224 lines
26 KiB
XML
224 lines
26 KiB
XML
<document id="AD40EA09BF6309D17E79EDB61C77E9ED" ID-CLB-Dataset="22249" ID-DOI="10.5852/ejt.2013.48" ID-GBIF-Dataset="02866e62-ab61-4cf4-9d98-0248e251084e" ID-ISSN="2118-9773" ID-Zenodo-Dep="3822836" ID-ZooBank="E7080722-E348-448D-96E5-D537F4865BB5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1589219740188" checkinUser="carolina" docAuthor="Thuy, Ben" docDate="2013" docId="9978976365058529D3B021D4FC369B36" docLanguage="en" docName="ejt-2013-48.pdf.imf" docOrigin="European Journal of Taxonomy 48" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Krohcoma ampla Thuy 2013, sp. nov." docType="treatment" docUuid="CE375CEA-89F3-4641-9C52-71B0F7E80517" docUuidSource="ZooBank" docVersion="10" lastPageNumber="83" masterDocId="6541EF1B6555857BD1302529FFF89F78" masterDocTitle="Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)" masterLastPageNumber="242" masterPageNumber="1" pageNumber="81" updateTime="1698839159859" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
|
|
<mods:mods id="032B0734F574F93E022FAB030D0C1DDF" xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo id="58FA08A21D4CFAC5B2896EE4E47B6891">
|
|
<mods:title id="E25BF4DEA77CFF9A783E1F3D543E181F">Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name id="89A707BD2A08322A226D7EED480A003F" type="personal">
|
|
<mods:role id="52B6C027BFDA37AF3767E862C0F4966B">
|
|
<mods:roleTerm id="8F3842F84E9A2A01AD9A3F8C529C1CF8">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="B7251A8417FA1892AEDFC9CA901C7819">Thuy, Ben</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource id="B11CFA9DB6F819ACD594FEE1E5E3E61A">text</mods:typeOfResource>
|
|
<mods:relatedItem id="37D699158CB0375F08591210349CFA42" type="host">
|
|
<mods:titleInfo id="D2FFDB2EABB2CDB8364CE6D000D9E835">
|
|
<mods:title id="8B0B906B84004222BB0F60D6321F63C6">European Journal of Taxonomy</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part id="18EDB3ACED1A561A1F1683652B841780">
|
|
<mods:date id="D3BEB398AED4B9973DE2A0087F10FA73">2013</mods:date>
|
|
<mods:detail id="07BB93AA54CCEBC0993CE93C9C709BF1" type="pubDate">
|
|
<mods:number id="B31ECF7DEBAEEADCC59519A8A2D1D254">2013-07-02</mods:number>
|
|
</mods:detail>
|
|
<mods:detail id="DECCCA1942BFAE95691884EAECDA17FF" type="issue">
|
|
<mods:number id="E9D43B5F6F44E9FC3C13D318051F4820">48</mods:number>
|
|
</mods:detail>
|
|
<mods:extent id="28287DE39ABD1CDED6C9DC7170905AFF" unit="page">
|
|
<mods:start id="262CD7C2981A8E62DDCD96F023AB37CA">1</mods:start>
|
|
<mods:end id="BBE8D948575064FE4B7BCBAF8685B030">242</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:classification id="DC06B4801206B2322A2F564FA2E94041">journal article</mods:classification>
|
|
<mods:identifier id="B3EF00F4F729F2A5104604D7771D3B60" type="CLB-Dataset">22249</mods:identifier>
|
|
<mods:identifier id="439EB7302EB335851420CAB96F16A543" type="DOI">10.5852/ejt.2013.48</mods:identifier>
|
|
<mods:identifier id="0D064FEA49F3D855BB96DDF0B524143E" type="GBIF-Dataset">02866e62-ab61-4cf4-9d98-0248e251084e</mods:identifier>
|
|
<mods:identifier id="DC2BDAC7980F86829FA281706267A066" type="ISSN">2118-9773</mods:identifier>
|
|
<mods:identifier id="F23B912861A4F65C1A38B44F96831148" type="Zenodo-Dep">3822836</mods:identifier>
|
|
<mods:identifier id="0F4EE180C1B1AF69597DB95B257A3234" type="ZooBank">E7080722-E348-448D-96E5-D537F4865BB5</mods:identifier>
|
|
</mods:mods>
|
|
<treatment id="9978976365058529D3B021D4FC369B36" ID-DOI="http://doi.org/10.5281/zenodo.3844325" ID-GBIF-Taxon="164044879" ID-Zenodo-Dep="3844325" LSID="urn:lsid:zoobank.org:act:CE375CEA-89F3-4641-9C52-71B0F7E80517" httpUri="http://treatment.plazi.org/id/9978976365058529D3B021D4FC369B36" lastPageId="82" lastPageNumber="83" pageId="80" pageNumber="81">
|
|
<subSubSection id="59CB75FE6505852BD3B021D4FC4C9A60" box="[640,948,1277,1304]" pageId="80" pageNumber="81" type="nomenclature">
|
|
<paragraph id="116E26756505852BD3B021D4FC4C9A60" blockId="80.[373,1214,1277,1379]" box="[640,948,1277,1304]" pageId="80" pageNumber="81">
|
|
<heading id="4A2691196505852BD3B021D4FC4C9A60" box="[640,948,1277,1304]" centered="true" fontSize="11" level="2" pageId="80" pageNumber="81" reason="2">
|
|
<taxonomicName id="D6D15DF66505852BD3B021D4FCAA9A60" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[640,850,1277,1304]" class="Ophiuroidea" family="Ophiacanthidae" genus="Krohcoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="80" pageNumber="81" phylum="Echinodermata" rank="species" species="ampla" status="sp. nov.">
|
|
<emphasis id="23A5FA676505852BD3B021D4FCAA9A60" bold="true" box="[640,850,1277,1304]" italics="true" pageId="80" pageNumber="81">Krohcoma ampla</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="3896471C6505852BD26921D7FC4C9A60" box="[857,948,1278,1304]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
|
|
</heading>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="59CB75FE6505852BD045200AFC9E9A1B" pageId="80" pageNumber="81" type="description">
|
|
<paragraph id="116E26756505852BD045200AFB469A46" blockId="80.[373,1214,1277,1379]" box="[373,1214,1315,1342]" pageId="80" pageNumber="81">
|
|
<uri id="65402A776505852BD045200AFB469A46" box="[373,1214,1315,1342]" pageId="80" pageNumber="81">
|
|
urn:lsid:zoobank.org:act:
|
|
<uuid id="65771CA06505852BD3A6200AFB469A46" box="[662,1214,1315,1342]" pageId="80" pageNumber="81">CE375CEA-89F3-4641-9C52-71B0F7E80517</uuid>
|
|
</uri>
|
|
</paragraph>
|
|
<paragraph id="116E26756505852BD3FE2060FC9E9A1B" blockId="80.[373,1214,1277,1379]" box="[718,870,1352,1379]" pageId="80" pageNumber="81">
|
|
<figureCitation id="89EA3AF06505852BD3FE2060FCDC9A1B" box="[718,804,1352,1379]" captionStart="Fig" captionStartId="76.[189,231,1613,1639]" captionTargetBox="[310,1278,265,1574]" captionTargetId="figure@76.[310,1278,265,1574]" captionTargetPageId="76" captionText="Fig. 15. Fossil lateral arm plates (LAPs) of ophiacanthid brittle stars in external (a) and internal (b) views. 1-3. Geromura teckliformis gen. et sp. nov. from the late Valanginian (Early Cretaceous) of the Ternberg Nappe, Austria. 1. NHMW 2012/0138/0005 (holotype), proximal LAP. 2. NHMW 2012/0138/0006 (paratype), median LAP. 3. NHMW 2012/0138/0007 (paratype), distal LAP. 4-6. Krohcoma mira gen. et sp. nov. from the late Sinemurian to early Pliensbachian (Early Jurassic) of the Glasenbach Gorge, Austria. 4. NHMW 2012/0137/0013 (holotype), proximal LAP. 5. NHMW 2012/0137/0014 (paratype), median LAP. 6. NHMW 2012/0137/0015 (paratype), distal LAP. 7-8. Krohcoma sp. nov. innom from the late Bathonian (Middle Jurassic) of Jumara, India. 7. GZG.INV.78574, proximal LAP. 8. GZG.INV.78575, median to distal LAP. 9-12. Krohcoma ampla gen. et sp. nov. from the early Kimmeridgian of the Pointe du Chay (9-10), France, and the late Kimmeridgian of Trancoso, Portugal (11-12). 9. GZG.INV.78577 (holotype), proximal LAP. 10. GZG.INV.78578 (paratype), median LAP. 11. GZG.INV.78579 (paratype), median LAP. 12. GZG.INV.78580 (paratype), distal LAP. One common scale bar per species is given." figureDoi="http://doi.org/10.5281/zenodo.3822871" httpUri="https://zenodo.org/record/3822871/files/figure.png" pageId="80" pageNumber="81">Fig. 15</figureCitation>
|
|
: 9-12
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="59CB75FE6505852BD18D20A5FDE699CF" pageId="80" pageNumber="81" type="diagnosis">
|
|
<paragraph id="116E26756505852BD18D20A5FECE9ADE" blockId="80.[189,310,1420,1446]" box="[189,310,1420,1446]" pageId="80" pageNumber="81">
|
|
<heading id="4A2691196505852BD18D20A5FECE9ADE" bold="true" box="[189,310,1420,1446]" fontSize="11" level="3" pageId="80" pageNumber="81" reason="3">
|
|
<emphasis id="23A5FA676505852BD18D20A5FECE9ADE" bold="true" box="[189,310,1420,1446]" pageId="80" pageNumber="81">Diagnosis</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph id="116E26756505852BD18D2094FDE699CF" blockId="80.[189,1399,1469,1719]" pageId="80" pageNumber="81">
|
|
Species of
|
|
<taxonomicName id="D6D15DF66505852BD0762094FE399AAF" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[326,449,1469,1495]" class="Ophiuroidea" family="Ophiacanthidae" genus="Krohcoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="80" pageNumber="81" phylum="Echinodermata" rank="genus" status="gen. nov.">
|
|
<emphasis id="23A5FA676505852BD0762094FE399AAF" box="[326,449,1469,1495]" italics="true" pageId="80" pageNumber="81">Krohcoma</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="3896471C6505852BD0FD2094FDC49AAF" box="[461,572,1469,1495]" pageId="80" pageNumber="81" rank="genus">gen. nov.</taxonomicNameLabel>
|
|
with large LAPs, displaying small, poorly defined, protruding spur in dorsal half of outer proximal edge, paralleled by more clearly defined, round spur in dorsal half of inner distal edges; up to six very large spine articulations; very strong dorsalward increase in size of spine articulations and of gaps separating them; ventral lobe of spine articulations connected with outer surface; row of spine articulations not protruding ventrally; tentacle notch relatively small, accounting for slightly more than one-quarter of the total ventral LAP edge width; ridge on inner side of LAP with widened, triangular dorsal tip.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="59CB75FE6505852BD18D23C8FCCF9829" pageId="80" pageNumber="81" type="etymology">
|
|
<paragraph id="116E26756505852BD18D23C8FEBC9983" blockId="80.[189,324,1761,1787]" box="[189,324,1761,1787]" pageId="80" pageNumber="81">
|
|
<heading id="4A2691196505852BD18D23C8FEBC9983" bold="true" box="[189,324,1761,1787]" fontSize="11" level="3" pageId="80" pageNumber="81" reason="3">
|
|
<emphasis id="23A5FA676505852BD18D23C8FEBC9983" bold="true" box="[189,324,1761,1787]" pageId="80" pageNumber="81">Etymology</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph id="116E26756505852BD18D223BFCCF9829" blockId="80.[189,1399,1809,1873]" pageId="80" pageNumber="81">
|
|
Name formed from
|
|
<emphasis id="23A5FA676505852BD098223BFE059854" box="[424,509,1810,1836]" italics="true" pageId="80" pageNumber="81">amplus</emphasis>
|
|
, Latin for “large”, “magnificent”, in reference to the large size of the LAPs of the species in comparison to those of its congeners.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="59CB75FE6505852AD18D2252FCAC9E15" lastPageId="81" lastPageNumber="82" pageId="80" pageNumber="81" type="materials_examined">
|
|
<paragraph id="116E26756505852BD18D2252FE9498EC" blockId="80.[189,364,1914,1941]" box="[189,364,1914,1941]" pageId="80" pageNumber="81">
|
|
<heading id="4A2691196505852BD18D2252FE9498EC" bold="true" box="[189,364,1914,1941]" fontSize="11" level="3" pageId="80" pageNumber="81" reason="3">
|
|
<emphasis id="23A5FA676505852BD18D2252FE9498EC" bold="true" box="[189,364,1914,1941]" pageId="80" pageNumber="81">Type material</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph id="116E26756505852BD1DA2285FE719893" blockId="80.[189,393,1964,2027]" pageId="80" pageNumber="81">
|
|
<materialsCitation id="A1B92C286505852BD1DA2285FE719893" ID-GBIF-Occurrence="2612041303" collectionCode="GZG" pageId="80" pageNumber="81" specimenCode="GZG.INV.78577" specimenCount="1" typeStatus="holotype">
|
|
<emphasis id="23A5FA676505852BD1DA2285FEA498BE" bold="true" box="[234,348,1964,1990]" pageId="80" pageNumber="81">
|
|
<typeStatus id="CE6A98D76505852BD1DA2285FEA498BE" box="[234,348,1964,1990]" pageId="80" pageNumber="81" type="holotype">Holotype</typeStatus>
|
|
</emphasis>
|
|
<specimenCode id="41778E0E6505852BD18D22F9FE7D9893" box="[189,389,2000,2027]" pageId="80" pageNumber="81">GZG.INV.78577</specimenCode>
|
|
.
|
|
</materialsCitation>
|
|
</paragraph>
|
|
<paragraph id="116E26756504852AD1DA2422FE9F9E5D" blockId="81.[189,1399,267,365]" box="[234,359,267,293]" pageId="81" pageNumber="82">
|
|
<emphasis id="23A5FA676504852AD1DA2422FE9F9E5D" bold="true" box="[234,359,267,293]" pageId="81" pageNumber="82">
|
|
<typeStatus id="CE6A98D76504852AD1DA2422FE9F9E5D" box="[234,359,267,293]" pageId="81" pageNumber="82" type="paratype">Paratypes</typeStatus>
|
|
</emphasis>
|
|
</paragraph>
|
|
<paragraph id="116E26756504852AD18D2406FCAC9E15" blockId="81.[189,1399,267,365]" pageId="81" pageNumber="82">
|
|
GZG.INV.78578, same locality as
|
|
<typeStatus id="CE6A98D76504852AD3582406FD2A9E31" box="[616,722,303,329]" pageId="81" pageNumber="82" type="holotype">holotype</typeStatus>
|
|
; GZG.INV.78579 and GZG.INV.78580 from the upper Kimmeridgian Amaral Formation of Trancoso,
|
|
<collectingCountry id="69C666E56504852AD3D9247AFCA89E15" box="[745,848,339,365]" name="Portugal" pageId="81" pageNumber="82">Portugal</collectingCountry>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="59CB75FE6504852AD18D24BCFEDA9D7A" pageId="81" pageNumber="82" type="distribution">
|
|
<paragraph id="116E26756504852AD18D24BCFE039ED7" blockId="81.[189,507,405,431]" box="[189,507,405,431]" pageId="81" pageNumber="82">
|
|
<heading id="4A2691196504852AD18D24BCFE039ED7" bold="true" box="[189,507,405,431]" fontSize="11" level="3" pageId="81" pageNumber="82" reason="3">
|
|
<emphasis id="23A5FA676504852AD18D24BCFE039ED7" bold="true" box="[189,507,405,431]" pageId="81" pageNumber="82">Type locality and horizon</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph id="116E26756504852AD18D24EDFEDA9D7A" blockId="81.[189,1398,452,514]" pageId="81" pageNumber="82">
|
|
Pointe du Chay near La Rochelle,
|
|
<collectingCountry id="69C666E56504852AD37924ECFD659EA7" box="[585,669,453,479]" name="France" pageId="81" pageNumber="82">France</collectingCountry>
|
|
;
|
|
<emphasis id="23A5FA676504852AD39724EDFCFE9EA6" box="[679,774,452,478]" italics="true" pageId="81" pageNumber="82">Achilles</emphasis>
|
|
Subzone,
|
|
<taxonomicName id="D6D15DF66504852AD24E24EDFBC69EA7" authority="Zone" authorityName="Zone" box="[894,1086,452,479]" class="Malacostraca" family="Sphaeromatidae" genus="Cymodoce" kingdom="Animalia" order="Isopoda" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="23A5FA676504852AD24E24EDFC029EA6" box="[894,1018,452,478]" italics="true" pageId="81" pageNumber="82">Cymodoce</emphasis>
|
|
Zone
|
|
</taxonomicName>
|
|
, early Kimmeridgian, Late Jurassic.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="59CB75FE6504852AD18D2702FAEE9DE0" pageId="81" pageNumber="82" type="materials_examined">
|
|
<paragraph id="116E26756504852AD18D2702FE4C9D3D" blockId="81.[189,436,555,581]" box="[189,436,555,581]" pageId="81" pageNumber="82">
|
|
<heading id="4A2691196504852AD18D2702FE4C9D3D" bold="true" box="[189,436,555,581]" fontSize="11" level="3" pageId="81" pageNumber="82" reason="3">
|
|
<emphasis id="23A5FA676504852AD18D2702FE4C9D3D" bold="true" box="[189,436,555,581]" pageId="81" pageNumber="82">Additional material</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph id="116E26756504852AD18D2773FAEE9DE0" blockId="81.[189,1398,602,665]" pageId="81" pageNumber="82">
|
|
GZG.INV.78581 (dissociated LAP) from the Kimmeridgian of Pointe du Chay,
|
|
<collectingCountry id="69C666E56504852AD5642772FB509D0D" box="[1108,1192,603,629]" name="France" pageId="81" pageNumber="82">France</collectingCountry>
|
|
; GZG.INV.78582 (12 dissociated LAPs) from the upper Kimmeridgian Amaral Formation of Trancoso,
|
|
<collectingCountry id="69C666E56504852AD59A2757FAE99DE0" box="[1194,1297,638,664]" name="Portugal" pageId="81" pageNumber="82">Portugal</collectingCountry>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="59CB75FE65048529D18D27E9FA8F9D58" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82" type="description">
|
|
<paragraph id="116E26756504852AD18D27E9FEB69DA2" blockId="81.[189,334,704,730]" box="[189,334,704,730]" pageId="81" pageNumber="82">
|
|
<heading id="4A2691196504852AD18D27E9FEB69DA2" bold="true" box="[189,334,704,730]" fontSize="11" level="3" pageId="81" pageNumber="82" reason="3">
|
|
<emphasis id="23A5FA676504852AD18D27E9FEB69DA2" bold="true" box="[189,334,704,730]" pageId="81" pageNumber="82">Description</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph id="116E26756504852AD1DA27D9FEA49C72" blockId="81.[189,1399,752,1354]" box="[234,348,752,778]" pageId="81" pageNumber="82">
|
|
<emphasis id="23A5FA676504852AD1DA27D9FEA49C72" bold="true" box="[234,348,752,778]" pageId="81" pageNumber="82">
|
|
<typeStatus id="CE6A98D76504852AD1DA27D9FEA49C72" box="[234,348,752,778]" pageId="81" pageNumber="82" type="holotype">Holotype</typeStatus>
|
|
</emphasis>
|
|
</paragraph>
|
|
<paragraph id="116E26756504852AD18D263DFACB9B36" blockId="81.[189,1399,752,1354]" pageId="81" pageNumber="82">GZG.INV.78577 is a very large, dissociated proximal LAP; at least twice higher than wide; dorsal edge nearly straight; distal edge obliquely convex; proximal edge irregularly concave, with small, poorly defined and weakly prominent, yet protruding spur; outer surface with finely meshed stereom, devoid of conspicuous ornament elements. Six very large, ear-shaped spine articulations arranged in dorsoproximally strongly receding row on elevated distal portion of LAP; very strong dorsalward increase in size of spine articulations and of gaps separating them; dorsal and ventral lobes of spine articulations thick, merged into continuous volute encompassing conspicuously large muscle opening; ventral lobe of spine articulations connected with finely meshed outer surface; row of spine articulations not protruding ventrally. Ventral edge of LAP irregularly convex, tentacle notch not discernible in external view.</paragraph>
|
|
<paragraph id="116E26756504852AD18D2171FAB89A32" blockId="81.[189,1399,752,1354]" pageId="81" pageNumber="82">Inner side of LAP with large, well-defined, prominent ridge; dorsal half nearly straight, oblique, with widened triangular dorsal tip, separated from ventral half by rounded kink; ventral half short, less welldefined than dorsal one, not connected with ventral edge of LAP; inner side of distal edge of LAP with small, moderately well-defined, slightly prominent spur composed of slightly more densely meshed stereom; inner side of tentacle notch large, accounting for slightly more than a quarter of the total ventral LAP edge width. Vertical row of small perforations distally bordering kink between dorsal and ventral portions of the ridge, with dorsalward increase in size of perforations and of gaps separating them.</paragraph>
|
|
<paragraph id="116E26756504852AD1DA205BFD489AF4" blockId="81.[189,1399,1394,1744]" box="[234,688,1394,1420]" pageId="81" pageNumber="82">
|
|
<emphasis id="23A5FA676504852AD1DA205BFD489AF4" bold="true" box="[234,688,1394,1420]" pageId="81" pageNumber="82">
|
|
<typeStatus id="CE6A98D76504852AD1DA205BFEA49AF4" box="[234,348,1394,1420]" pageId="81" pageNumber="82" type="paratype">Paratype</typeStatus>
|
|
supplements and variation
|
|
</emphasis>
|
|
</paragraph>
|
|
<paragraph id="116E26756504852AD18D20BFFCFC9964" blockId="81.[189,1399,1394,1744]" pageId="81" pageNumber="82">
|
|
GZG.INV.78578 is a dissociated median LAP; slightly higher than wide; well matching
|
|
<typeStatus id="CE6A98D76504852AD5FD20BFFACF9AC8" box="[1229,1335,1430,1456]" pageId="81" pageNumber="82" type="holotype">holotype</typeStatus>
|
|
; spur on proximal edge slighty better defined and more prominent. Five very large spine articulations similar to those observed on
|
|
<typeStatus id="CE6A98D76504852AD08320F7FDE59A80" box="[435,541,1502,1528]" pageId="81" pageNumber="82" type="holotype">holotype</typeStatus>
|
|
; connection between ventral lobe of spine articulations and finely meshed stereom of outer surface slightly more prominent.
|
|
</paragraph>
|
|
<paragraph id="116E26756504852AD18D230EFE2399A8" blockId="81.[189,1399,1394,1744]" pageId="81" pageNumber="82">Inner side of LAP largely obscured by sediment, only ventral portion of ridge discernible, separated from ventral edge of LAP; inner side of distal edge of LAP with small, poorly defined, round and weakly prominent spur composed of slightly more densely meshed stereom. Inner side of tentacle notch moderately large, distally bordered by moderately well-defined, oblique, prominent ridge. No perforations discernible.</paragraph>
|
|
<paragraph id="116E26756504852AD18D23D1FDDF98DB" blockId="81.[189,1399,1784,2027]" pageId="81" pageNumber="82">
|
|
GZG.INV.78579 is a dissociated median LAP; slightly higher than wide; dorsal edge concave; outer surface with slightly more coarsely meshed stereom than in
|
|
<typeStatus id="CE6A98D76504852AD2A72234FBF9984F" box="[919,1025,1821,1847]" pageId="81" pageNumber="82" type="holotype">holotype</typeStatus>
|
|
, probably as a result of poorer preservation. Four very large spine articulations similar to those observed in
|
|
<typeStatus id="CE6A98D76504852AD5052268FB649823" box="[1077,1180,1857,1883]" pageId="81" pageNumber="82" type="holotype">holotype</typeStatus>
|
|
but with less welldeveloped connection between outer surface and ventral lobe of spine articulations, again probably due to slightly poorer preservation.
|
|
</paragraph>
|
|
<paragraph id="116E267565048529D18D2284FE239E32" blockId="81.[189,1399,1784,2027]" lastBlockId="82.[189,1399,267,330]" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82">Inner side of LAP with well-defined ridge separated by round kink into dorsal portion with slightly widened dorsal tip, and much shorter ventral portion not connected with ventral edge of LAP; inner side of tentacle notch large, distally bordered by poorly defined, short, oblique ridge-like knob. No perforations discernible.</paragraph>
|
|
<paragraph id="116E267565078529D18D245AFD529EAF" blockId="82.[189,1399,371,544]" pageId="82" pageNumber="83">GZG.INV.78580 is a dissociated distal LAP; slightly wider than high; proximal edge slightly fragmentary. Three very large spine articulations, with very strong dorsalward increase in size; dorsal gap between spine articulations larger than ventral one.</paragraph>
|
|
<paragraph id="116E267565078529D18D24C8FA8F9D58" blockId="82.[189,1399,371,544]" pageId="82" pageNumber="83">Inner side of LAP with very poorly defined, oblique, weakly prominent ridge composed of more finely meshed stereom; dorsal half of inner side of distal LAP edge with small, round, well-defined, prominent spur.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="59CB75FE65078529D18D2761FCE09CA4" pageId="82" pageNumber="83" type="discussion">
|
|
<paragraph id="116E267565078529D18D2761FED79D1A" blockId="82.[189,303,584,610]" box="[189,303,584,610]" pageId="82" pageNumber="83">
|
|
<heading id="4A26911965078529D18D2761FED79D1A" bold="true" box="[189,303,584,610]" fontSize="11" level="3" pageId="82" pageNumber="83" reason="3">
|
|
<emphasis id="23A5FA6765078529D18D2761FED79D1A" bold="true" box="[189,303,584,610]" pageId="82" pageNumber="83">Remarks</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph id="116E267565078529D18D2751FCE09CA4" blockId="82.[189,1399,632,988]" pageId="82" pageNumber="83">
|
|
Although, at first sight, these LAPs seem to differ significantly from those assigned to
|
|
<taxonomicName id="D6D15DF665078529D5F22751FAC59DEA" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[1218,1341,632,658]" class="Ophiuroidea" family="Ophiacanthidae" genus="Krohcoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="82" pageNumber="83" phylum="Echinodermata" rank="genus" status="gen. nov.">
|
|
<emphasis id="23A5FA6765078529D5F22751FAC59DEA" box="[1218,1341,632,658]" italics="true" pageId="82" pageNumber="83">Krohcoma</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="3896471C65078529D4752750FF099DCF" pageId="82" pageNumber="83" rank="genus">gen. nov.</taxonomicNameLabel>
|
|
, similarities in spine articulation morphology and arrangement are striking. In fact, large spine articulations with thick dorsal and ventral lobes encompassing a conspicuously large muscle opening, arranged in a dorso-proximalwards receding row on the elevated distal portion of the LAPs are a highly distinctive feature which is not found in any other ophiacanthid LAP
|
|
<typeStatus id="CE6A98D765078529D2DF2622FBDD9C5D" box="[1007,1061,779,805]" pageId="82" pageNumber="83">type</typeStatus>
|
|
. There are admittedly strong differences between these LAPs and those of its congeners in terms of size of tentacle notch, connection between outer surface and ventral lobe of spine articulations and size and shape of the ridge on the inner side. These differences might, in fact, warrant separation at the generic level. On the basis of the striking similarities in spine articulation morphology and arrangement, however, the present LAPs are here considered as a new species of
|
|
<taxonomicName id="D6D15DF665078529D31B26EBFD5E9CA4" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[555,678,962,988]" class="Ophiuroidea" family="Ophiacanthidae" genus="Krohcoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="82" pageNumber="83" phylum="Echinodermata" rank="genus" status="gen. nov.">
|
|
<emphasis id="23A5FA6765078529D31B26EBFD5E9CA4" box="[555,678,962,988]" italics="true" pageId="82" pageNumber="83">Krohcoma</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="3896471C65078529D39E26EBFCE09CA4" box="[686,792,962,988]" pageId="82" pageNumber="83" rank="genus">gen. nov.</taxonomicNameLabel>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="59CB75FE65078529D18D212DFC369B36" box="[189,974,1028,1103]" pageId="82" pageNumber="83" type="distribution">
|
|
<paragraph id="116E267565078529D18D212DFEB79B66" blockId="82.[189,335,1028,1054]" box="[189,335,1028,1054]" pageId="82" pageNumber="83">
|
|
<heading id="4A26911965078529D18D212DFEB79B66" bold="true" box="[189,335,1028,1054]" fontSize="11" level="3" pageId="82" pageNumber="83" reason="3">
|
|
<emphasis id="23A5FA6765078529D18D212DFEB79B66" bold="true" box="[189,335,1028,1054]" pageId="82" pageNumber="83">Occurrence</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
<paragraph id="116E267565078529D18D211DFC369B36" blockId="82.[189,974,1076,1103]" box="[189,974,1076,1103]" pageId="82" pageNumber="83">
|
|
Early Kimmeridgian of
|
|
<collectingCountry id="69C666E565078529D0E6211CFDDE9B37" box="[470,550,1077,1103]" name="France" pageId="82" pageNumber="83">France</collectingCountry>
|
|
and late Kimmeridgian of
|
|
<collectingCountry id="69C666E565078529D253211DFC329B36" box="[867,970,1076,1102]" name="Portugal" pageId="82" pageNumber="83">Portugal</collectingCountry>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |