraph id="116E2675658885A6D18D24C5FE829DE3" blockId="221.[189,1399,267,667]" pageId="221" pageNumber="222">Inner side of LAP with well-defined, prominent, slender, evenly bent ridge with ventral part extending well onto the ventral portion of the LAP, and widened dorsal tip with ventro-proximalwards pointing extension; no spurs on inner side of distal edge of LAP; inner side of tentacle notch very weakly defined, almost indiscernible. Small, irregular perforations in very shallow, poorly defined vertical furrow distally bordering ridge.</paragraph>
</subSubSection>
<subSubSection id="59CB75FE658885A6D18D27EDFCE59A6B" pageId="221" pageNumber="222" type="discussion">
<paragraph id="116E2675658885A6D18D27EDFED79DA6" blockId="221.[189,303,708,734]" box="[189,303,708,734]" pageId="221" pageNumber="222">
<heading id="4A269119658885A6D18D27EDFED79DA6" bold="true" box="[189,303,708,734]" fontSize="11" level="3" pageId="221" pageNumber="222" reason="3">
<emphasis id="23A5FA67658885A6D18D27EDFED79DA6" bold="true" box="[189,303,708,734]" pageId="221" pageNumber="222">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="116E2675658885A6D18D27DCFCB39CB2" blockId="221.[189,1399,757,970]" pageId="221" pageNumber="222">
<bibRefCitation id="75405B84658885A6D18D27DCFE269C77" author="Kutscher M. &amp; Jagt J. W. M." box="[189,478,757,783]" pageId="221" pageNumber="222" pagination="45 - 107" refId="ref124720" refString="Kutscher M. &amp; Jagt J. W. M. 2000. Early Maastrichtian ophiuroids from Rugen (northeast Germany) and Mon (Denmark). In: Jagt J. W. M. 2000. Late Cretaceous-Early Palaeogene echinoderms and the K / T boundary in the southeast Netherlands and the northeast Belgium - Part 3: Ophiuroids. Scripta Geologica 121: 45 - 107." type="journal article" year="2000">Kutscher &amp; Jagt (2000)</bibRefCitation>
first described this species on the basis of dissociated LAPs and vertebrae from the early Maastrichtian of
<collectingCountry id="69C666E5658885A6D3042633FD599C4C" box="[564,673,794,820]" name="Germany" pageId="221" pageNumber="222">Germany</collectingCountry>
and
<collectingCountry id="69C666E5658885A6D3EA2633FCB39C4C" box="[730,843,794,820]" name="Denmark" pageId="221" pageNumber="222">Denmark</collectingCountry>
. As noted above, the original assignment to the ophiomyxid
<taxonomicName id="D6D15DF6658885A6D0642669FE129C22" authorityName="Matsumoto" authorityYear="1915" box="[340,490,832,858]" class="Ophiuroidea" family="Ophiomyxidae" genus="Ophiosmilax" kingdom="Animalia" order="Ophiurida" pageId="221" pageNumber="222" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658885A6D0642669FE129C22" box="[340,490,832,858]" italics="true" pageId="221" pageNumber="222">Ophiosmilax</emphasis>
</taxonomicName>
, even if tentative, is untenable. The LAPs in question clearly belong to the new ophiacanthid genus
<taxonomicName id="D6D15DF6658885A6D0EF264CFD889C07" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[479,624,869,895]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiojagtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="221" pageNumber="222" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="23A5FA67658885A6D0EF264CFD889C07" box="[479,624,869,895]" italics="true" pageId="221" pageNumber="222">Ophiojagtus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C658885A6D347264FFD1A9CF8" box="[631,738,870,896]" pageId="221" pageNumber="222" rank="genus">gen. nov.</taxonomicNameLabel>
Within this genus, they are unique in displaying three to four spine articulations, an extremely long ventral portion and a slender, evenly bent ridge on the inner side extending well onto the ventral portion of the LAP.
</paragraph>
<paragraph id="116E2675658885A6D18D26DAFCE59A6B" blockId="221.[189,1399,1009,1299]" pageId="221" pageNumber="222">
The LAPs described and illustrated as
<taxonomicName id="D6D15DF6658885A6D34A26DAFCE89B75" authorityName="Matsumoto" authorityYear="1915" box="[634,784,1011,1037]" class="Ophiuroidea" family="Ophiomyxidae" genus="Ophiosmilax" kingdom="Animalia" order="Ophiurida" pageId="221" pageNumber="222" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658885A6D34A26DAFCE89B75" box="[634,784,1011,1037]" italics="true" pageId="221" pageNumber="222">Ophiosmilax</emphasis>
</taxonomicName>
?
<taxonomicName id="D6D15DF6658885A6D21326DAFC629B75" authorityName="Kutscher &amp; Jagt, 2000 in Jagt" authorityYear="2000" box="[803,922,1011,1037]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiojagtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="221" pageNumber="222" phylum="Echinodermata" rank="species" species="alternatus">
<emphasis id="23A5FA67658885A6D21326DAFC629B75" box="[803,922,1011,1037]" italics="true" pageId="221" pageNumber="222">alternatus</emphasis>
</taxonomicName>
by
<bibRefCitation id="75405B84658885A6D2F326D8FBAD9B75" author="Storc R." box="[963,1109,1009,1037]" pageId="221" pageNumber="222" pagination="395 - 401" refId="ref128214" refString="Storc R. 2004. The Late Cretaceous brittle star Ophiosmilax? alternatus from Europe and northern Africa. Geobios 37 (3): 395 - 401. http: // dx. doi. org / 10.1016 / j. geobios. 2003.01.010" type="journal article" year="2004">Štorc (2004)</bibRefCitation>
from the Turonian of the
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and
<collectingCountry id="69C666E5658885A6D0802131FDFF9B4A" box="[432,519,1048,1074]" name="Tunisia" pageId="221" pageNumber="222">Tunisia</collectingCountry>
display much more coarsely meshed stereom on the outer surface and thus most probably belong to a different, still unnamed species of the present genus. The LAPs described by
<bibRefCitation id="75405B84658885A6D18D2148FE5C9B05" author="Storc R. &amp; Zitt J." box="[189,420,1121,1149]" pageId="221" pageNumber="222" pagination="123 - 140" refId="ref128259" refString="Storc R. &amp; Zitt J. 2008. Late Turonian ophiuroids (Echinodermata) from the Bohemian Cretaceous Basin, Czech Republic. Bulletin of Geosciences 83 (2): 123 - 140. http: // dx. doi. org / 10.3140 / bull. geosci. 2008.02.123" type="journal article" year="2008">Štorc &amp; Žítt (2008)</bibRefCitation>
from the Turonian of the
<collectingCountry id="69C666E5658885A6D3EB214AFC6F9B05" box="[731,919,1123,1149]" name="Czech Republic" pageId="221" pageNumber="222">Czech Republic</collectingCountry>
seem to display four to five, rather than three to four, spine articulations, which would challenge assignment to
<taxonomicName id="D6D15DF6658885A6D52421A0FADD9BDB" authorityName="Thuy" authorityYear="2013" baseAuthorityName="Kutscher &amp; Jagt" baseAuthorityYear="2000" box="[1044,1317,1161,1187]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiojagtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="221" pageNumber="222" phylum="Echinodermata" rank="species" species="alternatus" status="comb. nov.">
<emphasis id="23A5FA67658885A6D52421A0FADD9BDB" box="[1044,1317,1161,1187]" italics="true" pageId="221" pageNumber="222">Ophiojagtus alternatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C658885A6D41E21A1FF179BB0" pageId="221" pageNumber="222" rank="species">comb. nov.</taxonomicNameLabel>
In order to clarify the exact systematic position of these Turonian occurrences of
<taxonomicName id="D6D15DF6658885A6D5D52187FA8E9BB0" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[1253,1398,1198,1224]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiojagtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="221" pageNumber="222" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="23A5FA67658885A6D5D52187FA8E9BB0" box="[1253,1398,1198,1224]" italics="true" pageId="221" pageNumber="222">Ophiojagtus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C658885A6D18D21FDFED39B96" box="[189,299,1236,1262]" pageId="221" pageNumber="222" rank="genus">gen. nov.</taxonomicNameLabel>
, the original material needs to be re-examined, in particular the diagnostically important, but insufficiently described and unillustrated inner side.
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE658885A6D18D2015FBC49AFF" box="[189,1084,1340,1415]" pageId="221" pageNumber="222" type="distribution">
<paragraph id="116E2675658885A6D18D2015FEB79A2E" blockId="221.[189,335,1340,1366]" box="[189,335,1340,1366]" pageId="221" pageNumber="222">
<heading id="4A269119658885A6D18D2015FEB79A2E" bold="true" box="[189,335,1340,1366]" fontSize="11" level="3" pageId="221" pageNumber="222" reason="3">
<emphasis id="23A5FA67658885A6D18D2015FEB79A2E" bold="true" box="[189,335,1340,1366]" pageId="221" pageNumber="222">Occurrence</emphasis>
</heading>
</paragraph>
<paragraph id="116E2675658885A6D18D2044FBC49AFF" blockId="221.[189,1084,1389,1415]" box="[189,1084,1389,1415]" pageId="221" pageNumber="222">
Late Campanian of
<collectingCountry id="69C666E5658885A6D0952044FDEB9AFF" box="[421,531,1389,1415]" name="Germany" pageId="221" pageNumber="222">Germany</collectingCountry>
, early Maastrichtian of
<collectingCountry id="69C666E5658885A6D2192044FC6E9AFF" box="[809,918,1389,1415]" name="Germany" pageId="221" pageNumber="222">Germany</collectingCountry>
and
<collectingCountry id="69C666E5658885A6D2E02044FBC19AFF" box="[976,1081,1389,1415]" name="Belgium" pageId="221" pageNumber="222">Belgium</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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