369 lines
50 KiB
XML
369 lines
50 KiB
XML
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<mods:title id="8D196DFFD6D0959C2F26A8EB21ACA9F5">Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)</mods:title>
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<mods:namePart id="2926F7791B39A7D8CD3F3EE4A778464E">Thuy, Ben</mods:namePart>
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<mods:title id="2D1657393CB00E51CE9B6E4EFB3F474E">European Journal of Taxonomy</mods:title>
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<mods:date id="CD47763E90FDA7B52049A8B64E31621B">2013</mods:date>
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<treatment id="997897636527850ED0C42006FCE29B01" ID-DOI="http://doi.org/10.5281/zenodo.3844093" ID-GBIF-Taxon="164044858" ID-Zenodo-Dep="3844093" LSID="urn:lsid:plazi:treatment:997897636527850ED0C42006FCE29B01" httpUri="http://treatment.plazi.org/id/997897636527850ED0C42006FCE29B01" lastPageId="117" lastPageNumber="118" pageId="114" pageNumber="115">
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<subSubSection id="59CB75FE65278509D0C42006FBC79A32" box="[500,1087,1327,1354]" pageId="114" pageNumber="115" type="nomenclature">
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<paragraph id="116E267565278509D0C42006FBC79A32" blockId="114.[500,1087,1327,1389]" box="[500,1087,1327,1354]" pageId="114" pageNumber="115">
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<heading id="4A26911965278509D0C42006FBC79A32" box="[500,1087,1327,1354]" centered="true" fontSize="11" level="2" pageId="114" pageNumber="115" reason="2">
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<taxonomicName id="D6D15DF665278509D0C42006FC4E9A32" authority="(Hess, 1966)" authorityName="Thuy" authorityYear="2013" baseAuthorityName="Hess" baseAuthorityYear="1966" box="[500,950,1327,1354]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiogaleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="114" pageNumber="115" phylum="Echinodermata" rank="species" species="constrictus" status="comb. nov.">
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<emphasis id="23A5FA6765278509D0C42006FCE19A32" bold="true" box="[500,793,1327,1354]" italics="true" pageId="114" pageNumber="115">Ophiogaleus constrictus</emphasis>
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||
(
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<bibRefCitation id="75405B8465278509D2192019FC569A32" author="Hess H." box="[809,942,1327,1354]" pageId="114" pageNumber="115" pagination="1025 - 1063" refId="ref122420" refString="Hess H. 1966. Mikropalaontologische Untersuchungen an Ophiuren V: Die Ophiuren aus dem Argovien (unteres Ober-Oxford) vom Guldental (Kt. Solothurn) und von Savigna (Dept. Jura). Eclogae geologicae Helvetiae 59: 1025 - 1063." type="journal article" year="1966">Hess, 1966</bibRefCitation>
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||
)
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||
</taxonomicName>
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||
<taxonomicNameLabel id="3896471C65278509D28E2006FBC79A32" box="[958,1087,1327,1354]" pageId="114" pageNumber="115" rank="species">comb. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="59CB75FE65278509D396207AFC769A15" box="[678,910,1363,1389]" pageId="114" pageNumber="115" type="description">
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||
<paragraph id="116E267565278509D396207AFC769A15" blockId="114.[500,1087,1327,1389]" box="[678,910,1363,1389]" pageId="114" pageNumber="115">
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<figureCitation id="89EA3AF065278509D396207AFCFA9A15" box="[678,770,1363,1389]" captionStart="Fig" captionStartId="110.[189,231,1684,1710]" captionTargetBox="[285,1303,265,1642]" captionTargetId="figure@110.[285,1303,265,1642]" captionTargetPageId="110" captionText="Fig. 20. Fossil lateral arm plates (LAPs) of ophiacanthid brittle stars in external (a) and internal (b) views and articulated arm fragment. 1-2. Ophiogaleus dorecki (Hess, 1962) comb. nov. from the late Pliensbachian (Early Jurassic) of Seewen, Switzerland. 1. NHMB M11214, proximal LAP. 2. NHMB M11215, distal LAP. 3-5. Ophiogaleus stans sp. nov. from the early Bathonian (Middle Jurassic) of La Pouza, France. 3. GZG.INV.78615 (holotype), proximal LAP. 4. GZG.INV.78616 (paratype), median LAP. 5. GZG.INV.78617 (paratype), distal LAP. 6-7. Ophiogaleus sp. nov. innom 2 from the Callovian (Middle Jurassic) of Jumara, India. 6. GZG.INV.78619, proximal LAP. 7. GZG.INV.78620, distal LAP. 8-10. Ophiogaleus constrictus (Hess, 1966) comb. nov. from the late Oxfordian (Late Jurassic) of Savigna, France.8. GZG.INV.78624, proximal LAP. 9. GZG.INV.78625, distal LAP. 10. GZG.INV.78626, proximal arm fragment in ventral (a) and dorsal (b) views. One common scale bar per species except for 10." figureDoi="http://doi.org/10.5281/zenodo.3822881" httpUri="https://zenodo.org/record/3822881/files/figure.png" pageId="114" pageNumber="115">Figs 20</figureCitation>
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||
: 8-10; 21: 1
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="59CB75FE65278509D18D20BFFCAA99B0" pageId="114" pageNumber="115" type="reference_group">
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||
<paragraph id="116E267565278509D18D20BFFBDF9AD7" blockId="114.[189,1398,1429,1527]" box="[189,1063,1429,1456]" pageId="114" pageNumber="115">
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||
<treatmentCitationGroup id="31C1015B65278509D18D20BFFBDF9AD7" box="[189,1063,1429,1456]" pageId="114" pageNumber="115">
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||
<emphasis id="23A5FA6765278509D18D20BFFF119AD7" box="[189,233,1430,1455]" italics="true" pageId="114" pageNumber="115">p.p.</emphasis>
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<taxonomicName id="D6D15DF665278509D1C020BCFCDA9AC8" authorityName="Muller & Troschel" authorityYear="1842" box="[240,802,1429,1456]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="114" pageNumber="115" phylum="Echinodermata" rank="genus">
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||
<treatmentCitation id="9070006465278509D1C020BCFE719AD7" box="[240,393,1429,1455]" pageId="114" pageNumber="115">
|
||
<emphasis id="23A5FA6765278509D1C020BCFE719AD7" box="[240,393,1429,1455]" italics="true" pageId="114" pageNumber="115">Ophiacantha</emphasis>
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||
</treatmentCitation>
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||
?
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<emphasis id="23A5FA6765278509D0AE20BFFDB49AD7" box="[414,588,1430,1455]" italics="true" pageId="114" pageNumber="115">suprajurassica</emphasis>
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<bibRefCitation id="75405B8465278509D36420BFFCDA9AC8" box="[596,802,1429,1456]" pageId="114" pageNumber="115">Hess, 1965: 1065</bibRefCitation>
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||
</taxonomicName>
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||
, 1077, figs 12, 14, 43.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="116E267565278509D18D2090FB779A8F" blockId="114.[189,1398,1429,1527]" pageId="114" pageNumber="115">
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||
<treatmentCitationGroup id="31C1015B65278509D18D2090FB779A8F" pageId="114" pageNumber="115">
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||
<taxonomicName id="D6D15DF665278509D18D2090FCE99AAC" authorityName="Muller & Troschel" authorityYear="1842" box="[189,785,1465,1492]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="114" pageNumber="115" phylum="Echinodermata" rank="genus">
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<emphasis id="23A5FA6765278509D18D2090FEAE9AAB" box="[189,342,1465,1491]" italics="true" pageId="114" pageNumber="115">Ophiacantha</emphasis>
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||
?
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<emphasis id="23A5FA6765278509D05E2093FE1B9AAB" box="[366,483,1466,1491]" italics="true" pageId="114" pageNumber="115">constricta</emphasis>
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<treatmentCitation id="9070006465278509D0DE2090FD3C9AAB" author="Hess H." box="[494,708,1465,1491]" page="1031" pageId="114" pageNumber="115" year="1966">
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||
<bibRefCitation id="75405B8465278509D0DE2090FD3C9AAB" author="Hess H." box="[494,708,1465,1491]" pageId="114" pageNumber="115" pagination="1025 - 1063" refId="ref122420" refString="Hess H. 1966. Mikropalaontologische Untersuchungen an Ophiuren V: Die Ophiuren aus dem Argovien (unteres Ober-Oxford) vom Guldental (Kt. Solothurn) und von Savigna (Dept. Jura). Eclogae geologicae Helvetiae 59: 1025 - 1063." type="journal article" year="1966">Hess, 1966: 1031</bibRefCitation>
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||
</treatmentCitation>
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, 1054
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||
</taxonomicName>
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||
, figs 8, 11, 74-75, 78 [figs 76-77 show a different species which is here reinterpreted as
|
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<taxonomicName id="D6D15DF665278509D34A20F4FC759A8F" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[634,909,1501,1527]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiocamax" kingdom="Animalia" order="Ophiurida" pageId="114" pageNumber="115" phylum="Echinodermata" rank="species" species="dorotheae" status="sp. nov.">
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<emphasis id="23A5FA6765278509D34A20F4FC759A8F" box="[634,909,1501,1527]" italics="true" pageId="114" pageNumber="115">Ophiocamax dorotheae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3896471C65278509D2A420F4FC179A8F" box="[916,1007,1501,1527]" pageId="114" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
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(see below)].
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||
</treatmentCitationGroup>
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||
</paragraph>
|
||
<paragraph id="116E267565278509D18D2336FC089941" blockId="114.[189,1398,1566,1736]" box="[189,1008,1566,1593]" pageId="114" pageNumber="115">
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||
<treatmentCitationGroup id="31C1015B65278509D18D2336FC089941" box="[189,1008,1566,1593]" pageId="114" pageNumber="115">
|
||
<emphasis id="23A5FA6765278509D18D2336FE719940" box="[189,393,1566,1592]" italics="true" pageId="114" pageNumber="115">
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non
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<taxonomicName id="D6D15DF665278509D1C02337FE719940" authorityName="Muller & Troschel" authorityYear="1842" box="[240,393,1566,1592]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="114" pageNumber="115" phylum="Echinodermata" rank="genus">Ophiacantha</taxonomicName>
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</emphasis>
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? oder
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<taxonomicName id="D6D15DF665278509D0E92337FDA29940" authorityName="Muller & Troschel" authorityYear="1840" box="[473,602,1566,1592]" class="Ophiuroidea" family="Ophiotrichidae" genus="Ophiothrix" kingdom="Animalia" order="Ophiurida" pageId="114" pageNumber="115" phylum="Echinodermata" rank="genus">
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<emphasis id="23A5FA6765278509D0E92337FDA29940" box="[473,602,1566,1592]" italics="true" pageId="114" pageNumber="115">Ophiothrix</emphasis>
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||
</taxonomicName>
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||
? sp. –
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<treatmentCitation id="9070006465278509D39D2336FC9F9941" author="Hess H." box="[685,871,1566,1593]" page="417" pageId="114" pageNumber="115" year="1960">
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||
<bibRefCitation id="75405B8465278509D39D2336FC9F9941" author="Hess H." box="[685,871,1566,1593]" pageId="114" pageNumber="115" pagination="385 - 421" refId="ref122234" refString="Hess H. 1960. Ophiurenreste aus dem Malm des Schweizer Juras und des Departements Haut-Rhin. Eclogae geologicae Helvetiae 53: 385 - 421." type="journal article" year="1960">Hess 1960: 417</bibRefCitation>
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</treatmentCitation>
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, figs 39-40.
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</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="116E267565278509D18D236AFBD999F8" blockId="114.[189,1398,1566,1736]" pageId="114" pageNumber="115">
|
||
<treatmentCitationGroup id="31C1015B65278509D18D236AFBD999F8" pageId="114" pageNumber="115">
|
||
<emphasis id="23A5FA6765278509D18D236AFE769924" box="[189,398,1602,1628]" italics="true" pageId="114" pageNumber="115">
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||
non
|
||
<taxonomicName id="D6D15DF665278509D1C5236BFE769924" authorityName="Muller & Troschel" authorityYear="1842" box="[245,398,1602,1628]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="114" pageNumber="115" phylum="Echinodermata" rank="genus">Ophiacantha</taxonomicName>
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||
</emphasis>
|
||
? cf.
|
||
<emphasis id="23A5FA6765278509D0E2236AFDBF9924" box="[466,583,1603,1628]" italics="true" pageId="114" pageNumber="115">constricta</emphasis>
|
||
–
|
||
<treatmentCitation id="9070006465278509D35E236AFC219924" author="Hess H. & Holenweg H." box="[622,985,1602,1629]" page="164" pageId="114" pageNumber="115" year="1985">
|
||
<bibRefCitation id="75405B8465278509D35E236AFC219924" author="Hess H. & Holenweg H." box="[622,985,1602,1629]" pageId="114" pageNumber="115" pagination="141 - 177" refId="ref122839" refString="Hess H. & Holenweg H. 1985. Die Begleitfauna auf den Seelilienbanken im mittleren Dogger des Schweizer Juras. Tatigkeitsberichte Naturforschende Gesellschaft Baselland 33: 141 - 177." type="journal article" year="1985">Hess & Holenweg 1985: 164</bibRefCitation>
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||
</treatmentCitation>
|
||
, fig. 17 [material re-described by
|
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<bibRefCitation id="75405B8465278509D18D234FFE3A99F8" author="Thuy B. & Meyer C. A." box="[189,450,1638,1664]" pageId="114" pageNumber="115" pagination="5 - 21" refId="ref128841" refString="Thuy B. & Meyer C. A. 2013. The pitfalls of extrapolating present-day depth ranges to fossil communities: new insights from brittle stars (Echinodermata: Ophiuroidea) from the Middle Jurassic of Switzerland. Swiss Journal of Palaeontology 132 (1): 5 - 21. http: // dx. doi. org / 10.1007 / s 13358 - 012 - 0048 - 5" type="journal article" year="2013">Thuy & Meyer (2013)</bibRefCitation>
|
||
as
|
||
<taxonomicName id="D6D15DF665278509D0DA234FFBEE99F8" authority="Thuy & Meyer, 2013" authorityName="Thuy & Meyer" authorityYear="2013" box="[490,1046,1637,1664]" class="Ophiuroidea" family="Ophiacanthidae" genus="Hanshessia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="114" pageNumber="115" phylum="Echinodermata" rank="species" species="trochitophila">
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||
<emphasis id="23A5FA6765278509D0DA234FFCE899F8" box="[490,784,1638,1664]" italics="true" pageId="114" pageNumber="115">Hanshessia trochitophila</emphasis>
|
||
<bibRefCitation id="75405B8465278509D227234FFBEE99F8" author="Thuy B. & Meyer C. A." box="[791,1046,1637,1664]" pageId="114" pageNumber="115" pagination="5 - 21" refId="ref128841" refString="Thuy B. & Meyer C. A. 2013. The pitfalls of extrapolating present-day depth ranges to fossil communities: new insights from brittle stars (Echinodermata: Ophiuroidea) from the Middle Jurassic of Switzerland. Swiss Journal of Palaeontology 132 (1): 5 - 21. http: // dx. doi. org / 10.1007 / s 13358 - 012 - 0048 - 5" type="journal article" year="2013">Thuy & Meyer, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
].
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="116E267565278509D18D23A3FCAA99B0" blockId="114.[189,1398,1566,1736]" pageId="114" pageNumber="115">
|
||
<treatmentCitationGroup id="31C1015B65278509D18D23A3FCAA99B0" pageId="114" pageNumber="115">
|
||
<emphasis id="23A5FA6765278509D18D23A3FE6899DC" box="[189,400,1674,1700]" italics="true" pageId="114" pageNumber="115">
|
||
non
|
||
<taxonomicName id="D6D15DF665278509D1C723A3FE6899DC" authorityName="Muller & Troschel" authorityYear="1842" box="[247,400,1674,1700]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="114" pageNumber="115" phylum="Echinodermata" rank="genus">Ophiacantha</taxonomicName>
|
||
</emphasis>
|
||
?
|
||
<emphasis id="23A5FA6765278509D09C23A3FDD999DB" box="[428,545,1674,1699]" italics="true" pageId="114" pageNumber="115">constricta</emphasis>
|
||
–
|
||
<treatmentCitation id="9070006465278509D37D23A0FCBC99DB" author="Kutscher M." box="[589,836,1673,1700]" page="61" pageId="114" pageNumber="115" year="1987">
|
||
<bibRefCitation id="75405B8465278509D37D23A0FCBC99DB" author="Kutscher M." box="[589,836,1673,1700]" pageId="114" pageNumber="115" pagination="50 - 104" refId="ref124575" refString="Kutscher M. 1987 a. Die Echinodermen der Callovien-Geschiebe. Der Geschiebesammler 21: 50 - 104." type="journal article" year="1987">Kutscher 1987a: 61</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 3 figs 1-2 [material here redescribed as
|
||
<taxonomicName id="D6D15DF665278509D18D2384FE0299BF" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[189,506,1709,1735]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiomalleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="114" pageNumber="115" phylum="Echinodermata" rank="species" species="beneficarum" status="gen. et sp. nov.">
|
||
<emphasis id="23A5FA6765278509D18D2384FE0299BF" box="[189,506,1709,1735]" italics="true" pageId="114" pageNumber="115">Ophiomalleus beneficarum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3896471C65278509D3312387FD4999B0" box="[513,689,1710,1736]" pageId="114" pageNumber="115" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
(see below)].
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59CB75FE65278509D18D23C6FB779893" pageId="114" pageNumber="115" type="diagnosis">
|
||
<paragraph id="116E267565278509D18D23C6FECE9871" blockId="114.[189,310,1775,1801]" box="[189,310,1775,1801]" pageId="114" pageNumber="115">
|
||
<heading id="4A26911965278509D18D23C6FECE9871" bold="true" box="[189,310,1775,1801]" fontSize="11" level="3" pageId="114" pageNumber="115" reason="3">
|
||
<emphasis id="23A5FA6765278509D18D23C6FECE9871" bold="true" box="[189,310,1775,1801]" pageId="114" pageNumber="115">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="116E267565278509D18D2237FB779893" blockId="114.[189,1399,1822,2027]" pageId="114" pageNumber="115">
|
||
Species of
|
||
<taxonomicName id="D6D15DF665278509D0722237FE209840" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[322,472,1822,1848]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiogaleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="114" pageNumber="115" phylum="Echinodermata" rank="genus" status="gen. nov.">
|
||
<emphasis id="23A5FA6765278509D0722237FE209840" box="[322,472,1822,1848]" italics="true" pageId="114" pageNumber="115">Ophiogaleus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3896471C65278509D0D12236FDB69841" box="[481,590,1823,1849]" pageId="114" pageNumber="115" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
with relatively small LAPs displaying moderately coarsely meshed stereom on outer surface; up to ten spine articulations on strongly elevated distal edge proximally sharply bordered by undulose ridge; three poorly defined spurs on outer proximal edge paralleled by up to four well-defined spurs on inner distal edge in proximal LAPs; single large, well-defined spur on outer proximal and inner distal edges in median to distal LAPs; ridge on inner side of LAPs with strongly widened dorsal part with conspicuously pointed dorsal and ventro-proximal angles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59CB75FE65268508D18D2422FC359D4A" pageId="115" pageNumber="116" type="materials_examined">
|
||
<paragraph id="116E267565268508D18D2422FE529E5D" blockId="115.[189,426,267,293]" box="[189,426,267,293]" pageId="115" pageNumber="116">
|
||
<heading id="4A26911965268508D18D2422FE529E5D" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="115" pageNumber="116" reason="3">
|
||
<emphasis id="23A5FA6765268508D18D2422FE529E5D" bold="true" box="[189,426,267,293]" pageId="115" pageNumber="116">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="116E267565268508D18D2412FC359D4A" blockId="115.[189,1399,315,562]" pageId="115" pageNumber="116">
|
||
GZG.INV.78624, GZG.INV.78625, GZG.INV.78626, GZG.INV.78627, GZG.INV.78628 (7 dissociated LAPs) from sample S1 of
|
||
<bibRefCitation id="75405B8465268508D0DB2476FD819E02" author="Gale A. S." box="[491,633,351,378]" pageId="115" pageNumber="116" pagination="69 - 89" refId="ref121688" refString="Gale A. S. 2011. Asteroidea (Echinodermata) from the Oxfordian (Late Jurassic) of Savigna, Departement du Jura, France. Swiss Journal of Palaeontology 130 (1): 69 - 89. http: // dx. doi. org / 10.1007 / s 13358 - 010 - 0008 - x" type="journal article" year="2011">Gale (2011)</bibRefCitation>
|
||
; GZG.INV.78629 (14 dissociated LAPs) from sample S2a of
|
||
<bibRefCitation id="75405B8465268508D40F2476FEF69EE6" author="Gale A. S." pageId="115" pageNumber="116" pagination="69 - 89" refId="ref121688" refString="Gale A. S. 2011. Asteroidea (Echinodermata) from the Oxfordian (Late Jurassic) of Savigna, Departement du Jura, France. Swiss Journal of Palaeontology 130 (1): 69 - 89. http: // dx. doi. org / 10.1007 / s 13358 - 010 - 0008 - x" type="journal article" year="2011">Gale (2011)</bibRefCitation>
|
||
; GZG.INV.78630 (72 dissociated LAPs) from sample S2b of
|
||
<bibRefCitation id="75405B8465268508D2D424ADFB8C9EE6" author="Gale A. S." box="[996,1140,388,414]" pageId="115" pageNumber="116" pagination="69 - 89" refId="ref121688" refString="Gale A. S. 2011. Asteroidea (Echinodermata) from the Oxfordian (Late Jurassic) of Savigna, Departement du Jura, France. Swiss Journal of Palaeontology 130 (1): 69 - 89. http: // dx. doi. org / 10.1007 / s 13358 - 010 - 0008 - x" type="journal article" year="2011">Gale (2011)</bibRefCitation>
|
||
, and GZG.INV.78631 (4 articulated arm fragments) from sample S2b of
|
||
<bibRefCitation id="75405B8465268508D23A2480FC609EBB" author="Gale A. S." box="[778,920,425,451]" pageId="115" pageNumber="116" pagination="69 - 89" refId="ref121688" refString="Gale A. S. 2011. Asteroidea (Echinodermata) from the Oxfordian (Late Jurassic) of Savigna, Departement du Jura, France. Swiss Journal of Palaeontology 130 (1): 69 - 89. http: // dx. doi. org / 10.1007 / s 13358 - 010 - 0008 - x" type="journal article" year="2011">Gale (2011)</bibRefCitation>
|
||
, all from the upper Oxfordian
|
||
<emphasis id="23A5FA6765268508D5CD2480FA8E9EBB" box="[1277,1398,425,451]" italics="true" pageId="115" pageNumber="116">Bifurcatus</emphasis>
|
||
Zone of Savigna,
|
||
<collectingCountry id="69C666E565268508D0BA24E7FE269E90" box="[394,478,462,488]" name="France" pageId="115" pageNumber="116">France</collectingCountry>
|
||
; 22 dissociated LAPs from the upper Oxfordian
|
||
<emphasis id="23A5FA6765268508D53D24E7FB7E9E90" box="[1037,1158,462,488]" italics="true" pageId="115" pageNumber="116">Bifurcatus</emphasis>
|
||
Zone of Guldenthal,
|
||
<collectingCountry id="69C666E565268508D18D24DBFEB49D74" box="[189,332,498,524]" name="Switzerland" pageId="115" pageNumber="116">Switzerland</collectingCountry>
|
||
, the original material of
|
||
<bibRefCitation id="75405B8465268508D35224DAFD099D75" author="Hess H." box="[610,753,498,525]" pageId="115" pageNumber="116" pagination="1025 - 1063" refId="ref122420" refString="Hess H. 1966. Mikropalaontologische Untersuchungen an Ophiuren V: Die Ophiuren aus dem Argovien (unteres Ober-Oxford) vom Guldental (Kt. Solothurn) und von Savigna (Dept. Jura). Eclogae geologicae Helvetiae 59: 1025 - 1063." type="journal article" year="1966">Hess (1966)</bibRefCitation>
|
||
; 4 dissociated LAPs from the lower Oxfordian Renggeri Member of Chapois,
|
||
<collectingCountry id="69C666E565268508D086273EFDF19D49" box="[438,521,535,561]" name="France" pageId="115" pageNumber="116">France</collectingCountry>
|
||
, the original material of
|
||
<bibRefCitation id="75405B8465268508D21A273EFC319D4A" author="Hess H." box="[810,969,535,562]" pageId="115" pageNumber="116" pagination="1059 - 1082" refId="ref122348" refString="Hess H. 1965 a. Mikropalaontologische Untersuchungen an Ophiuren IV: Die Ophiuren aus dem Renggeri-Ton (Unter-Oxford) von Chapois (Jura) und Longecombe (Ain). Eclogae geologicae Helvetiae 58: 1059 - 1082." type="journal article" year="1965">Hess (1965a)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59CB75FE65268508D18D2773FA8F9893" pageId="115" pageNumber="116" type="description">
|
||
<paragraph id="116E267565268508D18D2773FEB69D0C" blockId="115.[189,334,602,628]" box="[189,334,602,628]" pageId="115" pageNumber="116">
|
||
<heading id="4A26911965268508D18D2773FEB69D0C" bold="true" box="[189,334,602,628]" fontSize="11" level="3" pageId="115" pageNumber="116" reason="3">
|
||
<emphasis id="23A5FA6765268508D18D2773FEB69D0C" bold="true" box="[189,334,602,628]" pageId="115" pageNumber="116">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="116E267565268508D18D27A3FEF099FB" blockId="115.[189,1399,650,1667]" pageId="115" pageNumber="116">Relatively small LAPs; at least twice higher than wide (proximal LAPs) to nearly as high as wide (distal LAPs); dorsal edge concave as a result of a well-developed constriction; distal edge convex; proximal edge concave in proximal LAPs, with three to four poorly defined, weakly prominent and very slightly protruding spurs, ventral one of which largest and best defined; proximal edge in median and distal LAPs irregularly undulose, with single large, moderately well-defined, prominent and strongly protruding spur probably corresponding to large, ventralmost spur in proximal LAPs; outer surface with moderately coarsely meshed stereom replaced by finely meshed stereom in very narrow band paralleling proximal edge of LAP; trabeculae of moderately coarsely meshed stereom proximally bordering spine articulations merged into very poorly developed vertical striation in a few LAPs; trabecular intersections slightly thickened into granules in ventral portion of distal LAPs. Ten (proximal LAPs) to six (distal ones) large, ear-shaped spine articulations freestanding on strongly elevated ridge; dorsal and ventral lobes of spine articulations merged into continuous volute; spine articulations proximally bordered by narrow, welldefined, undulose ridge; dorsalmost and ventralmost spine articulations slightly smaller than remaining, nearly equal-sized ones; gap separating spine articulations and distal edge very narrow in proximal LAPs, slightly wider and dorsally widening in median to distal LAPs; row of spine articulations strongly protruding ventralwards in proximal LAPs, slightly protruding in median ones and not protruding in distal ones. Ventral edge of LAPs nearly straight; tentacle opening generally invisible in external view. Inner side of LAP with very large, conspicuous, sharply defined, prominent ridge; relatively short, slender, oblique, straight central part of ridge, merged dorsally with very large, vertically elongate (in proximal to median LAPs), near-triangular dorsal part of ridge with concave edges and pointed dorsal and ventroproximal angles; central part of ridge merged ventrally with short, slender, ventro-proximally pointing part not merged with ventral portion of LAP; inner side of distal edge with three to four well-defined, weakly prominent, oval spurs generally with dorsalward increase in size and composed of more densely meshed stereom in proximal LAPs; single spur in median to distal LAPs similar to the ventralmost spur observed in proximal LAPs; inner side of tentacle notch relatively small, nearly vertical, moderately well defined laterally. Very shallow, narrow, moderately well-defined vertical furrow with small, irregularly spaced perforations dorsally bordering tentacle notch in proximal to median LAPs; no furrow in distal LAPs.</paragraph>
|
||
<paragraph id="116E267565268508D18D2382FA8F9893" blockId="115.[189,1399,1707,2027]" pageId="115" pageNumber="116">Articulated arm fragments: LAPs broadly in contact dorsally and ventrally in proximal to distal arm segments; dorsal arm plates relatively small, bell shaped, with slightly convex distal edge, pointed proximal angle with slightly concave edges; spine articulations meeting mid-radially forming continuous band at least in proximal segments, arm spines thus originally forming fan; ventral arm plates nearly T-shaped, weakly convex distal edge, deeply concave latero-proximal edges and obtuse proximal angle; proximal and latero-proximal edges of ventral arm plates slightly prominent; round, slightly swollen area of coarsely meshed stereom in centre of ventral arm plates; tentacle pores small, covered by single, spatulate, distally widening tentacle scale; arm spines slightly flattened, with coarsely reticulate stereom displaying small thorns, irregular longitudinal row of much larger thorns along both margins of the spines.</paragraph>
|
||
</subSubSection>
|
||
<caption id="45AE76FD6521850FD18D2359FB209893" ID-DOI="http://doi.org/10.5281/zenodo.3822883" ID-Zenodo-Dep="3822883" httpUri="https://zenodo.org/record/3822883/files/figure.png" pageId="116" pageNumber="117" startId="116.[189,231,1648,1674]" targetBox="[297,1290,265,1608]" targetPageId="116">
|
||
<paragraph id="116E26756521850FD18D2359FB209893" blockId="116.[189,1399,1648,2027]" pageId="116" pageNumber="117">
|
||
<emphasis id="23A5FA676521850FD18D2359FEE499F3" bold="true" box="[189,284,1648,1675]" pageId="116" pageNumber="117">Fig. 21.</emphasis>
|
||
Fossil lateral arm plates (LAPs) of ophiacanthid brittle stars in external (a) and internal (b) views and articulated arm fragment in ventral (a) and dorsal (b) views.
|
||
<emphasis id="23A5FA676521850FD2C023BDFBFE99D6" bold="true" box="[1008,1030,1684,1710]" pageId="116" pageNumber="117">1.</emphasis>
|
||
<taxonomicName id="D6D15DF66521850FD53C23BDFEF899AA" authority="(Hess, 1966)" authorityName="Thuy" authorityYear="2013" baseAuthorityName="Hess" baseAuthorityYear="1966" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiogaleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="116" pageNumber="117" phylum="Echinodermata" rank="species" species="constrictus" status="comb. nov.">
|
||
<emphasis id="23A5FA676521850FD53C23BDFADE99D6" box="[1036,1318,1684,1710]" italics="true" pageId="116" pageNumber="117">Ophiogaleus constrictus</emphasis>
|
||
(
|
||
<bibRefCitation id="75405B846521850FD40523BDFF0099AA" author="Hess H." pageId="116" pageNumber="117" pagination="1025 - 1063" refId="ref122420" refString="Hess H. 1966. Mikropalaontologische Untersuchungen an Ophiuren V: Die Ophiuren aus dem Argovien (unteres Ober-Oxford) vom Guldental (Kt. Solothurn) und von Savigna (Dept. Jura). Eclogae geologicae Helvetiae 59: 1025 - 1063." type="journal article" year="1966">Hess, 1966</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3896471C6521850FD03B239EFE7799AA" box="[267,399,1719,1746]" pageId="116" pageNumber="117" rank="species">comb. nov.</taxonomicNameLabel>
|
||
from the late Oxfordian (Late Jurassic) of Savigna, France; GZG.INV.78627 distal arm fragment.
|
||
<emphasis id="23A5FA676521850FD05A23F3FE78998C" bold="true" box="[362,384,1754,1780]" pageId="116" pageNumber="117">2.</emphasis>
|
||
<taxonomicName id="D6D15DF66521850FD0B723F3FDE3998C" authorityName="Thuy" authorityYear="2013" box="[391,539,1754,1780]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiogaleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="116" pageNumber="117" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="23A5FA676521850FD0B723F3FDE3998C" box="[391,539,1754,1780]" italics="true" pageId="116" pageNumber="117">Ophiogaleus</emphasis>
|
||
</taxonomicName>
|
||
sp. from the early Kimmeridgian (Late Jurassic) of Geisingen, Germany; GZG.INV.78632, proximal LAP.
|
||
<emphasis id="23A5FA676521850FD37E23D4FD84986F" bold="true" box="[590,636,1789,1815]" pageId="116" pageNumber="117">3-4.</emphasis>
|
||
<taxonomicName id="D6D15DF66521850FD3B823D4FB8A9860" authority="(Rasmussen, 1952)" authorityName="Thuy" authorityYear="2013" baseAuthorityName="Rasmussen" baseAuthorityYear="1952" box="[648,1138,1789,1816]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiogaleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="116" pageNumber="117" phylum="Echinodermata" rank="species" species="danicus" status="comb. nov.">
|
||
<emphasis id="23A5FA676521850FD3B823D4FC7B986F" box="[648,899,1789,1815]" italics="true" pageId="116" pageNumber="117">Ophiogaleus danicus</emphasis>
|
||
(
|
||
<bibRefCitation id="75405B846521850FD2A723D7FB929860" author="Rasmussen H. W." box="[919,1130,1789,1816]" pageId="116" pageNumber="117" pagination="47 - 57" refId="ref126877" refString="Rasmussen H. W. 1952. Cretaceous Ophiuroidea from Germany, Sweden, Spain and New Jersey. Meddelelser fra Dansk Geologisk Forening 12 (1951): 47 - 57." type="journal article" year="1952">Rasmussen, 1952</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3896471C6521850FD54E23D4FAFB9860" box="[1150,1283,1789,1816]" pageId="116" pageNumber="117" rank="species">comb. nov.</taxonomicNameLabel>
|
||
from the early Maastrichtian (Late Cretaceous) of Rügen, Germany.
|
||
<emphasis id="23A5FA676521850FD2422208FC709843" bold="true" box="[882,904,1825,1851]" pageId="116" pageNumber="117">3.</emphasis>
|
||
GZG.INV.78635, proximal LAP.
|
||
<emphasis id="23A5FA676521850FD4262208FAD39843" bold="true" box="[1302,1323,1825,1851]" pageId="116" pageNumber="117">4.</emphasis>
|
||
GZG. INV.78636, distal LAP.
|
||
<emphasis id="23A5FA676521850FD0FF226DFE059826" bold="true" box="[463,509,1860,1886]" pageId="116" pageNumber="117">5-6.</emphasis>
|
||
<taxonomicName id="D6D15DF66521850FD332226DFD709826" authorityName="Thuy & Meyer" authorityYear="2013" box="[514,648,1860,1886]" class="Ophiuroidea" family="Ophiacanthidae" genus="Hanshessia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="116" pageNumber="117" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="23A5FA676521850FD332226DFD709826" box="[514,648,1860,1886]" italics="true" pageId="116" pageNumber="117">Hanshessia</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. innom. 1 from the late Sinemurian to early Pliensbachian (Early Jurassic) of the Glasenbach Gorge, Austria.
|
||
<emphasis id="23A5FA676521850FD231224EFCEE98F9" bold="true" box="[769,790,1895,1921]" pageId="116" pageNumber="117">5.</emphasis>
|
||
NHMW 2012/0137/0021, proximal LAP.
|
||
<emphasis id="23A5FA676521850FD5CC224EFAE998F9" bold="true" box="[1276,1297,1895,1921]" pageId="116" pageNumber="117">6.</emphasis>
|
||
NHMW 2012/0137/0022, proximal LAP.
|
||
<emphasis id="23A5FA676521850FD37422A3FD8A98DC" bold="true" box="[580,626,1930,1956]" pageId="116" pageNumber="117">7-9.</emphasis>
|
||
<taxonomicName id="D6D15DF66521850FD34B22A3FCF998DC" authorityName="Thuy & Meyer" authorityYear="2013" box="[635,769,1930,1956]" class="Ophiuroidea" family="Ophiacanthidae" genus="Hanshessia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="116" pageNumber="117" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="23A5FA676521850FD34B22A3FCF998DC" box="[635,769,1930,1956]" italics="true" pageId="116" pageNumber="117">Hanshessia</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. innom. 2 from the early Bathonian (Middle Jurassic) of La Pouza, France.
|
||
<emphasis id="23A5FA676521850FD31D2287FDBB98B0" bold="true" box="[557,579,1966,1992]" pageId="116" pageNumber="117">7.</emphasis>
|
||
GZG.INV.78637, proximal LAP.
|
||
<emphasis id="23A5FA676521850FD2E62284FC1498BF" bold="true" box="[982,1004,1965,1991]" pageId="116" pageNumber="117">8.</emphasis>
|
||
GZG.INV.78638, proximal LAP.
|
||
<emphasis id="23A5FA676521850FD18D22F8FF2B9893" bold="true" box="[189,211,2001,2027]" pageId="116" pageNumber="117">9.</emphasis>
|
||
GZG.INV.78639, distal to median LAP. One common scale bar per species except for 1.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="59CB75FE6520850ED18D2422FB119B7B" pageId="117" pageNumber="118" type="discussion">
|
||
<paragraph id="116E26756520850ED18D2422FED79E5D" blockId="117.[189,303,267,293]" box="[189,303,267,293]" pageId="117" pageNumber="118">
|
||
<heading id="4A2691196520850ED18D2422FED79E5D" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="117" pageNumber="118" reason="3">
|
||
<emphasis id="23A5FA676520850ED18D2422FED79E5D" bold="true" box="[189,303,267,293]" pageId="117" pageNumber="118">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="116E26756520850ED18D2414FAD29DA0" blockId="117.[189,1399,317,728]" pageId="117" pageNumber="118">
|
||
The concept of
|
||
<taxonomicName id="D6D15DF66520850ED0492414FD619E2E" authorityName="Thuy" authorityYear="2013" baseAuthorityName="Hess" baseAuthorityYear="1966" box="[377,665,317,343]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiogaleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="117" pageNumber="118" phylum="Echinodermata" rank="species" species="constrictus" status="comb. nov.">
|
||
<emphasis id="23A5FA676520850ED0492414FD619E2E" box="[377,665,317,343]" italics="true" pageId="117" pageNumber="118">Ophiogaleus constrictus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3896471C6520850ED3922414FCDD9E2F" box="[674,805,317,343]" pageId="117" pageNumber="118" rank="species">comb. nov.</taxonomicNameLabel>
|
||
has repeatedly caused confusion in the past. This form was originally described by
|
||
<bibRefCitation id="75405B846520850ED351244DFD0C9E06" author="Hess H." box="[609,756,355,382]" pageId="117" pageNumber="118" pagination="1025 - 1063" refId="ref122420" refString="Hess H. 1966. Mikropalaontologische Untersuchungen an Ophiuren V: Die Ophiuren aus dem Argovien (unteres Ober-Oxford) vom Guldental (Kt. Solothurn) und von Savigna (Dept. Jura). Eclogae geologicae Helvetiae 59: 1025 - 1063." type="journal article" year="1966">Hess (1966)</bibRefCitation>
|
||
as
|
||
<taxonomicName id="D6D15DF66520850ED215244AFC469E05" authorityName="Muller & Troschel" authorityYear="1842" box="[805,958,355,381]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="117" pageNumber="118" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="23A5FA676520850ED215244AFC469E05" box="[805,958,355,381]" italics="true" pageId="117" pageNumber="118">Ophiacantha</emphasis>
|
||
</taxonomicName>
|
||
?
|
||
<emphasis id="23A5FA676520850ED2E8244DFBB59E05" box="[984,1101,356,381]" italics="true" pageId="117" pageNumber="118">constricta</emphasis>
|
||
when he noted that the type material from the Oxfordian of
|
||
<collectingCountry id="69C666E56520850ED3B124A0FCF59EDB" box="[641,781,393,419]" name="Switzerland" pageId="117" pageNumber="118">Switzerland</collectingCountry>
|
||
and
|
||
<collectingCountry id="69C666E56520850ED27E24A3FC669EDC" box="[846,926,394,420]" name="France" pageId="117" pageNumber="118">France</collectingCountry>
|
||
previously described as
|
||
<taxonomicName id="D6D15DF66520850ED5E024A0FA919EDB" authorityName="Muller & Troschel" authorityYear="1842" box="[1232,1385,393,419]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="117" pageNumber="118" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="23A5FA676520850ED5E024A0FA919EDB" box="[1232,1385,393,419]" italics="true" pageId="117" pageNumber="118">Ophiacantha</emphasis>
|
||
</taxonomicName>
|
||
?
|
||
<emphasis id="23A5FA676520850ED18D2498FE939EB2" box="[189,363,433,458]" italics="true" pageId="117" pageNumber="118">suprajurassica</emphasis>
|
||
Hess, 1965 included more than one type of LAP.
|
||
<bibRefCitation id="75405B846520850ED2EF2499FB899EB3" author="Hess H." box="[991,1137,432,459]" pageId="117" pageNumber="118" pagination="1025 - 1063" refId="ref122420" refString="Hess H. 1966. Mikropalaontologische Untersuchungen an Ophiuren V: Die Ophiuren aus dem Argovien (unteres Ober-Oxford) vom Guldental (Kt. Solothurn) und von Savigna (Dept. Jura). Eclogae geologicae Helvetiae 59: 1025 - 1063." type="journal article" year="1966">Hess (1966)</bibRefCitation>
|
||
based the distinction on the higher number of spine articulations and the lack of a vertical striation on the outer surface in
|
||
<taxonomicName id="D6D15DF66520850ED1EF24D4FE709D6F" authorityName="Thuy" authorityYear="2013" baseAuthorityName="Hess" baseAuthorityYear="1966" box="[223,392,509,535]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiogaleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="117" pageNumber="118" phylum="Echinodermata" rank="species" species="constrictus" status="comb. nov.">
|
||
<emphasis id="23A5FA676520850ED1EF24D4FE709D6F" box="[223,392,509,535]" italics="true" pageId="117" pageNumber="118">O. constrictus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3896471C6520850ED0A324D4FDE09D6F" box="[403,536,509,535]" pageId="117" pageNumber="118" rank="species">comb. nov.</taxonomicNameLabel>
|
||
These are, indeed, valid differences, yet the concept of
|
||
<taxonomicName id="D6D15DF66520850ED5FE24D4FA8F9D6F" authorityName="Thuy" authorityYear="2013" baseAuthorityName="Hess" baseAuthorityYear="1966" box="[1230,1399,509,535]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiogaleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="117" pageNumber="118" phylum="Echinodermata" rank="species" species="constrictus" status="comb. nov.">
|
||
<emphasis id="23A5FA676520850ED5FE24D4FA8F9D6F" box="[1230,1399,509,535]" italics="true" pageId="117" pageNumber="118">O. constrictus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3896471C6520850ED18D270AFEB99D46" box="[189,321,547,574]" pageId="117" pageNumber="118" rank="species">comb. nov.</taxonomicNameLabel>
|
||
was still not sufficiently specific because
|
||
<bibRefCitation id="75405B846520850ED20A270DFC329D46" author="Hess H." box="[826,970,547,574]" pageId="117" pageNumber="118" pagination="1025 - 1063" refId="ref122420" refString="Hess H. 1966. Mikropalaontologische Untersuchungen an Ophiuren V: Die Ophiuren aus dem Argovien (unteres Ober-Oxford) vom Guldental (Kt. Solothurn) und von Savigna (Dept. Jura). Eclogae geologicae Helvetiae 59: 1025 - 1063." type="journal article" year="1966">Hess (1966)</bibRefCitation>
|
||
also included LAPs which are here reinterpreted as
|
||
<taxonomicName id="D6D15DF66520850ED0B02763FD6E9D1C" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[384,662,586,612]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiocamax" kingdom="Animalia" order="Ophiurida" pageId="117" pageNumber="118" phylum="Echinodermata" rank="species" species="dorotheae" status="sp. nov.">
|
||
<emphasis id="23A5FA676520850ED0B02763FD6E9D1C" box="[384,662,586,612]" italics="true" pageId="117" pageNumber="118">Ophiocamax dorotheae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3896471C6520850ED3902763FD069D1C" box="[672,766,586,612]" pageId="117" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(see
|
||
<bibRefCitation id="75405B846520850ED2712763FC3A9D1C" author="Hess H." box="[833,962,586,612]" pageId="117" pageNumber="118" pagination="385 - 421" refId="ref122234" refString="Hess H. 1960. Ophiurenreste aus dem Malm des Schweizer Juras und des Departements Haut-Rhin. Eclogae geologicae Helvetiae 53: 385 - 421." type="journal article" year="1960">Hess 1960</bibRefCitation>
|
||
, figs 39-40;
|
||
<bibRefCitation id="75405B846520850ED56D2763FB259D1C" author="Hess H." box="[1117,1245,586,612]" pageId="117" pageNumber="118" pagination="1025 - 1063" refId="ref122420" refString="Hess H. 1966. Mikropalaontologische Untersuchungen an Ophiuren V: Die Ophiuren aus dem Argovien (unteres Ober-Oxford) vom Guldental (Kt. Solothurn) und von Savigna (Dept. Jura). Eclogae geologicae Helvetiae 59: 1025 - 1063." type="journal article" year="1966">Hess 1966</bibRefCitation>
|
||
, figs 76-77). Following this confusion,
|
||
<bibRefCitation id="75405B846520850ED0DE2759FD419DF2" author="Kutscher M." box="[494,697,624,650]" pageId="117" pageNumber="118" pagination="50 - 104" refId="ref124575" refString="Kutscher M. 1987 a. Die Echinodermen der Callovien-Geschiebe. Der Geschiebesammler 21: 50 - 104." type="journal article" year="1987">Kutscher (1987a)</bibRefCitation>
|
||
described dissociated LAPs from the Callovian of
|
||
<collectingCountry id="69C666E56520850ED4392759FA8E9DF2" box="[1289,1398,624,650]" name="Germany" pageId="117" pageNumber="118">Germany</collectingCountry>
|
||
as
|
||
<taxonomicName id="D6D15DF66520850ED1EF27BFFE809DC8" authorityName="Muller & Troschel" authorityYear="1842" box="[223,376,662,688]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="117" pageNumber="118" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="23A5FA676520850ED1EF27BFFE809DC8" box="[223,376,662,688]" italics="true" pageId="117" pageNumber="118">Ophiacantha</emphasis>
|
||
</taxonomicName>
|
||
?
|
||
<emphasis id="23A5FA676520850ED0BE27BEFDFB9DC8" box="[398,515,663,688]" italics="true" pageId="117" pageNumber="118">constricta</emphasis>
|
||
; these markedly differ from the
|
||
<typeStatus id="CE6A98D76520850ED2BB27BFFC0A9DC8" box="[907,1010,662,688]" pageId="117" pageNumber="118" type="holotype">holotype</typeStatus>
|
||
of the species but do have clear features in common with some of the LAPs erroneously included in that species by
|
||
<bibRefCitation id="75405B846520850ED5A42794FADE9DA0" author="Hess H." box="[1172,1318,701,728]" pageId="117" pageNumber="118" pagination="1025 - 1063" refId="ref122420" refString="Hess H. 1966. Mikropalaontologische Untersuchungen an Ophiuren V: Die Ophiuren aus dem Argovien (unteres Ober-Oxford) vom Guldental (Kt. Solothurn) und von Savigna (Dept. Jura). Eclogae geologicae Helvetiae 59: 1025 - 1063." type="journal article" year="1966">Hess (1966)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="116E26756520850ED18D262BFB119B7B" blockId="117.[189,1399,770,1027]" pageId="117" pageNumber="118">
|
||
A re-examination of the type specimens, complemented by new, partially articulated, equivalent finds from the Oxfordian of
|
||
<collectingCountry id="69C666E56520850ED0E52600FDD09C3B" box="[469,552,809,835]" name="France" pageId="117" pageNumber="118">France</collectingCountry>
|
||
, has enabled a consistent reinterpretation of
|
||
<taxonomicName id="D6D15DF66520850ED5792601FB1A9C3A" authorityName="Muller & Troschel" authorityYear="1842" box="[1097,1250,808,834]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="117" pageNumber="118" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="23A5FA676520850ED5792601FB1A9C3A" box="[1097,1250,808,834]" italics="true" pageId="117" pageNumber="118">Ophiacantha</emphasis>
|
||
</taxonomicName>
|
||
?
|
||
<emphasis id="23A5FA676520850ED5CA2600FA979C3A" box="[1274,1391,809,834]" italics="true" pageId="117" pageNumber="118">constricta</emphasis>
|
||
. Thanks to Hess’s (1966) careful choice of the
|
||
<typeStatus id="CE6A98D76520850ED3D62667FCB59C10" box="[742,845,846,872]" pageId="117" pageNumber="118" type="holotype">holotype</typeStatus>
|
||
(a single well-preserved proximal LAP, rather than a less distinctive median LAP or a series of LAPs), the species in question is now unambiguously identifiable. It is chosen as the type species of
|
||
<taxonomicName id="D6D15DF66520850ED3C026B2FC7E9CCD" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[752,902,923,949]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiogaleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="117" pageNumber="118" phylum="Echinodermata" rank="genus" status="gen. nov.">
|
||
<emphasis id="23A5FA676520850ED3C026B2FC7E9CCD" box="[752,902,923,949]" italics="true" pageId="117" pageNumber="118">Ophiogaleus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3896471C6520850ED2BF26B5FC079CCE" box="[911,1023,924,950]" pageId="117" pageNumber="118" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, an assumedly close relative of extant
|
||
<taxonomicName id="D6D15DF66520850ED03C26EBFE5D9CA4" authorityName="Muller & Troschel" authorityYear="1842" box="[268,421,962,988]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="117" pageNumber="118" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="23A5FA676520850ED03C26EBFE5D9CA4" box="[268,421,962,988]" italics="true" pageId="117" pageNumber="118">Ophiacantha</emphasis>
|
||
</taxonomicName>
|
||
(see above), since it is known from numerous well-preserved specimens, including articulated arm fragments, and most typically displays the distinctive features of the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59CB75FE6520850ED18D2104FCE29B01" box="[189,794,1069,1145]" pageId="117" pageNumber="118" type="distribution">
|
||
<paragraph id="116E26756520850ED18D2104FEB79B3F" blockId="117.[189,335,1069,1095]" box="[189,335,1069,1095]" pageId="117" pageNumber="118">
|
||
<heading id="4A2691196520850ED18D2104FEB79B3F" bold="true" box="[189,335,1069,1095]" fontSize="11" level="3" pageId="117" pageNumber="118" reason="3">
|
||
<emphasis id="23A5FA676520850ED18D2104FEB79B3F" bold="true" box="[189,335,1069,1095]" pageId="117" pageNumber="118">Occurrence</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="116E26756520850ED18D2176FCE29B01" blockId="117.[189,794,1119,1145]" box="[189,794,1119,1145]" pageId="117" pageNumber="118">
|
||
Early to Late Oxfordian of
|
||
<collectingCountry id="69C666E56520850ED0CD2176FDB59B01" box="[509,589,1119,1145]" name="France" pageId="117" pageNumber="118">France</collectingCountry>
|
||
and
|
||
<collectingCountry id="69C666E56520850ED3B62176FCEE9B01" box="[646,790,1119,1145]" name="Switzerland" pageId="117" pageNumber="118">Switzerland</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |