147 lines
16 KiB
XML
147 lines
16 KiB
XML
<document id="56EE18AF8C4B2B857B068C9F6DC13DBF" ID-CLB-Dataset="22249" ID-DOI="10.5852/ejt.2013.48" ID-GBIF-Dataset="02866e62-ab61-4cf4-9d98-0248e251084e" ID-ISSN="2118-9773" ID-Zenodo-Dep="3822836" ID-ZooBank="E7080722-E348-448D-96E5-D537F4865BB5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1589219740188" checkinUser="carolina" docAuthor="Thuy, Ben" docDate="2013" docId="99789763650A8524D350274CFE6C9893" docLanguage="en" docName="ejt-2013-48.pdf.imf" docOrigin="European Journal of Taxonomy 48" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Europacantha Thuy 2013, gen. nov." docType="treatment" docUuid="511AAE08-FCA9-4DA9-AFC4-6B2A2C39B2DB" docUuidSource="ZooBank" docVersion="10" lastPageNumber="96" masterDocId="6541EF1B6555857BD1302529FFF89F78" masterDocTitle="Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)" masterLastPageNumber="242" masterPageNumber="1" pageNumber="96" updateTime="1698839159859" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="735ABFCF7BAD87DBCEDA0B0A6B9E19A4">Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)</mods:title>
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<mods:namePart id="3DA08659F2CA01C14E30C2F1F4CCC9D9">Thuy, Ben</mods:namePart>
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<mods:typeOfResource id="33366C9EB42E8C96B6E9A5C1EDBF809E">text</mods:typeOfResource>
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<mods:title id="77707FDBC152CA0AA0B89B6B89F0C052">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="87B90E5358B0ACA9D23BB1F740609B10">
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<mods:date id="E53D5033155F37BB80BBDFE33E085603">2013</mods:date>
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<mods:detail id="52D49FC8204F55E21253B510E993D35C" type="pubDate">
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<mods:number id="BC1191A18BBB573D57E650070C6E93A1">2013-07-02</mods:number>
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<mods:detail id="BA4958B274861ABAEC4EB92B0D35E1E3" type="issue">
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<mods:number id="AE18536719C958152CCB6CC7C71453F2">48</mods:number>
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<mods:classification id="DDEFC4E27E4F0D7F8ED4CF90525F3D92">journal article</mods:classification>
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<mods:identifier id="8838F6BAD8A0FBF54123420A21CD5200" type="CLB-Dataset">22249</mods:identifier>
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<mods:identifier id="4E4ED8A625CB73222215548470A165C3" type="DOI">10.5852/ejt.2013.48</mods:identifier>
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<treatment id="99789763650A8524D350274CFE6C9893" ID-DOI="http://doi.org/10.5281/zenodo.3844213" ID-GBIF-Taxon="164044874" ID-Zenodo-Dep="3844213" LSID="urn:lsid:zoobank.org:act:511AAE08-FCA9-4DA9-AFC4-6B2A2C39B2DB" httpUri="http://treatment.plazi.org/id/99789763650A8524D350274CFE6C9893" lastPageNumber="96" pageId="95" pageNumber="96">
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<subSubSection id="59CB75FE650A8524D350274CFC2B9D07" box="[608,979,612,639]" pageId="95" pageNumber="96" type="nomenclature">
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<paragraph id="116E2675650A8524D350274CFC2B9D07" blockId="95.[349,1239,612,678]" box="[608,979,612,639]" pageId="95" pageNumber="96">
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<heading id="4A269119650A8524D350274CFC2B9D07" box="[608,979,612,639]" centered="true" fontSize="11" level="2" pageId="95" pageNumber="96" reason="2">
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Genus
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<taxonomicName id="D6D15DF6650A8524D383274DFC999D07" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[691,865,612,639]" class="Ophiuroidea" family="Ophiacanthidae" genus="Europacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="95" pageNumber="96" phylum="Echinodermata" rank="genus" status="gen. nov.">
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<emphasis id="23A5FA67650A8524D383274DFC999D07" bold="true" box="[691,865,612,639]" italics="true" pageId="95" pageNumber="96">Europacantha</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3896471C650A8524D258274CFC2B9D07" box="[872,979,613,639]" pageId="95" pageNumber="96" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="59CB75FE650A8524D06D27A2FB2F9DDE" box="[349,1239,651,678]" pageId="95" pageNumber="96" type="description">
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<paragraph id="116E2675650A8524D06D27A2FB2F9DDE" blockId="95.[349,1239,612,678]" box="[349,1239,651,678]" pageId="95" pageNumber="96">
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<uri id="65402A77650A8524D06D27A2FB2F9DDE" box="[349,1239,651,678]" pageId="95" pageNumber="96">
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urn:lsid:zoobank.org:act:
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<uuid id="65771CA0650A8524D34D27A2FB2F9DDE" box="[637,1239,651,678]" pageId="95" pageNumber="96">511AAE08-FCA9-4DA9-AFC4-6B2A2C39B2DB</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="59CB75FE650A8524D18D27F9FDC49C64" pageId="95" pageNumber="96" type="type_taxon">
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<paragraph id="116E2675650A8524D18D27F9FDD99D92" blockId="95.[189,545,720,746]" box="[189,545,720,746]" pageId="95" pageNumber="96">
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<heading id="4A269119650A8524D18D27F9FDD99D92" bold="true" box="[189,545,720,746]" fontSize="11" level="3" pageId="95" pageNumber="96" reason="3">
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<emphasis id="23A5FA67650A8524D18D27F9FDD99D92" bold="true" box="[189,545,720,746]" pageId="95" pageNumber="96">
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<typeStatus id="CE6A98D7650A8524D18D27F9FF029D92" box="[189,250,720,746]" pageId="95" pageNumber="96">Type</typeStatus>
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and sole known species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="116E2675650A8524D18D262BFDC49C64" blockId="95.[189,572,770,796]" box="[189,572,770,796]" pageId="95" pageNumber="96">
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<taxonomicName id="D6D15DF6650A8524D18D262BFE229C64" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[189,474,770,796]" class="Ophiuroidea" family="Ophiacanthidae" genus="Europacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="95" pageNumber="96" phylum="Echinodermata" rank="species" species="paciphila" status="sp. nov.">
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<emphasis id="23A5FA67650A8524D18D262BFE229C64" box="[189,474,770,796]" italics="true" pageId="95" pageNumber="96">Europacantha paciphila</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3896471C650A8524D0D1262BFDC49C64" box="[481,572,770,796]" pageId="95" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="59CB75FE650A8524D18D266FFB9F9B54" pageId="95" pageNumber="96" type="diagnosis">
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<paragraph id="116E2675650A8524D18D266FFECE9C18" blockId="95.[189,310,838,864]" box="[189,310,838,864]" pageId="95" pageNumber="96">
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<heading id="4A269119650A8524D18D266FFECE9C18" bold="true" box="[189,310,838,864]" fontSize="11" level="3" pageId="95" pageNumber="96" reason="3">
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<emphasis id="23A5FA67650A8524D18D266FFECE9C18" bold="true" box="[189,310,838,864]" pageId="95" pageNumber="96">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="116E2675650A8524D18D2651FB9F9B54" blockId="95.[189,1398,888,1068]" pageId="95" pageNumber="96">Ophiacanthid with relatively small, stout, trapezoid LAPs devoid of conspicuous outer surface ornament; small, inconspicuous, very weakly protruding spur on outer proximal edge; up to six large, ear-shaped spine articulations on strongly elevated distal portion of LAP, not bordered proximally by a ridge-like structure; relatively broad, gently proximally bent ridge on inner side with widened dorsal tip displaying a ventrally projecting extension; tentacle notch small, invisible in external view.</paragraph>
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</subSubSection>
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<subSubSection id="59CB75FE650A8524D18D217EFCF09B97" pageId="95" pageNumber="96" type="etymology">
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<paragraph id="116E2675650A8524D18D217EFEBC9B09" blockId="95.[189,324,1111,1137]" box="[189,324,1111,1137]" pageId="95" pageNumber="96">
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<heading id="4A269119650A8524D18D217EFEBC9B09" bold="true" box="[189,324,1111,1137]" fontSize="11" level="3" pageId="95" pageNumber="96" reason="3">
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<emphasis id="23A5FA67650A8524D18D217EFEBC9B09" bold="true" box="[189,324,1111,1137]" pageId="95" pageNumber="96">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="116E2675650A8524D18D21A0FCF09B97" blockId="95.[189,1399,1160,1263]" pageId="95" pageNumber="96">
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Name composed of
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<taxonomicName id="D6D15DF6650A8524D0F921A0FDD89BDA" authorityName="Thomson" authorityYear="1860" box="[457,544,1161,1186]" class="Insecta" family="Cerambycidae" genus="Europa" kingdom="Animalia" order="Coleoptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="genus">
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<emphasis id="23A5FA67650A8524D0F921A0FDD89BDA" box="[457,544,1161,1186]" italics="true" pageId="95" pageNumber="96">Europa</emphasis>
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</taxonomicName>
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, a Phoenician princess whose name is commonly translated into “perspicacious”, honouring the idea of a united Europe, and
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<taxonomicName id="D6D15DF6650A8524D2952186FBFF9BB1" authorityName="Laporte & Gory" authorityYear="1838" box="[933,1031,1199,1225]" class="Insecta" family="Buprestidae" genus="Acantha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="genus">
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<emphasis id="23A5FA67650A8524D2952186FBFF9BB1" box="[933,1031,1199,1225]" italics="true" pageId="95" pageNumber="96">Acantha</emphasis>
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</taxonomicName>
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, a Greek nymph whose name literally translates into “thorny”; gender feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="59CB75FE650A8524D18D2033FE6C9893" pageId="95" pageNumber="96" type="discussion">
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<paragraph id="116E2675650A8524D18D2033FED79A4C" blockId="95.[189,303,1306,1332]" box="[189,303,1306,1332]" pageId="95" pageNumber="96">
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<heading id="4A269119650A8524D18D2033FED79A4C" bold="true" box="[189,303,1306,1332]" fontSize="11" level="3" pageId="95" pageNumber="96" reason="3">
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<emphasis id="23A5FA67650A8524D18D2033FED79A4C" bold="true" box="[189,303,1306,1332]" pageId="95" pageNumber="96">Remarks</emphasis>
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</heading>
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</paragraph>
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<paragraph id="116E2675650A8524D18D2062FD859935" blockId="95.[189,1399,1355,1613]" pageId="95" pageNumber="96">
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The oldest known ophiacanthid record is here described in the form of rare dissociated LAPs from the Anisian (early Middle Triassic) of
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<collectingCountry id="69C666E5650A8524D363205BFD449AF4" box="[595,700,1394,1420]" name="Hungary" pageId="95" pageNumber="96">Hungary</collectingCountry>
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. The LAPs in question are unambiguously assignable to the
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<taxonomicName id="D6D15DF6650A8524D18D20B1FE8D9ACA" baseAuthorityName="Martynov" baseAuthorityYear="2010" box="[189,373,1432,1458]" class="Ophiuroidea" family="Ophiacanthidae" kingdom="Animalia" order="Ophiurida" pageId="95" pageNumber="96" phylum="Echinodermata" rank="family">Ophiacanthidae</taxonomicName>
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on account of the large, ear-shaped spine articulations with a typical sigmoidal fold, combined with several small perforations in a shallow vertical furrow on the inner side. Astonishingly, these Anisian LAPs display small tentacle notches, almost certainly reflecting originally small tentacle pores as defined by
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<bibRefCitation id="75405B84650A8524D0902325FD8B995E" author="Thuy B. & Ishida Y. & Doi E. & Kroh A." box="[416,627,1548,1574]" pageId="95" pageNumber="96" refId="ref128754" refString="Thuy B., Ishida Y., Doi E. & Kroh A. 2012. New ophiacanthid brittle stars (Echinodermata: Ophiuroidea) from the Upper Triassic of Japan: first insights into the origin and evolution of an extant deep-sea group. Journal of Systematic Palaeontology." type="book" year="2012">
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Thuy
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<emphasis id="23A5FA67650A8524D0D32324FDE5995E" box="[483,541,1548,1574]" italics="true" pageId="95" pageNumber="96">et al.</emphasis>
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(2012)
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</bibRefCitation>
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, which places them in the more derived small-pored ophiacanthids rather than the basal large-pored ones.
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</paragraph>
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<paragraph id="116E2675650A8524D18D235EFE6C9893" blockId="95.[189,1399,1655,2027]" pageId="95" pageNumber="96">
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Indeed, greatest similarities in LAP morphology are shared with the small-pored
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<taxonomicName id="D6D15DF6650A8524D59C235EFABD99E9" authorityName="Muller & Troschel" authorityYear="1842" box="[1196,1349,1655,1681]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="95" pageNumber="96" phylum="Echinodermata" rank="genus">
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<emphasis id="23A5FA67650A8524D59C235EFABD99E9" box="[1196,1349,1655,1681]" italics="true" pageId="95" pageNumber="96">Ophiacantha</emphasis>
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</taxonomicName>
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, on account of the number, size and position of the spine articulations, the absence of more than a single spur on the outer proximal edge, and the shape of the ridge on the inner side, as well as with
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<taxonomicName id="D6D15DF6650A8524D58523EDFEBC987C" authority="Thuy, 2011" authorityName="Thuy" authorityYear="2011" class="Ophiuroidea" family="Ophiacanthidae" genus="Inexpectacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="95" pageNumber="96" phylum="Echinodermata" rank="genus">
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<emphasis id="23A5FA67650A8524D58523EDFA8F99A6" box="[1205,1399,1732,1758]" italics="true" pageId="95" pageNumber="96">Inexpectacantha</emphasis>
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Thuy, 2011
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</taxonomicName>
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on account of the stout, trapezoid aspect, the proximally non-bordered spine articulations and the lack of a conspicuous outer surface ornament. Yet, in the LAPs of
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<taxonomicName id="D6D15DF6650A8524D56D2239FB0E9852" authorityName="Muller & Troschel" authorityYear="1842" box="[1117,1270,1808,1834]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="95" pageNumber="96" phylum="Echinodermata" rank="genus">
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<emphasis id="23A5FA67650A8524D56D2239FB0E9852" box="[1117,1270,1808,1834]" italics="true" pageId="95" pageNumber="96">Ophiacantha</emphasis>
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</taxonomicName>
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, the spine articulations are proximally sharply bordered by a ridge or the distalmost lamella of the outer surface striation, while in
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<taxonomicName id="D6D15DF6650A8524D0A52274FDAF980F" authorityName="Thuy" authorityYear="2011" box="[405,599,1885,1911]" class="Ophiuroidea" family="Ophiacanthidae" genus="Inexpectacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="95" pageNumber="96" phylum="Echinodermata" rank="genus">
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<emphasis id="23A5FA67650A8524D0A52274FDAF980F" box="[405,599,1885,1911]" italics="true" pageId="95" pageNumber="96">Inexpectacantha</emphasis>
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</taxonomicName>
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the ridge on the inner side of the LAPs lacks a ventrally projecting extension, and the spine articulations are more oblique, displaying a coarsely corrugated ventral lobe. On account of these incompatibilities,
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<taxonomicName id="D6D15DF6650A8524D3B32283FCD298BC" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[643,810,1962,1988]" class="Ophiuroidea" family="Ophiacanthidae" genus="Europacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="95" pageNumber="96" phylum="Echinodermata" rank="genus" status="gen. nov.">
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<emphasis id="23A5FA67650A8524D3B32283FCD298BC" box="[643,810,1962,1988]" italics="true" pageId="95" pageNumber="96">Europacantha</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3896471C650A8524D2012282FC6498BD" box="[817,924,1963,1989]" pageId="95" pageNumber="96" rank="genus">gen. nov.</taxonomicNameLabel>
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is thus introduced here to accommodate the Anisian LAPs.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |